http://www.collembola.org/taxa/arrhidae.htm - Last updated on 2019.05.24 by Frans Janssens
Checklist of the Collembola: Arrhopalitidae

Familia Arrhopalitidae Stach, 1956, sensu Bretfeld G, 1999:13 key

Genus Number
of species
Arrhopalites50
Pygmarrhopalites101
Troglopalites1
Arrhopalitidae152
Genera with number of species
  Syn.: ... Smynthuridae(sic) Lubbock, 1870, partim ISS
  Syn.: ... Smynthuridae(sic) Börner, 1906, partim ISS
  Syn.: Collembola Smynthuridae(sic) Lubbock, J, 1873:100 partim ISS
  Syn.: Collembola Sminthuri(sic) Folsom, JW, 1896:446 partim ISS


  Syn.: Katianninae Arrhopalini(sic) Richards, 1956 ISS
  Syn.: Katianninae Arrhopalini(sic) Richards in 0918:31 ISS
  Syn.: Sminthurinae Arrhopalini(sic) Christiansen, K, 1966:48 ISS
  Syn.: Symphypleona Arrhopalitdae(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:71 ISS
Arrhopalites hirtus from the USA
2005 © Cheung, D. & Schmidt, J.
Arrhopalites sp. from the USA
2006 © Bernard, E.




Genus Arrhopalites Börner, C, 1906:182 key  key  key  key  key

  Syn.: Smynthurus(sic) niveus Lebedinsky, 1904
  Syn.: Sminthuridae Pseudarrhopalites Stach, 1945
  Syn.: Arrhopalites Coecarrhopalites(sic) Yosii, 1954,nn IOS
  Syn.: Arrhopalites Coecarrhopalites(sic) Yosii, 1967 ISS
  Syn.: Arrhopalites binoculatus Yosii, 1955
  Syn.: Sphyrotheca binoculatus Scott, 1964 partim holotype
  Syn.: Sphyrotheca binoculatus Scott, 1964 partim paratypes
  Syn.: Sminthurinus reticulatus Falahati HosseinAbad, A, Kheirodin, A et Bretfeld, G, 2013:208
Sminthurinus reticulatus from Iran
After Falahati Hossein Abad A & al, 2013 Fig.3




  Syn.: Arrhopalites coecus(sic) Stach, J, 1945:7,gr ISS
  Syn.: Arrhopalites coecus(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:135,gr ISS
  Syn.: Arrhopalites coecus(sic) Zeppelini, D, 2006:22,gr ISS
  Syn.: Arrhopalites coecus(sic) Zeppelini, D, 2011:91,gr ISS
  Syn.: Arrhopalitidae Arrhropalites(sic) Palacios-Vargas, JG, 1999:102 ISS
  Syn.: Arrhopalitidae Arropalites(sic) Zeppelini, DF et Palacios-Vargas JG, 1999:80 ISS





Arrhopalites sp. from France
2007.10.07 © Lebeaux, P.
Arrhopalites sp. from Hungary
2010.03.15 © Pfliegler, W.
Arrhopalites sp. from Hungary
2010.03.15 © Pfliegler, W.
Arrhopalites sp. from Croatia
2010.11.19 © Lukic, M.




  Syn.: Arrhopalitidae Arrhoplaites(sic) Vargovitsh, RS et Busmachiu, G, 2015:539 ISS



Species sp.1 Stach, J, 1956:239

             Syn.: Smynthurus(sic) gracilis Absolon, K, 1901:209 ISS

Species sp.3 Stach, J, 1956:240

            Syn.: Smynthurus(sic) niveus Joseph, 1882

Species sp.4 Christiansen, K, 1966:49

Species sp.5 Christiansen, K, 1966:49

Species sp.6 Christiansen, K, 1966:49

Species sp.7 Zeppelini Filho, D et Castaño Meneses, G, 1995:10

Species sp.8 Zeppelini Filho, D, 1996:27

Species sp.9 Culik, de Souza & Ventura, 2002

Species sp.10 Zeppelini, D, 2011:95

Species sp.11 Zeppelini, D, 2011:95

Species sp.12 Janssens F & Dobbs B, 2016

Arrhopalites? sp.nov. from the USA
Ant.3 with basal swelling; ant.4 with 10 subdivisions
2016.12.20 © Dobbs, Br.
Arrhopalites? sp.nov. from the USA
Dorsal colouration
2016.12.20 © Dobbs, Br.

Species pilulifer (Schött, H, 1893:9) Zeppelini, D, 2006:35

            Syn.: Sminthurus piluliferus(sic) Schött, H, 1893:9 IOS


             Syn.: Sminthurus piluliferus(sic) Salmon, JT, 1964:597 ISS
             Syn.: Arrhopalites piluliferus(sic) Zeppelini, D, 2006:35 ISS



Species violaceus (Handschin, 1930) Stach, J, 1956:72, s.i.


Species-group caecus Vargovitsh RS, 2013:27
Species abchasicus Vargovitsh, RS, 2013:17

Arrhopalites abchasicus from Abkhazia
After Vargovitsh RS, 2013 Fig.23
Arrhopalites abchasicus from Abkhazia
After Vargovitsh RS, 2013 Fig.24
            Syn.: Arrhopalites caecus Barjadze & Djanashvili, 2008


Species antrobius Yosii, R, 1954:68

Species caecus (Tullberg, 1871:146 tt) Börner, C, 1906:182, t.t., B

            Syn.: Sminthurus cæcus(sic) Tullberg, 1871 tt IOS
            Syn.: Sminthurus carniolicus von Dalla Torre, KW, 1895:8
             Syn.: Sminthurus coecus(sic) Stach, 1956, nec Tullberg, 1871 ISS
            Syn.: Smynthurus(sic) benitus Folsom, 1896
             Syn.: Smynthurus(sic) banitus(sic) Zeppelini D, 2006:26 ISS
            Syn.: Sminthurinus aureus Tuxen, 1944


             Syn.: Sminthurus (Pilosi) cæcus(sic) Tullberg, T, 1872:33 ISS
             Syn.: Sminthurus coecus(sic) Reuter, OM, 1876:80 ISS
             Syn.: Sminthurus coecus(sic) Reuter, 1890 ISS
             Syn.: Sminthurus coecus(sic) Schäffer, 1896 ISS
             Syn.: Sminthurus coecus(sic) Scherbakov, 1898 ISS
             Syn.: Sminthurinus coecus(sic) Börner, 1901 ISS
             Syn.: Arrhopalites coecus(sic) Linnaniemi, WM, 1912:271 ISS
             Syn.: Arrhopalites coecus(sic) Houlbert, 1924 ISS
             Syn.: Arrhopalites coecus(sic) Brown, 1926 ISS
             Syn.: Arrhopalites coecus(sic) Handschin, 1926 ISS
             Syn.: Arrhopalites coecus(sic) Edwards, 1929 ISS
             Syn.: Arrhopalites coecus(sic) Handschin, 1929 ISS
             Syn.: Arrhopalites coecus(sic) Denis, 1932 ISS
             Syn.: Arrhopalites coecus(sic) Womersley, 1932 ISS
             Syn.: Arrhopalites coecus(sic) Turk, 1933 ISS
             Syn.: Arrhopalites coecus(sic) Davies, 1934 ISS
             Syn.: Arrhopalites coecus(sic) Dürkop, 1934 ISS
             Syn.: Arrhopalites coecus(sic) Stammer, 1936 ISS
             Syn.: Arrhopalites coecus(sic) Frenzel, 1937 ISS
             Syn.: Arrhopalites coecus(sic) Weidner, 1937 ISS
             Syn.: Arrhopalites coecus(sic) Denis, 1938 ISS
             Syn.: Arrhopalites coecus(sic) Muhlmann, 1942 ISS
             Syn.: Arrhopalites coecus(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Arrhopalites coecus(sic) Stach, J, 1945:9 ISS
             Syn.: Arrhopalites coecus(sic) Denis, 1948 ISS
             Syn.: Arrhopalites coecus(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:133 ISS
             Syn.: Arrhopalites coecus(sic) Weidner, 1953 ISS
             Syn.: Arrhopalites coecus(sic) Bellinger, 1954 ISS
             Syn.: Arrhopalites coecus(sic) Obenberger, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Stach, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Strenzke, 1955 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1956 ISS
             Syn.: Arrhopalites coecus(sic) Bödvarsson, H, 1957:38 ISS
             Syn.: Arrhopalites coecus(sic) Strebel, 1957 ISS
             Syn.: Arrhopalites coecus(sic) Grinbergs, 1958 ISS
             Syn.: Arrhopalites coecus(sic) von Törne, 1958 ISS
             Syn.: Arrhopalites coecus(sic) da Gama, 1959 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1959 ISS
             Syn.: Arrhopalites coecus(sic) Grinbergs, 1960 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1961 ISS
             Syn.: Arrhopalites coecus(sic) Paclt, 1962 ISS
             Syn.: Coecarrhopalites(sic) coecus(sic) Delamare Deboutteville & Massoud, 1963 ISS
             Syn.: Arrhopalites coecus(sic) Pomorski & Skarzynski, 1991 ISS
             Syn.: Arrhopalites coecus(sic) Palacios-Vargas & Zeppelini, [1996] ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, 1996 ISS
             Syn.: Arrhopalites (Arrhopalites) coecus(sic) Zeppelini Filho, D, 1996:57 ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, [1997] ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini Filho, D, 1997:122 ISS
             Syn.: Arrhopalites coecus(sic) Klag, J et Swiatek, P, 1999:161 ISS
             Syn.: Arrhopalites coecus(sic) Beruete, E, Baquero, E et Jordana, R, 2002:397 ISS
             Syn.: Arrhopalites coecus(sic) Zeppelini, D et Christiansen, K, 2003:37 ISS
Arrhopalites caecus from the UK
Mucro
2005 © Hopkin, S.P.

             Syn.: Arrhopalites ceacus(sic) Moore, JC, Saunders, P, Selby, G, Horton, H, Chelius, MK, Chapman, A et Horrocks, RD, 2005:116 ISS
Arrhopalites caecus from Hungary
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus ♀ from Hungary
Sub-anal appendage
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Pale colour form
2011.02.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Mandibula with apical teeth and molar plate
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus ♀ from Hungary
Sub-anal appendage
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Mucro with round tip (*), serrated edges and basal ventral spine (**)
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Undivided 4th antennal segment
2013.01.11 © Pfliegler, W.P.
Arrhopalites caecus from Hungary
Bothriotricha ABC sub-orthogonal
2013.01.11 © Pfliegler, W.P.

  Subspecies attenuatus Carpenter & Evans, 1899





Species coreanus Park & Kang, 2007

Species gul Yosii, R, 1966:559

Species incertus Zeppelini, D et Christiansen, K, 2003:36

Species karabiensis Vargovitsh, RS, 2009:10

Arrhopalites karabiensis from Ukraine
After Vargovitsh, R.S., 2009 Fig.2

Species loczyi Loksa, 1960

Species macronyx Vargovitsh, RS, 2012:2

Arrhopalites macronyx from Western Caucasus
After Vargotitsh RS, 2012 Fig.25
Arrhopalites macronyx from Western Caucasus
After Vargotitsh RS, 2012 Fig.26

Species millsi (Delamare Deboutteville, C & Massoud, Z, 1963:215) Betsch, 1980

            Syn.: Caecarrhopalites Millsi(sic) Delamare Deboutteville, C et Massoud, Z, 1963:215 IOS

Species minor Park & Kang, 2007

Species minutus Yosii, R, 1970:17

Species peculiaris Vargovitsh, RS, 2009:15

Species potapovi Vargovitsh, RS, 2015:102

Arrhopalites potapovi ♀ from Russia
After Vargovitsh RS, 2015 Fig.24

Species pukouensis Wu, M et Christiansen, K, 1997:266

Species tenuis Stach, J, 1945:14

            Syn.: Arrhopalites Remyi(sic) Denis, 1947


Species ulehlovae Rusek, 1970


Species-group diversus Vargovitsh RS, 2013:27
Species sp.1 Vargovitsh, RS et Busmachiu, G, 2015:548

Species acanthophthalmus Gisin, 1958

             Syn.: Arrhopalites acantophthalmus(sic) Dallai, R, 1973:560 ISS
             Syn.: Arrhopalites acantophthalmus(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895 ISS

Species antonioiGiuga & Jordana, 2013

Species baccettii Dallai, 1969

             Syn.: Arrhopalites (diversus-group) baccetii(sic) Vargovitsh, RS et Busmachiu, G, 2015:539 ISS

Species diversus Mills, HB, 1934:93

Arrhopalites diversus from the USA
Ant.3 with distinct basal swelling (*)
2018.01.17 © Dobbs, Br.
Arrhopalites diversus from the USA
Eyepatch with 1 ocellus
2018.01.17 © Dobbs, Br.
Arrhopalites diversus from the USA
Ant.1:2:3:4 = 1:2.2:3.75:6.25
Ant.4 not subdivided
2018.01.17 © Dobbs, Br.

Species ezoensis Zeppelini, D, 2004:4

Species microphthalmus Cassagnau, P et Delamare Deboutteville, C, 1953:133, B

             Syn.: Arrhopalites microphtalmus(sic) Zeppelini Filho, D, 1996:27 ISS
             Syn.: Arrhopalites (Arrhopalites) microphtalmus(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites microphtalmus(sic) Zeppelini Filho, D, [1997]:12 ISS
             Syn.: Arrhopalites microphtalmus(sic) Jacobs, W, 1997:71,B ISS
             Syn.: Arrhopalites microphtalmus(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:58,B ISS

Species nivalis Yosii, 1966

Species prutensis Vargovitsh, RS et Busmachiu, G, 2015:540

Arrhopalites prutensis ♀ from Moldova
After Vargovitsh RS & Busmachiu G, 2015 Fig.2

Species-group harveyi Vargovitsh RS, 2013:27
Species alambariensis Zeppelini, D, 2006:19

Species amorimi Palacios-Vargas, JG et Zeppelini, D, 1996:10

Species anulifer Nayrolles, 1990

Species botuveraensis Zeppelini, D, 2006:22

Species glabrofasciatus Zeppelini, D, Brito, RA et Lima, ECA, 2018:60

Arrhopalites glabrofasciatus ♀ from Brazil
After Zeppelini D & al, 2018 Fig.1

Species gnaspinii Palacios-Vargas, JG et Zeppelini, D, 1996:8

            Syn.: Arrhopalites (Coecarrhopalites(sic)) gnaspinius(sic) Palacios-Vargas, JG et Zeppelini, D, 1996:8 IOS


             Syn.: Arrhopalites gnaspinius(sic) Mari Mutt, JA et Bellinger, PF, 1996:172 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, 1996:29 ISS
             Syn.: Arrhopalites (Arrhopalites) gnaspinius(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, [1997]:12 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini Filho, D, 1997:122 ISS
             Syn.: Arrhopalites gnaspinius(sic) Zeppelini, DF et Palacios-Vargas JG, 1999:81 ISS
             Syn.: Arrhopalites gnaspinius(sic) Culik, MP et Zeppelini, DF, 2003:1131 ISS



Species harveyi Denis, 1933

             Syn.: Coecarrhopalites(sic) Harveyi(sic) Delamare Deboutteville & Massoud, 1963 ISS

Species heteroculatus Zeppelini, D, 2006:20

Species lawrencei Palacios-Vargas, JG et Zeppelini, D, 1996:12

             Syn.: Arrhopalites (Coecarrhopalites(sic)) lawrenci(sic) Palacios-Vargas, JG et Zeppelini, D, 1996:23 ISS

Species paranaensis Zeppelini, D, 2006:24

Species vazquezae Palacios-Vargas, JG et Zeppelini, D, 1996:14


Genus Pygmarrhopalites Vargovitsh, RS, 2009:20

  Syn.: Sminthurinus Articulati Börner, C, 1901:100
  Syn.: Collembola Smynthrus(sic) Christiansen, K, 1966:53 ISS
  Syn.: Arrhopalites Novarrhopalites Zeppelini Filho, D, 1996:57 =i
  Syn.: Arrhopalites pygmaeus-group Stach, J, 1945:7 ONB csyn Arrhopalites%{Katianninae}%2885:164%partim
  Syn.: Arrhopalitidae Arrhopalites Marx, MT et Weber, D, 2013:234


Arrhopalites sp. from Chile
After Jordana R & Baquero E, 2008 Fig.5
Remark by Zeppelini, D. 2011:96: The name Pygmarrhopalites has only a didactic meaning, being a convenience for referring to species in the pygmaeus-group, which follows the system proposed by Bretfeld (1999). Thus the generic name Pygmarrhopalites should be treated as a junior synonym of Arrhopalites s.l. Börner.


Species sp.1 Zeppelini Filho, D, 1996:27

Species sp.2 Vargovitsh, RS et Busmachiu, G, 2015:540

Species sp.3 Vargovitsh, RS et Busmachiu, G, 2015:540

Species aggtelekiensis (Stach, J, 1930:308) Vargovitsh, RS, 2009:26

             Syn.: Arrhopalites aggtelekensis(sic) Gisin, 1960 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Christiansen, 1966 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Loksa, I, 1967:294 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Zeppelini, 1996 ISS
             Syn.: Arrhopalites aggtelekensis(sic) Hubart, J-M et Dethier, M, 1999:168 ISS
Pygmarrhopalites aggtelekiensis from Slovakia
2012 © Kovác, L.



Species alticolus (Yosii, R, 1970:15) Vargovitsh, RS, 2009:36

             Syn.: Arrhopalites alticola(sic) Zeppelini, D, 2004:3 ISS
             Syn.: Pygmarrhopalites alticola(sic) Vargovitsh, RS, 2009:36 ISS
             Syn.: Pygmarrhopalites alticola(sic) Vargovitsh, RS, 2009:e18 ISS
             Syn.: Pygmarrhopalites alticola(sic) Jordana, R, Fadrique, F et Baquero, E, 2012:63 ISS

Species altus (Christiansen, K, 1966:51) Vargovitsh, RS, 2009:e18

Species amarus (Christiansen, K, 1966:62) Vargovitsh, RS, 2009:20

Species arcus (Zeppelini, D & Christiansen, K, 2003:38) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites arca(sic) Zeppelini, D et Christiansen, K, 2003:38 IOS
             Syn.: Pygmarrhopalites arca(sic) Zeppelini, D, Taylor, SJ et Slay, ME, 2009:1 ISS
             Syn.: Pygmarrhopalites (principalis-group) arca(sic) Vargovitsh, RS, 2017:33 ISS



Species ashcraftensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:13

Species ater (Christiansen, K & Bellinger, P, 1998:1247) Vargovitsh, RS, 2009:e18

Species bellingeri (Christiansen, K, 1966:51) Vargovitsh, RS, 2009:e18

Species benitus (Folsom, JW, 1896:446) Vargovitsh, RS, 2009:e18

Smynthurus(sic) benitus from the USA
Terminal antennal segment
After Folsom JW, 1896 Plate10 Fig.5

            Syn.: Sminthurus caecus Guthrie, 1903
            Syn.: Arrhopalites caecus Linnaniemi, 1907
            Syn.: Arrhopalites sub-binoculatus(sic) Denis, 1933 IOS
             Syn.: Arrhopalites sub-binoculatus(sic) Stach, J, 1956:70 ISS
            Syn.: Arrhopalites binoculatus Denis, 1931
            Syn.: Arrhopalites binoculatus Mills, 1934
             Syn.: ... bimaculatus(sic) Brimley, 1938 lapsus ISS
            Syn.: Arrhopalites binoculatus Maynard, 1951
            Syn.: Arrhopalites principalis Stach, 1945 =?




Species bifidus (Stach, J, 1945:34) Vargovitsh, RS, 2009:20, B

             Syn.: Arrhopalites bificus(sic) Salmon, JT, 1964:571 ISS


Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus from the UK
Sub-anal appendage
2012.08.19 © Murray, A.
Pygmarrhopalites bifidus ♀ from the UK
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus ♀ from the UK
Bifid sub-anal appendages (*)
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus juv. from the UK
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from the UK
Reddish eyepatches
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from the UK
Ant.4 with 5 subdivisions
2015.03.04 © Murray, A.
Pygmarrhopalites bifidus from Holland
Ant.4 with 5 subdivisions
2017.11.22 © Wieringa, P.
Pygmarrhopalites bifidus from Holland
2017.11.22 © Wieringa, P.
Pygmarrhopalites bifidus from Holland
2017.11.22 © Van duinen, J.

Species bimus (Christiansen, K, 1966:55) Vargovitsh, RS, 2009:e18

Species boneti (Stach, J, 1945:16) Vargovitsh, RS, 2009:31

             Syn.: Arrhopalites Boneti(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:135 ISS

Species buekkensis (Loksa, I, 1969:359) Vargovitsh, RS, 2009:26

Species buffaloensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:10

Species caedus (Christiansen, K & Bellinger, P, 1996:170) Vargovitsh, RS, 2009:e18

Species cantavetulae Jordana, R, Fadrique, F et Baquero, E, 2012:58

Species canzianus (Stach, J, 1945:28) Vargovitsh, RS, 2009:20

Species carolynae (Christiansen, K & Bellinger, P, 1996:171) Vargovitsh, RS, 2009:e18

             Syn.: Pygmarrhopalites carolinae(sic) Soto-Adames, FN et Taylor, SJ, 2013:173 ISS

Species carpathicus (Vargovich, RS, 1999:89) Vargovitsh, RS, 2009:20

Pygmarrhopalites carpathicus from the Carpathians
2011.03.11 © Vargovich, R.

Species changbaishanensis (Wu, M & Zhong, W-Y, 1997:215) Vargovitsh, RS, 2009:20

Species chiangdaoensis (Nayrolles, 1990) Vargovitsh, RS, 2009:20

Species chopardi (Cassagnau, P & Delamare Deboutteville, C, 1955:388) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites Chopardi(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:388 IOS

Species clarus (Christiansen, K, 1966:57) Vargovitsh, RS, 2009:e18

Species cochlearifer (Gisin, 1947) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites cochleafer(sic) Christiansen, 1966 ISS
             Syn.: Pygmarrhopalites cochlaerifer(sic) Zeppelini, D, 2011:95,gr ISS
Pygmarrhopalites cochlearifer from the UK
Dark colour form
2013.06.dd © Shaw, P.
Pygmarrhopalites cochlearifer from the UK
Ant.4 with 6 subsegments
2013.06.dd © Shaw, P.
Pygmarrhopalites cochlearifer from the UK
Distinctly strong anterior cephalic spines
2013.06.dd © Shaw, P.



Species commorus (Christiansen, K & Bellinger, P, 1996:172) Vargovitsh, RS, 2009:26

Species crepidinis Jordana R et Baquero E in Jordana, R, Barranco, P, Amezcua, A et Baquero, E, 2017:106

Pygmarrhopallites crepidinis from Spain
After Jordana R & Baquero E in Jordana R & al, 2017 Fig.2A

Species dbari Vargovitsh, RS, 2017:24

Pygmarrhopalites dbari from Abkhazia
After Vargovitsh RS, 2017 Fig.25

Species delamarei (Nosek & Paoletti, 1984:211) Vargovitsh, RS, 2009:e18

Species dubius (Christiansen, 1966:59) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites dubias(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS

Species dudichi (Loksa, I & Rubio, I, 1966:154) Vargovitsh, RS, 2009:20

Species elegans (Cassagnau, P & Delamare Deboutteville, C, 1953:138) Vargovitsh, RS, 2009:e18

Species fransjanssensi Soto-Adames, FN et Taylor, SJ, 2013:164

            Syn.: Pygmarrhopalites fransjanssens(sic) Soto-Adames, FN et Taylor, SJ, 2013:164 IOS
             Syn.: Pygmarrhopalites fransjanssnes(sic) Soto-Adames, FN et Taylor, SJ, 2013:173 ISS



Species furcatus (Stach, J, 1945:22) Vargovitsh, RS, 2009:20

Species giovannensis (Cassagnau, P & Delamare Deboutteville, C, 1953:136) Vargovitsh, RS, 2009:e18

Species gisini (Nosek, 1960) Vargovitsh, RS, 2009:e18

Species habei (Yosii, R, 1956:98) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites binoculatus Yosii, 1954
            Syn.: Arrhopalites binoculatus Yosii, 1955
            Syn.: Arrhopalites binoculatus Yosii, R, 1977:164
            Syn.: Arrhopalites spadica Salmon, 1963


             Syn.: Arrhopalites habet(sic) Yosii, R, 1956:99 ISS
             Syn.: Arrhopalites havei(sic) Christiansen, K, 1966:61 ISS
             Syn.: Arrhopalites Habei(sic) Zeppelini, D, 2004:13 ISS
Pymarrhopalites habei from Japan
2019.02.27 © Hara, Y.
Pymarrhopalites habei from Japan
2019.02.27 © Hara, Y.



Species hedrosensis (Selga, D, 1963:465) Vargovitsh, RS, 2009:e18

Species hirtus (Christiansen, K, 1966:61) Soto-Adames, FN & Taylor, SJ, 2013:148 lapsus

            Syn.: Pygmarrhopalites hirsuta(sic) Soto-Adames, FN et Taylor, SJ, 2013:148 lapsus


Arrhopalites hirtus from the USA
2005 © Cheung, D. & Schmidt, J.

Species hubbardi (Zeppelini, D & Christiansen, K, 2003:37) Vargovitsh, RS, 2009:e18

Arrhopalites hubbardi from the USA
2007.03.15 © Steinmann, D.

Species hungaricus (Loksa, I, 1967:294) Vargovitsh, RS, 2009:e18

Species incantator Soto-Adames, FN et Taylor, SJ, 2013:164

Species infrasecundarius (Loksa, I & Rubio, I, 1966:154) Vargovitsh, RS, 2009:36

Species intermedius (Loksa, I, 1969:357) Vargovitsh, RS, 2009:e18

Species japonicus (Yosii, R, 1956:99) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites binoculatus Yosii, 1954, nec Börner
            Syn.: Arrhopalites binoculatus Yosii, R, 1956:99


Species jay (Christiansen, K & Bellinger, P, 1996:173) Vargovitsh, RS, 2009:20

             Syn.: Arrhopalites jax(sic) Zeppelini Filho, D, 1996:27 ISS
             Syn.: Arrhopalites (Novarrhopalites) jax(sic) Zeppelini Filho, D, 1996:58 ISS
             Syn.: Arrhopalites jax(sic) Zeppelini Filho, D, [1997]:12 ISS

Species jeanneli (Cassagnau, P & Delamare Deboutteville, C, 1955:390) Vargovitsh, RS, 2009:e18

            Syn.: Arrhopalites Jeanneli(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:390 IOS

Species kaprusi Vargovitsh, RS, 2009:26

Species kolymensis (Tshelnokov, VG, 1990:351) Vargovitsh, RS, 2009:36

             Syn.: Arrhopalites kolymiensis(sic) Tshelnokov, VG, 1990:350 ISS

Species kovali Vargovitsh, RS, 2017:33

Pygmarrhopalites kvali from Abkhazia
After Vargovitsh RS, 2017 Fig.57

Species kristiani (Vargovich, RS, 2005:67) Vargovitsh, RS, 2009:20

Arrhopalites kristiani from Ukraine
After Vargovich RS, 2005 Fig.1

Species lacuna (Christiansen, K & Bellinger, P, 1996:174) Vargovitsh, RS, 2009:e18

Species leonardwoodensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:3

Species lewisi (Christiansen, K & Bellinger, P, 1998:1249) Vargovitsh, RS, 2009:e18

Species longicornis (Cassagnau, P & Delamare Deboutteville, C, 1953:141) Vargovitsh, RS, 2009:20

Species madonnensis (Zeppelini, D & Christiansen, K, 2003:41) Vargovitsh, RS, 2009:e18

Species maestrazgoensis Jordana, R, Fadrique, F et Baquero, E, 2012:52

Species marshalli (Christiansen, K & Bellinger, P, 1996:175) Vargovitsh, RS, 2009:e18

Species mauli (Delamare Deboutteville & Bassot, 1957) Vargovitsh, RS, 2009:e18

Species miravetensis (Baquero, E, Herrando-Pérez, S & Jordana, R, 2005:82) Vargovitsh, RS, 2009:20

Species nanjingensis (Lin & Chen, 1997) Vargovitsh, RS, 2009:20

Species nigripes (Park & Kang, 2007) Vargovitsh, RS, 2009:20

Species obtusus (Zeppelini, D & Christiansen, K, 2003:40) Vargovitsh, RS, 2009:e18

Species octacanthus (Yosii, R, 1970:14) Vargovitsh, RS, 2009:20

Species ornatus (Stach, J, 1945:24) Vargovitsh, RS, 2009:20

Species pavo (Christiansen, K & Bellinger, P, 1996:176) Vargovitsh, RS, 2009:e18

Species perezi Arbea, JI, 2013:37

Pygmarrhopalites perezi from Spain
After Arbea JI, 2013 Fig.16

Species plectrifer (Hüther, 1963) Vargovitsh, RS, 2009:20

Species plethorasari Zeppelini, D, Taylor, SJ et Slay, ME, 2009:6

Species postumicoides (Cassagnau, P & Delamare Deboutteville, C, 1953:137) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites postumicoïdes(sic) Cassagnau, P et Delamare Deboutteville, C, 1953:142 ISS

Species postumicus (Stach, J, 1945:25) Vargovitsh, RS, 2009:20

Species principalis (Stach, J, 1945:37) Vargovitsh, RS, 2009:9, B

            Syn.: Arrhopalites binoculatus Linnaniemi, 1912, nec Börner, 1901
            Syn.: Arrhopalites binoculatus f. principalis Linnaniemi, 1912, nec Börner, 1901
            Syn.: Sminthurinus binoculatus ab. ? Börner, 1903
            Syn.: Arrhopalites binoculatus Womersley, 1939, nec Börner, 1901


Arrhopalites principalis from the UK
Antenna
2005 © Hopkin, S.P.
Pygmarrhopalites principalis from Belgium
2011.09.06 © Henderickx, H.
Pygmarrhopalites principalis from Belgium
2011.09.06 © Henderickx, H.
Pygmarrhopalites principalis from the UK
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
4th antennal segment with 6 subdivisions
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
Anal segment
2012.11.22 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from the UK
2013.01.31 © Murray, A.
Pygmarrhopalites principalis from France
2013.06.02 © Alonso, C.
Pygmarrhopalites principalis from France
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
Ant.3 at basis enlarged (*)
2018.01.07 © Tiky, S.
Pygmarrhopalites principalis from France
2018.02.17 © Picard, J.
Pygmarrhopalites principalis from Denmark
2019.01.22 © Schlütter, P.

  Subspecies grisea Axelson, 1907

  Subspecies pallida Linnaniemi, WM, 1912:274

  Subspecies skelicus Vargovitsh, RS, 2009:36

Species pseudoappendices (Rusek, 1967:23) Vargovitsh, RS, 2009:e18

Species pseudoprincipalis Vargovitsh, RS, 2009:31

Species pygmaeus (Wankel, 1860:256 tt) Vargovitsh, RS, 2009:8, B

            Syn.: Smynthurus(sic) ferrugineus Packard, 1888
             Syn.: Arrhopalites ferruginaeus(sic) Christiansen, K, 1966:43 ISS
            Syn.: Smynthurus(sic) mammouthia Banks, 1897
             Syn.: Arrhopalites? mamouthia(sic) Bonet, F et Tellez, C, 1947:193 ISS
            Syn.: Sminthurinus affinis Börner, 1901
            Syn.: Sminthurinus binocolatus(sic) Börner, 1901 IOS
            Syn.: Sminthurinus (Articulati) binoculatus Börner, C, 1901:100 n
             Syn.: Arrhopalites binocolatus(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
            Syn.: Arrhopalites binoculatus var. cavernarum Bonet, 1928
            Syn.: Arrhopalites subboneti Cassagnau & Delamare Deboutteville, 1953
            Syn.: Arrhopalites sp. Stach, 1956
            Syn.: Sphyrotheca binoculata Scott, 1964, nec Börner, 1901


             Syn.: Dicyrtoma pigmaea(sic) Absolon, K, 1899:317 ISS
             Syn.: Arrhopalites pygmaea(sic) Bonet, 1931 ISS
             Syn.: Arrhopalites pygmaeeus(sic) Selga, D, 1963:454,B ISS
             Syn.: Arrhopalites pygmaeu(sic) Selga, D, 1963:455,B ISS
             Syn.: Arrhopalites pygmeus(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Arrhopalites pygmeus(sic) Skarzynski, D et Smolis, A, 2002:304 ISS


Arrhopalites pygmaeus from the UK
Antenna
2005 © Hopkin, S.P.
Arrhopalites pygmaeus (bar=2µm)
Aberrant epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.2D'
Arrhopalites pygmaeus (bar=500nm)
Epicuticular secondary granule
After Nickerl J & al, 2012 Fig.2D''
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
 
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
 
Pygmarrhopalites pygmaeus from the UK
2012.09.06 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
2012.09.07 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
2013.01.29 © Murray, A.
Pygmarrhopalites pygmaeus from the UK
Subdivisions of 4th antennal segment
2013.01.29 © Murray, A.
Pygmarrhopalites pygmaeus from Canada
2015.07.16 © Murray, A.
Pygmarrhopalites pygmaeus from Canada
2015.07.16 © Murray, A.
Pygmarrhopalites pygmaeus gr. from France
Ant.4 with 5 subdivisions
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
Spiky cephalic setae (*)
2018.05.02 © Picard, J.
Pygmarrhopalites pygmaeus gr. from France
Unguiculus (u) with long filament reaching beyond tip of unguis (U)
2018.05.02 © Picard, J.



Species ruseki (Nosek, J, 1975:45) Vargovitsh, RS, 2009:20

Species sacer (Christiansen, K & Bellinger, P, 1996:177) Vargovitsh, RS, 2009:26

Species salemensis Soto-Adames, FN et Taylor, SJ, 2013:167

             Syn.: Pygmarrhopalites salemsis(sic) Soto-Adames, FN et Taylor, SJ, 2013:174 ISS

Species sapo (Zeppelini, D & Christiansen, K, 2003:38) Vargovitsh, RS, 2009:e18

Species secundarius (Gisin, 1958) Vargovitsh, RS, 2009:36

Species sericus (Gisin, 1947) Vargovitsh, RS, 2009:e18, B

Species sextus (Zeppelini, D & Christiansen, K, 2003:40) Vargovitsh, RS, 2009:26

Species shoshoneiensis Zeppelini, D, Taylor, SJ et Slay, ME, 2009:11

Species silvus (Christiansen, K & Bellinger, P, 1996:177) Vargovitsh, RS, 2009:e18

Species slovacicus (Nosek, 1975) Vargovitsh, RS, 2009:20

Species spinosus (Rusek, 1967:23) Vargovitsh, RS, 2009:20

Species styriacus (Nosek & Neuherz, 1976) Vargovitsh, RS, 2009:20

Species subbifidus (Travé, J, Gadea, E & Delamare Deboutteville, C, 1954:213) Vargovitsh, RS, 2009:e18

Species subboneti (Cassagnau, P & Delamare Deboutteville, C, 1953:135) Vargovitsh, RS, 2009:e18

Species tauricus Vargovitsh, RS, 2009:20

Species terricola (Gisin, 1958) Vargovitsh, RS, 2009:e18

Species texensis (Christiansen, K & Bellinger, P, 1996:178) Vargovitsh, RS, 2009:e18

             Syn.: Arrhopalites texesensis(sic) Zeppelini Filho, D, [1997]:12 ISS

Species thermophilus (Loksa, I, 1964:96) Vargovitsh, RS, 2009:il

Species troglophilus (Palissa, 2000) Vargovitsh, RS, 2009:e19

Species uenoi (Yosii, R, 1956:100) Vargovitsh, RS, 2009:26

Species whitesidei (Jacot, 1938) Vargovitsh, RS, 2009:e19

Species yosiii (Zeppelini, D, 2004:8) Vargovitsh, RS, 2009:20

            Syn.: Arrhopalites yosii(sic) Zeppelini, D, 2004:8 IOS

Species youngsteadtorum Zeppelini, D, Taylor, SJ et Slay, ME, 2009:8

            Syn.: Pygmarrhopalites (pygmaeus-group ss) youngsteadtii(sic) Zeppelini, D, Taylor, SJ et Slay, ME, 2009:8 IOS

Species zlotensis Curcic, BPM et Lucic, LR, 1997:31P

            Syn.: Arrhopalites zloti(sic) Curcic, BPM et Lucic, LR, 1997:31P IOS
Arrhopalites zloti(sic) from East Serbia
After Curcic BPM & Lucic LR, 1997 Fig.1
Arrhopalites zloti(sic) from East Serbia
After Curcic BPM & Lucic LR, 1997 Fig.2

             Syn.: Arrhopalites zloti(sic) Jaksic, P, 2005:194 ISS





Genus Troglopalites Vargovitsh, RS, 2012:10

Species stygios Vargovitsh, RS, 2012:11, t.t.

Troglopalites stygios from Western Caucasus
After Vargotitsh RS, 2012 Fig.60
Troglopalites stygios from Western Caucasus
After Vargotitsh RS, 2012 Fig.61

References