http://www.collembola.org/taxa/bracidae.htm - Last updated on 2017.02.22 by Frans Janssens
Checklist of the Collembola: Brachystomellidae

Familia Brachystomellidae Stach, 1949 key
Genus Number
of species
Bellingeria1
Bonetella1
Brachystomella74
Brachystomellides4
Cassagnella4
Folsomiella6
Maricaella1
Massoudella1
Micronella4
Neorganella2
Parastomella1
Probrachystomellides1
Raponella1
Rapoportella15
Salvarella1
Setanodosa12
Subclavontella2
Winterella1
Brachystomellidae132
Genera with number of species
Brachystomella parvula from the UK
2012.06.11 © Murray, A.
Brachystomellidae from Tasmania
2014.04.11 © Murray, A.


Genus Brachystomella Ågren, 1903 key
F
  Syn.: ... Schöttella(sic) Schäffer, 1896 partim
  Syn.: Collembola Chondrachorutes Wahlgren, E, 1906:5
  Syn.: Achorutinae Chondachorutes(sic) Mills, HB, 1934:18 ISS
  Syn.: ... Guacharia Jackson, 1927
  Syn.: Neanurini Neanurodes Scott, HG, 1960:190
  Syn.: ... Brachygastrura Rapoport, 1962
  Syn.: Brachystomellidae Probrachystomella Salmon, JT, 1964:244 partim
  Syn.: Hypogastrurinae Neanura(sic) Christiansen, K et Bellinger, P, 1998:288 lapsus
  Syn.: Collembola Brachstomella(sic) Yosii, R, 1959:4 ISS
  Syn.: ... Brachistomella(sic) Cepeda & Whitford, 1990 ISS
  Syn.: ... Brachistomella(sic) Palacios-Vargas, JG, 1990:18 ISS
Brachystomella sp. from Vanuatu
2006 © Boldsystems.org
Brachystomella sp. from Spain
On Trichia decipiens
2012.12.15 © Jiménez, L.G.
Brachystomella neomexicana from the USA
Unguiculus absent (*)
2016.02.19 © Janssens, F.
Leg.: Vitanza, S.
Brachystomella neomexicana from the USA
Blunt-toothed maxilla
2016.02.19 © Janssens, F.
Leg.: Vitanza, S.

Species sp.1 Palacios-Vargas, 1979
Species sp.2 Christiansen, K et Bellinger, P, 1994:316
Species sp.3 Christiansen, K et Bellinger, P, 1994:316
Species sp.4 Christiansen, K et Bellinger, P, 1994:316
Species sp.5 González, V, Díaz, M et Palacios-Vargas, JG, 1997:45
Species sp.6 Culik, de Souza & Ventura, 2002
Species agrosa Wray, DL, 1953:148
            Syn.: Brachystomella maxima Cassagnau & Rapoport, 1962
Species allendei Hermosilla & Rubio, 1977
            Syn.: Brachystomella allendei Massoud & Rubio in Hermosilla & Rubio, 1976 nn
Species aspra Denis, 1931
             Syn.: Brachystomella aspera(sic) Bellinger, PF, 1996:il ISS
             Syn.: Brachystomella aspera(sic) Culik, MP et Zeppelini, DF, 2003:1122 ISS
Species baconaoensis Gruia, M, 1983:191
            Syn.: Brachystomella honda Christiansen & Bellinger, 1988
Species banksi (Maynard, EA, 1951:50)
             Syn.: Schoettella banski(sic) Díaz, A et Palacios-Vargas, JG, 1983:81,is ISS
Species barrerai Palacios-Vargas, JG et Najt, J, 1981:264
Species ceciliae Fernandes, LH et de Mendonça, MC, 2004:19
Species chilensis (Rapoport, EH & Rubio, IR, 1963:106) Rapoport, 1965
Organella chilensis from Chile
After Rapoport EH & Rubio IR, 1963 Fig.31
Brachystomella chilensis from Chile
After Weiner WM & Najt J, 2001 Fig.1

Species christianseni Massoud, 1965
Species clavigera (Schäffer, C, 1898:400) Stach, 1926
Species coatesi Weiner & Najt, 1991
             Syn.: Brachystomella coatsi(sic) Weiner, WM et Najt, J, 2001:405 ISS
Species contorta Denis, 1931
            Syn.: Brachystomella sabandinensis(sic) Winter, 1963 nn
            Syn.: Brachystomella subandinensis Massoud, Z, 1967:94
Brachystomella contorta from Costa Rica
After Weiner WM & Najt J, 2001 Fig.7

Species curvula Gisin, 1948
Species cyanea (Rapoport, 1962) Massoud, 1967:92
Brachystomella cyanea from Argentina
After Weiner WM & Najt J, 2001 Fig.14

Species desutterae Weiner, WM et Najt, J, 2001:393
Brachystomella desutterae from Peru
After Weiner WM & Najt J, 2001 Fig.20

Species dianae Greenslade, P et Najt, J, 1987:436
            Syn.: Brachystomella quadrituberculata Greenslade PJM & Greenslade P, 1973
Brachystomella dianae from Australia
2010.08.17 © Stevens, M.

Species disputa Greenslade, P et Najt, J, 1987:442
Species divisa Murphy, 1965
Species emder Greenslade P & Najt, 1987:440
Species fungicola Womersley, 1933
Species gabrielae Najt, J et Palacios-Vargas, JG, 1986:459
Brachystomella gabriela from Mexico
After Najt J & Palacios-Vargas JG, 1986 Fig.1

Species garayae Queiroz & Weiner, 2011
Species georgensis Weiner & Najt, 1991
Species globulosa Cassagnau & Rapoport, 1962
Brachystomella globulosa from Argentina
After Weiner WM & Najt J, 2001 Fig.26

Species grootaerti Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:156
             Syn.: grottaerti(sic) grottaerti(sic)} Peck, SB, 2001:106 Peck, SB, 2001:106 ISS
             Syn.: Brachystomella grottaerti(sic) Peck, SB et Jacquemart, J, 2011:4 ISS
Species hawaiiensis Yosii, 1965
Species heo Christiansen, K et Bellinger, P, 1992:43
Species hiemalis Yosii, R, 1956:20
Species insulae Najt, J et Thibaud J-M in Tillier, S, 1988:33
Species kahakai Christiansen, K et Bellinger, P, 1992:45
Species kiko Christiansen, K et Bellinger, P, 1992:45
Species koreana Weiner, WM et Najt, J, 1985:246
Species mataraniensis Weiner, WM et Najt, J, 2001:396
Brachystomella mataraniensis from Peru
After Weiner WM & Najt J, 2001 Fig.33

Species mauriesi Thibaud & Massoud, 1983
Species minimucronata Palacios-Vargas, JG et Najt, J, 1981:266
            Syn.: Brachystomella minimucronatus(sic) Palacios-Vargas, JG et Najt, J, 1981:266 IOS
Species momona Christiansen, K et Bellinger, P, 1992:48
Species montebella Najt, J et Palacios-Vargas, JG, 1986:461
Brachystomella montebella from Mexico
After Najt J & Palacios-Vargas JG, 1986 Fig.10

Species nana Rubio & Najt, 1979
Species neomexicana (Scott, HG, 1960:190) Bellinger, PF, 1985:79
Neanurodes neomexicanus from New Mexico
After Scott HG, 1960 Fig.5
            Syn.: Brachystomella arida Christiansen, K et Bellinger, P, 1980:268
             Syn.: Neanurodes Neomexicanus(sic) Massoud, Z, 1967:346 ISS
Brachystomella neomexicana from the USA
Mass aggregation
2015.09.20 © Vitanza, S.
Brachystomella neomexicana from the USA
Furcula: broad manubrium with 2 short dentes
2016.01.16 © Vitanza, S.
Brachystomella neomexicana from the USA
2016.02.16 © Janssens, F.
Leg.: Vitanza, S.
Brachystomella neomexicana from the USA
Post-antennal organ (PAO) with 5 lobes superequal to nearest ocellus
2016.02.19 © Janssens, F.
Brachystomella neomexicana from the USA
Trilobed apical bulb of ant.4
2016.02.19 © Janssens, F.
Leg.: Vitanza, S.

Species nubila Gisin, 1957
             Syn.: Brachystomella pubila(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:259 ISS
Species parvula (Schäffer, 1896:176) Stach, 1926, t.t., B
            Syn.: Schöttella(sic) minor Scherbakov, 1899
            Syn.: Schöttella(sic) media Axelson, 1900
            Syn.: Schöttella(sic) albomaculata Carl, J, 1901:254
            Syn.: Schötella(sic) crassicornis Schött, 1902
            Syn.: Brachystomella maritima Ågren, 1903
            Syn.: Chondrachorutes Wahlgreni(sic) Denis, 1924 IOS
            Syn.: Brachystomella terrafolia Massoud, Z, 1967:94 =?
Brachystomella parvula
After Potapov, M. in Babenko, A., 1988.
Brachystomella parvula from France
2012.01.15 © Lebeaux, P.
Brachystomella parvula from France
2012.01.23 © Lebeaux, P.
Brachystomella parvula from the UK
2012.06.11 © Murray, A.
Brachystomella parvula from Holland
2012.09.04 © van Duinen, J.
Brachystomella parvula from the UK
Large post-antennal organ
2012.09.19 © Nurcombe, E.
Brachystomella parvula from the UK
2012.09.19 © Nurcombe, E.
Brachystomella parvula from the UK
2012.09.19 © Nurcombe, E.
Brachystomella parvula from the UK
2012.09.19 © Nurcombe, E.
Brachystomella parvula from Portugal
2012.10.16 © Murray, A.
Brachystomella parvula juv. from the UK
2012.11.17 © Murray, A.
Brachystomella parvula from France
2012.12.01 © Sandra.
Brachystomella parvula from the UK
2013.01.04 © Nurcombe, E.

  Subspecies africana Yosii, R, 1959:4

            Syn.: Brachystomella parvula Womersley, 1933
            Syn.: Brachystomella parvula Womersley, 1934
            Syn.: Brachystomella parvula Womersley, 1937
            Syn.: Brachystomella parvula Womersley, 1939
Species pastoralis Greenslade P & Najt, 1987:444
Species pefauri Weiner, WM et Najt, J, 2001:398
Brachystomella pefauri from Venezuela
After Weiner WM & Najt J, 2001 Fig.40

Species perraulti Thibaud & Najt, 1993
Species pescotti (Womersley, H, 1942:23) Greenslade, P in Houston, WWK, 1994:44
Pseudachorutes pescotti from Australia
After Womersley H, 1942 Fig.1A
Remark by Janssens, F. 2014.02.14: It is clear from the drawing of Womersley (1942:Fig.1A) that the species does neither fit Pseudachorutes nor Brachystomella. IMHO, it fits Isotomidae. Possibly Proisotoma.

Species platensis Najt & Massoud, 1974
            Syn.: Brachystomella parvula King, Hutchinson & Greenslade P, 1976
Brachystomella platensis from Australia
After Greenslade P & Ireson JE, 1986 Fig.18

Species purma Weiner, WM et Najt, J, 2001:400
Brachystomella purma from Peru
After Weiner WM & Najt J, 2001 Fig.46

Species quadrituberculata Becker, 1905
Species quadrituberculata (Stach, 1964) Massoud, Z, 1967:92
Species ronderosi Najt, 1973
Species saladaensis Weiner, WM et Najt, J, 2001:401
Brachystomella saladaensis from Argentina
After Weiner WM & Najt J, 2001 Fig.52

Species septemoculata Denis, 1931
            Syn.: Brachystomella 7-oculata(sic) Denis, 1931 IOS
            Syn.: Brachystomella trinitata Weiner, WM et Najt, J, 2001:387 =?
             Syn.: Brachystomella 7-oculata(sic) Denis, 1933 ISS
             Syn.: Brachystomella 7-oculata(sic) Massoud & Bellinger, 1963 ISS
             Syn.: Brachystomella 7-oculata(sic) Winter, 1963 ISS
Brachystomella septemoculata from the Costa Rica
After Weiner WM & Najt J, 2001 Fig.59

Species sexoculata Massoud, Z, 1967:90
            Syn.: Brachystomella sexoculata Winter, 1963 nn
Species solidaria Greenslade, P et Najt, J, 1987:440
Species speciosa (Rainbow, WJ, 1907:314) Janssens F, 2014
Achorutes speciosus from Australia
After Rainbow WJ, 1907 Fig.51
Remark by Janssens, F. 2014.02.14: Given the species is described having 8 ocelli per eyepatch and the unguiculus is lacking, neither Hypogastrura nor Xenylla fits the species. The best tentative fit for now is Brachystomella.

Species stachi Mills, HB, 1934:19
Species subcrassa (Schäffer, 1897, i.s.) Denis, 1931
            Syn.: Pseudachorutes subcrassus Börner, 1903, nec Tullberg, 1871
Species surendrai Goto, 1961
Species taxcoana Palacios-Vargas & Najt, 1981:268
Species terrafolia Salmon, JT, 1944:135
Brachystomella terrafolia from India
After Bano R, 2016 p.58

Species trinitata (Jackson, 1927) Denis, 1931
Species tuberculata (Wahlgren, E, 1906:5) Stach, 1949
Brachystomella tuberculata from the Falkland Is
After Weiner WM & Najt J, 2001 Fig.65

Species unguilonga Najt & Thibaud in Tillier, 1988:33
            Syn.: Brachystomella unguilongus(sic) Najt & Thibaud in Tillier, 1988 IOS
             Syn.: Brachystomella unguilongus(sic) Fernandes, LH et de Mendonça, MC, 2004:19 ISS
Species ultima Greenslade, P et Najt, J, 1987:446
Species victoriensis de Izarra, DC, 1972:92
Brachystomella victoriensis from Argentina
After Weiner WM & Najt J, 2001 Fig.71

Species villalobosi Cassagnau & Rapoport, 1962
             Syn.: Brachystomella villa-lobosi(sic) Palacios-Vargas & Najt, 1981 ISS
Brachystomella villalobosi from Brasil
After Weiner WM & Najt J, 2001 Fig.77

Species zapatai Najt, J et Palacios-Vargas, JG, 1986:464
Brachystomella zapatai from Mexico
After Najt J & Palacios-Vargas JG, 1986 Fig.17

            Syn.: Brachystomella sp.nov. Palacios-Vargas, 1983
Species zerpa Weiner, WM et Najt, J, 2001:408
Brachystomella zerpa from Venezuela
After Weiner WM & Najt J, 2001 Fig.84


Genus Folsomiella Bonet, 1930
F
Species albida (Arlé, R, 1960:159) Salmon, JT, 1964:249
Species caeca (Folsom, JW, 1927:4) Stach, J, 1949:259, t.t.
Species intermedia (Arlé, 1939) Stach, J, 1949:306
Species polylepiana Massoud, Z, 1967:99
Species pseudocaeca de Mendonça, MC, Fernandes, LH et Abrantes, EA, 2005:16
Species trisetosa de Mendonça, MC, Fernandes, LH et Abrantes, EA, 2005:12

Genus Neorganella Rapoport, EH et Rubio, IR, 1963:108
F
Species nothofagutalis Rapoport, EH et Rubio, IR, 1963:109, t.t.
Neorganella nothofagutalis from Chile
After Rapoport EH & Rubio IR, 1963 Fig.49

Species rotundata Queiroz, GC et de Mendonça, MC, 2013:94
            Syn.: Neorganella rotundatae(sic) Queiroz, GC et de Mendonça, MC, 2013:94 IOS

Genus Setanodosa Salmon, 1942 key
F
Species afurcata (Womersley, 1933) Massoud, Z, 1967:104
Species capitata (Womersley, 1934) Massoud, Z, 1967:104
Species clavata (Schäffer, 1897)
Species decemoculata (Cassagnau & Rapoport, 1962) Massoud, Z, 1967:102
Species fueguensis Najt, 1973
Species granulata (Womersley, H, 1935:207) Massoud, Z, 1967:102
Species kanalua Christiansen, K et Bellinger, P, 1992:52
Species occidentalis (Arlé, R, 1960:160) Massoud, Z, 1967:104
Species rosasi (Bonet, 1934) Najt & Weiner, 1996
            Syn.: Anurida almanzori Bonet, 1930,nn
            Syn.: Setanodosa peruensis Massoud, Z, 1967:104
            Syn.: Salmonella peruensis Winter, 1963 nn
             Syn.: Brachystomella rosai(sic) Weiner, WM et Najt, J, 2001:387 ISS
             Syn.: Setanodosa rosai(sic) Weiner, WM et Najt, J, 2001:387 ISS
Setanodosa rosasi from Argentina
After Arbea J.I., 2007 Fig.1

Species serrata Massoud, Z, 1967:102
Species steineni (Schäffer, 1891) Wise, KAJ, 1970:183
Species tetrabrachta Salmon, 1942, t.t.

Genus Micronella Arlé, R, 1960:159 key
F
Species checayensis Massoud, 1967:105
            Syn.: Micronella checayensis Winter, 1963 nn
Species itacaman Queiroz, GC et de Mendonça, MC, 2013:84
Species longisensilla Queiroz, GC et de Mendonça, MC, 2013:87
Species porcus (Denis, 1933) Arlé, R, 1960:159, t.t.

Genus Winterella Massoud, Z, 1967:106
F
  Syn.: ... Organella Arlé, 1959 partim
Species arlesiana Massoud, Z, 1967:107, t.t.

Genus Maricaella de Mendonça, MC et Fernandes, LH, 1997:1

Species duna de Mendonça, MC et Fernandes, LH, 1997:2, t.t.

Genus Rapoportella Ellis, WN et Bellinger, PF, 1973:48
F
  Syn.: ... Probrachystomella Rapoport, 1962 u
  Syn.: Brachystomellidae Probrachystomella Salmon, JT, 1964:244 partim
  Syn.: Brachystomellinae Rapportella(sic) Najt, J et Palacios-Vargas, JG, 1986:468 ISS
Species sp.1 Najt, J et Palacios-Vargas, JG, 1986:466
            Syn.: Rapoportella rapoporti Thibaud & Massoud, 1983
Species sp.2 González, V, Díaz, M et Palacios-Vargas, JG, 1997:45
Species sp.3 Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:92
Species bonariensis (Rapoport, 1962) Najt, J & Palacios-Vargas, JG, 1986:466, t.t.
            Syn.: Probrachystomella rhodosoma Rapoport, 1962
Species boneti (Massoud, 1963) Najt, J & Palacios-Vargas, JG, 1986:466
Species karta Greenslade P & Najt, 1987
Species lowriei Najt, 1984
Species manzanoae Najt & Manzano in Manzano, 1990 nn
Species margaritae Najt, J et Palacios-Vargas, JG, 1986:467
Rapportella(sic) margaritae from Mexico
After Najt J & Palacios-Vargas JG, 1986 Fig.23

Species mucronata Najt & Massoud, 1974
Species pitomboi de Mendonça, MC et Fernandes, LH, 1995:1
Species punillensis (Izarra, 1973) Najt, J & Palacios-Vargas, JG, 1986:466
Species rapoporti (Massoud, 1963) Massoud & Thibaud, 1981
Species sigwalti Najt, J et Palacios-Vargas, JG, 1986:469
Rapoportella sigwalti from Mexico
After Najt J & Palacios-Vargas JG, 1986 Fig.31

Species yolandae (Rapoport & Maño, 1969) Najt, J & Palacios-Vargas, JG, 1986:466
            Syn.: Rapoportella rapoporti Thibaud, 1991
            Syn.: Rapoportella rapoporti Thibaud, 1993

Genus Salvarella Greenslade P & Najt, 1987

Species wallacei Greenslade P & Najt, 1987:118, t.t.
             Syn.: Salvarella Wallacei(sic) Greenslade P & Najt, 1987 ISS

Genus Brachystomellides Arlé, R, 1960:161 key
M
Species compositus Arlé, R, 1960:161, t.t.
Species micropilosus Cassagnau & Rapoport, 1962
Species navarinensis Weiner & Najt, 1997
Species neuquensis Cassagnau & Rapoport, 1962

Genus Massoudella Ellis, WN et Bellinger, PF, 1973:30
F
Species geniculata (Womersley, 1934) Ellis, WN & Bellinger, PF, 1973:30, t.t.

Genus Subclavontella Stach, 1949
F
Species acantha (Womersley, 1933) Massoud, 1967:116, t.t.
Species subacantha Massoud, 1967:116

Genus Bonetella Stach, 1949
F
Species terricola (Womersley, 1934) Massoud, 1967:117, t.t.

Genus Cassagnella Najt & Massoud, 1974
F
Species sp. Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:68
Species alba Najt & Massoud, 1974, t.t.
Species anomala (Womersley, 1933, s.i.) Greenslade P in Houston, 1994:45
Species sergioi (Najt, 1973) Najt & Weiner, 2001

Genus Raponella Najt, 1988

Species dodecophthalma (Rapoport, EH & Rubio, IR, 1963:108) Najt, 1988
             Syn.: Probrachystomella dodecophtalma(sic) Massoud, Z, 1967:110 ISS

Genus Parastomella Rapoport & Rubio, 1968
F
Species mylodontis Rapoport & Rubio, 1968, t.t.

Genus Probrachystomellides Weiner & Najt, 1991
M
Species nicolaii Weiner & Najt, 1991 tt

Genus Bellingeria Christiansen, KA et Pike, E, 2002:178, -

Species cornua (Christiansen, KA & Pike, E, 2002:178, -) Christiansen, KA & Pike, E, 2002:178, -, t.t.



References