http://www.collembola.org/taxa/bouridae.htm - Last updated on 2017.06.29 by Frans Janssens
Checklist of the Collembola: Bourletiellidae

Familia Bourletiellidae Börner, 1912, sensu Bretfeld, 1994
Genus Number
of species
A1
Acutoempodialis1
Adisianus3
Anarmatus5
Aneuempodialis2
Anjavidiella5
Arlesminthurus4
B1
Bourletides1
Bourletiella31
Bourletiellitas1
Bovicornia7
Cassagnaudiella8
Corynephoria7
Cyprania2
Deuterosminthurus26
Diksamella1
Ellisiella1
Fasciosminthurus28
Heterosminthurus26
Karakumiella1
Kaszabellina3
Koupelia1
Madecassiella1
Massoudia1
Nasosminthurus4
Navarrella1
Orenius2
Parabourletiella1
Paulianitas1
Prorastriopes31
Pseudobourletiella1
Rastriopes15
Sanaaiella5
Stenognathriopes (Stenognathriopes)6
Stenognathriopes (Tenentiella)2
Stenognathriopes8
Tritosminthurus1
Vatomadiella3
Bourletiellidae241
Genera with number of species
Deuterosminthurus sp. from France
Courtship dance
2010.06.04 © Elpachato.
Bourletiella sp. from Australia
2012.07.03 © Hort, J.

  Syn.: ... Smynthuridae(sic) Lubbock, 1862, partim Lubbock, 1870
  Syn.: Sminthuridæ(sic) Smynthurus(sic) Packard, 1873 partim
  Syn.: Collembola Boerletiellidae(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:111 ISS

Genus A Bretfeld, G, 2005:50
Species spec Bretfeld, G, 2005:50
Gen. spec. A from Yemen
After Bretfeld G, 2005 Fig.101


Genus B Bretfeld, G, 2005:53
Species spec Bretfeld, G, 2005:53

Subfamilia Parabourletiellinae Betsch J-M, 1974:181

Genus Bourletiellitas Betsch, J-M, 1974:554
Species imerinensis Betsch, J-M, 1974:547, t.t.

Genus Anjavidiella Betsch, J-M, 1974:560
Species andoharianensis Betsch, J-M, 1980:182
Species andringitrensis Betsch, J-M, 1980:182
Species anjavidilavensis Betsch, J-M, 1980:182
Species ankaratrensis Betsch, J-M, 1974:554, t.t.
Species vohidrayensis Betsch, J-M, 1980:184

Genus Vatomadiella Betsch, J-M, 1974:567
Species descarpentriesi Betsch, J-M, 1980:185
Species pauliani Betsch, J-M, 1974:560, t.t.
Species peyrierasi Betsch, J-M, 1980:184

Genus Parabourletiella Betsch, J-M, 1975:477
Species mahafalensis Betsch, J-M, 1975:478, t.t.

Subfamilia Bourletiellinae Börner, 1912, sensu Betsch J-M, 1974:181
Bourletiellinae from Hungary
2008.06.07 © Keresztes, G.


Genus Acutoempodialis Bretfeld, 2000
Species lineatus Bretfeld, 2000, t.t.

Genus Adisianus Bretfeld, G, 2003:551
Species fuscus (Bretfeld, 2002) Bretfeld, G, 2003:551
Species maassius (Palacios-Vargas, JG & González, V, 1995:290) Bretfeld, G, 2003:551
Deuterosminthurus maassius from Cuba
After Palacios-Vargas JG & González V, 1995 Fig.17
Deuterosminthurus maassius from Cuba
After Palacios-Vargas JG & González V, 1995 Fig.18

Species maculatus (Bretfeld, 2002) Bretfeld, G, 2003:551, t.t.

Genus Anarmatus Bretfeld, 2000 key
Species diksamanus Bretfeld, G, 2005:2
Anarmatus diksamanus from Yemen
After Bretfeld G, 2005 Fig.1

Species goeehensis Bretfeld, G, 2005:4
Anarmatus goeehensis from Yemen
After Bretfeld G, 2005 Fig.5

Species irregularius Bretfeld, G, 2005:6
Anarmatus irregularius from Yemen
After Bretfeld G, 2005 Fig.9

Species longisetus Bretfeld, G, 2005:8
Species taizzus Bretfeld, 2000, t.t.

Genus Sanaaiella Bretfeld, 2000
Species circumscripta Bretfeld, 2000
Species dorsopallida Bretfeld, 2000
Species multisensillata Bretfeld, 2000
Species quadripunctata Bretfeld, 2000
Species septemlineata Bretfeld, G, 2005:19
Sanaaiella septemlineata from Yemen
After Bretfeld G, 2005 Fig.34


Genus Rastriopes Börner, C, 1906:182
Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2014.08.11 © Zosterops, K.
Rastriopes sp. from Tasmania
2016.04.14 © Zosterops, K.

Species sp.1 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species aculeatus (Börner, C, 1906:182) Stach, 1956, t.t.
Species amphigyus Börner, 1906
Species deformis (Nicolet, H, 1847:359) Stach, J, 1956:238, i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species dromedarius (Womersley, H, 1932:28) Greenslade, P, 1974:259
Rastriopes nr dromedarius from Australia
2015.11.09 © Murray, A.

Species echidnus Yoshii, R et Suhardjono, YR, 1992:92
Species fulvipes (Nicolet, H, 1847:359) Stach, J, 1956:238, i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species fuscus Yosii, 1960
Species gisinianus Murphy, 1960
Species handschini (Womersley, H, 1932:28) Greenslade, P in Houston, WWK, 1994:129
            Syn.: Deuterosminthurus (Rastriopes) handschini var. atratus Womersley, H, 1932:28
Rastriopes handschini from Australia
2012.07.15 © Hort, J.
Rastriopes handschini from Australia
2016.11.20 © MacAulay, I.
Rastriopes handschini from Australia
2016.11.20 © MacAulay, I.

Species lineatus Womersley, 1931
            Syn.: Rastriopes lineata(sic) Womersley, 1931 IOS
            Syn.: Rastriopes schultzei Paclt, 1959
             Syn.: Rastriopes lineata(sic) Salmon, JT, 1964:610 ISS
             Syn.: Rastriopes lineata(sic) Janion-Scheepers, C, Deharveng, L, Bedos, A et Chown, L, 2015:73 ISS
Species obscurus (Womersley, H, 1931:7) Greenslade, P, 1977:247
Deuterosminthurus obscurus from Tasmania
After Womersley, 1931 Pl.I Fig.4

            Syn.: Deuterosminthurus (Deuterosminthurus) obscurus var. maculatus Womersley, H, 1939:322
Species schultzei Börner, C in Schultze, 1908:61
Species tricuspidatus (Börner, 1907) Mari Mutt & Bellinger, 1990:151
            Syn.: Sminthurus pallipes Parona, 1895
Species vexillarius (Schäffer, C, 1898:422) Stach, 1956

Genus Deuterosminthurus Börner, 1901 nc Opinion 1042, 1976 key
Remark by ICZN. 2007:241: Placed on the Official List of Names in Zoology. Opinion 1042.

Deuterosminthurus sp. from France
2010.05.24 © Elpachato.
Deuterosminthurus sp. from France
2010.05.24 © Elpachato.
Deuterosminthurus sp. from France
2010.05.24 © Elpachato.
Deuterosminthurus sp. from France
2010.06.04 © Elpachato.
Caudal view
Deuterosminthurus sp. from France
2010.06.16 © Elpachato.
Deuterosminthurus sp. juv. from Ibiza
2014.01.09 © Beer, R.
Deuterosminthurus sp. from Ibiza
2014.02.03 © Beer, R.

Species bellingeri Nayrolles, 1997
            Syn.: Deuterosminthurus maculatus Nayrolles, 1996, nec Womersley, 1939
Deuterosminthurus bellingeri from the UK
2009.07.10 © Ardron, P.A.
Deuterosminthurus bellingeri ♂ from the UK
2009.07.10 © Ardron, P.A.

Species bicinctus (Koch in Herrich-Schäffer, 1840) Linnaniemi, 1912, t.t., B
            Syn.: ... pallipes Bourlet
            Syn.: Sminthurus pallipes Tullberg, 1871
            Syn.: Deuterosminthurus bicinctus var. pallipes Linnaniemi, 1912
            Syn.: Smynthurus(sic) luteus Lubbock, J, 1870:281 partim
            Syn.: Sminthurus luteus Tullberg, 1872
            Syn.: Sminthurus cinctus Tullberg, 1871
            Syn.: Sminthurus lupulinae Lubbock, 1873
            Syn.: Sminthurus speciosus Krausbauer, 1898
            Syn.: Sminthurus novemlineatus Börner, 1900, partim
            Syn.: Sminthurus (Deuterosminthurus) oblongus ssp. minor Börner, C, 1901:110
            Syn.: Sminthurus (Deuterosminthurus) oblongus ssp. minor var. puncteola Börner, C, 1901:112
Sminthurus oblongus var. puncteola
from Germany
After Börner C, 1901 Tafel I Fig.5
Deuterosminthurus bicinctus var. puncteola
from Poland
After Stach J, 1956 Plate XX Fig.3
Deuterosminthurus bicinctus from Poland
Courtship with female var. puncteola
2002 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus var. puncteola juv. from the UK
2012.05.14 © Nurcombe, E.
Deuterosminthurus bicinctus var. puncteola ♀ from the UK
2012.06.20 © Nurcombe, E.
Deuterosminthurus bicinctus from the UK
Courtship with female var. puncteola
2012.06.20 © Nurcombe, E.

            Syn.: Burletiella(sic) sulphurea Börner, 1908
            Syn.: ... sulphureus Collinge & Shoebotham, 1910
            Syn.: Deuterosminthurus bicinctus f. repandus Womersley, H, 1931:6
Deuterosminthurus bicinctus f. repandus from Tasmania
After Womersley, 1931 Pl.I Fig.2

             Syn.: Deuterosminthurus bicinctus f. repanda(sic) Womersley, H, 1931:6 ISS
Deuterosminthurus bicinctus
subsp. repandus from New Zealand
After Salmon JT, 1941 Pl.72 Fig.520

            Syn.: Bourletiella (Deuterosminthurus) flava Gisin, 1946
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, 1970 ISS
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, 1996 ISS
             Syn.: Deuterosminthurus bicinctus f. flava(sic) Bretfeld, G, 1999:236 ISS
Deuterosminthurus bicinctus from the USA
Deuterosminthurus bicinctus f. flavus ♀ & ♂
Deuterosminthurus bicinctus ♂
2007.06.04 © Reed, D.
Deuterosminthurus bicinctus ♂ & f. flavus ♀ from the UK
2010.05.22 © Michaelson, J.

             Syn.: Deuterosminthurus bicinctus var. flava(sic) Berg, M et den Ouden, A, 2013:5 ISS
Deuterosminthurus bicinctus from Russia
After Potapov M. in Babenko A.B., 1988
Deuterosminthurus bicinctus from Poland
2002 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus
2005 © Kozlowski, M.W. & Aoxiang, S.
Deuterosminthurus bicinctus
'Premating-dance'
2005 © Kozlowski MW & Aoxiang S.
Deuterosminthurus bicinctus
Sperm deposition and take-up
2005 © Kozlowski MW & Aoxiang S.
Deuterosminthurus bicinctus ♂ from the USA
2007.06.02 © Reed, D.
 
 
Deuterosminthurus bicinctus ♂ from the USA
2007.06.04 © Reed, D.
 
Deuterosminthurus bicinctus from the USA
2009.05.20 © Loser, I.
Deuterosminthurus bicinctus ♂ from the UK
2012.06.20 © Nurcombe, E.
 
Deuterosminthurus bicinctus juv. from the UK
2012.06.26 © Murray, A.
 
Deuterosminthurus bicinctus ♀ from the UK
2012.06.27 © Murray, A.
Deuterosminthurus bicinctus from the USA
2013.07.04 © Eiseman, C.
Deuterosminthurus bicinctus from the USA
Ant.4 with 7 subsegments
2013.07.04 © Eiseman, C.
Deuterosminthurus bicinctus ♂ & ♀ from Germany
Courtship
2014.06.20 © Beck, K.

Species bisetosus Baquero, E, Moraza, ML et Jordana, R, 2003:9
Deuterosminthurus bisetosus from Spain
After Baquero E, Moraza ML & Jordana R, 2003 Fig.29
Deuterosminthurus bisetosus from Spain
Generalised colouration
After Baquero E, Moraza ML & Jordana R, 2003 Fig.37
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.36
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.39
Deuterosminthurus bisetosus from Spain
Pale specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.42
Deuterosminthurus bisetosus from Spain
Orange-red specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.38
Deuterosminthurus bisetosus from Spain
Red coloured specimen
After Baquero E, Moraza ML & Jordana R, 2003 Fig.40
Deuterosminthurus bisetosus from Spain
Dark pattern
After Baquero E, Moraza ML & Jordana R, 2003 Fig.41

Species caeruleacaudus (Scott, 1965)
            Syn.: Bourletiella (Deuterosminthurus) sp. Bellinger, PF, 1985:78
Species delatorrei Palacios-Vargas, JG et González, V, 1995:287
Deuterosminthurus delatorrei from Mexico
After Palacios-Vargas JG & González V, 1995 Fig.1
Deuterosminthurus delatorrei from Mexico
After Palacios-Vargas JG & González V, 1995 Fig.2
Deuterosminthurus delatorrei from Peru
Cephalic branching trachea
2016.06.09 © Palacios-Vargas, J.G.

Species doriae (Parona, 1884) Stach, J, 1956:238, i.s.
            Syn.: Smynthurus(sic) Doriæ(sic) Parona, 1884 IOS
             Syn.: Sminthurus Doriæ(sic) Schött, H, 1893:4 ISS
Species ezoensis Yosii, R, 1972:95
Species fenyesi Stach, 1926
            Syn.: Deuterosminthurus beckeri Coineau & Delamare Deboutteville, 1961
            Syn.: Deuterosminthurus pallipes ssp. fenyesi f. pallida Ellis, WN, 1974:142
Deuterosminthurus fenyesi f. pallida from Greece
2008.01.14 © Markopoulos, S.

             Syn.: Deuterosminthurus Fényesi(sic) Stach, J, 1930:271 ISS
Species kugleri Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species luridus (Snider, 1978) Palacios-Vargas & González, 1995:293
             Syn.: Deuterosminthurus lurida(sic) Palacios-Vargas & González, 1995 ISS
             Syn.: Deuterosminthurus lurida(sic) Vázquez, M et Palacios-Vargas, JG, 2004:105 ISS
             Syn.: Deuterosminthurus lurida(sic) Kang, J-S, Lee, B-H et Park, K-H, 2005:238 ISS
             Syn.: Deuterosminthurus lurida(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:111 ISS
Species mediterraneus Ellis, 1974
Species nonfasciatus (Snider, 1978) Palacios-Vargas, JG & González, V, 1995:293
Deuterosminthurus nonfasciatus from Canada
2007.05.14 © Pilon, C.
Deuterosminthurus nonfasciatus from Canada
2007.05.14 © Pilon, C.
Deuterosminthurus nonfasciatus from the USA
'Mating dance'
2007.05.25 © Reed, D.
Deuterosminthurus nonfasciatus from the USA
Male and female specimen
2007.05.25 © Reed, D.
Deuterosminthurus nonfasciatus from Canada
Female and male specimen
2007.05.27 © Marsman, I.
Deuterosminthurus nonfasciatus from the USA
Female and male specimen
2007.06.02 © Reed, D.
Deuterosminthurus nonfasciatus from the USA
2009.05.18 © Loser, I.
Deuterosminthurus nonfasciatus from Canada
2009.08.21 © Erbland, M.
Deuterosminthurus nonfasciatus from the USA
2012.04.07 © Kouri, J.

Species okinawanus Yosii, 1965
Species pallipes (Bourlet, 1843) Handschin, E in Dahl, F, 1929:117, B
            Syn.: Smynthurus(sic) echii Koch, 1840
            Syn.: Sminthurus ochraceus Koch, 1840
            Syn.: Smynthurus(sic) sulphureus Koch, 1840
            Syn.: Smynthurus(sic) lupulinae Bourlet, 1842
            Syn.: Sminthurus luteus Lubbock, 1867
            Syn.: Sminthurus atratus Schäffer, 1896
            Syn.: Sminthurus speciosus Krausbauer, 1898
            Syn.: Sminthurus repandus Ågren, 1903
             Syn.: Deuterosminthurus bicinctus f. repanda(sic) Handschin, E in Dahl, F, 1929:117 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 1999:239 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, 2000 ISS
Deuterosminthurus pallipes f. repandus from the UK
2001 © Hopkin, S.P.

             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, 2002 ISS
             Syn.: Deuterosminthurus repanda(sic) Baquero, E, Moraza, ML et Jordana, R, 2003:15 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895 ISS
Deuterosminthurus pallipes f. repandus from the UK
2005 © Hopkin, S.P.
Deuterosminthurus pallipes f. repandus from Belgium
2007.05.03 © Gielen, K.
Deuterosminthurus pallipes f. repandus from the UK
2008.07.11 © Barton, T.
Deuterosminthurus pallipes f. repandus from the UK
2008.07.13 © Michaelson, J.

             Syn.: Deuterosminthurus pallipes f. repanda(sic) Bretfeld, G, 2010:303 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Petersen, H, 2011:472 ISS
             Syn.: Deuterosminthurus pallipes f. repanda(sic) Petersen, H, 2011:476,cf ISS
Deuterosminthurus pallipes f. repandus from Ibiza
Grooming with everted vesicle of collophore
2011.02.11 © Beer, R.
Deuterosminthurus pallipes f. repandus from Ibiza
Mating ritual
2011.12.14 © Beer, R.
Deuterosminthurus pallipes f. repandus from the UK
2012.07.01 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus from the UK
2012.07.01 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
2012.07.01 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
2012.07.09 © Nurcombe, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
Subdivided 4th antennal segment
2013.02.21 © Phillips, E.
Deuterosminthurus pallipes f. repandus juv. from the UK
'Pearly' composed eye
2013.02.21 © Phillips, E.
Deuterosminthurus pallipes f. repandus from Russia
Predated by pseudoscorpion
2013.10.05 © Haritonoff.
Deuterosminthurus pallipes f. repandus ♀ & ♂ from Holland
2014.05.01 © van Duinen, J.
Deuterosminthurus pallipes f. repandus from Belgium
Ant.4 with 8 subdivisions
2016.07.09 © Huskens, M.-L.
Deuterosminthurus pallipes f. repandus from Belgium
♂ guides ♀ to spermatophore
2017.06.20 © Huskens, M.-L.


            Syn.: Bourletiella (Deuterosminthurus) mixta Gisin, 1946
Smynthurus(sic) pallipes from the UK
After Lubbock J, 1870 Pl.XXI Fig.13
Smynthurus(sic) pallipes from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.14
Smynthurus(sic) pallipes from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.15
Smynthurus(sic) pallipes from the UK
After Lubbock J., 1873 Pl.4.
Deuterosminthurus pallipes from the UK
2005 © Hopkin, S.P.
Deuterosminthurus pallipes from the UK
2006 © Barton, T.
Deuterosminthurus pallipes from the UK
2006 © Barton, T.
Deuterosminthurus pallipes ♂ from the UK
2007.04.28 © Barton, T.
Deuterosminthurus pallipes from the UK
2007.06.03 © Saul, R.
Deuterosminthurus pallipes from the UK
2007.06.20 © Barton, T.
Deuterosminthurus pallipes from the UK
2008.06.02 © Farrell, S.
Deuterosminthurus pallipes juv. from the UK
2009.04.24 © Valentine., B
Deuterosminthurus pallipes from the UK
2009.05.03 © Barton, T.
Deuterosminthurus pallipes from the UK
2009.05.03 © Barton, T.
Deuterosminthurus pallipes ♂ from the UK
2009.07.16 © Dawes, D.
Deuterosminthurus pallipes from the UK
Male with female forma repanda
2010.05.17 © Barton, T.
Deuterosminthurus pallipes from the UK
2010.05.17 © Barton, T.
Deuterosminthurus pallipes ♂ & ♀ from the UK
2012.05.12 © Murray, A.
Deuterosminthurus pallipes from the UK
2012.05.13 © Murray, A.
 
Deuterosminthurus pallipes from the UK
Pale colour form
2012.05.13 © Murray, A.
Deuterosminthurus pallipes ♀ & ♂ from the UK
2012.05.29 © Nurcombe, E.
Deuterosminthurus pallipes ♂ & ♀ from the UK
2012.06.01 © Nurcombe, E.
Deuterosminthurus pallipes ♂ from the UK
Raised genital papilla
2012.06.01 © Nurcombe, E.
Deuterosminthurus pallipes ♂ f. repanda & ♀ from the UK
2012.06.15 © Nurcombe, E.
Deuterosminthurus pallipes from the UK
2013.07.11 © Nurcombe, E.
Deuterosminthurus pallipes ♀ from the UK
2014.04.26 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from the UK
Ant.4 with 8 subsegments
2014.04.26 © Henton-Jones, D.
Deuterosminthurus pallipes ♂ & ♀ from the UK
Courtship
2014.05.03 © Henton-Jones, D.
Deuterosminthurus pallipes ♀ from the UK
Eyepatch with 3 large inner ocelli
2014.05.03 © Henton-Jones, D.

Species pleuracanthus Nayrolles, 1996
Species polenus (Christiansen, K et Bellinger, P, 1992:360)
Species propallipes (Börner, C, 1901:107) Stach, 1956
Species prospathaceus (Rapoport in Delamare Deboutteville & Massoud, 1963) Mari Mutt & Bellinger, 1990, n.n.
Species quadrangulatus (Loksa, I & Bogojevic,J, 1970:140) Ellis, 1974
Bourletiella (Deuterosminthurus) quadrangulata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.61

Species russatus (Maynard, EA, 1951:258) Stach, J, 1956:160
Bourletiella (Deuterosminthurus) russata from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.406
             Syn.: Deuterosminthurus russata(sic) Snider, RJ, 1967:228 ISS
Species separatus Arlé, R, 1943:112
Species serratus Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:238
Deuterosminthurus serratus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.2A

Species sp. nov. González, V, Díaz, M et Palacios-Vargas, JG, 1997:48
Species sulphureus (Koch, 1840) Stach, 1956:159, B
            Syn.: Smynthurus(sic) lupulinae Bourlet, 1841
             Syn.: Smynthurus(sic) lupulinæ(sic) Lubbock, J, 1870:296 ISS
            Syn.: Smynthurus(sic) baulastinus Harvey, 1848
            Syn.: Smynthurus(sic) solani Curtis, 1860
            Syn.: Smynthurus(sic) luteus Lubbock, J, 1870:296
Smynthurus(sic) luteus from the UK
After Lubbock J, 1870 Pl.XXI Fig.4
Smynthurus(sic) luteus from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.5
Smynthurus(sic) luteus from the UK
Foot with 2 tenent hairs
After Lubbock J, 1870 Pl.XXI Fig.6
Smynthurus(sic) luteus from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.7
Smynthurus(sic) luteus from the UK
After Lubbock J, 1873 Pl.3

             Syn.: Sminthurus luteo(sic) Reuter, OM, 1876:84 ISS
             Syn.: Bourletiella (Bourletiella) luteus(sic) Kloet & Hincks, 1964 ISS
             Syn.: Bourletiella luteus(sic) Janssens, F, 1995:19,B ISS
            Syn.: Smynthurus(sic) vitis Lichtenstein, 1883
            Syn.: Sminthurus fulvus Lie-Pettersen, 1896
            Syn.: Sminthurus repandus Ågren, 1902
            Syn.: Corynephoria gibbera Salmon, JT, 1941:416
Corynephoria gibbera from New Zealand
After Salmon JT, 1941 Pl.72 Fig.521

             Syn.: Corynephoria gibbosa(sic) Stach, J, 1956:133 ISS
            Syn.: Deuterosminthurus Gisini(sic) Stach, 1956
Neozygites sp. from Denmark. Bar=10µm
Conidia infecting Deuterosminthurus sulphureus
After Dromph & al, 2001 Fig.5
Deuterosminthurus sulphureus from the UK
Annulated fourth antennal segment
2004 © Hopkin, S.P.

             Syn.: Deuterosminthurus sulphereus(sic) Berg, M et den Ouden, A, 2013:5 ISS
  Subspecies mediterraneus Ellis, 1974
Species tristani Denis, 1933
Species yumanensis Wray, 1967

Genus Karakumiella Bretfeld, G, 2010:310
Species kaplini (Martynova in Martynova & al, 1979 tt) Bretfeld, G, 2010:310

Genus Arlesminthurus Bretfeld, G, 1999:219
Species aueti (Arlé, R, 1961:37) Bretfeld, G, 1999:219
Species franzkafkai Palacios-Vargas, JG et Cabrera, M, 2015:63
Arlesminthurus franzkafkai ♀ from Nicaragua
After Palacios-Vargas JG & Cabrera M, 2015 Fig.1
Arlesminthurus franzkafkai ♂ from Nicaragua
After Palacios-Vargas JG & Cabrera M, 2015 Fig.2

Species richardsi (Arlé, 1971) Bretfeld, G, 1999:219, t.t.
Arlesminthurus nr richardsi ♂ from Costa Rica
2013.07.21 © Kouri, J.D.

Species salinensis (Arlé, 1971) Bretfeld, G, 1999:219

Genus Ellisiella Bretfeld, 1999
Species pandayi (Ellis, WN, 1974:143) Bretfeld, 1999, t.t.

Genus Corynephoria Absolon, 1907
Species absoloni Womersley, 1939
Corynephoria absoloni ♀ from Australia
With dorsal digitation
2015.11.09 © Murray, A.
Corynephoria absoloni ♂ from Australia
With dorsal digitation
2015.11.09 © Murray, A.
Corynephoria absoloni from Australia
2015.11.09 © Murray, A.

Species albidorsalis Womersley, H, 1932:36
Corynephoria albidorsalis from Australia
After Womersley H, 1932 Fig.14a
             Syn.: Corynephoria albodorsalis(sic) Stach, J, 1956:133 ISS
Species cassida Womersley, H, 1932:34
Corynephoria cassida from Australia
After Womersley H, 1932 Fig.12a

Species dubia Womersley, H, 1932:35
Species jacobsoni Absolon, 1907, t.t.
Species quadrimaculata Womersley, H, 1942:31
Corynephoria quadrimaculata from Australia
After Womersley H, 1942 Fig.8A

Species reticulata Salmon, 1963

Genus Bourletiella Banks, 1899, n.c. Opinion 291
  Syn.: Sminthuridæ(sic) Sminthurus Schött, H, 1891:7 partim
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
Remark by ICZN. 2007:112: Placed on the Official List of Names in Zoology. Opinion 291.

  Syn.: Bourletiellidae Borletiella(sic) Nickerl, J, Helbig, R, Schulz, H-J, Werner, C et Neinhuis, C, 2012:7 ISS
Bourletiella sp. from Australia
2015.11.22 © Steve & Alison.
Bourletiella sp. from Australia
2015.11.22 © Steve & Alison.

Species arvalis (Fitch, 1862) Scott, HG, 1963:107
            Syn.: Smynthurus(sic) luteus Lubbock, 1867
            Syn.: Smynthurus(sic) roseus Packard, AS, 1873:43
            Syn.: Bourletiella hortensis f. hispanica Denis, 1930
            Syn.: Deuterosminthurus neopandus Wray, 1950
            Syn.: Bourletiella (Bourletiella) savona Maynard, EA, 1951:249
             Syn.: Bourletiella repandus(sic) Scott, HG, 1963:107 ISS
Bourletiella arvalis from Tasmania
After Womersley, 1931 Pl.I Fig.5
Bourletiella arvalis from India
After Denis JR, 1936 Fig.49
Bourletiella arvalis from the USA
After Maynard, E.A, 1951 Plate XXIII Fig.407
Remark by Stach, J. 1956:147-148: In fig.567 given by Maynard, in, 1951, is the crest [of the male clasping organ] entirely wanting.

Bourletiella arvalis from the USA
2006 © McClarin, J.
Bourletiella arvalis from France
2007.04.16 © Lebeaux, P.
Bourletiella arvalis from Canada
2007.05.19 © Pilon, C.
Bourletiella arvalis from France
2007.06.18 © Lebeaux, P.
Bourletiella arvalis from Germany
2007.08.15 © Maier, T.
Bourletiella arvalis from the UK
2007.10.18 © Campbell, A.
Bourletiella arvalis from the USA
2008.05.11 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.16 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.18 © Thombs, D.
Bourletiella arvalis from the USA
2008.05.18 © Thombs, D.
Bourletiella arvalis from the USA
2008.06.02 © Thombs, D.
Bourletiella arvalis from the USA
Abdomen, lateral aspect
2008.06.16 © Thombs, D.
Bourletiella arvalis from the USA
2008.06.18 © Cowen, R.
 
Bourletiella arvalis from France
2008.10.06 © Lebeaux, P.
Bourletiella arvalis from France
2008.10.06 © Lebeaux, P.
Bourletiella arvalis from France
2008.10.19 © Lebeaux, P.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.04.12 © Justis, S.
Bourletiella arvalis from the USA
2009.06.15 © Thombs, D.
Bourletiella arvalis from Tasmania
2009.08.16 © Daley, A.
Bourletiella arvalis from the USA
2011.08.23 © Carr, J.
Bourletiella arvalis from the UK
2012.08.06 © Nurcombe, E.
Bourletiella arvalis from Holland
2012.09.04 © van Duinen, J.
Bourletiella arvalis from New Zealand
2013.12.28 © Sullivan, J.
Bourletiella arvalis from Germany
2014.05.05 © Beck, K.
Bourletiella arvalis from Germany
Ant.4 with 7 annulations
2014.05.06 © Beck, K.
Bourletiella arvalis from the UK
2014.08.22 © Nurcombe, E.

Species captis Baijal, 1969
Species christianseni Snider, 1978
Bourletiella (Bourletiella) christianseni from the USA
After Christiansen K & Bellinger P, 1981 Fig.963A

Species drachi Delamare Deboutteville & Massoud, 1963:263
            Syn.: Bourletiella Drachi(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species friedlaenderi (Börner, 1903) Stach, 1956:141
Species gibbonsi Snider, RJ, 1981:93
Bourletiella (Bourletiella) gibbonsi from the USA
After Snider RJ, 1981 Fig.1
Bourletiella (Bourletiella) gibbonsi from the USA
After Snider RJ, 1981 Fig.2

Species guevarai Rapoport, 1970
Species hortensis (Fitch, 1862) Brown, 1918, t.t., B
            Syn.: Podura signata Fabricius, 1783
             Syn.: Smynthurus(sic) Signata(sic) Templeton, 1835 ISS
            Syn.: Smynthurus(sic) sp. Harris, 1844
            Syn.: Smynthurus(sic) fuliginosus Nicolet, 1847
             Syn.: Smynthurella fuliginosus(sic) Houlbert, C, 1924:159 ISS
            Syn.: Smynthurus(sic) solani Curtis, 1860
            Syn.: Smynthurus(sic) apicalis Fitch, 1863
            Syn.: Smynthurus(sic) basalis Fitch, 1863
            Syn.: Smynthurus(sic) dorsalis Fitch, 1863
            Syn.: Smynthurus(sic) juvenilis Fitch, 1863
            Syn.: Smynthurus(sic) maturus Fitch, 1863
            Syn.: Smynthurus(sic) cucumeris Harris, 1869
            Syn.: Sminthurus pruinosus Tullberg, 1871
Sminthurus pruinosus from Sweden
After Schött H, 1893 Taf.II Fig.13

            Syn.: Bourletiella pruinosa Folsom, 1899
Sminthurus pruinosus from Europe
After Folsom in Escherich K, 1923 Abb.1b

            Syn.: Sminthurus pruinosus Matsumura & Ishida, 1931
            Syn.: Bourletiella pruinosa Kinoshita, 1932
            Syn.: Bourletiella pruinosa Uchida, 1940
            Syn.: Bourletiella pruinosa Kinoshita & Uchida, 1950
            Syn.: Sminthurus pruinosus Yosii, R, 1977:164
            Syn.: Bourletiella pruinosa Yosii, R, 1977:164
            Syn.: Smynthurus(sic) luteus Lubbock, 1873
            Syn.: Smynthurus(sic) 4-signatus(sic) Packard, AS, 1873:44 IOS
            Syn.: Sminthurus lineatus Reuter, OM, 1876:84
            Syn.: Smynthurus frontalis Uzel, 1890
            Syn.: Smynthurus(sic) albamaculatus Harvey, 1896
            Syn.: Sminthurus atratus Schäffer, 1896
            Syn.: Sminthurus pruinosus var. sabulosa Skorikov, 1899
            Syn.: Sminthurus pruinosus var. stscherbakowi Skorikov, 1899
            Syn.: Sminthurus pruinosus var. viridula Skorikov, 1899
            Syn.: Bourletiella hortensis f. Linnaniemi(sic) Denis, 1930
            Syn.: Bourletiella hortensis f. Folsomi(sic) Denis, 1930
            Syn.: Bourletiella insula Folsom, JW, 1932:73
            Syn.: Bourletiella viridis Greenslade, 2002
Sminthurus hortensis from Japan
After Folsom JW, 1899 Pl.3 Fig.37
Bourletiella hortensis from Tasmania
After Womersley, 1931 Pl.I Fig.6
Bourletiella hortensis from Australia
After Greenslade P & Ireson JE, 1986 Fig.9
Bourletiella hortensis from Sweden
2005 © Hall, K.
Bourletiella hortensis from the USA
2006 © McClarin, J.
Bourletiella hortensis from Canada
2007.05.03 © Pilon, C.
Bourletiella hortensis from Canada
2007.05.04 © Pilon, C.
Bourletiella hortensis from the USA
2007.05.06 © Schimming, L.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from the USA
2007.05.10 © Murray, T.
Bourletiella hortensis from France
2007.05.14 © Lebeaux, P.
Bourletiella hortensis from the USA
2008.04.23 © Bradford, A.
Bourletiella hortensis from Hungary
2008.06.21 © Keresztes, G.
Bourletiella hortensis from Antarctica
2009.03.20 © Key, R.S.
Det. Greenslade P.
Bourletiella hortensis from Antarctica
2009.03.20 © Key, R.S.
Det. Greenslade P.
Bourletiella hortensis from the USA
2009.05.06 © Murray, T.
Bourletiella hortensis from the USA
2009.05.18 © Murray, T.
Bourletiella hortensis from the USA
2009.05.18 © Murray, T.
Bourletiella hortensis from Belgium
2009.06.18 © Hermans, R.
Bourletiella hortensis from the UK
2010.06.26 © Michaelson, J.
Bourletiella hortensis from the UK
Subdivided 4th antennal segment
2010.07.26 © Robertson, A.
Bourletiella hortensis from the UK
Footcomplex with clavate tenent hairs
2010.07.26 © Robertson, A.
Bourletiella hortensis from France
2011.08.29 © Lebeaux, P.
 
Borletiella(sic) hortensis (bar=2µm)
Rhombic epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.6E
Bourletiella hortensis from the UK
2012.05.28 © Jones, A.
Bourletiella hortensis from the UK
Pale colour form
2012.06.21 © Murray, A.
Bourletiella hortensis juv. from the UK
2012.06.24 © Nurcombe, E.
 
Bourletiella hortensis from the UK
With parasitic larva of Acari:Erythraeoidea
2012.06.27 © Nurcombe, E.
Bourletiella hortensis from the UK
With parasitic larva of Acari:Erythraeoidea
2012.06.27 © Nurcombe, E.
Bourletiella hortensis from Belgium
Mucro spoonshaped
2012.10.29 © Vannieuwenhuyse., B
Bourletiella hortensis from the UK
Grooming using drop of saliva on front claw and everted tube-like vesicle of collophore
2013.05.25 © Nurcombe, E.
Bourletiella hortensis from the UK
2014.05.04 © Evans, M.

Species insula Folsom, JW, 1932:73
Species juanitae Maynard, EA, 1951:249
Bourletiella juanitae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.387
Bourletiella juanitae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.388

            Syn.: Bourletiella (Deuterosminthurus) fallonae Maynard, EA, 1951:255
Bourletiella fallonae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.404
Bourletiella fallonae from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.405

             Syn.: Bourletiella jaunitae(sic) Richards, WR, 1968:33 ISS
Bourletiella juanitae ♀ from the USA
2007.08.30 © Murray T.
Bourletiella juanitae ♀ from the USA
2007.08.30 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.
Bourletiella juanitae ♂ from the USA
2007.09.05 © Murray T.

Species khenifrana Bretfeld, 2001
Species melanocephala Dallai, 1970
Species millsi Pedigo, 1968
Species pisifera Fjellberg, 1984
Species pistilla Gisin, 1946
            Syn.: Bourletiella pistillum(sic) Gisin, 1946 IOS
            Syn.: Bourletiella pruinosa Ågren, 1903
            Syn.: Bourletiella agreni Stach, J, 1956:140
            Syn.: Bourletiella (Deuterosminthurus) cruciata Haybach, 1972
            Syn.: Bourletiella mongolica Betsch, J-M, 1977:77
            Syn.: Bourletiella pistillum(sic) ssp. incisa Ellis, WN, 1975:77
             Syn.: Bourletiella pistillum(sic) Stach, J, 1956:140 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) Gisin, H, 1960:286 ISS
             Syn.: Bourletiella pistillum(sic) Ellis, WN, 1975:76 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) Ellis, WN, 1975:78 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) ssp. pistillum(sic) Ellis, WN, 1975:79 ISS
             Syn.: Bourletiella pistillum(sic) ssp. pistillum(sic) Ellis, WN, 1975:81 ISS
             Syn.: Bourletiella (Cassagnaudiella) pistillum(sic) Klaver, E, 1975:185 ISS
             Syn.: Bourletiella pistillum(sic) Rusek, J in Dlabola, J, 1977:16 ISS
             Syn.: Bourletiella pistillum(sic) Fjellberg, A, 1984:87 ISS
             Syn.: Bourletiella pistillum(sic) Nayrolles, P, 1993:661 ISS
             Syn.: Bourletiella pistillum(sic) Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:14 ISS
             Syn.: Bourletiella pistillum(sic) Nayrolles, P, 1995:47 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) ssp. pistillum(sic) Bretfeld, G, 1999:226 ISS
             Syn.: Bourletiella pistillum(sic) Auclerc, A, Ponge, JF, Barot, S et Dubs, F, 2009:1602 ISS
             Syn.: Bourletiella pistillum(sic) Dunger, W et Voigtländer, K, 2009:18 ISS
             Syn.: Bourletiella (Bourletiella) pistillum(sic) Schulz, H-J, 2015:17 ISS
Species pluslineata (Stach, 1922) Gisin, 1960:289, s.d.
Species radula Gisin, 1946
            Syn.: Bourletiella rustica Cassagnau, 1968 lapsus
Species rapoporti Delamare Deboutteville & Massoud, 1963:259
            Syn.: Bourletiella Rapoporti(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species rustica Maynard, EA, 1951:248
Bourletiella rustica from the USA
After Maynard EA PlateXXII Fig.373
Bourletiella rustica from the USA
After Maynard EA PlateXXX Fig.568
Bourletiella rustica from the USA
After Maynard EA PlateXXX Fig.569
Bourletiella rustica from the USA
2006 © Gross, J.
Bourletiella rustica from the USA
2008.08.20 © Montgomery, G.
Bourletiella nr rustica from Argentina
2009.06.11 © Gonzalez-Carducci, E.
Bourletiella rustica from the USA
2009.06.25 © Melder, R.
Bourletiella rustica from the USA
2010.08.16 © Zungri, D.
Bourletiella rustica from the USA
2010.08.18 © Zungri, D.
Bourletiella rustica from the USA
2010.08.21 © Zungri, D.
Bourletiella rustica from the USA
2010.08.21 © Zungri, D.
Bourletiella rustica from the USA
2011.06.18 © Schneider, J.
Bourletiella rustica from the USA
Subdivided 4th antennal segment
2011.06.18 © Schneider, J.
Bourletiella rustica from the USA
2013.06.01 © Heo, C.-C.
Bourletiella rustica from the USA
2013.06.01 © Heo, C.-C.
Bourletiella rustica juv. from the USA
2014.05.29 © Dankowicz, E.
Bourletiella rustica from the USA
2014.08.24 © Dankowicz, E.
Bourletiella rustica from the USA
2015.08.21 © Rosenfeld, J.
Bourletiella rustica from the USA
2015.11.24 © Snyder, S.L.
Bourletiella rustica from the USA
2015.11.24 © Snyder, S.L.
Bourletiella rustica from the USA
2016.08.07 © Bailey, J.
Bourletiella rustica from the USA
2016.08.07 © Bailey, J.
Bourletiella rustica from the USA
2016.12.13 © Snyder, S.L.

Species savona Maynard, EA, 1951:249
Bourletiella savona from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.384.

Species sicki Bretfeld, 1994
Species sp. Maes & Palacios-Vargas, 1988:7
Species sp. Palacios-Vargas & Gómez-Anaya, 1993:16
Species sp. Palacios-Vargas & Gómez-Anaya, 1995:29
Species sp. Brand, RH et Dunn, CP, 1998:238
Species sp. Christiansen & Bellinger, 1998
Species sp. Culik, de Souza 1 Ventura, 2002
Species spectabilis Handschin, 1930
Species tullbergi (Reuter, OM, 1876:85) Stach, J, 1956:241, i.s.
            Syn.: Sminthurus Tullbergi(sic) Reuter, OM, 1876:85 IOS
Species venusta Bretfeld, 2001
Species viridescens Stach, 1920
            Syn.: Bourletiella arvalis var. fusca Womersley, H, 1932:27
            Syn.: Bourletiella arvalis ssp. dorsobscura Salmon, JT, 1941:415
Bourletiella arvalis
subsp. dorsobscura from New Zealand
After Salmon JT, 1941 Pl.71 Fig.516
Remark by Stach, J. 1956:148: Fig.515 of Bourletiella arvalis dorsobscura Salmon, 1941, from New Zealand, without crest [of male clasping organ].

Bourletiella viridescens f. dorsobscura from New Zealand
2007.12.29 © van de Vyver, L.

            Syn.: Bourletiella lutea Gisin, 1946, nec Lubbock, 1867
            Syn.: Bourletiella hortensis Stach, 1956, nec Fitch, 1863
Bourletiella viridescens from Australia
After Greenslade P & Ireson JE, 1986 Fig.7
Bourletiella viridescens from Tasmania
After Ireson JE, 1990 Fig.42
Bourletiella viridescens from New Zealand
2007.12.21 © van de Vyver, L.
Bourletiella viridescens from New Zealand
2007.12.21 © van de Vyver, L.
Bourletiella viridescens from New Zealand
2007.12.21 © van de Vyver, L.
Bourletiella nr viridescens from the USA
2009.01.02 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.02 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.02 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.24 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.24 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.24 © Justis, S.
Bourletiella nr viridescens from the USA
2009.01.24 © Justis, S.
Bourletiella nr viridescens from the USA
2009.02.07 © Justis, S.
Bourletiella nr viridescens from the USA
2009.02.24 © Justis, S.
Bourletiella nr viridescens from the USA
2009.02.24 © Justis, S.
Bourletiella viridescens from the UK
2009.08.20 © Ransom, T.
Bourletiella viridescens from the UK
2009.08.24 © Ransom, T.
Bourletiella nr viridescens from the USA
2009.11.22 © Gledhill, L.
Bourletiella nr viridescens from the USA
2009.11.25 © Gledhill, L.
Bourletiella viridescens from the UK
2013.08.01 © Garlick, T.
Bourletiella viridescens juv. from Holland
2015.08.21 © van Duinen, J.
Bourletiella viridescens from Holland
2015.08.28 © van Duinen, J.
Bourletiella viridescens from Holland
2015.08.28 © Wieringa, P.
Bourletiella viridescens from Holland
2015.08.28 © Wieringa, P.
Bourletiella viridescens from Holland
2015.09.18 © van Duinen, J.
Bourletiella viridescens from Holland
2016.09.03 © van Duinen, J.
Bourletiella viridescens from Greece
2017.03.04 © Dimitrakopoulos, C.


Genus Diksamella Bretfeld, G, 2005:13
Species pohli Bretfeld, G, 2005:13, t.t.
Diksamella pohli from Yemen
After Bretfeld G, 2005 Fig.22


Genus Nasosminthurus Stach, 1955
Species analis (Womersley, H, 1939:251) Greenslade P in Houston, 1994:127
Species dimorphus (Womersley, H, 1932:31) Stach, J, 1956:134, t.t.
Species nigridorsalis (Womersley, H, 1932:30) Greenslade, P in Houston, WWK, 1994:127
Species sigmoides (Womersley, 1933) Greenslade P in Houston, 1994:127

Genus Navarrella Bretfeld & Arbea, 2000
Species apicalis Bretfeld & Arbea, 2000, t.t.

Genus Orenius Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species maculatus Bretfeld, G, 2005:16
Orenius maculatus from Yemen
After Bretfeld G, 2005 Fig.29

Species parvus Bretfeld in Bretfeld, Poliakov & Broza, 2000, t.t.

Genus Pseudobourletiella Stach, J, 1956:142
  Syn.: Thysanura Smynthurus(sic) MacGillivray, AD, 1893:127 partim ISS
  Syn.: Smynthuri(sic) Smynthurus(sic) Folsom, JW, 1896:449 partim ISS
  Syn.: ... Symnthurus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:54 lapsus ISS
Species spinata (MacGillivray, AD, 1893:127) Stach, J, 1956:142, t.t.
            Syn.: Smynthurus(sic) spinatus MacGillivray, AD, 1893:127,tt IOS
            Syn.: Pseudobourletiella chandleri Pedigo, 1968
             Syn.: Bourletiella spinatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:54 ISS
Pseudobourletiella spinata from the USA
2006 © Web, S.
Pseudobourletiella spinata from Canada
2006.06.23 © Pilon, C.
Pseudobourletiella spinata ♂ from Canada
2006.07.23 © Pilon, C.
Pseudobourletiella spinata ♂ from Canada
2006.09.06 © Pilon, C.
Pseudobourletiella spinata from Canada, juvenile
2007.07.07 © Pilon, C.
Pseudobourletiella spinata from Canada
2007.07.18 © Pilon, C.
 
Pseudobourletiella spinata ♀ from the USA
2008.06.07 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.06.07 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.06.13 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2008.06.13 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2008.08.15 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2008.12.07 © Fleming, K.
Pseudobourletiella spinata ♀ from the USA
2009.02.01 © Fleming, K.
Pseudobourletiella spinata ♀ from the USA
2009.07.28 © Murray, T.
Pseudobourletiella spinata ♀ from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.15 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.20 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.20 © Justis, S.
Pseudobourletiella spinata juv. from the USA
2010.05.24 © Justis, S.
Pseudobourletiella spinata ♀ from the USA
2011.01.21 © Montgomery, G.
Pseudobourletiella spinata ♀ from the USA
2011.05.27 © Carr, J.
Pseudobourletiella spinata ♀ from the USA
2011.05.29 © Murray, T.
Pseudobourletiella spinata ♂ from the USA
2011.08.04 © Eiseman, C.
Pseudobourletiella spinata ♀ from the USA
2011.08.04 © Eiseman, C.
Pseudobourletiella spinata ♀ from the USA
2012.01.01 © Justis, S.
Pseudobourletiella spinata ♀ from the USA
2012.01.01 © Justis, S.
Pseudobourletiella spinata ♂ & ♀ from Canada
Courtship
2015.06.05 © Murray, A.
Pseudobourletiella spinata juv. ♂ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♂ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
With dorsal thoracic bulge (*)
2015.06.07 © Murray, A.
Pseudobourletiella spinata ♀ from Canada
2015.06.07 © Murray, A.


Genus Heterosminthurus Stach, 1955
  Syn.: Collembola Heteromurus(sic) Hopkin, SP, 1997:82 lapsus
Heterosminthurus insignis from the UK
Empodium of metaleg with distinct lamella
2008.06.30 © Ardron, P.A. (Det.)

Species aquaticus (Maynard, EA, 1951:259)
Bourletiella (Deuterosminthurus) aquatica
from the USA, ♀
After Maynard, E.A, 1951 Plate XXIII Fig.408
Bourletiella (Deuterosminthurus) aquatica
from the USA, ♂
After Maynard, E.A, 1951 Plate XXIII Fig.409

            Syn.: Bourletiella (Heterosminthurus) insignis American authors
            Syn.: Deuterosminthurus insignis Mills, 1939
            Syn.: Deuterosminthurus insignis Hammer, M, 1953:59
            Syn.: Deuterosminthurus insignis Snider, RJ, 1969:358
Heterosminthurus aquaticus? from the USA
2017.06.10 © Dobbs, Br.
Heterosminthurus aquaticus? from the USA
Ant.4 with 8 subdivisions
2017.06.10 © Dobbs, Br.

Species bilineatus (Bourlet, 1842) Stach, J, 1956:182, B
            Syn.: Smynthurus(sic) Bourletii(sic) Gervais in Walkenaër, 1844 IOS
             Syn.: Smynthurus(sic) Bourletii(sic) Nicolet, H, 1847:359 ISS
             Syn.: Smynthurus(sic) Bourletii(sic) Lubbock, J, 1870:297 ISS
Smynthurus(sic) Bourletii(sic) from the UK
After Lubbock J, 1870 Pl.XXI Fig.8
Smynthurus(sic) Bourletii(sic) from the UK
Antenna
After Lubbock J, 1870 Pl.XXI Fig.9
Smynthurus(sic) Bourletii(sic) from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.10

             Syn.: Smynthurus(sic) Bourletii(sic) Lubbock, J, 1873:110 ISS
Smynthurus(sic) Bourletii(sic) from the UK
After Lubbock, J, 1873 Pl.5.

            Syn.: Smynthurus(sic) oblongus Nicolet, H, 1842:81
            Syn.: Deuterosminthurus coronatus Bagnall, 1921
            Syn.: Heterosminthurus cornutus Stach, J, 1956:190
Heterosminthurus cornutus from Poland
After Stach, J, 1956 Pl.XXII Fig.7

Heterosminthurus bilineatus from Poland
After Stach, J, 1956 Pl.XXII Fig.2
Heterosminthurus bilineatus from Poland
After Stach, J, 1956 Pl.XXII Fig.1
Heterosminturus bilineatus from Germany
Mating ritual
After Bretfeld G, 1973 Fig.1B
             Syn.: Bourletiella (Bourletiella) bilineatus(sic) Skidmore, 1995 ISS
Heterosminthurus bilineatus from the UK
Footcomplex, empodium with lamella
2004 © Hopkin, S.P.
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK, ♀
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from the UK, ♂
Male interocular sinusoid setae
2008.07.13 © Ardron, P.A. (Det.)
Heterosminthurus bilineatus from France
2009.09.dd © Lebeaux, P.
Heterosminthurus bilineatus from Holland
2015.05.04 © van Duinen, J.
Heterosminthurus bilineatus from Holland
2015.06.01 © van Duinen, J.

  Varietas nigriceps Schille
Species borealis Bretfeld & Zoellner, 2000
Species caucasicus Stach, 1956
Species chaetocephalus Hüther, 1971
Heterosminthurus chaetocephalus from Germany
Mating ritual
After Bretfeld G, 1976

Species claviger (Gisin, 1958) Bretfeld, G, 1971:150, B
            Syn.: Bourletiella craggi Murphy, 1960
Bourletiella (Heterosminthurus) clavigera
from Süd-Bayern
After Gisin H, 1960 Abb.521
Heterosminthurus claviger from the UK
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♀
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♀
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♀
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♂
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♂
Male interocular paddle-like setae
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from the UK, ♂
Male interocular paddle-like setae
2008.07.10 © Ardron, P.A. (Det.)
Heterosminthurus claviger from Holland
2013.08.09 © den Ouden, A.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2014.10.06 © van Duinen, J.
Heterosminthurus claviger from Holland
2015.07.26 © van Duinen, J.
Heterosminthurus claviger from the UK
2015.09.10 © Boardman, P.
Heterosminthurus claviger from Holland
2016.09.03 © van Duinen, J.

Species diffusus (Gisin, H, 1962:23) Bretfeld, 1999
Species dreisbachi (Snider, RJ, 1969:360)
Deuterosminthurus dreisbachi from the USA
After Snider RJ, 1969 Fig.1

Species ihu Christiansen, K et Bellinger, P, 1992:365
Species insignis (Reuter, OM, 1876:83) Stach, J, 1956:181, t.t., B
            Syn.: Sminthurus novemlineatus var. palustris Becker, 1902
            Syn.: Sminthurus repandus Ågren, 1902
            Syn.: Bourletiella (Deuterosminthurus) aquatica Maynard, EA, 1951:259

Heterosminthurus insignis from Poland, ♂
Male with 4 distincly longer interocular setae
After Stach, J, 1956 Pl.XXV Fig.1
Heterosminthurus insignis from Germany
Mating ritual
After Bretfeld G, 1976
Heterosminthurus insignis from the UK, ♂
Some interocular setae are longer than the others
2005 © Hopkin, S.P.
Heterosminthurus insignis from France
2007.05.14 © Lebeaux, P.
Heterosminthurus insignis from France
2007.05.14 © Lebeaux, P.
Heterosminthurus insignis from France
2007.06.18 © Lebeaux, P.
Heterosminthurus insignis from France
2007.06.18 © Lebeaux, P.
Heterosminthurus insignis from the UK, ♀
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Det. Ardon, P.A.
2008.06.21 © Ardron, P.A.
Heterosminthurus insignis from the UK
Male chasing female
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
Male rejected by female
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK, ♀
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
2008.06.30 © Ardron, P.A. (Det.)
Heterosminthurus insignis from the UK
2008.08.06 © Robertson, A.
Heterosminthurus insignis from France
2009.09.dd © Lebeaux, P.
Heterosminthurus insignis from France
2009.09.dd © Lebeaux, P.

             Syn.: Heterosminthurus insigris(sic) Nickerl, J, Helbig, R, Schulz, H-J, Werner, C et Neinhuis, C, 2012:7 ISS
Heterosminthurus insignis juv. from the UK
2012.04.15 © Nurcombe, E.
Heterosminthurus insignis juv. from the UK
2012.04.15 © Nurcombe, E.
Heterosminthurus insignis from the UK
2012.05.09 © Murray, A.
Heterosminthurus insignis from France
2012.07.26 © Lebeaux, P.
Heterosminthurus insignis juv. from France
2012.07.31 © Lebeaux, P.
Heterosminthurus insignis ♀ & ♂ from France
Courtship ritual
2012.07.31 © Lebeaux, P.
Heterosminthurus insignis from Holland
2013.08.22 © van Duinen, J.
Heterosminthurus insignis from the UK
2014.06.13 © Michaelson, J.
Heterosminthurus insignis ♀ from Holland
2016.08.28 © van Duinen, J.
Heterosminthurus insignis ♂ from Holland
2016.08.28 © van Duinen, J.

Species kiianus Yoshii & Sawada, 1997
Species koontzi Snider, RJ et Calandrino, FJ, 1987:10
Bourletiella (Heterosminthurus) koontzi from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.14
Bourletiella (Heterosminthurus) koontzi from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.15

Species linnaniemii (Stach, 1920) Stach, 1955
            Syn.: Bourletiella nigriceps Latzel, R, 1917:250
            Syn.: Deuterosminthurus linnaniemii var. decolorata Stach, 1919
Heterosminthurus linnanieimii var. decolorata from Poland
After Stach J, 1956 PlateXXIII Fig.4

            Syn.: Heterosminthurus linnaniemii f. principalis Stach, J, 1956:189
            Syn.: Heterosminthurus linnaniemii f. rubra Bretfeld, 2000
Heterosminthurus linnanieimii ♀ from Poland
After Stach J, 1956 PlateXXIII Fig.1
Heterosminthurus linnanieimii ♂ from Poland
After Stach J, 1956 PlateXXIII Fig.3

             Syn.: Deuterosminthurus linnaniemi(sic) Szeptycki & Weiner in Razowski, 1990 ISS
Species multiornatus Bretfeld & Zoellner, 2000
Species nonlineatus (Gisin, 1946)
Species novemlineatus (Tullberg, 1871) Stach, J, 1956:193
            Syn.: Sminthurus novem-lineatus(sic) Tullberg, 1871 IOS
            Syn.: Sminthurus pilosicauda Reuter, OM, 1876:84
Heterosminthurus novemlineatus from Poland
Varietas pilosicauda
After Stach, J, 1956 Pl.XXIV Fig.5

            Syn.: Heterosminthurus novemlineatus f. bifasciata Stach, J, 1956:196
Heterosminthurus novemlineatus from Poland
Forma bifasciata
After Stach, J, 1956 Pl.XXIV Fig.2

            Syn.: Heterosminthurus novemlineatus f. malopigmentata Stach, J, 1956:196
Heterosminthurus novemlineatus from Poland
Forma malopigmentata
After Stach, J, 1956 Pl.XXIV Fig.1

            Syn.: Heterosminthurus novemlineatus f. unilineata Stach, J, 1956:195
Heterosminthurus novemlineatus f. unilineata from Poland
After Stach J, 1956 Pl.XXIV Fig.4

             Syn.: Sminthurus (Setosi) novem-lineatus(sic) Tullberg, T, 1872:30 ISS
             Syn.: Sminthurus novemlineato(sic) Reuter, OM, 1876:84 ISS
             Syn.: Sminthurus 9-lineatus(sic) f. principalis Börner, 1900 ISS
Heterosminthurus novemlineatus from Poland
After Stach, J, 1956 Pl.XXIV Fig.6
Heterosminthurus novemlineatus from Poland
After Stach, J, 1956 Pl.XXIV Fig.3
Heterosminthurus novemlineatus from France
2012.07.26 © Lebeaux, P.
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
2015.05.13 © Beck, K.
Heterosminthurus novemlineatus from Poland
Caudal aspect
With faecal pellet
2015.05.13 © Beck, K.

Species nymphes Yosii, 1970
Species pirika Yosii, R, 1972:96
Species punctatus Bretfeld, 1996
Species putoranae Bretfeld, 2000
Species quadristrigatus Bretfeld, 2000
Species setiger Gisin, 1961
Species stebaevae Bretfeld, 1996
Species umbonicus Bretfeld, 2000
Species undulans Yosii, R et Lee, C-E, 1963:11
Heterosminthurus undulans from Korea
After Yosii R & Lee C-E, 1963 Fig.6A

            Syn.: Heterosminthurus undulatus Yosii, R et Lee, C-E, 1963:12 =?
            Syn.: Heterosminthurus insignis Yosii, 1954 =?
            Syn.: Heterosminthurus insignis Yosii, R, 1977:164 =?
Species undulatus (Hammer, 1953) Stach, 1956:180

Genus Aneuempodialis Stach, 1955
Species cinereus (Womersley, H, 1932:31) Stach, 1955 tt
Deuterosminthurus cinereus from Australia
After Womersley H, 1932 Fig.10d

            Syn.: Deuterosminthurus (Deuterosminthurus) cinereus var. quadripunctatus Womersley, H, 1932:31
Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.07.10 © Steve & Alison.
Aneuempodialis cinereus from Australia
2016.08.07 © Steve & Alison.

Species maculatus (Womersley, H, 1939:322) Greenslade, P, 1977:249

Genus Bovicornia Delamare Deboutteville, 1947
Bovicornia sp. from Botswana
2015.01.06 © Dabek, L.

Species bidoma Mari Mutt, JA, 1978:166
Species coronata Delamare Deboutteville, 1947, t.t.
Species greensladei Massoud & Delamare Deboutteville, 1967
Species paulani Massoud & Delamare Deboutteville, 1964
Species rhinoceros Massoud & Delamare Deboutteville, 1964
Species somala Dallai, R, 1981:242
Species uniformis Murphy, 1960

Genus Prorastriopes Delamare Deboutteville, 1947
  Syn.: Bourletiellidae Andiella Stach, 1955
Species sp.1 Bellini, BC et Zeppelini, D, 2004:588
Species sp.2 Janssens F & Takahashi M, 2009
Prorastriopes sp. from the USA
2009.08.25 © Takahashi, M.
Prorastriopes sp. from the USA
Fourth antennal segment with 12 subsegments
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.

Species sp.3 Janssens F & Takahashi M, 2009
Prorastriopes sp. from the USA
2009.08.25 © Takahashi, M.
Prorastriopes sp. from the USA
Fourth antennal segment with 12 subsegments
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.
Prorastriopes sp. from the USA
2009.08.27 © Takahashi, M.

Species sp.4 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species alatipes Murphy, 1960
Species azuladus Bretfeld, 1992
Species barnardi (Womersley, 1931)
Species carrerensis (Rapoport & Bianco, 1962) Betsch, 1980
Species cingulus (Bonet, 1933) Bretfeld, 1992
            Syn.: Deuterosminthurus cingula(sic) Bonet, 1933 IOS
             Syn.: Deuterosminthurus cingula(sic) Stach, 1956 ISS
Prorastriopes cingulus ♂ from Peru
2009.03.23 © Contreras Valdez, A.

Species coalingaensis Snider, 1978
             Syn.: Prorastriopes coalingensis(sic) Snider, RJ et Loring, SJ, 1984:37 ISS
Species domeykoi (Stach, 1955) Betsch, 1977
Species izarrae Najt, 1967
Species lippsoni Snider, 1978
Species marginatus Bretfeld, 1992
Species marmoratus (Womersley, 1931)
Species olivieri Delamare Deboutteville & Massoud, 1963:267
            Syn.: Bourletiella Olivieri(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species pampeanus Rapoport & Bianco, 1962
Species paryskii (Stach, 1955) Betsch, 1977
Species patagonicus (Delamare Deboutteville & Massoud, 1963) Betsch, 1980
Species pauliani Massoud & Delamare Deboutteville, 1964
Species pulcher Delamare Deboutteville, 1947, t.t.
            Syn.: Prorastriopes pulchra(sic) Delamare Deboutteville, 1947,tt IOS
             Syn.: Prorastriopes pulchrus(sic) Salmon, JT, 1964:610 ISS
  Subspecies punctatus Delamare Deboutteville, 1947
Species risbeci Delamare Deboutteville, 1947
Species savannus Murphy, 1960
Species schultzei (Börner, C in Schultze, 1908:61)
Species spathaceus (Börner, 1907)
Species validentatus (Snider, 1978)
            Syn.: Bourletiella hoffi Scott, 1965 =?
            Syn.: Bourletiella multimaculata Scott, 1965 =?
Species venezuelae Bretfeld, 1992
Species violaceus Murphy, 1960
Species webbi Paclt, 1964
Species wexfordensis (Snider, RJ, 1969:362)
Deuterosminthurus wexfordensis from the USA
After Snider RJ, 1969 Fig.23

Species xeromorphus Snider, 1978

Genus Bourletides Betsch, J-M et Massoud, Z, 1972:234
Species wallacei Betsch, J-M et Massoud, Z, 1972:225, t.t.

Genus Cassagnaudiella Ellis, WN, 1975:78 key
Species caeruleovernalis Nayrolles, P, 1995:48
            Syn.: Cassagnaudiella cæruleovernalis(sic) Nayrolles, P, 1995:48 IOS
             Syn.: Bourletiella (Cassagnaudiella) coeruleovernalis(sic) Schulz, H-J et van Harten, A, 2013:13 ISS
Bourletiella (Cassignaudiella) coeruleovernalis(sic) from the UAE
After Schulz H-J & van Harten A, 2013 Pl.2

Species cadiensis Nayrolles, P, 1995:51
Cassagnaudiella cadiensis from Spain
After Nayrolles P, 1995 Fig.15

Species canigouensis Nayrolles, P, 1995:56
Bourletiella (Cassignaudiella) cf. canigouensis from the UAE
After Schulz H-J & van Harten A, 2013 Pl.1

Species cruciata (Haybach, 1972)
            Syn.: Cassagnaudiella trichroma Nayrolles, P, 1995:56
Cassagnaudiella trichroma from France
After Nayrolles P, 1995 Fig.27

Bourletiella (Cassignaudiella) cruciata from the UAE
After Schulz H-J & van Harten A, 2013 Pl.3

Species gamae (Bretfeld, 1994)
Species lanceolata Bretfeld, 2001
Species luteovernalis Nayrolles, P, 1995:51
Species pruinosa (Tullberg, 1871) Boguleanu, 1982
            Syn.: Sminthurus luteo(sic) Reuter, OM, 1876:84
            Syn.: Smynthurus(sic) frontalis Uzel, 1890
            Syn.: Smynthurus fulvus Lie-Petterson, 1896 =?
            Syn.: Bourletiella radula Cassagnau, 1964 partim
            Syn.: Bourletiella radula Raynal, 1973
            Syn.: Bourletiella radula Raynal, G, 1974:182
            Syn.: Bourletiella (Cassignaudiella) radula Ellis, WN, 1975:78
            Syn.: Bourletiella (Cassagnaudiella) radula Klaver, E, 1975:185
            Syn.: Bourletiella radula Nayrolles, 1995

Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.16
Lowland colour form
Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.17
Medium mountain colour form
Cassagnaudiella pruinosa from France
After Nayrolles P, 1995 Fig.18
High mountain colour form
Cassagnaudiella pruinosa ♂ from Holland
2013.09.19 © van Duinen, J.
Cassagnaudiella pruinosa ♂ from Holland
2013.09.19 © van Duinen, J.
Cassagnaudiella pruinosa ♂ from Holland
Abd.5 with modified setae
2013.09.19 © van Duinen, J.


Genus Cyprania Bretfeld, 1992
Species gisae Bretfeld, 1992
Species inopinata Bretfeld, 1996

Genus Fasciosminthurus Gisin, H, 1960:281
Species albanicus (Stach, J, 1956:164) Bretfeld, 1992
Deuterosminthurus albanicus from Poland
After Stach J, 1956 Plate XXI Fig.6
Deuterosminthurus albanicus from Poland
After Stach J, 1956 Plate XXI Fig.7

  Subspecies angulipunctatus Loksa, I et Bogojevic,J, 1970:138
Bourletiella (Fasciosminthurus) albanica ssp. angulipunctata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.51

Species badikahanus Bretfeld, 2001
Species bedosae Nayrolles, P, 1994:326
Fasciosminthurus bedosae from Spain
After Nayrolles P, 1994 Fig.21

Species canariensis (Paclt, 1964)
Species cassagnaui Nayrolles, P, 1994:330
Fasciosminthurus cassagnaui from Spain
After Nayrolles P, 1994 Fig.30
Fasciosminthurus cassagnaui from Ibiza
2015.09.15 © Beer, R.

Species circumfasciatus (Stach, J, 1956:176) Bretfeld, 1992
Deuterosminthurus circumfasciatus from Poland
After Stach J, 1956 Plate XXI Fig.1
Deuterosminthurus circumfasciatus from Poland
After Stach J, 1956 Plate XXI Fig.2

  Subspecies maculatus Betsch, J-M, 1977:85
Prorastriopes maculatus from Mongolia
After Betsch J-M, 1977 Fig.2D

Species coronatus (Betsch, J-M, 1977:85)
Species cugnyi (Nayrolles, 1987) Nayrolles, P, 1993:671
Species dictyostigmatus Nayrolles, P, 1993:669
Fasciosminthurus dictyostigmatus from France
After Nayrolles P, 1993 Fig.10
Fasciosminthurus dictyostigmatus from France
After Nayrolles P, 1993 Fig.9
Fasciosminthurus dictyostigmatus from Spain
2012.04.09 © Joaquim, F.P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.
Fasciosminthurus dictyostigmatus from France
2014.05.06 © Lebeaux, P.

Species lacazei (Denis, JR, 1924:199) Delamare Deboutteville, C & Massoud, Z, 1963:256
            Syn.: Deuterosminthurus Lacazei(sic) Denis, JR, 1924:199 IOS
             Syn.: Fasciosminthurus Lacazei(sic) Delamare Deboutteville, C et Massoud, Z, 1963:256 ISS
Fasciosminthurus lacazei from Spain
After Nayrolles P, 1993 Fig.1

Species lacunifer Bretfeld, 2001
Species longisetus Nayrolles, P, 1994:322
Fasciosminthurus longisetus ♀ from Spain
After Nayrolles P, 1994 Fig.7
Fasciosminthurus longisetus ♀ from Spain
After Nayrolles P, 1994 Fig.8

Species mascaraeus Bretfeld, 2001
Species mertensi Bretfeld, 2001
Species nairicus Bretfeld, 2001
Species niger Bretfeld, 2001
Species obtectus Bretfeld, 1992
Species perforatus Bretfeld, 2001
Species pontignanoi Bretfeld, 1992
Species pseudovirgulatus Nayrolles, 1996
Species quinquefasciatus (Krausbauer, 1898) Salmon, 1964, t.t., B
             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, H, 1931:7 ISS
Deuterosminthurus cinquefasciatus(sic) from Tasmania
After Womersley, 1931 Pl.I Fig.3

             Syn.: Deuterosminthurus 5-fasciatus(sic) Womersley, H, 1931:7 ISS
             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, 1932 ISS
             Syn.: Deuterosminthurus cinquefasciatus(sic) Womersley, 1939 ISS
             Syn.: Prorastriopes cinquefasciatus(sic) Greenslade, P et Ireson, JE, 1986:282 ISS
Prorastriopes cinquefasciatus(sic) from Australia
After Greenslade P & Ireson JE, 1986 Fig.1

             Syn.: Prorastriopes cinquefasciatus(sic) Ireson, 1993 ISS
Fasciosminthurus quinquefasciatus from Portugal
2012.05.15 © Jacinto, V.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from France
2012.09.16 © Lebeaux, P.
Fasciosminthurus quinquefasciatus from Holland
2013.08.11 © den Ouden, A.
Fasciosminthurus quinquefasciatus juv. from Holland
After Berg M, 2013 Fig.2 lower
Fasciosminthurus quinquefasciatus juv. from Holland
After Berg M, 2013 Fig.2 upper
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.06.24 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Holland
2014.08.14 © Wieringa, P.
Fasciosminthurus quinquefasciatus from Holland
2014.08.14 © van Duinen, J.
Fasciosminthurus quinquefasciatus ♀ from Holland
Sub-anal appendage (s.a.a.)
2014.08.14 © van Duinen, J.
Fasciosminthurus quinquefasciatus from Canada
2015.06.18 © Murray, A.
Fasciosminthurus quinquefasciatus from Canada
2015.06.18 © Murray, A.
Fasciosminthurus quinquefasciatus from Canada
Walking
2015.06.18 © Murray, A.

Species raynalae (Nayrolles, 1987) Nayrolles, P, 1993:659
Species saportae Nayrolles, 1996
Species sauteri Nayrolles & Lienhard, 1990
Fasciosminthurus sauteri from France
2017.04.22 © Tiky, S.
Fasciosminthurus sauteri from France
2017.04.22 © Tiky, S.
Fasciosminthurus sauteri from France
2017.04.22 © Tiky, S.

Species strasseni (Paclt, 1964) Bretfeld, 1999
Species strigatus (Stach, 1922) Rusek, J in Dlabola, J, 1977:16
  Subspecies sajanensis Bretfeld, 1996
Species tarianus Bretfeld, 2001
Species virgulatus (Skorikov, 1899) Delamare Deboutteville, C & Massoud, Z, 1963:256
            Syn.: Deuterosminthurus bilineatus var. plusilineata Stach, 1922
            Syn.: Deuterosminthurus bilineatus ab. prodromata Stach, 1922
Bourletiella (Fasciosminthurus) virgulata from Jugoslavia
After Loksa I & Bogojevic J, 1970 Abb.39
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.08.27 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2010.09.14 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2014.10.14 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.10 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.10 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.11 © Beer, R.
Fasciosminthurus virgulatus from Ibiza
2015.09.14 © Beer, R.
Fasciosminthurus virgulatus from France
2017.04.22 © Tiky, S.
Fasciosminthurus virgulatus from France
2017.04.22 © Tiky, S.


Genus Kaszabellina Betsch, J-M, 1977:79
Species major Betsch, J-M, 1977:83
Species minima Betsch, J-M, 1977:84
Species variabilis Betsch, J-M, 1977:79, t.t.

Genus Koupelia Bretfeld, 2001
Species foliata Bretfeld, 2001, t.t.

Genus Madecassiella Betsch & Waller, 1996
Species ibityensis Betsch & Waller, 1996, t.t.

Genus Massoudia Betsch, 1975
Species griveaudi Betsch, 1975

Genus Paulianitas Betsch, 1978
Species vietti Betsch, 1978

Genus Stenognathriopes Betsch, J-M et Lasebikan, BA, 1979:166
  Syn.: Sminthuridae(lapsus) Senognathriopes(sic) Vázquez-G, M et Tun Osorio, R, 2004:283 ISS
Subgenus Stenognathriopes Betsch, J-M et Lasebikan, BA, 1979:166
Species sp. Bretfeld, G, 2005:45
Species huetheri Betsch, J-M et Lasebikan, BA, 1979:166
            Syn.: Stenognathriopes hütheri(sic) Betsch, J-M et Lasebikan, BA, 1979:166 IOS
             Syn.: Stenognathriopes hutheri(sic) Zeppelini, D et da Silva, DD, 2012:52 ISS
Species interpositus (Hüther, 1967) Betsch, J-M & Lasebikan, BA, 1979:166
Species rastrifer (Denis, 1948) Betsch, J-M & Lasebikan, BA, 1979:166
Species vilhenai (Delamare Deboutteville, C & Massoud, Z, 1964:73) Betsch, J-M & Lasebikan, BA, 1979:166
Species yemenensis Bretfeld, G, 2005:42
Subgenus Tenentiella Palacios-Vargas, JG et Vázquez, M, 1997:285
Species janssensi Zeppelini, D et da Silva, DD, 2012:52
Stenognathriopes (Tenentiella) janssensi ♀ from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.1A
Stenognathriopes (Tenentiella) janssensi ♀ from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.1B
Stenognathriopes (Tenentiella) janssensi from Brazil
After Zeppelini D & da Silva DD, 2012 Fig.2

Species siankaana Palacios-Vargas, JG et Vázquez, M, 1997:286, t.t.

Genus Tritosminthurus Snider, RJ, 1988:260
Species schuhi Snider, RJ, 1988:260, t.t.

References