http://www.collembola.org/taxa/cyphoder.htm - Last updated on 2019.09.08 by Frans Janssens
Checklist of the Collembola: Cyphoderinae

Subfamilia Cyphoderinae Börner C, 1906:162 sensu Janssens F, 2018 key

Genus Number
of species
Calobatinus4
Campylothorax14
Cephalophilus3
Cyphoda10
Cyphoderinus1
Cyphoderodes7
Cyphoderopsis17
Cyphoderus71
Delamarerus2
Megacyphoderus5
Mimoderus4
Paracyphoderus1
Paronella3
Pseudocyphoderus4
Serroderus25
Troglobius4
Troglopedetes31
Trogolaphysa77
Cyphoderinae284
Genera with number of species



  Syn.: Entomobryidae Cyphoderitae(sic) Delamare Deboutteville, C, 1945:48 ISS
  Syn.: Mydoniidae Cyphodurinae(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
Cyphoderus albinus from the UK
With Myrmica rubra
2007 © Cornwall, N.J.
Cyphoderus albinus from France
With Pheidole pallidula
2008.03.15 © Lebeaux, P.

  Syn.: Lepidocyrtidae Cyphoderidae(sic) Castaño-Meneses, G, Palacios-Vargas, JG, Delabie, JHC, Zeppelini, D et Mariano, CSF, 2017:741 ISS
Remark by Janssens, F. 2018.11.04: We propose to cluster Paronellini and Cyphoderini in Cyphoderinae, under Lepidocyrtidae, in line with the molecular phylogeny of Zhang et al., 2015, and the morphological study of Zeppelini & Oliveira, 2016.


Tribus Paronellini Börner, 1906:162 sensu Zhang F et al, 2015:7 key

  Syn.: Cyphoderinae Troglopedetini Börner, C, 1913:321


Paronellini from Indonesia
Composed eye with Extra Ocular Structure (EOS)
2013.09.21 © Bertner, P.
Paronellini from Indonesia
Hyaline oval apically rounded scales
2013.09.21 © Bertner, P.
Paronellini from Malaysia
Composed eye with Extra Ocular Structure (EOS)
2014.02.12 © Bertner, P.
Paronellini from Malaysia
Apically knobbed dorsal thoracic macrosetae
2014.02.12 © Bertner, P.
Paronellini from Malaysia
Collophore with extended vesicles pressed in the surface waterfilm (drinking)
2014.02.12 © Bertner, P.
Remark by Zhang, F & al. 2015:7: Molecular phylogeny supports the closer relationship of Cyphoderini and Troglopedetini [= Paronellini], both of which have been clustered with Lepidocyrtinae.
Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The presence of pseudopores behind the posterior row of setae on the fourth abdominal segment, as seen in Troglopedetes, Trogolaphysa, Cyphoderus and Troglobius, corroborates the close relationship of Cyphoderus ([Cyphoderini]) to Trogolaphysa (Paronellini).


Genus Paronella Schött, H, 1893:14 F


  Syn.: Paronellidae Trogonella Massoud in Wilson JM, 1982:218
  Syn.: Paronellidae Trogonella Wilson, JM, 1982:218
  Syn.: ... Trichorypha Thibaud, J-M et Najt, J, 1988:722
  Syn.: ... Trichorypha Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15
  Syn.: ... Microparonella Salmon, 1964


Remark by ICZN. 2007:608: Placed on the Official List of Names in Zoology. Opinion 291.

Species fusca Schött, H, 1893:16, t.t.

Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1A
Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1B
Paronella fusca
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.29

Species montana Delamare Deboutteville, 1950

Species purpurea Barra, 1969



Genus Campylothorax Schött, H, 1893:18 key


  Syn.: Dicranocentroides Dicranocentropha Wray, DL, 1953:144


Campylothorax
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.33

  Syn.: Paronellini Campylothorx(sic) Mitra, SK, 1993:65 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:19 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:20 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:21 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:24 ISS
Campylothorax sp. from Mexico
2016.10.06 © Murray, A.
Campylothorax sabanus from Brazil
Modified after Mari Mutt, 1987 in Bellini BC & Cipola NG, 2017 Fig.2


  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:165 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:166 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:167 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:168 ISS


Species camelinus Womersley, 1930

            Syn.: Campylothorax camelina(sic) Womersley, 1930 IOS


             Syn.: Campylothorax camelina(sic) Mitra, SK et Dallai, R, 1980:275 ISS



Species cassagnaui Mitra, SK et Dallai, R, 1980:313

            Syn.: Campylothorax cassagnaus(sic) Mitra, SK et Dallai, R, 1980:313 IOS
            Syn.: Campylothorax cinctus Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:131
             Syn.: Campylothorax cincta(sic) Soto-Adames, FN, 2016:2 ISS


             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & Najt, 1995 ISS
             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1996:171 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & al., 1997 ISS
             Syn.: Campylothorax cassagnaus(sic) Culik, MP et Zeppelini, DF, 2003:1129 ISS
Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Leg. McAlister, E.; Det. Culik, M.
Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Mucro
Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.
Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.

             Syn.: Campylothorax cassagnaus(sic) da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:1 ISS
             Syn.: Campylothorax cassagnaus(sic) Brito, RA, Lima, ECA, Ferreira, AS, de Oliveira, JVLC, Amorim, TPL, Machado, TG, Lopes, BCH, Costa, IA, Alves, JLS, Brito, NP, Zeppelini, D, 2017:11 ISS




Species cinctus Schött, 1927

Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.45A
Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.46A

Species cubanus Gruia, M, 1983:203

            Syn.: Campylothorax cubana(sic) Gruia, M, 1983:203 IOS
             Syn.: Campylothorax cubana(sic) Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:94 ISS
             Syn.: Camphylotorax(sic) cubana(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS
Campylothtorax(sic) cubanus from Haiti
After Soto-Adames FN, 2016 Fig.12A
Campylothtorax(sic) cubanus from Hispaniola
After Soto-Adames FN, 2016 Fig.12B





Species dominicanus Soto-Adames, FN, 2016:20

Campylothtorax(sic) dominicanus from Hispaniola
After Soto-Adames FN, 2016 Fig.10
             Syn.: Campylothorax domincanus(sic) Soto-Adames, FN, 2016:28 ISS


Species hexosetosus Soto-Adames, FN, 2016:16

Campylothorax hexosetosus from Hispaniola
After Soto-Adames FN, 2016 Fig.7
             Syn.: Campylothorax hexosetosa(sic) Soto-Adames, FN, 2016:16 ISS


Species longicornis Schött, H, 1893:20, t.t.

Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.1A
From syntype
Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.20
Mucro

Species melanocephalus Mitra, SK et Dallai, R, 1980:298

Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.29
Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.30

Species mitrai Bellini & Meneses, 2012

Species notidanus Soto-Adames, FN, 2016:11

Campylothorax notidanus from Hispaniola
After Soto-Adames FN, 2016 Fig.4A

Species plagatus Cipola, NG et de Lima Oliveira, FG, 2016:488

Campylothorax plagatus from Brazil
After Cipola NG & de Lima Oliveira FG, 2016 Fig.1

Species sabanus (Wray, DL, 1953:144) Mitra, 1975

            Syn.: Dicranocentroides (Dicranocentropha) sabana(sic) Wray, DL, 1953:144 IOS
            Syn.: Dicranocentroides fasciculatus Rapoport, 1971 =lapsus
            Syn.: Dicranocentroides fasciculatus Mari Mutt, JA et Bellinger, PF, 1990:125


             Syn.: Campylothorax sabana(sic) Mitra, 1975 ISS
             Syn.: Campylothorax sabana(sic) Mitra, SK et Dallai, R, 1980:300 ISS
Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.32A
Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.33A

             Syn.: Campylothorax sabana(sic) Cipola, NG et de Lima Oliveira, FG, 2016:494 ISS




Species schaefferi Börner, C, 1906:179

            Syn.: Campylothorax schäfferi(sic) Börner, C, 1906:179 IOS
             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:2 ISS
             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:12 ISS



Species viruaensis Santos IP, Cipola NG et Bellini BC in da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:2

Campylothorax viruaensis from Brazil
After da Silva Santos IP, Cipola NG & Bellini BC in da Silva Santos IP & al, 2016 Fig.1


Genus Trogolaphysa Mills, HB in Pearse, AS, 1938:184 key


  Syn.: Paronellinae Dicranocentruga Wray, DL, 1953:141
  Syn.: ... Dicranocentrus(sic) Mari Mutt, JA, 1979:60 ISS
  Syn.: Paronellini Diranocentruga(sic) Mitra, SK, 1993:42 ISS
Dicranocentruga
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.31

  Syn.: Paronellini Dicraanocentruga(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellinae Dicranocentuga(sic) Mitra, SK, 2002:109 ISS
  Syn.: Paronellinae Metaparonella Mitra, 1972 u



Trogolaphysa
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.40

  Syn.: Troglopedetini Trogolaphyusa(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellidae Troglolaphysa(sic) Palacios-Vargas, JG et Zeppelini, D, 1995:173 ISS
  Syn.: Paronellini Trogolpahysa Soto-Adames, FN, 2015:23 ISS


Species sp.1 (Mari Mutt, JA, 1987:403)

Species sp.2 Palacios-Vargas, JG et Thibaud, J-M, 1997:328

Species sp.3 Bellini, BC et Zeppelini, D, 2004:588

Species sp.4 (Bellini, BC et Zeppelini, D, 2004:588)

Species sp.5 Palacios-Vargas, JG, 2008:46

Species sp.6 Bellini, 2014

Species sp.7 de Oliveira, MPA, de Oliveira Bernardi, LF, Zeppelini, D et Ferreira, RL, 2016:133

Trogolaphysa new sp. from Brazil
Parasitised by mite (Parasitengona, Microtrombidiidae)
After de Oliveira MPA & al, 2016 Fig.2A

Species sp.8 Mari Mutt & al, 2016

Species sp.9 Soto-Adames, FN et Anderson, EW, 2017:40

Species aelleni Yoshii, R, 1988:453

             Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:36 ISS
             Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:39 ISS

Species balteata Soto-Adames, FN, 2015:27

Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4C
Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4D

Species beguei (Delamare Deboutteville, 1951) Nguyen, M & Soto-Adames, FN, 2018:182

            Syn.: Paronella beguei Delamare Deboutteville, 1948 nn
Trogolaphysa beguei from the Ivory Coast
After Nguyen M & Soto-Adames FN, 2018 Fig.5J



Species belizeana Palacios-Vargas, JG et Thibaud, J-M, 1997:326

Species berlandi (Denis, 1925) Thibaud, J-M & Najt, J, 1988:726

             Syn.: Paronella Berlandi(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:134 ISS

Species bessoni Thibaud, J-M et Najt, J, 1988:724

Species caripensis (Gruia, M, 1987:155) Thibaud, J-M & Najt, J, 1988:723

Species carpenteri (Denis, 1925) Yoshii, 1988

            Syn.: Dicranocentruga geminata Mari Mutt, JA, 1987:402 =?
            Syn.: Dicranocentruga jataca Wray, DL, 1953:143 =?


             Syn.: Paronella Carpenteri(sic) Cassagnau, 1963 ISS

Species clarencei Soto-Adames, FN, 2015:21

Trogolaphysa clarencei from Mexico
After Soto-Adames FN, 2015 Fig.3E

Species cotopaxiana Thibaud, J-M et Najt, J, 1988:726

Species dimorphica Soto-Adames, FN, 2015:14

Trogolaphysa dimorphica from Mexico
After Soto-Adames FN, 2015 Fig.3B

Species distinguenda (Denis, 1931) Thibaud, J-M & Najt, J, 1988:726

Species ecuatorica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:22) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa ecuatoricus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa ecuatoricus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS
             Syn.: Trogolaphysa ecuatoriana(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 ISS

Species entreriosensis Soto-Adames, FN, 2015:23

Trogolaphysa entreriosensis from Argentina
After Soto-Adames FN, 2015 Fig.3F

Species ernesti Cipola NG et Bellini BC in Bellini, BC et Cipola, NG, 2017:170

Trogolaphysa ernesti from Brazil
After Bellini BC & Cipola NG, 2017 Fig.62

Species formosensis da Silva & Bellini, 2015

Species fuelleborni (Börner, 1903) Nguyen, M & Soto-Adames, FN, 2018:182

            Syn.: Paronella fülleborni(sic) Börner, 1903 IOS
             Syn.: Paronella fülleborni(sic) Wahlgren, 1908 ISS
             Syn.: Paronella fülleborni(sic) Handschin, 1929 ISS
             Syn.: Paronella fülleborni(sic) Denis, 1933 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:441 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:471 ISS
             Syn.: Paronella fülleborni(sic) Absolon, K et Ksenemann, M, 1942:13 ISS



Species geminata (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species ghesquierei (Marlier, 1945) Nguyen, M & Soto-Adames, FN, 2018:182

             Syn.: Dicranocentruga ghesourieri(sic) Mitra, SK, 2002:114 ISS
             Syn.: Trogolaphysa ghesquieri(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS
             Syn.: Dicranocentruga ghesouerei(sic) Nguyen, M et Soto-Adames, FN, 2018:183 ISS

Species giordanoae Soto-Adames, FN et Taylor, SJ, 2013:40

Trogolaphysa giordanoae from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.1

Species grassei (Delamare Deboutteville, 1952) Nguyen, M & Soto-Adames, FN, 2018:183

            Syn.: Paronella Grassei(sic) Delamare Deboutteville, 1951 nn IOS

Species guacharo Yoshii, R, 1988:455

Species guayabambensis (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species haitica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:16) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa haiticus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa haiticus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS

Species hauseri Yoshii, R, 1988:457

Species hirtipes (Handschin, 1924) Thibaud, J-M & Najt, J, 1988:725

            Syn.: Paronella carpenteri Denis, 1925 =?


Species hondurasensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa honduraensis(sic) Soto-Adames, FN, 2015:36 ISS

Species innominata (Denis, 1933) Thibaud, J-M & Najt, J, 1988:726

            Syn.: Paronella innominata f. I Denis, 1933
            Syn.: Paronella innominata f. II Denis, 1933
            Syn.: Paronella innominata f. III Denis, 1933


Remark by Mitra, S.K. 2009.07.31:in litt.: Originally, Denis (1933) described 3 forms of "innominata": C.R.I, II, and III. They were considered as different species by Salmon (1964:544), but are infraspecific morphs or varieties.

Species jacobyae Soto-Adames, FN et Taylor, SJ, 2013:47

            Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 IOS


Trogolaphysa jacobyi(sic) from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.24
Trogolaphysa jacobyi(sic) from Belize
Ant.4 with constriction
After Soto-Adames FN & Taylor SJ, 2013 Fig.25

             Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:2 ISS
             Syn.: Trogolaphysa jacobyi(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS




Species jamaicana (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:19) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa jamaicanus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species jataca (Wray, DL, 1953:143) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Dicranocentruga jacata(sic) Mari Mutt, JA, 1979:60 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:28 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas et al., 1986 ISS

Species judithnajtae Nguyen, M et Soto-Adames, FN, 2018:184

Trogolaphysa judithnajtae from Madagascar
After Nguyen M & Soto-Adames FN, 2018 Fig.1

Species laterolineata Soto-Adames, FN, 2015:16

Trogolaphysa laterolineata from Mexico
After Soto-Adames FN, 2015 Fig.3C

Species luquillensis (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species marielouiseae Soto-Adames, FN, 2015:19

             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:1 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:2 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:20 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:10 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:17 ISS
Trogolaphysa marielouisea(sic) from Mexico
After Soto-Adames FN, 2015 Fig.3D



Species marimutti (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

Species maya Mills, HB in Pearse, AS, 1938:184, t.t.

            Syn.: Troglopedetes delamarei Massoud, Z et Gruia, M, 1973:339
Troglopedetes delamarei from Cuba
After Massoud Z & Gruia M, 1973 Fig.7E




Species millsi Arlé, 1939

Species monomaculata (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

Species nacionalica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa nacionalicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species neoannulicornis (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species nigromaculata (Schött, 1903) Nguyen, M & Soto-Adames, FN, 2018:189

             Syn.: Paronella fülleborni(sic) Philiptschenko, 1926 ISS
             Syn.: Paronella fülleborni(sic) Martynova, 1961 ISS
            Syn.: Neophorella dubia Paclt, 1959


  Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Pauliani, 1952

            Syn.: Paronella nigromaculata ssp. pallida Delamare Deboutteville, 1951 nn

Species ocellata Soto-Adames, FN, 2015:25

Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4A
Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4B

Species octosetosa Soto-Adames, FN, 2015:32

Trogolaphysa octosetosa from Mexico
After Soto-Adames FN, 2015 Fig.4F

Species oztotlica (Ojeda, M & Palacios-Vargas, JG, 1984:16) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetes sp. Palacios-Vargas, 1982


             Syn.: Trogolaphysa oztotlicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species palaciosi Soto-Adames, FN, 2015:32

Species paracarpenteri Soto-Adames, FN, 2015:29

Trogolaphysa paracarpenteri from Mexico
After Soto-Adames FN, 2015 Fig.4E
             Syn.: Trogolaphysa paracapenteri(sic) Soto-Adames, FN, 2015:31 ISS


Species penicillata (Schött, 1926) Nguyen, M & Soto-Adames, FN, 2018:190

  Subspecies obscura Delamare Deboutteville, 1950

            Syn.: Paronella penicillata ssp. obscura Delamare Deboutteville, 1951 nn

  Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Paulian, 1952



Species quinquesetosa (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species quisqueyana Soto-Adames, FN, Jordana, R et Baquero, E, 2014:10

Trogolaphysa quisqueyana from the Dominican Republic
After Soto-Adames & al., 2014 Fig.43

Species relicta (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:26) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa relictus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species riopedrensis (Mari Mutt, JA, 1987:403)

Species sauroni Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12

            Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12 IOS
Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 left
Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 right
Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 left
Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 right

             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:17 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:29 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN et Anderson, EW, 2017:40 ISS




Species separata (Denis, 1933) Thibaud, J-M & Najt, J, 1988:726

            Syn.: Dicranocentruga riopedrensis Mari Mutt, JA, 1987:403 =?


Species sexsetosa (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species stannardi Soto-Adames, FN, 2015:8

Trogolaphysa stannardi from Mexico
After Soto-Adames FN, 2015 Fig.3A

Species strinatii Yoshii, R, 1988:452

Species subterranea (Mari Mutt, JA, 1987:403) Mari Mutt, JA & Bellinger, PF, 1996:171

            Syn.: Dicranocentruga subterranean(sic) Mari Mutt, JA, 1987:403 IOS

Species tijucana (Arlé & Guimaraes, 1979) Mari Mutt, JA & Bellinger, PF, 1990:124

             Syn.: Troglopedetes tijucana(sic) Palacios-Vargas et al., 1989 ISS
             Syn.: Trogolaphysa tijuacana Soto-Adames, FN, 2015:39 ISS

Species toroi (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:12) Thibaud, J-M & Najt, J, 1988:723

Species trioculata Soto-Adames, FN, 2015:35

Trogolaphysa trioculata from Mexico
After Soto-Adames FN, 2015 Fig.3G

Species trisetosa (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species variabilis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

Species villiersi (Delamare Deboutteville, 1950) Soto-Adames, FN, 2014:il

            Syn.: Paronella villiersi Delamare Deboutteville & Paulian, 1947 nn
             Syn.: Paronella villersi(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

  Subspecies anomala Delamare Deboutteville, 1951

  Subspecies pallida Delamare Deboutteville, 1950



Species vivieni (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

             Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:181 ISS
             Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

Species wahlgreni (Mitra, SK, 2002:109) Nguyen, M & Soto-Adames, FN, 2018:192

            Syn.: Paronella nigromaculata Wahlgren, 1908


Dicranocentruga wahlgreni from Kenia
After Mitra SK, 2002 Fig.1A
             Syn.: Dicranocentruga whalgreni(sic) Nguyen, M et Soto-Adames, FN, 2018:189 ISS
             Syn.: Trogolaphysa wahlbergi(sic) Nguyen, M et Soto-Adames, FN, 2018:194 lapsus ISS


Species xtolokensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:10) Thibaud, J-M & Najt, J, 1988:723

Species yoshiii (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:14) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetes yoshiius(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:14 IOS


             Syn.: Trogolaphysa yoshius(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa yoshiia(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Trogolaphysa yoshiia(sic) Palacios-Vargas, JG, 2006:10 ISS
             Syn.: Trogolaphysa yoshiia(sic) Soto-Adames, FN et Taylor, SJ, 2013:37 ISS





Genus Troglopedetes Joseph, 1872 M


  Syn.: Paronellidae Cyphoderopsis Bonet, 1931
  Syn.: Paronellidae Cyphoderopsis Wilson, JM, 1982:211
  Syn.: Paronellidae Cypherodopsis(sic) Wilson, JM, 1982:214 ISS


  Syn.: Paronellidae Troglopedetes Wilson, JM, 1982:211 nec Joseph, 1872
Remark by Paclt, 1947, cited from Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Joseph, 1872 is a nomen nudum.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:57: The opinion of Paclt, 1947, that Troglopedetes Joseph is a nomen nudum cannot be upheld, however meagre the description may be.



Species albus Joseph, 1872, t.t.

            Syn.: Troglopedetes pallidus Paclt, 1946 =?
            Syn.: Troglopedetes pallidus Wilson, JM, 1982:211
            Syn.: Troglopedetes pallidus ssp. distinctus Wilson, JM, 1982:211



Genus Troglopedetes Absolon, 1907 nec Joseph, 1872 M


  Syn.: Paronellini Microparonella Carpenter, GH, 1916:43 partim
  Syn.: Troglopedetinae Cyphoderopsis Bonet, F, 1931:362
  Syn.: Paronellidae Cyphoderopsis Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5
  Syn.: Paronellinae Trogolaphysa Bonet, F, 1942:58
  Syn.: Paronellidae Trogolaphysa Salmon, 1964
  Syn.: Paronellidae Trogolaphysa Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5
  Syn.: Troglopedetini Troglopedetina Delamare Deboutteville, C, 1945:40
  Syn.: Paronellidae Dicranocentruga Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5


Remark by Paclt, 1947, cited from Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Absolon, 1907 might be identical with Troglopedetes Joseph, 1872.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Absolon, 1907 is a junior homonym of Troglopedetes Joseph, 1872.

  Syn.: Paronellidae Trolgopedetes(sic) Greenslade, P et Yoshii, R, 2000:139 ISS

Species sp.1 Deharveng, L, 1990:281

Species absoloni Bonet, F, 1931:363

            Syn.: Troglopedetes sp. Arbea & Baena, 2004
            Syn.: Troglopedetes sp. Pérez Fernández & Tinaut, 2005
            Syn.: Troglopedetes sp. Pérez Fernández & Pérez Ruiz, 2006


             Syn.: Troglopedetes Absoloni(sic) Absolon, K et Ksenemann, M, 1942:17 ISS
             Syn.: Troglopedetes Absoloni(sic) Delamare Deboutteville, C, 1945:46 ISS
Troglopedetes absoloni from Spain
After Arbea JI, 2013 Fig.15
Troglopedetes absoloni from Spain
Ant.4 subdivided in 2 parts
After Soto-Adames & al., 2014 Fig.6



Species caeruleus (Carpenter, GH, 1916:43) Deharveng, L & Bedos, A, 1995:22

             Syn.: Paronella coerulea(sic) Handschin, 1929 ISS
             Syn.: Microparonella caeruleus(sic) Yoshii, R et Suhardjno, YR, 1992:48 ISS

Species calvus Deharveng, L et Gers, C, 1993:112

Species canis Christiansen, K, 1957:86

Species cavernicolus Delamare Deboutteville, 1944

             Syn.: Troglopedetes cavernicola(sic) Delamare Deboutteville, 1944 ISS
            Syn.: Troglopedetes wichmanni Delamare Deboutteville, 1950 nn
            Syn.: Troglopedetes wichmanni Wilson, JM, 1982:214 nn
            Syn.: Troglopedetes cretensis Wilson, JM, 1982:214 =?


             Syn.: Troglopedetes cavernicola(sic) Thibaud, J-M et Najt, J, 1988:720 ISS
             Syn.: Troglopedetes cavernicola(sic) da Gama & Ferreira, 2000 ISS
             Syn.: Troglopedetes cavernicola(sic) Wilson, JM, 1982:214 ISS

Species centralis Deharveng, L et Gers, C, 1993:109

Species churchillatus Wilson in Durrant, Smart, Turner & Wilson, 1979

Troglopedetes churchillatus from Nepal
After Wilson JM, 1982 Fig.1A

Species convergens Deharveng, L et Gers, C, 1993:111

Species cretensis Ellis, WN, 1976:306

            Syn.: Troglopedetes wichmanni Delamare Deboutteville, 1950 =?
            Syn.: Troglopedetes wichmanni Wilson, JM, 1982:214 =?


Species dispersus Deharveng, L et Gers, C, 1993:110

Species fredstonei Deharveng, L, 1988:95

Species ildumensis Soto-Adames, FN, Jordana, R et Baquero, E, 2014:8

Species jeanneli (Delamare Deboutteville, C, 1945:41) Thibaud, J-M & Najt, J, 1988:720

            Syn.: Troglopedetina Jeanneli(sic) Jeannel, R, 1945:15 IOS

Species leclerci Deharveng, L, 1990:279

Species longicornis Deharveng, L et Gers, C, 1993:109

Species machadoi Delamare Deboutteville, 1946

             Syn.: Troglopedetes rnachadoi(sic) da Gama, MM, 1988:85 ISS

Species maffrei Deharveng, L et Gers, C, 1993:108

Species maungonensis Deharveng, L et Gers, C, 1993:108

Species microps Deharveng, L et Gers, C, 1993:108

Species minor Hüther, 1983

Species multispinosus Deharveng, L et Gers, C, 1993:110

Species orientalis Cassagnau, P et Delamare Deboutteville, C, 1955:385

Species pallidus Absolon, 1907 tt

             Syn.: Troglopedetes pallidus var. distincta(sic) Absolon, K et Ksenemann, M, 1942:6 ISS


Troglopedetes pallidus from Croatia
2010.11.19 © Lukic, M.
Troglopedetes pallidus from Croatia
2010.11.19 © Lukic, M.

Species paucisetosus Deharveng, L et Gers, C, 1993:111

Species rasendrans Bhattacharjee, 1985

Species ruffoi Delamare Deboutteville, 1951

Species schalleri (Hüther, 1962) Thibaud, J-M & Najt, J, 1988:720

Species tridentatus (Salmon, 1954) Thibaud, J-M & Najt, J, 1988:720

             Syn.: Troglopedetes tridentata(sic) Thibaud, J-M et Najt, J, 1988:720 ISS

Species vandeli Cassagnau, P et Delamare Deboutteville, C, 1955:387

            Syn.: Troglopedetes Vandeli(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:387 IOS

Species vanharteni Barra, J-A, 2006:69


Genus Cyphoderopsis Carpenter, 1917


  Syn.: ... Trogonella Delamare Deboutteville, 1951
  Syn.: Paronellidae Trogonella Thibaud, J-M et Najt, J, 1988:721
  Syn.: Paronellinae Trogonella Soto-Adames, FN, Jordana, R et Baquero, E, 2014:1



Species-group to be defined

Species sp. Deharveng & Bedos, 2000

Species lindbergi (Stach, J, 1960:546) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetina Lindbergi(sic) Stach, J, 1960:546 IOS

Species-group ceylonica Yosii, 1985

Species ceylonica Yosii, 1966

             Syn.: Troglopedetes ceylonica(sic) Wilson, JM, 1982:214 ISS

Species gorumaraensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2019:148

Cyphoderopsis gorumaraensis from India
After Mandal GP & al, 2019 Fig.5

Species nayakensis (Stach, J, 1960:543) Thibaud, J-M & Najt, J, 1988:721

Species nepalensis (Wilson, JM, 1982:221) Thibaud, J-M & Najt, J, 1988:721

Troglopedetes nepalensis from Nepal
After Wilson JM, 1982 Fig.5A

Species sexocellatus Yosii, 1966

            Syn.: Cyphoderopsis 6-ocellata(sic) Yosii, 1966 IOS
             Syn.: Troglopedetes sexoculata(sic) Wilson, JM, 1982:214 ISS
             Syn.: Cyphoderopsis (ceylonica-group) sexoculata(sic) Thibaud, J-M et Najt, J, 1988:721 ISS
             Syn.: Cyphoderopsis sixocellata(sic) Mandal, GP, 2018:124 ISS

Species-group kempi Yosii, 1985



Species cavicola

Species decemoculatus Prabhoo, 1971

             Syn.: Troglopedetes decemoculata(sic) Wilson, JM, 1982:214 ISS
             Syn.: Troglopedetes decemoculata(sic) Wilson, JM, 1982:219 ISS

Species gracilis Carpenter, 1924

Cyphoderopsis gracilis from India
After Mandal GP & Hazra AK, 2009 Fig.42a
             Syn.: Cyphoderopsis gracilils(sic) Mandal, GP et Hazra, AK, 2009:41 ISS


Species kempi Carpenter, 1917 tt

             Syn.: Cyphoderopsis Kempi(sic) Delamare Deboutteville, C, 1945:46 ISS
Cyphoderopsis kempi from India
After Mandal GP & Hazra AK, 2009 Fig.44a



Species khaophang

Species lamottei Delamare Deboutteville, 1950

Trogolaphysa lamottei from Guinea
After Nguyen M & Soto-Adames FN, 2018 Fig.5I

Species madagascarensis (Wilson, JM, 1982:222) Thibaud, J-M & Najt, J, 1988:721

Troglopedetes madagascarensis from Madagascar
After Wilson JM, 1982 Fig.6

Species pauliani (Delamare Deboutteville, 1951) Thibaud, J-M & Najt, J, 1988:721

Species phangnga

Species thachana


Tribus Cyphoderini Börner C, 1906:162 key


Genus Cyphoderus Nicolet H, [1842:63], n.c. Opinion 291 key  key M


  Syn.: ... Podura Fabricius, 1783 partim
  Syn.: ... Lepidocyrthus(sic) Bourlet ISS
  Syn.: Podurelles(sic) Cyphodurus Nicolet, H in Desor, E, 1841:384 nn
  Syn.: Podurelles (Corps linéaires) Cyphodeirus(sic) Nicolet H, 1842:63, nr Opinion291 IOS
  Syn.: Thysanura Beckia Lubbock, J, 1870:279
  Syn.: ... Cyphoderella Börner, 1913 nn
  Syn.: ... Borecus Folsom, 1923
  Syn.: Troglopedetini Camponotus(sic) Salmon, JT, 1964:533 lapsus ISS
  Syn.: Collembola Cyphodems(sic) da Gama, MM, 1988:74 ISS
  Syn.: Collembola Cyphodem(sic) da Gama, MM, 1988:84 ISS
  Syn.: Cyphoderinae Cyphoda Soto-Adames & al, 2008
  Syn.: Cyphoderinae Cyphoda de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47


Remark by ICZN. 2007:226: Emendation of Cyphodeirus. Placed on the Official List of Names in Zoology. Opinion 291.

  Syn.: Cyphoderidae Cyphyoderus(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS



Species sp.1 Zeppelini Filho, D et Castaño Meneses, G, 1995:10

Species sp.2 Bellini, BC et Zeppelini, D, 2004:588

Species adelaideae Womersley, 1934

Species affinis Giard, 1895

Species africanus Womersley, 1929

Species agnotus Börner, C, 1906:180

             Syn.: Beckia albinos(sic) Parona, 1895 =: ISS


             Syn.: Cyphoderus aenotus(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:48 ISS
Cyphoderus cf. agnotus from the Galapagos
After Katz AD & al, 2016 Fig.21B



Species albinus Nicolet, H, 1842:67, t.t., B

            Syn.: Cyphodeirus(sic) albinos(sic) Nicolet, H, 1842:67,tt IOS
Cyphodeirus(sic) albinos(sic) from Switzerland
After Nicolet H., 1842 Pl.7 Fig.7

            Syn.: crystalpoduren Fabricius O, 1783
            Syn.: Lepidocyrtus argentatus Bourlet, C, 1845:67
            Syn.: Beckia argentea Lubbock, J, 1870:279,tt
Beckia argentea from the UK
After Lubbock J., 1873 Pl.23

             Syn.: Beckia argenta(sic) Tömösváry, 1883 ISS
             Syn.: Beckia argenta(sic) Dalla Torre, 1888 ISS
             Syn.: Cyphodeirus(sic) argentus(sic) Vellay, 1899 ISS
             Syn.: Cyphodeirus(sic) (Beckia) argenteus Vellay, 1900 ISS
            Syn.: Cyphoderus affinis Giard, 1895
            Syn.: Tullbergia immaculata Lie-Pettersen, 1896
            Syn.: Cyphodeirus(sic) albus Carl, 1899



             Syn.: Lepidocyrtus albinos(sic) Gervais in Walckenaër, 1844 ISS
             Syn.: Cyphodeirus(sic) Albinos(sic) Bourlet, C, 1845:67 ISS
             Syn.: Cyphoderus albinos(sic) Nicolet, H, 1847:383 ISS
             Syn.: Lepidocyrtus albinos(sic) Lubbock, J, 1870:301 ISS
             Syn.: Beckia albinos(sic) Lubbock, J, 1870:279 ISS
             Syn.: Beckia albinos(sic) Lubbock, J, 1873:149 ISS
Beckia albinos(sic) from the UK
After Lubbock J, 1873 Pl.24
Beckia albinos(sic) from the UK
After Beck J in Lubbock J, 1873 Pl.72

             Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 ISS
             Syn.: Beckia albinos(sic) Parona, 1878 ISS
             Syn.: Beckia albinos(sic) Parona, 1882 ISS
             Syn.: Beckia albinos(sic) Tömösváry, 1883 ISS
             Syn.: Beckia albinos(sic) Brook, 1884 ISS
             Syn.: Beckia (Cyphoderus) albinos(sic) Reuter, 1886 ISS
             Syn.: Cyphoderus albinos(sic) Oudemans, 1887 ISS
             Syn.: Beckia albinos(sic) Parona, 1887 ISS
             Syn.: Beckia albinos(sic) Dalla Torre, 1888 ISS
             Syn.: Beckia albinos(sic) Parona, 1888 ISS
             Syn.: Cyphoderus albinos(sic) Uzel, 1890 ISS
             Syn.: Cyphoderus albinos(sic) Moniez, 1891 ISS
             Syn.: Cyphoderus albinos(sic) Moniez, 1891 ISS
             Syn.: Cyphoderus albinos(sic) Uzel, 1891 ISS
             Syn.: Beckia albinos(sic) Moniez, 1894 ISS
             Syn.: Beckia albinos(sic) Parona, 1895 ISS
             Syn.: Cyphoderus albinos(sic) Lameere, A, 1895:591,B ISS
             Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1896 ISS
             Syn.: Cyphoderus albinos(sic) Poppe & Schäffer, 1897 ISS
             Syn.: Beckia albinos(sic) Parona, 1897 ISS
             Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1898 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 ISS
             Syn.: Cyphoderus albinos(sic) Carl, J, 1899:336 ISS
             Syn.: Cyphoderus albinos(sic) Carl, 1900 ISS
             Syn.: Cyphoderus albinos(sic) Börner, C, 1901:71 ISS
             Syn.: Cyphoderus albinos(sic) Börner, C, 1901:136,B ISS
             Syn.: Cyphoderus albinos(sic) Carl, 1901 ISS
             Syn.: Cyphoderus albinos(sic) Ågren, 1903 ISS
             Syn.: Cyphoderus albinos(sic) Axelson, 1907 ISS
             Syn.: Cyphoderus albinos(sic) Evans, 1908 ISS
             Syn.: Cyphoderus albinos(sic) ssp. borneri(sic) Schille, 1908 ISS
             Syn.: Cyphoderus albinos(sic) Collinge, WE et Shoebotham, JW, 1909:89 ISS
             Syn.: Cyphoderus albinos(sic) Bagnall, 1910 ISS
             Syn.: Cyphoderus albinos(sic) Bagnall, 1910 ISS
             Syn.: Cyphoderus albinos(sic) Axelson, 1911 ISS
             Syn.: Cyphoderus albinos(sic) Carpenter, GH, 1916:47 ISS
             Syn.: Cyphoderus albinos(sic) Womersley, 1923 ISS
             Syn.: Cyphoderus albinos(sic) Denis, 1924 ISS
             Syn.: Cyphoderus albinos(sic) Brown, 1926 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Coutelen, F, 1928:854 ISS
             Syn.: Cyphoderus albinos(sic) Turk, 1938 ISS
             Syn.: Cyphoderus albinos(sic) Geyskens, J, 1945:16,B ISS
             Syn.: Cyphodurus albinos(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
             Syn.: Cyphoderus albinos(sic) Chopard, 1955 ISS
             Syn.: Camponotus(sic) albinos(sic) Salmon, JT, 1964:533 ISS
             Syn.: Cyphoderus albidus(sic) Berg, MP, 2001:xiv ISS
             Syn.: Cyphoderus albinos(sic) Hopkin, SP, 2002:1 ISS
             Syn.: Cyphoderus albinos(sic) Henderickx, H et Vets, V, 2003:9 ISS
Cyphoderus albinus from the UK
2007 © Cornwall, N.J.
Cyphoderus albinus from the UK
2007 © Cornwall, N.J.
Cyphoderus albinus from France
2008.03.15 © Lebeaux, P.
Cyphoderus albinus from Czechia
2008.03.28 © Krásenský, P.
Cyphoderus albinus from Germany
With Platyarthrus hoffmannseggi
2009.04.22 © Schlüter, H.
Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.
 
Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.
Cyphoderus albinus from Sweden
2011.04.11 © Engström, A.
Cyphoderus albinus from France
2012.08.07 © Quintin, C.
Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.
Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.
Cyphoderus albinus from the UK
2012.11.27 © Murray, A.
Cyphoderus albinus from the UK
2012.11.27 © Murray, A.
Cyphoderus albinus juv. from the UK
2012.11.28 © Murray, A.
Cyphoderus albinus from the UK
With Lasius flavus
2012.11.28 © Murray, A.
Cyphoderus albinus from France
Ant.3 as 'pearly bead'
2013.05.12 © Quintin, C.
Cyphoderus albinus from France
Head in dorsal aspect semi-circular
2014.07.28 © Quintin, C.

             Syn.: Cyphoderus albinos(sic) Manu, M, Szekely, L, Oromulu, LV, Barbuceanu, D, Honciuc, V, Maican, S, Fiera, C, Purice, D et Ion, M, 2015:299 ISS
Cyphoderus albinus from Holland
2015.11.14 © van Leeuwen, J.

             Syn.: Cyphoderus albinos(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45,tt ISS
Cyphoderus albinus from Spain
2016.04.22 © Tizon, J.
Cyphoderus albinus from Belgium
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from Belgium
Ant.3 shaped as a 'pearly bead' (*)
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from Belgium
Frontal aspect
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from France
With Lasius flavus
2018.12.19 © Lebeaux, P.
Cyphoderus albinus from France
2019.01.14 © Tiky, S.

  Subspecies apatelus Börner, 1913


             Syn.: Cyphoderus albinus ssp. apaletus(sic) Jacquemart, S(+), 1980:2 ISS

  Subspecies boerneri Schille, 1906


            Syn.: Cyphoderus albinus ssp. Börneri(sic) Schille, 1906 IOS
            Syn.: Cyphoderus albinus Stach, 1919
            Syn.: Cyphoderus albinus Stach, 1922


             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1908 ISS
             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1912 ISS
             Syn.: Cyphoderus Börneri(sic) Börner, 1913 ISS
             Syn.: Cyphoderus albinus ssp. Börneri(sic) Delamare Deboutteville, C, 1948:342 ISS
             Syn.: Cyphoderus albinus f. Boerneri(sic) Cassagnau, 1963 ISS
             Syn.: Cyphoderus albinus ssp. börneri(sic) Jacquemart, S(+), 1980:2 ISS



Species amaranus Brown, JM, 1926:215

Species antennatus Delamare Deboutteville, C, 1945:39

Species antigone Fernando, 1959

Species arcuatus Wahlgren, E, 1906:20

Species arlei Cassagnau, 1963:138

            Syn.: Cyphoderus Arlei(sic) Cassagnau, 1963 IOS

Species asiaticus Yosii, R, 1959:56

Species assimilis Börner, C, 1906:181

            Syn.: Cyphoderus simulans Imms, 1912
            Syn.: Cyphoderus similis Denis, 1931
            Syn.: Cyphoderus subsimilis Paclt, 1959
            Syn.: Cyphoderus assimilis ssp. empodialis Nuñez, 1962


Species bidenticulatus (Parona, 1888) Börner, 1903

            Syn.: Cyphoderus bidenticulatus var. veneris Denis, 1924
            Syn.: Cyphoderus jugoslavicus Denis, 1936


             Syn.: Cyphoderus bidenticulata(sic) Handschin, 1942 ISS
             Syn.: Cyphoderus bidenticulati(sic) Delamare Deboutteville, 1948 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Jantarit & al, 2014 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:263,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:640,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:641 ISS

Species bidenticulatus Womersley, 1934, nec Parona, 1888 s.d.

Species buxtoni Brown, 1920

            Syn.: Cyphoderus Buxtoni(sic) Brown, 1920 IOS


             Syn.: Cyphoderus Buxtoni(sic) Brown, 1926 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, C, 1945:36 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, 1948 ISS



Species caetetus Zeppelini, D et Oliveira, JVLC, 2016:561

Cyphoderus caetetus from Brazil
After Zeppelini D & Oliveira JVLC & al2016 Fig.2

Species canariensis da Gama, MM, 1988:82

Species cuthbertsoni Womersley, 1929

             Syn.: Cyphoderus Cuthbertsoni(sic) Delamare Deboutteville, C, 1945:36 ISS

Species dorsti Jacquemart, 1974 in Peck SB & Jacquemart J, 2011:1 u

Species empodialis Rapoport, 1962

Species folsomi Handschin, 1927

            Syn.: Cyphoderus Folsomi(sic) Handschin, 1927 IOS
            Syn.: Cyphoderus Folsomi(sic) Delamare Deboutteville, 1948 IOS

Species galapagoensis Jacquemart, S, 1976:153

Species ganeensis Tyagi & Baijal, 1979

Species genneserae Carpenter, 1913

Species gisini Gruia, 1967

Species handschini (Delamare Deboutteville, C, 1948:346) Dallai, R, 1973:536

            Syn.: Cyphoderus Handschini Delamare Deboutteville, C, 1948:346 IOS
            Syn.: Cyphoderus assimilis Handschin, 1926




Species holonycha Silvestri, 1918

Species hrdyi Rusek, 1971

Species inaequalis Folsom, JW, 1927:12

Species indicus Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45

Cyphoderus indicus from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.45 Img.1

Species innominatus Mills, HB in Pearse, AS, 1938:186

             Syn.: Cyphoderus inomminatus(sic) Zeppelini, D et Oliveira, JVLC, 2016:568 ISS

Species insularum Carpenter, GH, 1916:47

             Syn.: Cyphoderus bidenticula Handschin, 1942, partim ISS

Species javanus Börner, C, 1906:180

            Syn.: Cyphoderus agnotus Handschin, 1942
            Syn.: Cyphoderus assimilis Yosii, 1953
            Syn.: Cyphoderus assimilis Yosii, R, 1977:162
            Syn.: Cyphoderus javanus f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143
            Syn.: Cyphoderus borneensis Yoshii, 1980


             Syn.: Cyphoderus javanicus(sic) Schött, H, 1917:51 ISS
Cyphoderus javanus from India
After Mandal GP & Hazra AK, 2009 Fig.41a



Species jharkhandensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:47

Cyphoderus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.47 Img.1

Species kawari Jacquemart, S, 1974:33

Species khaochakanus Jantarit, Satasook & Deharveng, 2014

             Syn.: Cyphoderus (bidenticulati(sic)-group) khaochanus(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:58 ISS

Species komareki Rusek, 1961

Species laticlavatus (Stach, J, 1960:545) Wilson, JM, 1982:218

             Syn.: Cyphoderopsis lavaticus(sic) Massoud in Wilson JM, 1982:218 lapsus ISS
             Syn.: Cyphoderopsis lavaticus(sic) Wilson, JM, 1982:218 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:218 ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:219 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Thibaud, J-M et Najt, J, 1988:724 lapsus ISS

Species manuneru Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:259

Species mitumbis Tshelnokov, 1977

Species monopterus Rusek, 1981

Species mucrominimus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:48

Species mucrostrimenus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:54

Species napoensis Thibaud, J-M et Najt, J, 1987:939

Species nigeri Jacquemart, 1980:1

Species omoensis Delamare Deboutteville, C, 1945:35

Species paralbinus Jacquemart, S(+), 1980:3

Species pinnatus (Folsom, 1923) Folsom, JW, 1927:13

Species propinquus Stach, 1963

Species pseudalbinus Schött, H, 1917:50

Species reuteri Loksa, I, 1978:61

Species rubiae Baijal, HN, 1955:536

Species saheli Jacquemart, 1980:3

Species sarojinii Bhattacharjee, 1985

Species similis Folsom, JW, 1927:12

            Syn.: Cyphoderus assimilis Folsom, JW, 1932:71
            Syn.: Cyphoderus similis f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143
            Syn.: Cyphoderus javanus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 =?


Cyphoderus similis from Canada
2007.04.29 © Luk, S.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.09.03 © McClarin, J.
Cyphoderus similis from the USA
2007.09.03 © McClarin, J.
Cyphoderus similis from the USA
2007.09.03 © McClarin, J.
Cyphoderus similis from the USA
2008.04.05 © Murray, T.
Cyphoderus similis from the USA
2008.05.17 © Montgomery, G.
Cyphoderus similis from the USA
2009.12.06 © Justis, S.
Cyphoderus similis from the USA
2009.12.06 © Justis, S.
Cyphoderus similis from the USA
On Camponotus casteneus
2010.10.31 © Hart, J.
Cyphoderus similis from the USA
On Camponotus casteneus
2010.10.31 © Hart, J.
Cyphoderus similis from the USA
2011.12.11 © Kouri, J.
Cyphoderus similis from the USA
2013.05.12 © Liberta, A.
Cyphoderus similis from the USA
In nest of Tapinoma sessile
2015.12.07 © Keim., B
Cyphoderus similis from Canada
With Lasius sp.
2017.06.04 © Guay, P.

Species songkhlaensis Jantarit, Satasook & Deharveng, 2014

Species squamidives Silvestri, 1918

Species subserratus Delamare Deboutteville, 1948

Species subsimilis Delamare Deboutteville, C, 1948:346

Cyphoderus subsimilis from the Ivory Coast
2016.05.10 © Parmentier, T.
Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.
Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.
Distal dental scale as long as bidentate mucro
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Two unpair ungual teeth (*)
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Straight bidentate mucro
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Dens with 4 internal and 6 external scales

Species sudanensis Wahlgren, E, 1906:18

Species sumatranus Yoshii, 1987

Species termitellus Delamare Deboutteville, C, 1948:339

Species trinervoides Paclt, 1965

Species variabilis Jacquemart, 1980:2

Species veneris Delamare Deboutteville, 1923

Species virungiensis Tshelnokov, 1977

Species yoshiiorum Thibaud, J-M et Najt, J, 1987:937

Species yugoslavicus Denis, 1933

Species zoroastris Yosii, 1963


Genus Cyphoda Delamare Deboutteville, C, 1948:360 F


Species sp. Arbea, JI et Blasco-Zumeta, J, 2001:47

Species colura (Börner, C in Schultze, 1908:57) Delamare Deboutteville, 1948

             Syn.: Cyphoda colurus(sic) Delamare Deboutteville, 1948 ISS

Species grassei Cassagnau, P et Delamare Deboutteville, C, 1955:384

            Syn.: Cyphoda Grassei(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:384 IOS

Species heymonsi (Börner, 1906) Delamare Deboutteville, 1948

            Syn.: Cyphoderus Heymonsi(sic) Börner, 1906 IOS
            Syn.: Cyphoderus Heymonsi(sic) Börner, 1913 IOS
            Syn.: Cyphoda Heymonsi(sic) Delamare Deboutteville, 1948 IOS

Species lantohi (Yoshii, 1987)

Species limboxiphia (Börner, 1913)

            Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Rapoport & Izarra, 1962 IOS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Bernava-Laborde & Palacios-Vargas, 2008 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Salazar Martínez, AE, Narváez Beinhorn, MJ et Bernava Laborde, V, 2017:110 ISS




Species maroccana Delamare Deboutteville, 1948:361, t.t.

            Syn.: Cyphoda marocanus(sic) Delamare Deboutteville, 1948 IOS
             Syn.: Cyphoda marocanus(sic) Jacquemart, 1974 ISS
             Syn.: Cyphoderus marocanus(sic) Jacquemart, 1974 ISS



Species natalensis (Börner, 1913) Delamare Deboutteville, 1948

Species nichollsi (Womersley, 1934) Delamare Deboutteville, 1948:364

            Syn.: Cyphoderus Nichollsi(sic) Womersley, 1934 IOS

Species nitricola Rapoport, 1962


Genus Pseudocyphoderus Imms, 1912 M


  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:132 ISS

Species annandalei Imms, 1912, t.t.

            Syn.: Pseudocyphoderus Annandalei(sic) Imms, 1912,tt IOS
             Syn.: Pseudocyphoderus Annandalei(sic) Börner, 1913 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Schött, H, 1917:2 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Delamare Deboutteville, C, 1948:391 ISS
Pseudocyphoderus annandalei from India
After Mandal GP & Hazra AK, 2009 Fig.45a

             Syn.: Pseudocyphoderus annendalei(sic) Mandal, GP et Hazra, AK, 2009:42 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:132 ISS




Species melittophilus Mari Mutt, JA et Bellinger, PF, 1990:130

             Syn.: Pseudocyphoderus melittophilus(sic) Mari Mutt, JA et Bellinger, PF, 1990:130 ISS
             Syn.: Pseudocyphoderus mellitophilous(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:52 ISS

Species squamicaudus Silvestri, 1917

Species wasmanni Börner, 1913

             Syn.: Pseudocyphoderus Wassmanni(sic) Delamare Deboutteville, 1948 ISS



Genus Cyphoderodes Silvestri, 1910 M


Species ceylonicus Silvestri, 1910

Species dubius Börner, 1913

Species mitrai Yoshii, 1987

Species punjabicus Yosii & Ashraf, 1964

Species singaporicus Murphy & Yoshii, 1992

Species xenopus (Börner, 1913) Culik, MP & Zeppelini, DF, 2003:1130

Species yayukae (Yoshii, 1987)


Genus Paracyphoderus Delamare Deboutteville, C, 1948:365 M


Species dimorphus (Silvestri, 1910) Delamare Deboutteville, 1948, t.t.


Genus Megacyphoderus Delamare Deboutteville, C, 1948:365 M


Species gallicus Delamare Deboutteville, 1948:368

Species oraniensis Delamare Deboutteville, 1948:366

Species orientalis (Folsom, 1924) Castaño-Meneses, G, Palacios-Vargas, JG & do Carmo, AFR, 2015:394

Species silvestrii (Denis, J-R, 1924:199) Delamare Deboutteville, C, 1948:367

            Syn.: Cyphoderus Silvestrii(sic) Denis, J-R, 1924:199 IOS


Species termitum (Wahlgren, E, 1906:19) Delamare Deboutteville, 1948


Genus Serroderus Delamare Deboutteville, C, 1948:369 M


Species acanthotermiensis Delamare Deboutteville, 1948:382

Species alfredi (Yosii, R, 1959:56) Yoshii, 1987

Species alvaradoi Selga, 1962

Species aurantius Delamare Deboutteville, 1948

Species bilobatus Delamare Deboutteville, 1948:375

Species brevimucronatus (Silvestri, 1917)

Species delamarei Cassagnau, 1963:147

Species dicuspiditermitis (Yoshii, 1980) Yayuk, 1989:18

Species distinctus (Denis, 1942) Delamare Deboutteville, 1948, t.t.

Species durio Yoshii, 1987

Species guanobia Paulian & Delamare Deboutteville, 1948

Species hozawai (Kinoshita, 1917) Yosii, R, 1977:162

Serroderus hozawai from Japan
2014.10.02 © Hara, Y.

Species humilis Delamare Deboutteville, 1948:371

  Varietas guanobia Delamare Deboutteville & Paulian, 1948

  Varietas aurantia Delamare Deboutteville, 1948

Species interpositus (Denis, 1942) Delamare Deboutteville, 1948

  Forma termitophila Delamare Deboutteville, 1948

Species mango Yoshii, 1992

Species multidentatus Delamare Deboutteville, 1948:356

Species protermiensis Delamare Deboutteville, 1948:381

Species quadridentatus (Delamare Deboutteville, 1948:355)

             Syn.: Cyphoderus quadridentalus Jacquemart, 1980 ISS

Species sabahnus (Yoshii, 1980) Yayuk, 1989:18

Species socialis Delamare Deboutteville, 1948:379

Species spinatus Christiansen, 1957:79

Species spinicaudus Delamare Deboutteville, 1948

Species sublimis Delamare Deboutteville, 1948:374

Species termitophilus Delamare Deboutteville, 1948

Species tridenticulatus Denis, 1948


Genus Cephalophilus Delamare Deboutteville, C, 1948:382 M


Species anoplotermiensis Barra, 1969

Species protermiensis Delamare Deboutteville, 1948:386

Species rectangularis Delamare Deboutteville, 1948:383, t.t.


Genus Calobatinus Silvestri, 1918 M


  Syn.: Calobatina Denis, 1942 Calobatina Denis, 1942 Calobatina Denis, 1942 ISS

Species erythraeus Silvestri, 1918

Species grassei (Denis, 1942) Delamare Deboutteville, 1948

            Syn.: Calobatina Grassei Denis, 1942 IOS
            Syn.: Calobatinus Grassei Delamare Deboutteville, 1948 IOS

Species occidentalis Silvestri, 1918

Species rhadinopus (Börner, 1913) Silvestri, 1918, t.t.


Genus Cyphoderinus Denis, 1942 M


Species fraterculus Denis, 1942, t.t.


Genus Mimoderus Yoshii, 1980 M


Species diusi Yoshii, 1980

Species novaeguinae (Uchida, 1949) Yayuk, RS (sic), 1989:18)

            Syn.: Cyphoderus novaguinae(sic) Uchida, 1949 IOS
            Syn.: Mimoderus novaguinae(sic) Yayuk, RS(sic), 1989:18) IOS

Species saikehi Yoshii, 1980

Species serratus (Schött, H, 1917:52) Greenslade, P in Houston, WWK, 1994:109


Genus Delamarerus Mitra, SK, 1976:645 M


  Syn.: Cyphoderidae Delamareus(sic) Mari Mutt, JA, 1977:230 ISS

Species immsi Mitra, SK, 1976:647, t.t.

Delamarerus immsi from India
After Mitra SK, 1976 Fig.1A
Delamarerus immsi from India
After Mandal GP & Hazra AK, 2009 Fig.46a

Species machadoi (Delamare Deboutteville, 1958) Mari Mutt, JA, 1977:234


Genus Troglobius Palacios-Vargas, JG et Wilson, J, 1990:68 key M


Species albertinoi Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2016:659

Troglobius albertinoi from Brazil
After Cipola NG & Bellini BC in Cipola NG & al, 2016 Fig.1
             Syn.: Troglobius albertonoi(sic) Cipola, NG, de Morais, JW et Bellini, BC, 2016:658 ISS


Species brasiliensis Palacios-Vargas, JG et Zeppelini, D, 1995:173

Species coprophagus Palacios-Vargas, JG et Wilson, J, 1990:69, t.t.

Species ferroicus Zeppelini, D, da Silva, DD et Palacios-Vargas, JG, 2014:3

Troglobius ferroicus from Brazil
Candle-shaped antennal sensilla
After Zeppelini D & al, 2014 Fig.1-2

References