http://www.collembola.org/taxa/heteinae.htm - Last updated on 2017.01.24 by Frans Janssens
Checklist of the Collembola: Heteromurinae

Subfamilia Heteromurinae Absolon & Kseneman, 1942 sensu Zhang F et Deharveng L, 2015:307 key  key
Genus Number
of species
Alloscopus10
Dicranocentrus61
Dicranorchesella4
Genusnov1
Heteromurtrella20
Heteromurus18
Mastigoceras1
Pseudodicranocentrus3
Verhoeffiella8
Heteromurinae126
Genera with number of species
Heteromurinae from Australia
2009.09.25 © Daley, A.


  Syn.: Entomobryidae Heteromurininae(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:153 ISS

Tribus Heteromurini Absolon & Kseneman, 1942 sensu Zhang F & Deharveng L, 2015:309 key
  Syn.: ... Verhoeffiella Bellini, 2014

Genus Dicranorchesella Mari Mutt, JA, 1977:377 key
Species boneti Mari Mutt, JA, 1977:381, t.t.
Species fina Mari Mutt, JA, 1981:21
Species occulta Mari Mutt, JA, 1978:127
Species seminuda Mari Mutt, JA, 1984:819

Genus Dicranocentrus Schött, H, 1893:21 key
  Syn.: Dicranocentrus Dicranocentrella Wray, DL, 1953:140
  Syn.: Heteromuricinae Heteromuricus Imms, 1912
  Syn.: Heteromurus Hetermuricus(sic) Mari Mutt, JA, 1979:544 ISS
Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B
Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B
Dicranocentrus from Peru
2009.01-03.dd © Damiano-Palomino., B

Species amazonicus Bellini, Morais & Oliveira, 2013
Species antillensis Mari Mutt, JA, 1979:57
  Subspecies jamaicensis Mari Mutt, JA, 1981:99
Species assimilis Schött, 1927
Species bicolor Handschin, 1924
Species bidentatus Mari Mutt, JA, 1983:154
Species biseriatus Mari Mutt, JA, 1981:97
            Syn.: Dicranocentrus millsi Mari Mutt, JA, 1979:61
Species capitaneus Mari Mutt, JA, 1985:297
Species celatus Mari Mutt, JA, 1985:105
            Syn.: Dicranocentrus sp. Mari Mutt, 1977
            Syn.: Dicranocentrus millsi Mari Mutt, 1981
             Syn.: Dicranocentrus celarus(sic) Mari Mutt, JA, 1985:109 ISS
Species cercifer (Imms, 1912) Mari Mutt, JA, 1979:543
             Syn.: Heteromuricus crucifer(sic) Schött, 1925 ISS
Heteromuricus cercifer from South-India
After Handschin E, 1929 Fig.48
Heteromuricus cercifer from India
After Mandal GP & Hazra AK, 2009 Fig.78a
Dicranocentrus cercifer from India
Abd.5 with dorsal median cercus
After Mandal GP & Suman KK, 2015 Fig.p.524

Species chenae Ma, Y-T, Chen, J-X et Soto-Adames, FN, [2007]:485
Species chimborazoensis Najt, J, Thibaud, J-M et Mari Mutt, JA, 1988:553
Dicranocentrus chimborazoensis
After Thibaud J-M & D'Haese C, 2010 Fig.9

Species colombiensis Mari Mutt, JA, 1979:23
            Syn.: Dicranocentrus gracilis Winter, 1963 =?
Species deharvengi Mari Mutt, JA, 1981:493
Species delamarei Mari Mutt, JA, 1981:160
Species eburneus Delamare Deboutteville, 1951 nn
Species edicitae Jacquemart, S, 1980:2
Species fasciatus Yosii, 1961
Species fraternus Mari Mutt, JA et Bhattacharjee, RK, 1980:162
Species gapudi Mari Mutt, JA, 1985:303
Species gemellus Mari Mutt, JA, 1985:321
Species gracilis Schött, H, 1893:23, t.t.
Species halophilus Mari Mutt, JA, 1985:320
Species heloisae Arlé & Mendonça, 1982
Dicranocentrus heliosae from Brazil
After Xisto T & de Mendonça MC, 2014 Fig.1

Species indecisus Mari Mutt, JA, 1985:301
Species indicus Bonet, 1930
Dicranocentrus indicus from India
After Mandal GP & Hazra AK, 2009 Fig.36a

Species inermis Schött, 1927
            Syn.: Dicranocentrus insignis(sic) Yosii, 1961 lapsus
Species inermodentes (Uchida, 1944) Mari Mutt, JA, 1979:67
Species janetscheki Yosii, 1971
Species javanus Yoshii & Yayuk, 1989:62
  Subspecies salatiganus Yoshii, R et Suhardjono, YR, 1989:62
Species lawrencei Mari Mutt, JA, 1979:66
Species linnaniemii Börner, 1912
             Syn.: Dicranocentrus Linnaniemii(sic) Delamare Deboutteville, C, 1945:33 ISS
Species litoreus Mari Mutt, JA, 1985:318
Species liuae Xu, G-L et Zhang, F, 2014:3
Dicranocentrus liuae from China
After Xu G-L & Zhang F, 2014 Fig.2

Species longicornis (Carpenter, GH, 1916:36) Handschin, 1929
Species luzonensis Mari Mutt, JA, 1985:317
Species marias Wray, DL, 1953:141
            Syn.: Dicranocentrus silvestrii Denis, 1931
            Syn.: Dicranocentrus silvestrii Denis, 1933
Species meruensis Wahlgren, 1908
             Syn.: Heteromurus (Heteromuricus) Stachi(sic) Denis, 1925 =? ISS
            Syn.: Dicranocentrus aethiopicus Handschin, 1929
            Syn.: Dicranocentrus Jeanneli(sic) Delamare Deboutteville, C, 1945:31 IOS
            Syn.: Dicranocentrus Jeanneli(sic) Jeannel, R, 1945:18 IOS
Species millsi Mari Mutt, JA, 1979:61
            Syn.: Dicranocentrus millsi ssp. cubensis Mari Mutt, JA, 1979:62
            Syn.: Dicranocentrus sp. Mari Mutt, 1978
Species nepalensis Mari Mutt JA in Mari Mutt, JA et Bhattacharjee, RK, 1980:165
Species nigritus Mari Mutt, JA, 1979:23
Species orellanae Jacquemart, S, 1980:6
            Syn.: Dicranocentrus orellanae ssp. cuzcoensis Jacquemart, S, 1980:8
Species orientalis Mari Mutt, JA, 1979:67
Species paramoensis Mari Mutt, JA, 1983:149
            Syn.: Dicranocentrus paramoense(sic) Mari Mutt, JA, 1983:149 IOS
             Syn.: Dicranocentrus paramoense(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:158 ISS
Species peckorum Najt, J, Thibaud, J-M et Mari Mutt, JA, 1988:556
Species platensis (Izarra, 1972, i.s.) Mari Mutt, JA, 1977:248
            Syn.: Tomocerus plumbeus Parona, 1895 =?
            Syn.: Dicranocentrus silvestrii Bonet, 1934
Species problematicus Denis, 1948
Species reunionensis Mari Mutt, JA, 1985:299
Species schoetti Mari Mutt, JA, 1979:56
            Syn.: Dicranocentrus maculatus Winter, 1963 =?
            Syn.: Dicranocentrus muyensis Winter, 1963 =?
            Syn.: Dicranocentrus silvestrii var. circulata Winter, 1963 =?
Species silvestrii Absolon, K, 1903:105
            Syn.: Dicranocentrus Silvestrii(sic) Absolon, K, 1903:105 IOS
            Syn.: Tomocerus plumbeus Parona, 1895
            Syn.: Dicranocentrus silvestrii var. nigrescens Absolon, 1903
            Syn.: Dicranocentrus silvestrii var. annulata Börner, C, 1906:176
            Syn.: Dicranocentrus bicolor Handschin, 1924 =?
            Syn.: Dicranocentrus termitophilus Arlé & Mendonça, 1982 =?
             Syn.: Dicranocentrus Silvestrii(sic) Cassagnau, P in Delamare Deboutteville, C et Rapaport, E, 1963:127 ISS
             Syn.: Dicranocentrus silvestri(sic) Abrantes, EA, Bellini, BC, Bernardo, AN, Fernandes, LH, Mendonça, MC, Oliveira, EP, Queiroz, GC, Sautter, KD, Silveira, TC et Zeppelini, D, 2010:2 ISS
Species simplex Yosii, R, 1959:41
Species singularis Mari Mutt, JA et Bhattacharjee, RK, 1980:168
Species solomonensis Mari Mutt, JA, 1979:68
Species spinosus Prabhoo, 1971
Species stachi (Denis, 1925) Handschin, 1929
             Syn.: Heteromurus major ssp. Stachi(sic) Stach, J, 1930:294 =? ISS
             Syn.: Dicranocentrus? Stachi(sic) Delamare Deboutteville, C, 1945:34 ISS
Species sundanensis Schött, 1925
             Syn.: Dicranocentrus sudanensis(sic) Jacquemart, S, 1980:5 ISS
Species termitophilus Handschin, 1924
Species thaicus Yosii, 1961
Species violaceus Mari Mutt, JA, 1981:496
Species wangi Ma, YT et Chen, J-X, 2007:?
Species yoshiius Mari Mutt, JA, 1988:179
Species zebratus Mari Mutt, JA, 1985:106
            Syn.: Dicranocentrus sp. Mari Mutt, 1977
            Syn.: Dicranocentrus millsi Mari Mutt, 1981

Genus Genusnov Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:154
Species pilosus (Mari Mutt JA in Mari Mutt, JA & Bhattacharjee, RK, 1980:168) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:154

Genus Pseudodicranocentrus Mari Mutt, JA, 1981:91 key
  Syn.: Dicranocentrus circulatus Mari Mutt, JA, 1979:6,gr
Species christianseni (Mari Mutt, JA, 1979:21) Mari Mutt, JA, 1981:93
Species circulatus (Mari Mutt, JA, 1979:21) Mari Mutt, JA, 1981:93, t.t.
Species niger Mari Mutt, JA, 1984:816

Genus Heteromurus Wankel, 1860 nc Opinion 1064, 1976
  Syn.: Podura Ptenura Templeton, 1844
  Syn.: ... Templetonia Lubbock, 1862
  Syn.: Entomobryinae Typhlopodura Absolon, K, 1900:428
  Syn.: ... Heteromurodes Absolon, 1901
  Syn.: Heteromurini Heteromuroder(sic) Mari Mutt, JA, 1980:456 ISS
  Syn.: ... Euheteromurus Börner, 1901
  Syn.: ... Propemesira Salmon, 1942
  Syn.: ... Troglosinella Salmon, 1955, nec Delamare Deboutteville, 1949
  Syn.: ... Heteromurtella Mari Mutt, 1979
  Syn.: Collembola Heteromuris(sic) Christiansen, K, 1958:475 ISS
Remark by ICZN. 2007:384: Placed on the Official List of Names in Zoology. Opinion 1064.

Heteromurus sp. from Russia
2008.10.21 © Macroclub.ru
Heteromurus major from the UK
With 5 secondary antennal segments
2009.07.26 © Barton, T.


  Syn.: Collembola Hetermurus(sic) Mejía-Recamier, BE, Palacios-Vargas, JG et Franco, A, 2012:27 ISS
  Syn.: Heteromurini Heterumurus(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:155 ISS
Heteromurus sp. from the USA
Abd.4 shorter than 2 times abd.3
2016.11.28 © Dobbs, Br.

Species sp.1 Shaw P, 2012:il
Heteromurus nr nitidus from the UK
Descaled revealing intrinsic pigmentation
2012.04.23 © Shaw, P.
Heteromurus nr nitidus from Hungary
Dark intrinsic pigmentation
2012.05.21 © Lásló, D.

Species sp.2 Janssens F & Dobbs B, 2016
Heteromurus sp. from the USA
2016.11.28 © Dobbs, Br.

Species brevicornis Bonet, 1934
Species caophongensis Nguyen, 1995
Species constantinellus Curcic BPM et Lucic LR in Lucic, LR, Curcic, BPM et Mitic, BM, 2007:67P
Species dallaii Nosek & Paoletti, 1981
Species dogieli (Philiptschenko, 1926) Denis, 1933
Species fuscatus (Koch, CL & Berendt, GC in Berendt, GC, 1854:119) -
Podura fuscata from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.127

Species gigans Mari Mutt, JA et Stomp, N, 1980:183
Species major (Moniez, 1889) Handschin, E, 1942:426, B
            Syn.: Podura teres Linnæus, C, 1746:342
            Syn.: Podura plumbea Linnaeus, C, 1761:473
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1762:610
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1764:610
            Syn.: Podura plumbeo(sic) de Geer, C, 1778:31
Podura corpore teriti nigrocaeruleo plumbeo
from Sweden
After de Geer, C., 1778 Tom.VII Pl.3 Fig.1.

            Syn.: pondura(sic) plumbea de Bomare, V, 1780:296
            Syn.: Podura plumbea Fabricius, JC, 1793:66
            Syn.: Podura plumbea Fabricius, JC, 1796:138
            Syn.: Podura (Longæ) plumbea Geoffroy, M, 1799:610
            Syn.: Podura plumbea Walckenær, CA, 1802:4
            Syn.: Podura plumbea Latreille, PA, 1806:166
            Syn.: Podura (Podura) plumbea Latreille, X in Cuvier, G, 1817:162
            Syn.: Podura plumbea de Lamarck, M, 1818:21
            Syn.: Podura plumbea Latreille, Le Peletier de Saint-Fargeau, Serville et Guérin in Latreille, M, 1825:167
            Syn.: Podura (Longæ) violacea Geoffroy, M, 1762:611
            Syn.: Heteromurus caerulescens Börner, 1903
             Syn.: Heteromurus coerulescens(sic) Handschin, E, 1942:426 ISS
             Syn.: Heteromurus Coerulescens(sic) Handschin, E, 1942:426 ISS
            Syn.: Heteromurus melitensis Stach, J, 1924:115
Heteromurus melitensis from Malta
After Stach J, 1924 Taf.I Fig.13

             Syn.: Heteromurus melittensis(sic) Handschin, E, 1942:425 ISS
             Syn.: Heteromurus Melittensis(sic) Handschin, E, 1942:426 ISS
            Syn.: Heteromurus mexicanus Handschin, 1928
            Syn.: Lepidocyrtus lundbladi Agrell, 1939
            Syn.: Heteromurus caucasicus Tshelnokov in Martynova, Tshelnokov & Rasulova, 1974
Heteromurus major from Malta
After Stach J, 1967 Pl.XXXIII Fig.1
Heteromurus (Heteromurus) major
After Mari Mutt, J.A., 1980 Fig.2
Heteromurus (Heteromurus) major
After Mari Mutt, J.A., 1980 Fig.3
Heteromurus major from the UK
2004 © Hopkin, S.P.
Heteromurus major from the UK
2004 © Hopkin, S.P.
Heteromurus major from the UK
2004 © Hopkin, S.P.
Heteromurus major from the UK
2006 © Barton, T.
Heteromurus major from France
2006 © Boldsystems.org
Heteromurus major from the UK
2007 © Cornwall, N.J.
Heteromurus major from Portugal
2007.02.25 © Almeida, J.
Heteromurus major from the UK
2007.03.14 © Barton, T.
Heteromurus major from the UK
2007.09.03 © Barton, T.
Heteromurus major from the UK, juvenile
2007.09.13 © Barton, T.
Heteromurus major from the UK, juvenile
2008.04.27 © Barton, T.
Heteromurus major from the Netherlands
2008.09.16 © Baas, A.H.
Heteromurus major from the Netherlands
2008.10.03 © Baas, A.H.
Heteromurus major from Hungary
2008.10.23 © Keresztes, G.
Heteromurus major from Holland
2011.08.01 © van Duinen, J.
Heteromurus major from Canada
2011.09.20 © Hochhalter, R.
Heteromurus major from the UK
2012.01.25 © Barton, T.
Heteromurus major from the UK
2012.01.25 © Barton, T.
Heteromurus major from La Palma
2012.12.02 © Barton, T.
Heteromurus major from France
2012.12.30 © Sandra.
Heteromurus major juv. from Holland
2013.02.17 © van Duinen, J.
Heteromurus major juv. from Holland
2013.02.25 © van Duinen, J.
Heteromurus major from France
Annulated apical antennal segment
2013.03.16 © Sandra.
Heteromurus major from the UK
Ant.1 subdivided in two parts (1a-1b)
2014.11.14 © Barton, T.
Heteromurus major from France
Relative short 4th abdominal segment
2014.11.23 © Quintin, C.
Heteromurus major from Belgium
2016.01.28 © Huskens, M.-L.
Heteromurus major from Greece
2016.12.16 © Dimitrakopoulos, C.
Heteromurus major from Spain
Ant.1 with basal subdivision (*)
2017.01.01 © Tizon, J.
Heteromurus major from Spain
2017.01.01 © Tizon, J.
Heteromurus major from Belgium
Body scales apically rounded
2017.01.22 © Huskens, M.-L.
Heteromurus major from Belgium
Eyepatch with 4+2 ocelli
2017.01.22 © Huskens, M.-L.

Species nitidus (Templeton, R in Templeton, R & Westwood, JO, 1836:94) Megnin, 1888, B
Podura nitida from Ireland
After Templeton, R, 1836 Pl.XI Fig.5

            Syn.: Podura crystallina Müller OF, 1776
             Syn.: Heterotoma cristallina(sic) Bourlet, C, 1839:396 ISS
             Syn.: Podura cristallina(sic) Bourlet, C, 1839:396 ISS
Templetonia crystallina from the UK
After Lubbock, J., 1873 Pl.20.
Templetonia crystallina from the UK
After Beck J in Lubbock J, 1873 Pl.69


            Syn.: Degeeria margaritacea Nicolet, H, 1842:75
            Syn.: Heteromurus nitidus f. margaritarius Wankel, 1860
             Syn.: Heteromurus nitidus ssp. margaritaria(sic) Jordana, R et Baquero, E, 2010:364 ISS
            Syn.: Lepidocyrtus Albinos(sic) McIntire, SJ, 1867:57
Templetonia nitida from the UK
After McIntire SJ, 1867 Fig.46
After McIntire SJ, 1868
Templetonia nitida from the UK
After McIntire SJ, 1867 Fig.47
After McIntire SJ, 1868


            Syn.: Lepidocyrtus marmoratus Packard, AS, 1873:36
            Syn.: Heteromurus albus Joseph, 1881
            Syn.: Tomocerus niveus Joseph, 1882,sd
            Syn.: Tomocerus Doderii(sic) Parona, 1888,sd
             Syn.: ... doderoi(sic) Gisin, 1960 ISS
            Syn.: Templetonia americana Harvey, 1892
            Syn.: Sinella cavernicola Carpenter, 1895
            Syn.: Templetonia quadrioculata Schött, 1896
            Syn.: Heteromurus hirsutus Absolon, K, 1899:495
            Syn.: Heteromurus margaritatus Carpenter, 1902
            Syn.: Heteromurus tetrophthalmus Börner, 1903
            Syn.: Heteromurus tetrophthalmus Stach, J, 1924:109
             Syn.: Heteromurus tatraphthalmus(sic) Denis, 1930 ISS
            Syn.: Heteromurus nitidus var. paucodentatus(sic) Stach, J, 1930:271 IOS
            Syn.: Heteromurus nitidus var. paucidentatus Stach, J, 1930:295
            Syn.: Heteromurus gradgensis Denis, 1933
            Syn.: Heteromurus nitidus ssp. albicans Durkop, 1934
            Syn.: Heteromurus nitidus ssp. brevicornis Bonet, 1934
            Syn.: Heteromurus nitidus f. quadriocellata Kseneman, M, 1935:14
             Syn.: Pseudosinella Kemneri(sic) Agrell, I, 1936:27 ISS
            Syn.: Pseudosinella Kemneri(sic) var. tridentata Agrell, I, 1936:29
            Syn.: Propemesira duo-oculata(sic) Salmon, 1942
             Syn.: Propemesira duoculata(sic) Mari Mutt, JA, 1980:37 ISS
            Syn.: Heteromurus nitidus ssp. armapollerae Massera, 1952
            Syn.: Heteromurus nitidus var. callaticola Gruia, 1965
             Syn.: Isotoma nitidens(sic) Kolenati, 1858 ISS
            Syn.: Heteromurus nitidus Stach, J, 1924:109
             Syn.: Heteromurus nitida(sic) Salmon, JT, 1945:70 ISS
             Syn.: Heteromurus nididus(sic) Godissart, J et Delhez, F, 1969:38,B ISS
             Syn.: Heteromurus nitida(sic) Poole in Poole & Gentili, 1997 ISS
Heteromurus nitidus from the UK
2004 © Hopkin, S.P.
Heteromurus nitidus from Belgium
2004 © Vuijlsteke, M.
Heteromurus nitidus from Belgium
2006 © Vuijlsteke, M.
Heteromurus nitidus from Germany
2006.12.26 © Kohl, F.
Heteromurus nitidus from Germany
2006.12.26 © Kohl, F.
Heteromurus nitidus from the UK
2007.11.10 © Valentine., B
Heteromurus nitidus from the UK
2008.01.17 © Valentine., B
Heteromurus nitidus from the UK
2008.01.17 © Valentine., B
Heteromurus nitidus from the UK
2008.04.09 © Spears, G.
Heteromurus nitidus from Belgium
2008.06.09 © Vuijlsteke, M.
Heteromurus nitidus from the UK
2008.11.15 © Valentine., B
Heteromurus nitidus from the UK
2008.11.19 © Valentine., B
Heteromurus nitidus from the UK
2009.09.26 © Valentine., B
Heteromurus nitidus from the UK
2011.02.26 © Valentine., B
Heteromurus nitidus from the UK
2012.06.12 © Murray, A.
Heteromurus nitidus from the UK
Small reddish eyepatch
2012.07.18 © Jones, A.
Heteromurus nitidus from the UK
Antenna
2012.07.30 © Murray, A.
Heteromurus nitidus from the UK
2012.08.23 © Murray, A.
Heteromurus nitidus from the UK
2012.09.12 © Jones, A.
Heteromurus nitidus from the UK
Ribbed scales
2012.12.20 © Prior, F.
Heteromurus nitidus from the UK
2013.01.23 © Cobby, P.
Heteromurus nitidus from Slovakia
Annulated apical antennal segment
2013.11.05 © Rahmé, N.
Heteromurus nitidus from Spain
Ant.1 subdivided; reddish eyes
2015.06.30 © Miralles, A.
Heteromurus nitidus from Holland
Predated by Acari
2015.10.11 © Heemskerk, R.
Heteromurus nitidus from the USA
Ribbed scales
2016.07.07 © Raymond-Léonard, J.L.
Heteromurus nitidus from the USA
Mucro with back-tilted subapical tooth, and basal spine
2016.07.08 © Raymond-Léonard, J.L.
Heteromurus nitidus from Belgium
2016.12.04 © Huskens, M.-L.

Species noseki Mari Mutt, JA et Stomp, N, 1980:186
Heteromurus noseki from Portugal
After Mari Mutt JA & Stomp N, 1980 Fig.3C

Species peyerimhoffi Denis, 1937
Species pulcher (Koch, CL & Berendt, GC in Berendt, GC, 1854:119) -
Podura pulchra from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.128

Species sexoculatus Brown, JM, 1926:213
            Syn.: Heteromurus hexophthalmus Denis, 1948
Species sinensis Liu & Li, 1999
Species uzicensis (Lucic & Curcic, 2002) Lucic & al., 2002
Species variabilis Martynova in Martynova, Tshelnokov & Rasulova, 1974
Heteromurus (Heteromurus) variabilis
After Mari Mutt, J.A., 1980 Fig.9



Genus Verhoeffiella Absolon, K, 1900:429
  Syn.: Entomobryinae Typhlopodura Absolon, K, 1900:428
Verhoeffiella sp. from Croatia
2010.11.19 © Lukic, M.

Species absoloni Kseneman, 1938
Species anagastumensis Lucic LR in Lucic, LR, Tomic, VT, Brajkovic, MM et Curcic, SB, 2008:298
Heteromurus (Verhoeffiella) anagastumensis from Montenegro
After Lucic LR et al, 2008 Fig.1

Species cavicola Absolon, K, 1900:430, t.t.
Species constantius (Curcic BPM & Lucic LR in Lucic, LR, Curcic, BPM & Tomic, VT, 2007:72P) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:155
Heteromurus (Verhoeffiella) constantius from Herzegovina
After Lucic LR & al, 2007 Fig.1

Species dallaii (Nosek, J & Paoletti, MG, 1985:271) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:155
             Syn.: Verhoeffiella dallai(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:155 ISS
Species hispanicus (Bonet, F, 1931:354) Salmon, JT, 1964:478
            Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Bonet, F, 1931:354 IOS
             Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Lucic, LR, Curcic, BPM et Tomic, VT, 2007:71P ISS
             Syn.: Heteromurus (Verhoeffiella) hispanica(sic) Lucic, LR, Tomic, VT, Brajkovic, MM et Curcic, SB, 2008:298 ISS
Species longicornis (Absolon, K, 1900:428) Thibaud, J-M & Deharveng, L in Jubertie, C & Decu, V, 1994:268
Heteromurus (Verhoeffiella) longicornis from Austria
After Absolon K, 1900
Verhoeffiella longicornis from Croatia
2010.11.19 © Bedek, J.
Verhoeffiella longicornis from Croatia
2010.11.19 © Lucic, M.
Verhoeffiella longicornis from Austria
After Absolon K, 1900 in Cipola NG & al, 2016 Fig.2

Species medius Loksa & Bogojevic, 1967


Genus Alloscopus Börner, C, 1906:177 key  key
  Syn.: ... Indoscopus Prabhoo, 1971
Species aspinosus (Prabhoo, 1971) Yoshii, R & Suhardjono, YR, 1989:64
Species deharvengi (Mari Mutt, JA, 1985:338) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:160
            Syn.: Alloscopus tetracanthus Yosii, 1959 =?
Species fallax Yoshii, R et Suhardjono, YR, 1992:35
Species multispinatus (Mari Mutt, JA, 1982:86) Yoshii, R & Suhardjono, YR, 1992:34
            Syn.: Heteromurus (Alloscopus) tenuicornis Handschin, 1925 =?
Species spinosus (Prabhoo, 1971) Yoshii, R & Suhardjono, YR, 1989:64
Species strebeli Winter, 1966
            Syn.: Alloscopus strebeli Winter, 1963 nn
Species tenuicornis (Börner, C, 1906:177) Handschin, 1928, t.t.
            Syn.: Templetonia sp. Oudemans, 1890 =?
            Syn.: Alloscopus tetracanthus Yosii, 1960 =?
            Syn.: Alloscopus tetracanthus Gapud, 1971
            Syn.: Heteromurus yosiius Yoshii, R et Suhardjono, YR, 1989:64
            Syn.: Heteromurus (Alloscopus) tenuicornis Stach, J, 1924:109
Species tetracanthus (Börner, C, 1906:177) Handschin, 1928
            Syn.: Heteromurus (Alloscopus) tetracantha(sic) Börner, C, 1906:177 IOS
            Syn.: Templetonia sp. Oudemans, JT in Weber, M, 1890:88
             Syn.: Heteromurus tetracantha(sic) Imms, 1912 ISS
             Syn.: Heteromurus (Alloscopus) tetracantha(sic) Stach, J, 1924:109 ISS
             Syn.: Alloscopus tetracantha(sic) Prabhoo, 1969 ISS
             Syn.: Alloscopus tetracantha(sic) Yoshii & Yayuk, 1989 ISS
             Syn.: Heteromurus (Alloscopus) tetracantha(sic) Mandal, GP et Hazra, AK, 2009:68 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:68 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:72 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:76 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:80 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:125 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:127 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:128 ISS
             Syn.: Alloscopus tetracantha(sic) Mandal, GP et Hazra, AK, 2009:205 ISS
Alloscopus tetracantha(sic) from India
After Mandal GP & Hazra AK, 2009 Fig.75a

Species thailandensis (Mari Mutt, JA, 1985:336) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:161
Species yosiius (Mari Mutt, JA, 1985:336) Yayuk, RS (sic), 1989:19

Genus Heteromurtrella Mari Mutt, JA, 1979:214 key  key
  Syn.: Heteromurini Alloscopus Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:157 lapsus
Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.
Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.
Heteromurtrella sp. from the Philipinnes
2009.03.26 © Tamayo, A.

  Syn.: Heteromurini Heterumurtrella(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:165 ISS
Species abrupta (Salmon, 1951) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:165
            Syn.: Heteromurus? sp. Yayuk, RS(sic), 1989:19
Species affinis (Mari Mutt, JA, 1987:575) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166
Species anae Cipola NG in Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:175
Heteromurtrella anae from Brazil
After Cipola NG in Cipola NG & al, 2016 Fig.22

Species barrai (Mari Mutt, JA, 1979:221) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166
            Syn.: Heteromurus mirificus Barra, 1968 nec Salmon, 1954
Species caeca (Najt, J, Thibaud, J-M & Mari Mutt, JA, 1988:560) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:166
Species diommata (Denis, 1931) Palacios-Vargas, 1992
Species dubia (Delamare Deboutteville in Delamare Deboutteville & Paulian, 1952) Castaño-Meneses, G, Palacios-Vargas, JG & do Carmo, AFR, 2015:392
            Syn.: Heteromurus dubius Delamare Deboutteville, 1948a nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1948b nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1951a nn
            Syn.: Heteromurus dubius Delamare Deboutteville, 1951b nn
             Syn.: Ptenura dubius(sic) Salmon, JT, 1964:479 ISS
Species echinata (Mari Mutt, JA, 1983:158) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:165
             Syn.: Heteromurtrella echinatus(sic) Cipola, NG, de Lima Oliveira, FG, de Morais, JW et Bellini, BC, 2016:167 ISS
Species greensladeae (Mari Mutt, JA, 1987:572) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:167
Species livida (Salmon, 1954) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172
Species mirifica (Salmon, 1954) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172
Species nitens (Yosii, R, 1964:14) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172
Heteromurus (Heteromurtrella) nitens from the Galagapos
After Katz AD et al, 2016 Fig.5A

Species pruinosa (Mari Mutt, JA, 1983:163) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:172
Species puertoricensis (Mari Mutt, JA, 1979:220) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173, t.t.
Heteromurtrella puertoricensis from Puerto Rico
After Mari Mutt JA, 1979 in Cipola NG & al, 2016 Fig.3

Species schoetti (Denis, 1931) Palacios-Vargas, 1992
            Syn.: Heteromurus schoetti f. tridentata Denis, 1931
            Syn.: Heteromurus schoetti f. quadridentata Denis, 1931
            Syn.: Heteromurus bidentatus Denis, 1931
            Syn.: Heteromurus nitidus Winter, 1963
Species similis (Barra, 1968) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173
Species stannardi (Mari Mutt, JA, 1976:326) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:173
Species subdubia (Barra, 1968) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174
             Syn.: Heteromurus subduvius(sic) Mari Mutt, JA, 1976:326 ISS
Species tihuiensis (Mari Mutt, JA, 1985:145) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174
Species zairensis (Chelnokov, 1977) Cipola, NG, de Lima Oliveira, FG, de Morais, JW & Bellini, BC, 2016:174

Tribus Mastigocerini Mari Mutt, 1980

Genus Mastigoceras Handschin, 1924
Species camponoti Handschin, 1924, t.t.



References