http://www.collembola.org/taxa/frieinae.htm - Last updated on 2017.06.20 by Frans Janssens
Checklist of the Collembola: Frieseinae

Subfamilia Frieseinae Massoud, 1967
Genus Number
of species
Friesea181
Gisinea2
Halofriesea2
Tremoisea3
Frieseinae188
Genera with number of species

  Syn.: Collembola Friesinae(sic) Babenko, AB, 1998:606 ISS
Friesea sp. from Belgium
2010.01.05 © Soors, J.


Genus Friesea von Dalla Torre, 1895 key
  Syn.: ... Triaena Tullberg, 1871 nec Hübner, 1818
  Syn.: Poduridæ(sic) Triæna(sic) MacGillivray, AD, 1893:318 ISS
  Syn.: Thysanura Triæna(sic) Cockerell, TDA, 1894:116 ISS
  Syn.: ... Macgillivraya Grote, 1894 nec Forbes, 1852
  Syn.: ... Polyacanthella Schäffer, 1897
  Syn.: Pseudachorutini Polycanthella(sic) Mills, HB, 1934:20 ISS
  Syn.: ... Polycanthella(sic) Bagnall, 1939 ISS
  Syn.: Collembola Polycanthella(sic) Bagnall, RS, 1940:169 ISS
  Syn.: Pseudachorutinae Polycanthella(sic) Kloet, GS et Hincks, WD, 1945:4 ISS
  Syn.: ... Achorutoides Willem, 1901
  Syn.: ... Achorutoïdes(sic) Willem, V, 1901:260 ISS
  Syn.: Neanuridae Achorutoïdes(sic) Willem, V, 1902:8 ISS
  Syn.: Achorutinae Subantarctica Salmon, JT, 1949:12
  Syn.: ... Prospinanura Wray, 1952
  Syn.: ... Wrayella Salmon, 1956
  Syn.: Hypogastrurinae Neoprospinanura Wray, DL, 1963:101
  Syn.: Anuridae Pseudocolonavis Salmon, JT, 1964:114
  Syn.: ... Spinanurida Salmon, 1969
  Syn.: Anuridae Fresea(sic) Snider, RJ, 1967:188 ISS
  Syn.: Neanuridae Frisea(sic) Palacios-Vargas & Mejía, 1988 ISS
Remark by ICZN. 2007:326: Placed on the Official List of Names in Zoology. Opinion 1049.


  Syn.: Hypogastruridae Frisea(sic) Mandal, GP et Hazra, AK, 2009:85 ISS
  Syn.: Collembola Friese(sic) Chauvat, M, Trap, J, Perez, G, Delporte, P et Aubert, M, 2011:411 ISS
  Syn.: Neanuridae Friesa(sic) Nickerl, J, Helbig, R, Schulz, H-J, Werner, C et Neinhuis, C, 2012:6 ISS
Species acuminata (Denis, 1925) Goto & Lawrence in Kloet & Hincks, 1964:5
            Syn.: Polycanthella(sic) thalassophila Bagnall, 1939
Friesea acuminata from the UK
2016.09.23 © Shaw, P.
Leg;: Trewhella, S.
Friesea acuminata from the UK
2016.09.23 © Trewhella, S.
Det.: Fjellberg, A.
Friesea acuminata from the UK
2016.09.24 © Trewhella, S.

Species aeolica Dallai, R, 1973:491
Species africana Delamare Deboutteville, 1953
Species afurcata (Denis, 1926) Denis, 1927
Species alaskella Fjellberg, 1985
             Syn.: Friesea allaskella(sic) Gao, Y et Palacios-Vargas, JG, 2010:442 ISS
Species albida Stach, 1949
Species albithorax Massoud & Thibaud, 1981
Species anophthalma (Thibaud & Weiner, 1997) Thibaud, J-M & Weiner, WM in Najt, J & Matile, L, 1997:69
Species araucana de Izarra, DC, 1972:94
Species arlei Massoud & Bellinger, 1963
Species arnei Weiner, WM et Najt, J, 1985:248
Species atypica Cassagnau, 1958
Species australica Greenslade, P et Deharveng, L, 1997:323
Species baltica Szeptycki, 1964
Species bispinosa Deharveng, 1981
            Syn.: Friesea simplex Greenslade P, 1990
            Syn.: Friesea sp. Greenslade P, 1990
Species bodenheimeri Börner, 1927
Species bonariensis Izarra, 1965
             Syn.: Friesea bonarensis(sic) Palacios-Vargas, JG, 2005:94 ISS
Species brevicaudata (Schäffer, 1897) Wahlgren, 1906
             Syn.: Friesea brevicauda(sic) Palacios-Vargas, 1986 ISS
Species buyuni Gao, Y et Palacios-Vargas, JG, 2010:441
Friesea buyuni from China
After Gao Y & Palacios-Vargas JG, 2010 Fig.3

Species carlota Christiansen, K et Bellinger, P, 1988:222
Species cassagnaui Simón-Benito, JC et Deharveng L, 1997:453
Species cauchoisi Delamare Deboutteville, 1951
Species cera Christiansen, K et Bellinger, P, 1973:389
Species chiangdaoensis Deharveng & Bedos, 1991
Species christiani Thibaud, 1995
Species claviseta Axelson, 1900:112
            Syn.: Friesea caldaria Guthrie, JE, 1903:95
            Syn.: Friesea sublimis Macnamara, 1921
            Syn.: Friesea claviseta ssp. emucronata Stach, 1922
            Syn.: Friesea bodenheimeri Börner, 1927
Species coiffaiti Cassagnau, P et Delamare Deboutteville, C, 1955:372
            Syn.: Friesea Coiffaiti(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:372 IOS
Species colorata Cassagnau, P, 1958:22
            Syn.: Friesea gadeae(sic) Selga, 1958 IOS
             Syn.: Friesea (Polyacanthella) gadeae(sic) Gisin, H, 1960:74 ISS
             Syn.: Friesea gadeae(sic) Cassagnau, 1961 ISS
             Syn.: Friesea gadeae(sic) Selga, 1962 ISS
             Syn.: Friesea gadeae(sic) Salmon, JT, 1964:263 ISS
             Syn.: Friesea (Polyacanthella) gadeae(sic) Selga, 1971 ISS
             Syn.: Friesea gadeae(sic) Christiansen, 1986 ISS
             Syn.: Friesea gadeae(sic) Jordana & al, 1990 ISS
             Syn.: Friesea gadeae(sic) Arbea, JI et Jordana, R in Ramos, MA et al, 1997:310 ISS
            Syn.: Friesea nietonis Selga, 1958
             Syn.: Friesea nietoi(sic) Selga, 1966 ISS
             Syn.: Friesea nietoi(sic) Selga, 1967 ISS
             Syn.: Friesea (Polyacanthella) nietoi(sic) Selga, 1971 ISS
             Syn.: Friesea nietoi(sic) Franz, 1975 ISS
             Syn.: Pseudocolonavis fagei ssp. Colorata(sic) Salmon, JT, 1964:268 ISS
Species cruchagae Arbea, JI et Jordana, R, 1989:128
Species cubensis Potapov & Banasko, 1985
Species daliensis Tamura, H et Yue, Q, 1998:623
Species danica Fjellberg, A, 1998:66
Species decemoculata Börner, 1903
Species decipiens Steiner, 1958
             Syn.: Friesea decipines(sic) Palacios-Vargas, JG et Traser, G, 1997:104 ISS
Species deharvengi Izarra, 1980
Species denisi Kseneman, 1936
  Varietas anophthalma Kseneman, 1938:461
Species duodecimoculata Denis, 1926
            Syn.: Friesea 12-oculata(sic) Denis, 1926 IOS
             Syn.: Friesea 12-oculata(sic) Massoud, Z, 1967:131 ISS
Species emucronata Stach, 1922
Species espunaensis Arbea, JI et Jordana, R, 1993:9
Species excelsa Denis, JR, 1936:261
Species fagei Denis, 1932
Species fara Christiansen, K et Bellinger, P, 1973:393
Species filii Palacios-Vargas, JG et Mejía, BE, 1988:21
            Syn.: Friesea filli(sic) Palacios-Vargas, JG et Mejía, BE, 1988:21 IOS
             Syn.: Friesea filli(sic) Palacios-Vargas & Gómez-Anaya, 1993 ISS
             Syn.: Friesea fili(sic) Mari Mutt & Bellinger, 1996 ISS
             Syn.: Friesea fili(sic) Palacios-Vargas, JG, 2000:28 ISS
Species flava (Salmon, JT, 1949:13) Massoud, Z, 1967:136
Species florifera Greenslade, P et Deharveng, L, 1997:325
Species furculata Deharveng & Bedos, 1991
Species geminioculata Loksa, I, 1964:85
Species gigliolii Murphy, 1965
Species gracilispina Deharveng & Bedos, 1991
Species grandis Mills, HB, 1934:21
Species grisea (Schäffer, 1891) Wahlgren, E, 1906:6
            Syn.: Achorutoïdes(sic) antarcticus Willem, V, 1901:260
Achorutoïdes(sic) antarcticus from Antarctica
After Willem V, 1902 Pl.II Fig.3

Friesea grisea from Antarctica
2008 © Univ of Waikato.

Species guarinoi Giuga & Jordana, 2013
Species haldanei Rapoport & Maño, 1969
             Syn.: Friesea haldeni(sic) Palacios-Vargas, JG et Traser, G, 1997:103 ISS
Species hammeni Massoud, 1965
Species handschini Kseneman, M, 1938:461
Friesea handschini from Romania
After Dányi L et al., 2010 Fig.1

Species hnaeu Weiner WM, Bedos A et Deharveng L, 2009:509
Friesea hnaeu from New Caledonia
After Weiner WM et al., 2009 Fig.1A

Species hoffmannorum Palacios-Vargas, JG, 1982:46
             Syn.: Friesea hoffmanorum(sic) Mari Mutt & Bellinger, 1990 ISS
Species inermis (Cassagnau P, 1958:21) Massoud, 1967:136
Species inthanonensis Deharveng & Bedos, 1991
Species isabelae Arbea, JI et Jordana, R, 1993:11
Species jaliscoensis Palacios-Vargas, JG et Mejía, BE, 1988:23
Species japonica Yosii, 1954
             Syn.: Friesea Japonica(sic) Gao, Y et Yin, WY, 2005:63 ISS
Friesea (Friesea) japonica from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo13

Species jeanneli Denis, 1947
Species josei Palacios-Vargas, JG, 1986:16
Species judithae Palacios-Vargas, JG, 1986:7
Species jugoslavica da Gama, 1963
Species kanchanaburiensis Deharveng & Bedos, 1991
Species kardosia (Wray, 1952) Christiansen & Bellinger, 1980:282
             Syn.: Friesea kardiosa(sic) Greenslade, P et Deharveng, L, 1997:330 ISS
Species kariae Smolis, A, 2010:37
Friesea kariae from the USA
After Smolis A, 2010 Fig.1
             Syn.: Friesea karie(sic) Smolis, A, 2010:40 ISS
Species kymgangsani Weiner, WM et Najt, J, 1985:250
Species ladeiroi da Gama, 1959
             Syn.: Friesea laderoi(sic) Massoud, Z, 1967:135 ISS
             Syn.: Friesea laderoi(sic) Simón-Benito, JC, 2005:329 ISS
Species lagrecai Dallai, R, 1973:491
Species landwehri Christiansen, K et Bellinger, P, 1980:283
Species laouina Deharveng, L et Hamra-Kroua, S, 2004:141
Species laszloi Palacios-Vargas, JG et Traser, G, 1997:101
Species leei Thibaud & Kim, 1995
Species lijiangensis Tamura, H et Yue, Q, 1998:626
Species lisu Deharveng & Bedos, 1991
Species litoralis Wise, 1964
             Syn.: Friesea littoralis Massoud, 1967 ISS
Species lobulata Palacios-Vargas & Díaz A, 1986:86
Species loicmatilei Weiner & Najt, 2001
Species macuillimitl Palacios-Vargas, 1986:14
Species magnicornis Denis, 1931
             Syn.: Friesea magnicoris(sic) Mari Mutt & Bellinger, 1996 ISS
Species mahajanga Thibaud, J-M, 2008:506
Species major Hamra-Kroua, S, Jordana, R et Deharveng, L, 2009:65
Friesea major from Algeria
After Hamra-Kroua S et al., 2009 Fig.1
Friesea major from Algeria
Hamra-Kroua, S. ©, 2009.06.01

Species malvinensis Najt & Rubio, 1979
Species mandibulata (Salmon, 1969) Bedos, 1994
Species marianoius Palacios-Vargas, JG, 2005:91
Species maritima da Gama, 1964
             Syn.: Friesea maritina(sic) Simón-Benito, JC, 2005:328 ISS
Species massoudi Rusek, 1973
Species mauriesi Cassagnau, P, 1958:22
            Syn.: Friesea Mauriesi(sic) Cassagnau, P, 1958:22 IOS
Species maxima Baijal, 1957, s.d.
Species microphthalma Deharveng & Bedos, 1991
Species millsi Christiansen & Bellinger, 1973:389
Species mirabilis (Tullberg, 1871) Kseneman, 1937, t.t., B
            Syn.: Friesea mirabilis ssp. claviseta Axelson, 1900
            Syn.: Friesea mirabilis f. quadrispina Linnaniemi, 1912
            Syn.: Friesea mirabilis ab. anophthalma Kseneman, 1937
            Syn.: Friesea poseidonis Bagnall, 1939
            Syn.: Friesea poseidonis f. inermis Bagnall, 1939
            Syn.: Friesea poseidonis var. tetracanthella Bagnall, 1939
            Syn.: Friesea poseidonis var. tumidicornis Bagnall, 1939
            Syn.: Friesea mirabilis f. principalis Gisin, 1944
            Syn.: Friesea mirabilis ab. alba Stach, 1949
Friesea mirabilis from Canada
1999 © Janssens, F.
Friesea mirabilis from Canada
1999 © Janssens, F.

             Syn.: Friesea irabilis(sic) Simón-Benito, JC, 2005:330 ISS
Species mistralae Rapoport, EH et Rubio, IR, 1963:101
            Syn.: Friesea mistrali(sic) Rapoport, EH et Rubio, IR, 1963:101 IOS
Friesea mistrali(sic) from Chile
After Rapoport EH & Rubio IR, 1963 Fig.14

             Syn.: Friesea mistrali(sic) Massoud, Z, 1967:137 ISS
             Syn.: Friesea mistralis(sic) Izarra, 1972 ISS
             Syn.: Friesea mistrali(sic) Christiansen, K et Bellinger, P, 1973:392 ISS
             Syn.: Friesea mistrali(sic) Hermosilla & Rubio, 1976 ISS
             Syn.: Friesea mistrali(sic) Hermosilla & Rubio, 1977 ISS
             Syn.: Friesea mistrali(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Friesea mistralis(sic) Izarra & Boo, 1980 ISS
Species monoculata Dunger, 1972
Species montana Cassagnau, P, 1958:24
Species montechristii Dallai, 1969
             Syn.: Friesea albida f. montecristi(sic) Dallai, R, 1973:491 ISS
Species monteiroi Rapoport, 1962
             Syn.: ... monteroi(sic) auctt. ISS
            Syn.: Friesea monteiroi ssp. simplex Cassagnau & Rapoport, 1962
             Syn.: Friesea Monteiroi(sic) Rapoport, EH et Rubio, IR, 1963:101 ISS
Species mucumontana Palacios-Vargas, JG et Díaz, A, 1986:85
Species multispinosa Denis, 1947
            Syn.: Friesea tilbrooki Wise, 1970 =?
Species najtae D'Haese, CA, Stevens, MI et Weiner, WM, 2017:22
Friesea najtae from Australia
After D'Haese CA, Stevens MI & Weiner WM, 2017 Fig.2

Species nauimetztli Palacios-Vargas, JG et Acosta VV, 1994:292
             Syn.: Friesea nahuimetztli(sic) Palacios-Vargas, JG, 2000:28 ISS
             Syn.: Friesea nauimetzli(sic) Gao, Y et Palacios-Vargas, JG, 2010:442 ISS
Species neocaledonica Palacios-Vargas, JG in Tillier, S, 1988:39
Species neptunia Greenslade, P et Deharveng, L, 1997:328
Species nigrimontana Cassagnau, P, 1964:470
            Syn.: Friesea handschini Selga, 1958
            Syn.: Friesea handschini Jordana & al, 1990
Species nigrissima (Salmon, 1954) Massoud, 1967:135
Species nigroviolacea Enderlein, 1909, s.i.
Species ninau Christiansen, K et Bellinger, P, 1992:57
Species normae Weiner, WM et Najt, J, 1985:255
Species octooculata Stach, 1949
             Syn.: Friesea octoculata Rusek, 1977 ISS
             Syn.: Friesea octoculata Szeptycki & Weiner in Razowski, 1990 ISS
Species oleia Christiansen, K et Bellinger, P, 1992:59
Species oligorhopala Caroli, 1914
             Syn.: Friesea oligorhophala(sic) Simón-Benito, JC, 2005:330 ISS
Species onphoriensis Weiner, WM et Najt, J, 1985:252
Species oshoro Uchida & Tamura, 1966
Species oteruelensis Simón-Benito, JC, 2005:327
Species oxfordi (Wray, 1953) Christiansen, K & Bellinger, P, 1980:274
Species pacifica (Yosii, R, 1958:681) Massoud, Z, 1967:133
            Syn.: Conotelsa pacyfica(sic) Yosii, R, 1958:681 IOS
Species paitooni Deharveng & Bedos, 1991
Species palaciosi Deharveng & Bedos, 1991
Species palafoxaliciae Palacios-Vargas, JG, 2005:88
Species parva (Womersley, H, 1936:317) Massoud, Z, 1967:135
Species pati Fjellberg, 1984
Species paula Yosii, 1966
Species pentacantha Mills, HB, 1934:20
             Syn.: Friesea dentacantha(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS
Species peruensis Smolis, A et Skarzynski, D, 2006:44
Species petiti (Delamare Deboutteville, C & Massoud, Z, 1964:382) Massoud, Z, 1967:133
Conotelsa petiti from Madagascar
After Delamare Deboutteville C & Massoud Z, 1964 Fig.1A

Species pins Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:69
Species polla Christiansen & Bellinger, 1973:394
Species pseudodecipiens Arbea, JI et Jordana, R in Ramos, MA et al, 1997:295
Species pyrenaica Cassagnau, 1958
Species quadriocellata Izarra, 1980
Species quadrispina (Cassagnau & Rapoport, 1962) Rapoport & Rubio, 1963
Species quadrispinensis Gao, Y et Yin, WY, 2005:?
             Syn.: Friesea quadrispiensis(sic) Gao, Y et Palacios-Vargas, JG, 2010:440 ISS
Species quinquespinosa (Wahlgren, 1900) Cassagnau, P, 1958:26
            Syn.: Friesea Nauroisi(sic) Cassagnau, P, 1958:26 IOS
Friesea quinquespinosa from Norway
2011 © Fjellberg, A.

Species quinta Christiansen & Bellinger, 1973:392
Species reducta Denis, 1931
            Syn.: Friesea reducta ssp. bonariensis Izarra, 1965
Species rothi Christiansen & Bellinger, 1988:224
Species rubeni Deharveng & Bedos, 1991
Species salmoni Massoud, Z, 1967:134
Species sandzuensis Weiner, WM et Najt, J, 1985:253
Species salti (Salmon, 1954) Massoud, Z, 1967:135
Species santo Weiner WM, Bedos A et Deharveng L, 2009:513
Friesea santo from Vanuatu
After Weiner WM et al., 2009 Fig.3A

Species sensillata Palacios-Vargas & Díaz A, 1986:87
Species septem Weiner WM, Bedos A et Deharveng L, 2009:515
Friesea septem from Vanuatu
After Weiner WM et al., 2009 Fig.4A

Species sexoculata Cassagnau, P, 1958:26
            Syn.: Friesea Cauchoisi(sic) f. 6-oculata(sic) Cassagnau, P, 1958:26 IOS
Species shaanxiensis Gao, Y et Yin, WY, 2005:?
Species simoni Arbea, JI et Jordana, R, 1993:7
Species sp. Vázquez & Palacios-Vargas, 1990:13
Species sp. Palacios-Vargas, 1978
Species sp. nov Palacios-Vargas, 1979
Species sp. nov Palacios-Vargas, 1981:89
Species sphaerulaphora Barra, 1970
Species stachi Kseneman, 1936
Species steineri Simón, 1973
Species stepposa Tshelnokov, 1990
Species sublimis Macnamara, 1921:128
             Syn.: Friesea sublimes(sic) Culik, MP, Martins, DS et Ventura, JA, 2006:4 ISS
Species subterranea Cassagnau, P, 1958:25
            Syn.: Friesea (Polyacanthella) caeca Steiner
            Syn.: Friesea subterranea f. coeca(sic) Cassagnau, 1958 IOS
             Syn.: Friesea subterranea f. coeca(sic) Rubio, 1974 ISS
             Syn.: Friesea subterranea f. coeca(sic) Jordana, R et Asiain, A, 1981:56 ISS
             Syn.: Friesea subterranea f. coeca(sic) Mari Mutt, JA et Bellinger, PF, 1990:36,is ISS
  Subspecies bioculata Jordana, R et Asiain, A, 1981:59
  Subspecies tolosana Cassagnau, 1958
  Subspecies troglophila Cassagnau, 1958
Species tatrica Nosek, 1965
Species tecala (Wray, DL, 1963:101) Christiansen & Bellinger, 1980, i.s.
Species tepetlana Palacios-Vargas, 1986:9
Species tibiotarsalis Deharveng & Bedos, 1991
Species tilbrooki Wise, KAJ, 1970:190
            Syn.: Friesea viennei Deharveng, 1981
Friesea tilbrooki from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.47.

Species topo Greenslade, P, 1995:1996:309
Species tourratensis Cassagnau, P, 1958:23
Species travei Deharveng, 1981
Species truncata Cassagnau, P, 1958:20, B
            Syn.: Friesea mirabilis f. reducta Stach, 1949
            Syn.: Friesea mirabilis Selga, 1971
Species truncatopilosa Rusek, 1971
Species tzontli Palacios-Vargas, JG et Acosta VV, 1994:293
Species valeriae Thibaud, 1993
Species versabilis Barra, 1995:130
Species villanuevai Arbea, JI et Jordana, R, 1991:63
  Subspecies populi Arbea, JI et Jordana, R, 1991:65
Species voisini Deharveng, 1981
Species vtorovi Tshelnokov, 1977
Species wabao Weiner WM, Bedos A et Deharveng L, 2009:511
Friesea wabao from New Caledonia
After Weiner WM et al., 2009 Fig.2A

Species wilkeyi Christiansen & Bellinger, 1973:393
             Syn.: Friesea wilkei(sic) Christiansen & Bellinger, 1973 ISS
             Syn.: Friesea wilkelyi(sic) Palacios-Vargas & Mejía, 1988 ISS
Species woyciechowskii Weiner, 1980
             Syn.: Friesea woyciechowski(sic) Greenslade P, [1996] ISS
Species xitlensis Palacios-Vargas, JG et Acosta VV, 1994:292
Species yosii Prabhoo, 1971
Species yupanquii Izarra, 1973
             Syn.: Friesea yupanquil(sic) Palacios-Vargas, JG et Acosta VV, 1994:292 ISS

Genus Gisinea Massoud, Z, 1965:431
Species delhezi Massoud, Z, 1965:432, t.t., B
             Syn.: Gisinea delhesi(sic) Hubart, J-M, 1982:343,B ISS
Species tiliae Babenko, AB, 1998:606

Genus Halofriesea Yoshii & Sawada, 1997
Species kai (Christiansen, K & Bellinger, P, 1992:55) Yoshii & Sawada, 1997
Species kuroshio Yoshii & Sawada, 1997

Genus Tremoisea Cassagnau, 1973
Species enigmatica Cassagnau, 1973, t.t.
Species israelica Gruia, Poliakov & Broza, 1999
Species ossica Cassagnau, 1973



References