http://www.collembola.org/taxa/isotinae.htm - Last updated on 2019.02.27 by Frans Janssens
Checklist of the Collembola: Isotominae

Subfamilia Isotominae Schäffer, 1896

Genus Number
of species
Aackia1
Acanthomurus6
Agrenia13
Antarcticinella1
Araucanocyrtus1
Axelsonia8
Azoritoma1
Cheirotoma1
Chionobora1
Desoria102
Folsomotoma9
Gnathisotoma5
Halisotoma8
Heteroisotoma6
Hydroisotoma1
Isotoma76
Isotomedia1
Isotomurus70
Kaylathalia1
Marisotoma3
Metisotoma5
Misturasotoma1
Mucronia2
Myopia1
Najtia1
Paracerura10
Parisotoma27
Procerura13
Protodesoria1
Protoisotoma2
Psammisotoma6
Pseudisotoma8
Pseudosorensia4
Pteronychella4
Rhyniella1
Semicerura3
Sericeotoma1
Setocerura6
Skadisotoma1
Spinocerura2
Tibiolatra1
Tomocerura6
Vertagopus28
Isotominae449
Genera with number of species
  Syn.: Entomobryidae Isotominini(sic) Börner, C, 1901:43 ISS
  Syn.: Collembola Isotomiens(sic) Denis, J-R, 1924:231 ISS
Isotominae from Belgium
2006 © Vuijlsteke, M.
Isotoma viridis from the USA
2009.03.10 © Cowen, R.





Genus Protodesoria Christiansen, KA et Nascimbene, P, 2006:346, -

Species granda Christiansen, KA et Nascimbene, P, 2006:346, -


Genus Tibiolatra Salmon, JT, 1941:320


Tibiolatra latronigra from New Zealand
After Salmon, J.T. 1941 Pl.49 Fig.165.
Species latronigra Salmon, JT, 1941:320, t.t.


Genus Acanthomurus Womersley, 1934

Acanthomurus sp. from Tasmania
2014.04.29 © Murray, A.

Species sp.nov. Greenslade, P in Dallai, R, 1989:116

            Syn.: Isotomurus chiltoni Womersley, 1934
            Syn.: Isotomurus chiltoni Womersley, 1937
            Syn.: Isotomurus chiltoni Norris, 1938
            Syn.: Isotomurus chiltoni Womersley, H, 1939:149
            Syn.: Isotomurus chiltoni Salmon, 1949 partim
            Syn.: Isotomurus chiltoni Stach, 1949 partim
            Syn.: Isotomurus chiltoni Salmon, JT, 1964:361 partim
            Syn.: Isotomurus chiltoni Wallace & Mackerras, 1970
            Syn.: Isotomurus chiltoni Greenslade, P in Houston, WWK, 1994:64
            Syn.: Isotoma chiltoni Greenslade, P in Houston, WWK, 1994:74


Acanthomurus sp. from Tasmania
2017.08.22 © Ellingsen, K.

Species alpinus Salmon, JT, 1941:317

Acanthomurus alpinus from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.169.

  Subspecies obscuratus Salmon, JT, 1943:381

Acanthomurus alpinus
subsp. obscuratus from New Zealand
After Salmon J.T., 1943 Pl.41 Fig.34.

Species plumbeus Womersley, 1934, t.t.

  Subspecies lineatus Womersley, 1934

Species rivalis Wise, 1964

Species setosus Salmon, JT, 1941:319

Acanthomurus setosus from New Zealand
After Salmon JT, 1941 Pl.49 Fig.162
Acanthomurus setosus? from New Zealand
2018.09.09 © Schulz, K.

  Subspecies violaceus Salmon, JT, 1941:320

Species womersleyi Salmon, JT, 1941:318

Acanthomurus womersleyi from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.176.


Genus Axelsonia Börner, 1906

Species crassicornis (Schött, 1893) Yayuk, RS (sic), 1989:8

Species johnstoni Jordana, 1997

Species littoralis (Moniez, 1890) Denis, 1923

            Syn.: Isotoma nitida Handschin, E in Dahl, F, 1929:60 =?
            Syn.: Axelsonia thalassophila Handschin, E in Dahl, F, 1929:60
            Syn.: Axelsonia australis Bagnall, 1939


             Syn.: Isotoma litoralis(sic) Moniez, 1894 ISS
             Syn.: Axelsonia litoralis(sic) Handschin, E in Dahl, F, 1929:60 ISS
             Syn.: Axelsonia litoralis(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
Axelsonia littoralis from Holland
2018.09.30 © van Leeuwen, J.
Axelsonia littoralis from Holland
2018.09.30 © van Leeuwen, J.
Axelsonia littoralis from Portugal
2019.01.06 © Almeida, J.
Axelsonia littoralis from Portugal
2019.01.06 © Almeida, J.
Axelsonia littoralis from Portugal
2019.01.09 © Almeida, J.



Species nitida (Folsom, JW, 1899:264) Börner, 1906

Isotoma nitida from Japan
After Folsom JW, 1899 Pl.1 Fig.14

            Syn.: Axelsonia littoralis Yosii, 1955
            Syn.: Axelsonia littoralis Yosii, R, 1977:155
            Syn.: Isotoma (Isotoma) pteromucronata Uchida, H, 1965:88
            Syn.: Isotoma (Isotoma) pteromucronata Yosii, R, 1977:155



Species sarahae Christiansen, K et Bellinger, P, 1992:151

Species thalassophila Börner, 1906, t.t.

Species tubifera Strenzke, 1958

Species yinii Huang & Liang, 1992


Genus Isotomurus Börner, 1903 key  key  key

  Syn.: Collembola Istoma(sic) Childs, GH, 1915:21 ISS
  Syn.: ... Isotama(sic) Stach, J, 1924:114 ISS
  Syn.: ... Proisotomurus Womersley, 1934


  Syn.: ... Isotomorus(sic) Stach, J, 1924:115 ISS
  Syn.: Isotomidae Isotomourus(sic) Baijal, HN, 1958:355 ISS
  Syn.: Isotomidae Isotomorus(sic) Poinsot, N, 1972:687 ISS
  Syn.: Isotomidae Isomurys(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS
Remark by ICZN. 2007:420: Placed on the Official List of Names in Zoology. Opinion 291.

Isotomurus sp. from the USA
2007 © McClarin, J.
Isotomurus sp. from Norway
Triangular head shape
After Fjellberg A, 2010 p.2
Isotomurus sp. from Norway
3+3 macrosetae on abd.4 in triangle
After Fjellberg A, 2010 p.2
Isotomurus palustris from the UK
Triangular head shape
2010.02.08 © Robertson, A.
Isotomurus graminis from Sweden
Difference between bothriotrichum (bot) and macroseta (mac)
2011.09.24 © Hall, K.
Isotomurus sp. from the UK
2011.12.20 © Kent, J.J.
Isotomurus sp. from the UK
Pigmented clypeus
2012.12.18 © Phillips, E.

  Syn.: Isotomidae isotomurus(sic) Glime, JM, 2015:26 ISS
Isotomurus sp. juv. from Holland
2015.01.20 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.
Isotomurus pseudopalustris from Belgium
Macrosetae only present on posterior body segments
2016.01.04 © Huskens, M.-L.
Isotomurus pseudopalustris from Belgium
Macrosetae on abd.4 in triangular constellation
2016.01.04 © Huskens, M.-L.
Isotomurus graminis from the UK
Mucro with 4 teeth (outer aspect)
2016.01.26 © Shepherd, M.J.
Isotomurus maculatus? from Belgium
Ovoid post-antennal organ (PAO)
2016.02.25 © Huskens, M.-L.
Isotomurus sp. from Holland
Embryo in egg
2017.04.28 © van Duinen, J.
Isotomurus sp. from the UK
Ovoid post-antennal organ (PAO)
2017.12.23 © Cann, A.J.
Isotomurus sp. from Belgium
Ocelli B-D-E in a straight line
2018.02.13 © Huskens, M.-L.
Isotomurus sp. from France
Abd.4 with 3 dorsolateral macrosetae in triangular constellation
2018.11.25 © Tiky, S.
Isotomurus pseudopalustris f. aquatilis from the UK
Mucro with 3 teeth + lateral ridge (inner aspect)
2019.01.21 © Cann, A.J.
Isotomurus nr pseudopalustris from Japan
With Sminthurides sp. juv.
2019.01.20 © Hara, Y.



Species aetnensis Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:116

Isotomurus aetnensis
2001 © Carapelli, A. et al.
Isotomurus nr aetnensis from the Azores
2010.09.22 © E.D.E.N.

Species afghanicus Yosii, 1966

Isotomurus afghanicus from Afghanistan
After Potapov M, 2001 Fig.374

Species alticolus (Carl, J, 1899:313) Handschin, 1919

             Syn.: Isotomurus alticola(sic) Kloet, GS et Hincks, WD, 1945:7 ISS
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate L Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.3.

             Syn.: Isotomurus alticola(sic) Rusek, 1977 ISS
             Syn.: Isotomurus alticola(sic) Dallai & Malatesta in Calderini, 1995 ISS
Isotomurus alticolus from Italy
2001 © Carapelli, A. et al.



Species annectens Yosii, 1963

Species antennalis Bagnall, 1940, B

             Syn.: Isotoma antenalis(sic) Camousseight, A, 1999:6 ISS
Isotomurus antennalis
2007 © Fjellberg, A.
Isotomurus antennalis from the UK
2010.04.04 © Botting, J.



Species atreus Christiansen & Bellinger, 1980

Species balteatus (Reuter, OM, 1876:86) Handschin, E in Dahl, F, 1929:73

Isotoma palustris var. balteata from Sweden
After Schött H, 1893 Taf.V Fig.10
Isotomurus palustris f. balteata from Poland
After Stach, J., 1947 Plate XLVIII Fig.6
Isotomurus balteatus from Korea
After Yosii R & Lee C-E, 1963 Fig.2A
Isotomurus palustris f. balteata from Poland
After Potapov M, 2001 Fig.371
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus from India
After Mandal GP & Hazra AK, 2009 Fig.17a
             Syn.: Isotomurus palustris var. balteata(sic) Pfliegler, WP et Janssens, F, 2016:202 ISS
             Syn.: Isotomurus balteaus(sic) Yadav, RS, 2017:11058 ISS
Isotomurus balteaus(sic) from India
After Yadav RS, 2017 Img.2



Species beskidensis Rusek, 1963

Species bimaculatus Ågren, 1903

Species bimus Christiansen, K et Bellinger, P, 1980:679

Isotomurus (Isotomurus) bimus from the USA
After Christiansen K & Bellinger P, 1980 Fig.541H
            Syn.: Isotomurus palustris var. balteatus Folsom, 1937
             Syn.: Isotomurus palustris ssp. balteata(sic) Brimley, 1938 ISS
             Syn.: Isotomurus palustris ssp. balteata(sic) Mills, 1938 ISS
            Syn.: Isotomurus palustris f. balteatus Maynard, EA, 1951:123
Isotomurus palustris f. balteatus from the USA
After Maynard EA PlateXXII Fig.222

            Syn.: Isotomurus palustris ssp. balteatus Snider, RJ, 1967:213
Isotomurus palustris ssp. balteatus from the USA
After Snider RJ, 1967 Backcover

            Syn.: Isotomurus tricolor Hart, 1974 =?



Isotomurus bimus from the USA
2007.03.31 © Babin, P.
Isotomurus bimus from the USA
2009.07.05 © Zungri, D.
Isotomurus bimus juv. from the USA
2009.07.06 © Zungri, D.
Isotomurus bimus from the USA
2010.08.03 © Zungri, D.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.11.01 © Emile.
Isotomurus bimus from the USA
2015.07.22 © Dankowicz, E.
Isotomurus bimus from the USA
2016.12.24 © Montgomery, G.
Isotomurus bimus from the USA
2017.08.13 © Lee, C.
Isotomurus nr bimus from Holland
2017.09.21 © Wieringa, P.
Isotomurus nr bimus from Holland
With incomplete pigmentation pattern
2017.09.21 © van Duinen, J.
Isotomurus nr bimus from Holland
With incomplete pigmentation pattern
2017.09.21 © van Duinen, J.
Isotomurus nr bimus from Holland
2017.10.01 © van Duinen, J.
Isotomurus nr bimus from Holland
2017.10.01 © van Duinen, J.
Isotomurus bimus from the USA
2017.12.26 © Abbott, J.

Species cassagnaui Deharveng & Lek, 1993

Isotomurus cassagnaui
2001 © Carapelli, A. et al.

Species chaos Potapov & Babenko, 2011

Species chiltoni (Carpenter, GH, 1925:9) Womersley, H, 1936:321

Species cibus Christiansen & Bellinger, 1980

Species denticulatus Kos, 1937

Species dhanbadensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:100

Isotomurus dhanbadensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.2 Fig.1

Species festus Potapov, M, Porco, D et Deharveng, L, 2018:385

Isotomurus festus from FE Russia
After Potapov M & al, 2018 Fig.1
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.18
Isotomurus festus from FE Russia
Dark form
After Potapov M & al, 2018 Fig.19
Isotomurus festus from FE Russia
Pale banded form
After Potapov M & al, 2018 Fig.17
Isotomurus festus from FE Russia
Dark banded form
After Potapov M & al, 2018 Fig.20
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.26low
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.25low
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.25mid
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.26up
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.25up

Species fucicolus (Schött, H, 1893:66) Axelson, 1905

Isotoma palustris var. fucicola from Sweden
After Schött H, 1893 Taf.V Fig.6
            Syn.: Isotoma fucicola Reuter, 1891 il
            Syn.: Isotoma janthina Vogler, 1896
            Syn.: Isotomurus italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:120
            Syn.: Isotomurus italicus Potapov, M, 2001:331



             Syn.: Isotomurus palustris ssp. fucicola(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris f. fascicola(sic) Womersley, H, 1939:149 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Stach, J, 1947:460 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Yosii, R, 1956:65 ISS
             Syn.: Isotomurus palustris var. fucicola(sic) Gisin, H, 1960:214 ISS
Isotomurus fucicolus
2001 © Carapelli, A. et al.


             Syn.: Isotomurus fucicola(sic) Fanciulli, PP, Summa, D, Dallai, R et Frati, F, 2001:251 ISS
Isotomurus fucicola(sic) from Scandinavia
2007 © Fjellberg, A.

             Syn.: Isotomurus fucicola(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
Isotomurus fucicolus from Belgium
2017.06.12 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.06.29 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.07.07 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
Bluish antennae
2017.07.07 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
Facial pattern
2017.08.04 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
Cephalic pattern
2017.08.05 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
Abdominal dorsal pattern
2017.08.05 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
2017.08.10 © Huskens, M.-L.
Isotomurus fucicolus from France
2017.09.29 © Tiky, S.
Isotomurus fucicolus from France
2018.05.21 © Tiky, S.



Species fucicolus2 da Gama, 1964

            Syn.: Isotomurus (gallicus-group) fucicola(sic) da Gama, 1964 IOS
             Syn.: Isotomorus(sic) fucicola(sic) Poinsot, N, 1972:688 ISS
             Syn.: Isotomurus fucicola(sic) Poinsot, N, 1972:689 ISS
Isotomorus(sic) fucicola(sic) from France
After Poinsot N, 1972 Fig.6D





Species fuscus (Salmon, JT, 1944:145) Greenslade, P, 2003:1, s.i.

Species gallicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:120

Isotomurus gallicus from France
2001 © Carapelli, A. et al.
             Syn.: Isotomurus palustris var. balteata(sic) Handschin, E, 1942:445 ISS
            Syn.: Isotomurus balteatus Poinsot, N, 1972:689
            Syn.: Isotomurus balteatus Ellis, WN, 1974:121
            Syn.: Isotomurus balteatus Poinsot-Balaguer & Ferard, 1983
            Syn.: Isotomurus balteatus Deharveng & Lek, 1993
            Syn.: Isotomurus balteatus Sousa & al., 1997



Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from Norway
2010 © Fjellberg, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.18 © Murray, A.
Isotomurus gallicus from the UK
2013.06.12 © Murray, A.
Isotomurus gallicus from Holland
Diffuse pigmentation
2016.02.16 © Wieringa, P.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
Diffuse pigmentation
2016.02.19 © Wieringa, P.
Isotomurus gallicus from Holland
2017.10.12 © van Duinen, J.

Species ghibellinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94

Isotomurus ghibellinus
2001 © Carapelli, A. et al.

Species graminis Fjellberg, A, 2007:96

            Syn.: Isotomurus palustris var. pallida Schäffer, 1896 =?
            Syn.: Isotomurus palustris var. prasinus Deharveng & Lek, 1993
            Syn.: Isotomurus prasinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127
Isotomurus prasinus
2001 © Carapelli, A. et al.
Isotomurus prasinus from Sweden
2005 © Hall, K.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.




Isotomurus graminis
2007 © Fjellberg, A.
Isotomurus graminis from the UK
2007.03.18 © Cornwall, N.J.
Isotomurus graminis from the UK
2007.04.01 © Cornwall, N.J.
Isotomurus graminis from the UK
2009.03.29 © Michaelson, J.
Isotomurus graminis from the UK
2009.03.29 © Robertson, A.
Isotomurus graminis from the UK
2009.09.30 © Robertson, A.
Isotomurus graminis from the UK
2009.11.15 © Robertson, A.
Isotomurus graminis from the UK
2009.11.22 © Robertson, A.
Isotomurus graminis from the UK
Abd.3 with 4 bothriotricha
2009.11.26 © Robertson, A.
Isotomurus graminis from Sweden
2011.09.24 © Hall, K.
Isotomurus graminis from the UK
2012.04.21 © Murray, A.
Isotomurus graminis from the UK
2012.11.21 © Murray, A.
Isotomurus graminis from the UK
2013.08.10 © Fenwick, D.
Isotomurus graminis from Holland
2014.01.14 © van Duinen, J.
Isotomurus graminis from Belgium
2014.01.18 © Wallays, H.
Isotomurus graminis ♂ from Holland
2014.02.24 © van Duinen, J.
Isotomurus graminis ♂ from Holland
Spiny setae of male
2014.02.24 © van Duinen, J.
Isotomurus graminis from Germany
2014.09.20 © Beck, K.
Isotomurus graminis from Germany
2014.11.09 © Landgraf, A.
Isotomurus graminis from the UK
2015.01.15 © Thomas, Br.
Isotomurus graminis from Holland
2015.02.06 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.
Isotomurus graminis from Germany
Macrosetae on abd.4 in triangular constellation
2015.02.14 © Kluck, A.
Isotomurus graminis from Belgium
2016.01.10 © Huskens, M.-L.
Isotomurus graminis from the UK
2016.01.26 © Shepherd, M.J.
Isotomurus graminis from Belgium
2016.11.23 © Huskens, M.-L.
Isotomurus graminis from Belgium
2017.01.31 © Huskens, M.-L.

Species griseus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:282

Species hadriaticus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:121

Isotomurus hadriaticus
2001 © Carapelli, A. et al.

Species indicus Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:102

Isotomurus indicus from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.3 Fig.1

Species infuscatus Yosii, 1963

            Syn.: Isotomurus palustris ssp. fucicolus Yosii, 1955
            Syn.: Isotomurus palustris ssp. fucicolus Yosii, R, 1977:156


Species italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94

Isotomurus italicus
2007 © Fjellberg, A.


Species jharkhandensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:103

Isotomurus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.4 Fig.1
Isotomurus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.4 Fig.2


Species lapidosus (Salmon, JT, 1949:25) Greenslade, P, 2003:1, s.i.

Species lateclavus Kos, 1937

Species lineatus (Salmon, JT, 1941:322) Christiansen, K & Bellinger, P, 2000:114

  Subspecies violaceus Salmon, JT, 1941:323

Species louisiana (Scott, 1962) Waltz, RD, 1998:195

Species maculatus (Schäffer, 1896) Ågren, 1903, B

            Syn.: Isotoma bifasciata Bourlet, C, 1839:401
            Syn.: Desoria (Première division) fulvomaculata Nicolet, H, 1842:59
             Syn.: Isotoma fulvo-maculata(sic) Nicolet, 1847 ISS
            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
Desoria virescens from Switzerland
After Nicolet, H, 1842 Pl.5 Fig.12.

            Syn.: Desoria tigrina Bourlet, C, 1845:67
            Syn.: Desoria tigrina Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura viridis Bourlet, 1843 partim Schött, 1891
            Syn.: Isotoma lineata Lubbock, 1862
            Syn.: Isotoma palustris f. principalis Tullberg, 1872, partim Schött, 1891
            Syn.: Desoria riparia Lubbock, J, 1873:170
            Syn.: Isotomurus palustris var. texensis Folsom, JW, 1937:76
            Syn.: Isotomurus palustris f. hortorum Cassagnau, 1987
            Syn.: Isotomurus palustris Greenslade, 2002



             Syn.: Isotomurus palustris var. maculata(sic) Stach, J, 1924:113 ISS
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.9
Isotomurus palustris var. maculata(sic) from Malta
After Stach J, 1924 Taf.II Fig.10
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.11
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.12

             Syn.: Isotomurus palustris f. maculata(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Kseneman, M, 1932:20 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Stach, J, 1947:459 partim ISS
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.1.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.4.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.5.

             Syn.: Isotomurus palustris var. maculata(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Stach, J, 1967:401 ISS
Isotomurus palustris ssp. maculatus from France
After Poinsot N, 1972 Fig.6C

             Syn.: Isotomurus palustris f. maculata(sic) Verlinden, A, 1977:114,B ISS
             Syn.: Isotomurus maculata(sic) Deharveng, 1981 gr ISS
             Syn.: Isotomurus maculata(sic) Gabriel & al, 1990 gr ISS
Isotomurus maculatus
2001 © Carapelli, A. et al.
Isotomurus maculatus from Italy
After Potapov M, 2001 Fig.368-369
Isotomurus maculatus from the UK
2005 © Hopkin, S.P.
Isotomurus maculatus from the UK
2005 © Brocklehurst, K.
Isotomurus maculatus from Scandinavia
After Fjellberg, A, 2006 Fig.4
Isotomurus maculatus from the UK
2006.04.08 © Read, D.
Isotomurus maculatus from the UK
2007 © Cornwall, N.J.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus maculatus
2007 © Fjellberg, A.
Isotomurus maculatus from Italy
2007 © Bartolini, L.
Isotomurus maculatus from the UK
2007.12.01 © Warner, P.
Isotomurus palustris f. maculatus from the USA
2008.05.09 © Zungri, D.
Isotomurus maculatus from Hungary
2008.11.08 © Keresztes, G.
Isotomurus maculatus from the Netherlands
2008.11.14 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from the Netherlands
2008.11.19 © Baas, A.H.
Isotomurus maculatus from France
2009.02.25 © Lefebvre, V.
Isotomurus maculatus from Belgium
2009.03.08 © De Groot, S.
Isotomurus maculatus from the UK
2009.04.06 © Robertson, A.
Isotomurus maculatus from Tasmania
2009.08.16 © Daley, A.

             Syn.: Isotomurus maculata(sic) Greenslade, 2010 gr ISS
Isotomurus maculatus from Belgium
2010.01.31 © Wallays, H.

             Syn.: Isotomurus maculata(sic) Greenslade, P et Convey, P, 2012:411,gr ISS
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2013.12.01 © van Duinen, J.
Isotomurus maculatus from the UK
Pigmented clypeus and labrum
2015.02.05 © Horton, M.
Isotomurus maculatus from the UK
Pre-jump posture
2015.02.05 © Horton, M.

             Syn.: Isotomurus palustris var. maculata(sic) Pfliegler, WP et Janssens, F, 2016:202 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Pfliegler, WP et Janssens, F, 2016:204 ISS
Isotomurus maculatus from Belgium
2016.01.11 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Unguis with basal lateral teeth (*)
2016.03.08 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.04.24 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.09.04 © Huskens, M.-L.
Isotomurus palustris f. maculatus from the USA
2017.01.22 © Dobbs, Br.
Isotomurus palustris f. maculatus from the USA
2017.02.06 © Dobbs, Br.
Isotomurus maculatus from Belgium
2017.02.07 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Dark clypeus
2017.02.08 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Ventral aspect; with open valves of collophore
2017.12.08 © Huskens, M.-L.
Isotomurus maculatus from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomururus palustris f. maculatus from the USA
2017.12.27 © Dobbs, Br.
Isotomururus palustris f. maculatus from the USA
2017.12.27 © Dobbs, Br.
Isotomurus maculatus ♂ from Belgium
Ventral aspect; roundish genital plate
2018.01.11 © Huskens, M.-L.
Isotomurus maculatus ♂ from Belgium
Latero-ventral aspect; circumgenital setae
2018.01.11 © Huskens, M.-L.
Isotomurus maculatus from the Canary Islands
2018.03.09 © Huskens, M.-L.
Isotomururus palustris f. maculatus from the USA
2018.12.16 © Duncan, W.L.



Species mankyungensis Kim, JT et Lee, B-H, 2000:181

Species martiniquensis Stach, J, 1947:449

Isotomurus tricuspis ssp. martiniquensis from Martinique
After Stach, J., 1947 Plate XLIX Fig.8.

Species mauretanicus (Handschin, 1926, s.d.) Handschin, 1928

Species myohyangsanus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285

Species nebulosus Lek, S et Carapelli, A, 1998:125

Isotomurus nebulosus from France
After Lek & Carapelli, 1998 Fig.1
Isotomurus nebulosus
2001 © Carapelli, A. et al.
Isotomurus ?nebulosus? from France
Dark form
2018.04.05 © Garcelon, P.
Isotomurus ?nebulosus? from France
Pale form
2018.04.06 © Garcelon, P.

Species nicolae Stomp, 1983

Species novaezealandiae (Salmon, JT, 1941:321) Christiansen, K & Bellinger, P, 2000:114

            Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1941:321 IOS
Proisotomurus novae-zealandiae(sic) from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.179




             Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1964:359 ISS



Species obscurus Carpenter, GH, 1916:34

Species opala Christiansen, K et Bellinger, P, 1992:155

Isotomurus opala from Hawaii
Type 1
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.C
Isotomurus opala from Hawaii
Type 2
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.D
Isotomurus opala from Hawaii
Intermediate between type 3 & 4
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.A
Isotomurus opala from Hawaii
Type 4
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.B

            Syn.: Isotomurus palustris var. balteatus Folsom, 1932



Isotomurus opala? from Hawaii
2017.10.03 © Nelson, S.
Isotomurus opala? from Hawaii
2017.10.06 © Nelson, S.
Isotomurus opala? from Hawaii
2017.10.10 © Nelson, S.

Species palliceps (Uzel, 1891) Stach, 1929

Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.1.
Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.2.

Species palustris (Müller OF, 1776) Houlbert, C, 1924:106, t.t., B

            Syn.: Podura aquatica cinerea de Geer, C, 1740:63
"Graue Insekten die unter den vorigen schwarzen [Podura aquatica] zusammen auf den Wasser sitzen und etwas grösser sind" from Sweden
After de Geer, C., 1749 Tab.IV Fig.1.

            Syn.: Podura aquatilis Müller OF, 1776
            Syn.: Podura aquatica grisea de Geer, C, 1778:28
Podura aquatica grisea from Sweden
After de Geer, C., 1778 Tom.VII Pl.2 Fig.19.

             Syn.: Podura Grisea(sic) Herbst, JFW, 1787:350 ISS
Podura Grisea(sic) from Germany
After Herbst JFW, 1787 Taf.LXXV Fig.4ab

            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
            Syn.: Podura Stagnorum(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:93
Podura Stagnorum(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.4

            Syn.: Podura armata Nicolet, 1842
            Syn.: Aetheocerus aquaticus Bourlet, 1842
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
            Syn.: Isotoma spilosoma Gervais in Walckenaër, 1844
Isotome spilosome from France
Isotoma spilosoma
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.9


             Syn.: Isotoma Spilosoma(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Isotoma palustris var. spilosoma Parfitt, 1891
             Syn.: Isotomurus spilosoma(sic) Houlbert, C, 1924:106 ISS
            Syn.: Desoria riparia Bourlet, C, 1845:67
            Syn.: Desoria riparia Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura psi Herklots, 1857
            Syn.: Degeeria pi Herklots, 1858
            Syn.: Isotoma tricolor MacGillivray, 1891
            Syn.: Isotoma tricolor MacGillivray, 1896
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Womersley, 1934
            Syn.: Isotomurus tricolor Womersley, 1934
            Syn.: Isotoma tricolor Stach, J, 1947:422
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotomurus tricolor Hart, 1974 =?
            Syn.: Isotomurus tricolor Fjellberg, 1986
            Syn.: Isotoma balteata Reuter, 1876
            Syn.: Isotoma Stuxbergii(sic) Tullberg, 1876 IOS
            Syn.: Isotoma Tullbergi(sic) Moniez, 1889
            Syn.: Isotoma aequalis MacGillivray, 1896
            Syn.: Isotoma palustris f. pallida Schäffer, 1896
             Syn.: Isotomurus palustris ssp. pallida(sic) Salmon, JT, 1964:368 ISS
            Syn.: Isotoma maculata Börner, 1901
            Syn.: Isotoma palustris f. cincta Krausbauer, 1902
            Syn.: Isotoma palustris ab. bimaculata Agren, 1903
            Syn.: Isotomurus palustris ssp. tricuspis Börner, C, 1906:173
            Syn.: Proisotomurus papillatus Womersley, 1934
            Syn.: Isotomurus antennalis Poinsot, N, 1972:687
            Syn.: Isotomurus palustris f. catalana Cassagnau, 1987
Isotomurus palustris f. catalana from France
After Potapov M, 2001 Fig.370

            Syn.: Isotomurus independente Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotomurus independente
2001 © Carapelli, A. et al.

            Syn.: Isotomurus sp. Dethier, M et Hubart, J-M, 2000:29,B



Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.9
             Syn.: Isotoma pulustris(sic) Stach, J, 1924:114 ISS
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.1
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.2
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.3
Isotomurus palustris ssp. palustris from the USA
After Snider RJ, 1967 Cover
Isotomurus palustris ssp. palustris from France
After Poinsot N, 1972 Fig.6A


             Syn.: Isotomurus palustis(sic) Heungens, A et Van Daele, E, 1973:141,B ISS
             Syn.: Isotomurus pallustris(sic) Greenslade, P in Dallai, R, 1989:111 ISS
Isotomurus palustris from Hawaii
After Christiansen K & Bellinger P, 1992 Plate 65 Fig.A

Isotomurus palustris
2001 © Carapelli, A. et al.
Isotomurus palustris from Italy
After Potapov M, 2001 Fig.372

Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from Belgium
2006 © Vuijlsteke, M.
Isotomurus palustris from the UK
2007 © Valentine., B
Isotomurus palustris from the USA
2007.02.20 © Moorehead, C.
Isotomurus palustris
2007 © Fjellberg, A.
Isotomurus palustris from the UK
2007.10.16 © Campbell, A.
Isotomurus palustris from the UK
2007.10.19 © Campbell, A.
Isotomurus palustris from the Netherlands
2007.11.08 © Baas, A.H.
Isotomurus palustris from the UK
2007.11.24 © Campbell, A.
Isotomurus palustris from the USA
2008.01.12 © Moorehead, C.
Isotomurus palustris from the UK
2008.01.12 © Farrell, S.
Isotomurus palustris from the Netherlands
2008.01.26 © Baas, A.H.
Isotomurus palustris from Germany
2008.04.17 © Peters, J.
Isotomurus palustris from the Netherlands
2008.11.14 © Baas, A.H.
Isotomurus palustris
2008.12.25 © onefivenine.


             Syn.: Isotomurus palustres(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Cutz-Pool, LQ, 2009:205 ISS
Isotomurus palustris from the USA
2009.01.07 © Fleming, K.
Isotomurus palustris from the USA
2009.01.31 © Justis, S.
Isotomurus palustris from the USA
2009.02.01 © Justis, S.
Isotomurus palustris from France
2009.02.25 © Lefebvre, V.
Isotomurus palustris from the USA
2009.03.03 © Justis, S.
Isotomurus palustris from the UK
2009.03.07 © Barton, T.
Isotomurus palustris from the Netherlands
2009.03.27 © Baas, A.H.
Isotomurus palustris from the Netherlands
2009.03.27 © Baas, A.H.
Isotomurus palustris from the UK
2009.12.26 © Barton, T.
Isotomurus palustris from the UK
2009.12.26 © Barton, T.
Isotomurus palustris from the UK
Post-antennal organ
2012.01.03 © Kent, J.J.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from La Palma
2012.12.01 © Barton, T.
Isotomurus palustris from Holland
2014.11.07 © van Duinen, J.
Isotomurus palustris from the UK
Phoretic curled up nematode
2015.02.24 © Thomas., B
Isotomurus palustris from the UK
2015.03.23 © Barton, T.
Isotomurus palustris from the UK
2015.03.23 © Barton, T.
Isotomurus palustris from the UK
Aberrant abdominal pigmentation
2015.12.04 © Barton, T.
Isotomurus palustris from Belgium
2016.01.04 © Huskens, M.-L.
Isotomurus palustris from the UK
2016.07.22 © David W.
Isotomurus palustris from the USA
2016.11.28 © Dobbs, Br.

             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:101,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:103,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:104,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:105,gr ISS
Isotomurus palustris from the USA
2017.01.20 © Fleming, K.
Isotomurus palustris juv. from the UK
2018.02.01 © Horton, M.





Species palustroides Folsom, JW, 1937:77

Species paraciliatus Cardoso, MA, 1970:109

Species philfrancisi Bernard, 2006

Isotomurus philfrancisi
2006 © Bernard, E.

Species plumosus Bagnall, 1940, B

            Syn.: Isotomurus palustroides ssp. subciliatus Stach, J, 1947:461
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.1
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.2

             Syn.: Isotomurus palustroides ssp. subcialiatus(sic) Rusek, J in Dlabola, J, 1977:15 ISS



Isotomurus plumosus from Belgium
2001 © Hopkin, S.P.
Isotomurus plumosus from Poland
After Potapov M, 2001 Fig.373
Isotomurus plumosus from Sweden
2006 © Hall, K.
Isotomurus plumosus
2007 © Fjellberg, A.
Isotomurus plumosus from the UK
2007.04.09 © Cornwall, N.J.
Isotomurus plumosus from Belgium
2007.03.28 © Lindsley, J.
Isotomurus plumosus juv. from France
2008.03.02 © Lebeaux, P.
Isotomurus plumosus from the UK
2010.02.06 © Talbot, M.E.
Isotomurus plumosus from Denmark
2012.03.19 © Bildstrup, O.
Isotomurus plumosus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus plumosus from the UK
2013.06.24 © Fenwick, D.
Isotomurus plumosus from the UK
2013.08.10 © Fenwick, D.
Isotomurus plumosus juv. from the UK
2014.09.16 © Garlick, T.
Isotomurus plumosus juv. from the UK
2014.09.16 © Garlick, T.
Isotomurus plumosus juv. from the UK
2014.10.24 © Garlick, T.
Isotomurus plumosus from Czechia
With Sminthurinus elegans f. ornata
2015.03.05 © Krásenský, P.
Isotomurus plumosus juv. from Holland
2015.07.02 © van Leeuwen, J.
Isotomurus plumosus from Holland
2015.12.04 © van Duinen, J.
Isotomurus plumosus from Belgium
2016.03.13 © Janssens, F.
Isotomurus plumosus juv. from Belgium
2016.03.13 © Janssens, F.
Isotomurus plumosus juv. from Belgium
2016.04.17 © Huskens, M.-L.
Isotomurus plumosus from Belgium
2016.07.10 © Janssens, F.
Isotomurus plumosus from Belgium
Postantennal organ (PAO) elliptical with central constriction
2016.08.10 © Huskens, M.-L.
Isotomurus plumosus from the UK
2017.02.01 © David W.
Isotomurus plumosus juv. from Belgium
2017.09.08 © Huskens, M.-L.
Isotomurus plumosus from France
Head without interocular stripe
2017.09.10 © Picard, J.
Isotomurus plumosus juv. from Belgium
Different juvenile stages
+ Sminthurides malmgreni ♀
2019.02.15 © Janssens, F.
Isotomurus plumosus juv. from Belgium
+ Podura aquatica
2019.02.15 © Janssens, F.
Isotomurus plumosus from the UK
2019.02.23 © Smart, J.

Species pseudopalustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127

            Syn.: Podura palustris ssp. aquatilis Müller OF, 1776
Isotoma aquatilis from the UK
After Lubbock, J., 1873 Pl.37.

             Syn.: Isotoma aquatelis(sic) Parfitt, 1891 ISS
Isotoma palustris var. aquatilis from Sweden
After Schött H, 1893 Taf.V Fig.7
Isotomurus pseudopalustris nr f. aquatilis from France
2013.01.13 © Quintin, C.
Isotomurus pseudopalustris f. aquatilis from the UK
2013.01.25 © Cobby, P.
Isotomurus pseudopalustris nr f. aquatilis from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus pseudopalustris nr f. aquatilis from France
2018.02.24 © Tiky, S.
Isotomurus pseudopalustris f. aquatilis from the UK
2018.12.28 © Marquis, A.
Isotomurus pseudopalustris f. aquatilis from the UK
Abd.4 pattern
2019.01.21 © Cann, A.J.


            Syn.: Isotoma trifasciata Bourlet, C, 1839:402
Isotoma trifasciata from the UK
After McIntire SJ, 1867 Fig.37
After McIntire SJ, 1868
Isotoma trifasciata from the UK
After Lubbock J, 1870 Pl.XXII Fig.20
Isotoma trifasciata from the UK
Foot of anterior leg
After Lubbock J, 1870 Pl.XXII Fig.21

            Syn.: Isotomurus palustris f. trimaculata(sic) Womersley, H, 1939:149 lapsus
            Syn.: Isotoma Gervaisii(sic) Nicolet, H, 1847:374 IOS
Isotoma Gervaisii(sic) from Switzerland
After Nicolet,, 1847 Pl.6 Fig.9


             Syn.: Isotoma Gervaisii(sic) Brauer, 1855 ISS
             Syn.: Isotoma Gervaisii(sic) Kolenati, FA, 1858:133 ISS
             Syn.: Isotoma Gervaisii(sic) Lubbock, J, 1873:176 ISS
             Syn.: Isotoma Gervaisii(sic) Stach, J, 1947:367 ISS
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2005.06.02 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2008.12.11 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from the UK
2011.08.26 © Barton, T.
Isotomurus pseudopalustris f. gervaisi from Holland
2014.02.11 © van Duinen, J.
Isotomurus pseudopalustris f. gervaisi from the UK
2014.12.20 © Evans, K.
Isotomurus pseudopalustris f. gervaisi from Belgium
2016.05.27 © Huskens, M.-L.
Isotomurus pseudopalustris f. gervaisi from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus pseudopalustris f. gervaisi from Belgium
2018.12.27 © Huskens, M.-L.


            Syn.: Isotoma tricolor Schött, H, 1891:22
            Syn.: Isotoma tricolor MacGillivray, AD, 1894:105
            Syn.: Isotoma palustris forma principalis Schött, H, 1893:63 partim
Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.8
Isotomurus palustris forma principalis from Germany
After Heymons R & Heymons H, 1909 Fig.11


            Syn.: Isotomurus palustris f. maculata Stach, J, 1947:459 partim
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.2.
Isotomurus palustris f. maculata from Poland
After Stach, J., 1947 Plate XLVIII Fig.3.


            Syn.: Isotomurus palustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:87



Isotomurus pseudopalustris from Italy
After Potapov M, 2001 Fig.366-367
Isotomurus pseudopalustris from the UK
2006 © Brocklehurst, K.
Isotomurus pseudopalustris from the UK
2006.06.24 © Botting, J.
Isotomurus palustris f. pseudopalustris from the USA
2007 © Moorehead, C.
Isotomurus pseudopalustris from Germany
2008.04.17 © Peters, J.
Isotomurus pseudopalustris from the UK
2008.09.24 © Robertson, A.
Isotomurus palustris f. pseudopalustris from the USA
2010.02.07 © Zungri, D.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.11.03 © Nurcombe, E.
Isotomurus pseudopalustris from France
2012.11.05 © Quintin, C.
Isotomurus pseudopalustris from Germany
2015.01.20 © Landgraf, A.
Isotomurus pseudopalustris from Holland
2015.02.24 © van Duinen, J.
Isotomurus pseudopalustris from Holland
Two pale dots on abd.4
2015.10.19 © van Duinen, J.
Isotomurus pseudopalustris from the UK
2015.11.25 © Evans, K.
Isotomurus pseudopalustris from the UK
2015.12.08 © Horton, M.
Isotomurus pseudopalustris from the UK
2015.12.28 © Evans, K.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
Unguiculus with distinct inner tooth
2016.02.13 © Phillips, E.
Isotomurus pseudopalustris from Belgium
2016.04.09 © Huskens, M.-L.
Isotomurus palustris f. pseudopalustris from the USA
2016.04.17 © Dobbs, Br.
Isotomurus palustris f. pseudopalustris from the USA
2016.04.21 © Dobbs, Br.
Isotomurus pseudopalustris from Belgium
2016.05.08 © Huskens, M.-L.
Isotomurus pseudopalustris from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus pseudopalustris? from Australia
2018.09.28 © Buck, D.
Isotomurus pseudopalustris from Holland
2018.11.23 © Kamsteeg, G.
Isotomurus palustris f. pseudopalustris from the USA
2019.01.04 © Pearson, R.R.
Isotomurus pseudopalustris juv. from the UK
2019.02.20 © Jeffery, J.



Species pseudosensillatus de Mendonça, 1990

Species punctiferus Yosii, 1963

             Syn.: Isotomurus punctatus(sic) Yosii, R, 1967:2 lapsus ISS
             Syn.: Isotomurus punctiferous(sic) Gupta & Mukharji, 1977 ISS
Isotomurus punctiferus from Japan
After Potapov M, 2001 Fig.375
Isotomurus nr punctiferus from Norway
2007.03.29 © Fauskanger, I.
Isotomurus nr punctiferus from France
2008.11.08 © Jacquemin, G.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.

             Syn.: Isotomurus punctiferous(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:104 ISS


Species qinpuensis Chen, 1985

Species quadrisetosus Rusek, 1971

Species rabili Deharveng & Lek, 1993

Isotomurus rabili
2001 © Carapelli, A. et al.

Species retardatus Folsom, JW, 1937:71

Species riparius de Mendonça, 1990

Species sahebganjensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:105

Isotomurus sahebganjensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.5 Fig.1

Species sensillatus Winter, 1967

Species stepposus Potapov, MB et Starostenko, EV, 2002:331

Isotomurus stepposus from Ukraine
After Potapov MB & Starostenko EV, 2002 Fig.1

Species stuxbergi (Tullberg, T, 1877:35) Babenko & Bulavintsev, 1993

            Syn.: Isotoma Stuxbergii(sic) Tullberg, T, 1877:35 IOS
            Syn.: Isotoma Stuxbergi(sic) var. prasina Reuter, 1891
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1905 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Wahlgren, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Collinge & Shoebotham, 1910 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1911 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1912 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Shoebotham, 1914 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Brown, 1918 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1920 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Denis, 1921 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1923 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Thompson, 1924 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stirnimann, 1926 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Jackson, 1928 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1929 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1930 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1930 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Kseneman, M, 1932:20 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Turk, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills, 1934 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1935 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Drenowski, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Folsom, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1939 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Szent-Ivany, 1940 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1941 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Stach, J, 1947:459 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1949 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills & Richards, 1953 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, 1954 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Goto, 1955 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Grinbergs, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) von Torne, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus (Isotomurus) palustris var. prasina(sic) Grinbergs, 1960 ISS
             Syn.: Isotomurus palustris ssp. prasina(sic) Salmon, JT, 1964:368 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Poinsot-Balaguer, 1976 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, R, 1977:156 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Fjellberg, A, 2007:98=? ISS
            Syn.: Isotomurus palustris ssp. palustris Brimley, 1942
            Syn.: Isotomurus ciliatus Stach, 1947
            Syn.: Isotomurus tricolor Babenko & Bulavintsev, 1993 =?


Isotomurus stuxbergi
2007 © Fjellberg, A.



Species subterraneus Stach, 1946

Species takahashii (Yosii, R, 1940:115) Yosii, 1963

Isotoma takhaashii(sic) from Formosa
After Yosii R, 1940 Fig.7
            Syn.: Isotomurus alticulus ssp. japonicus Yosii, R, 1956:64
            Syn.: Isotomurus alticolus ssp. japonicus Yosii, R, 1977:156



             Syn.: Isotoma takhaashii(sic) Yosii, R, 1940:116 ISS

Species texensis Folsom, JW, 1937:76

Species tricolor (Packard, AS, 1873:34) Lippert & Butler, 1976

            Syn.: Isotoma palustris Folsom, JW, 1901:159
            Syn.: Isotoma palustris Guthrie, JE, 1903:70
            Syn.: Isotoma palustris f. principalis Guthrie, JE, 1903:70
            Syn.: Isotoma palustris Folsom, JW, 1919:11A
            Syn.: Isotomurus palustris Mills, HB, 1934:54
            Syn.: Isotomurus palustris Maynard, EA, 1951:122
            Syn.: Isotomurus palustris Salmon, JT, 1964:362 partim
            Syn.: Isotomurus palustris Salmon, JT, 1964:363 partim
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1980:687
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1998:756


Isotomurus (Isotomurus) tricolor from the USA
After Folsom JW in Christiansen K & Bellinger P, 1980 Fig.546A
Isotomurus tricolor from the USA
2006 © Murray, T.
Isotomurus tricolor from the USA
2007.05.18 © Cowen, R.
Isotomurus tricolor from the USA
2008.01.06 © Moorehead, C.
Isotomurus tricolor from the USA
2009.03.04 © Murray, T.
Isotomurus tricolor from the USA
2009.08.19 © Zungri, D.
Isotomurus tricolor from the USA
2010.01.23 © Justis, S.
Isotomurus tricolor from the USA
2010.02.08 © Zungri, D.
Isotomurus tricolor from the USA
2010.01.21 © Murray, T.
Isotomurus tricolor from the USA
2012.01.14 © Justis, S.
Isotomurus tricolor from the USA
2012.09.22 © Murray, T.

Species tricuspis Börner, C, 1906:173

Species unifasciatus (Börner, C, 1901:50) Stach, J, 1947:459, B

            Syn.: Isotoma palustris f. principalis Schött partim


Isotomurus unifasciatus
2001 © Carapelli, A. et al.
Isotomurus unifasciatus from the UK
2005 © Hopkin, S.P.
Isotomurus unifasciatus from the UK
2007 © Barton, T.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus
2007 © Fjellberg, A.
Isotomurus unifasciatus from the UK
2007 © Spears, G.
Isotomurus unifasciatus from Norway
2009.11.29 © Haraldseide, H.
Isotomurus unifasciatus from the UK
2012.01.26 © Kent, J.J.
Isotomurus unifasciatus from the UK
2012.11.02 © Murray, A.
Isotomurus unifasciatus juv. from the UK
2013.01.18 © Nurcombe, E.
Isotomurus unifasciatus from Holland
2013.02.07 © van Duinen, J.
Isotomurus nr unifasciatus from New Zealand
2014.06.01 © Murray, A.
Isotomurus unifasciatus from Germany
2015.01.05 © Beck, K.
Isotomurus unifasciatus juv. from Germany
2015.03.28 © Kluck, A.
Isotomurus unifasciatus from the UK
2018.12.04 © Smart, J.

Species venustus Barra, J-A, 2006:67

Species viridipalustris Kos, 1937

Species yamaquizuensis Winter, 1967

            Syn.: Isotomurus yamaquizuensis Winter, 1963 nn


Isotomurus cf. yamaquizuensis from Colombia
After Mari Mutt JA, 1987 Fig.11


Genus Chionobora Greenslade, P et Potapov, M, 2015:334

Species amila Greenslade, P et Potapov, M, 2015:334, t.t.

Chionobora amila from Tasmania
After Greenslade P & Potapov M, 2015 Fig.1
Chionobora amila ♂ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1A
Chionobora amila ♀ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1B



Genus Kaylathalia Stevens, MI et D'Haese, CA, 2016:3

Species klovstadi (Carpenter, 1902) Stevens, MI & D'Haese, CA, 2016:3, t.t.

            Syn.: Isotoma Klovstadi(sic) Carpenter, 1902 IOS


             Syn.: Isotoma Klovstadi(sic) Enderlein, 1909 ISS
             Syn.: Isotoma Klovstadi(sic) McNamara, 1919 ISS
             Syn.: Isotoma Klovstadi(sic) Gressitt & Weber, 1959 ISS
Desoria klovstadi from Antarctica
2008.05.13 © O'Brien., B

             Syn.: Chionobora klovsatdi(sic) Stevens, MI et D'Haese, CA, 2016:3 ISS






Genus Hydroisotoma Stach, J, 1947:249

Species schaefferi (Krausbauer, 1898) Stach, J, 1947:250, t.t.

            Syn.: Isotoma schäfferi(sic) Krausbauer, 1898,tt IOS
            Syn.: Isotoma rivularis Latzel, 1921
            Syn.: Archisotoma dimorpha Denis, 1922
            Syn.: Proisotoma Schäfferi(sic) ssp. aculeata Stach, J, 1930:289
             Syn.: Proisotoma schaefferi ssp. aculata(sic) Wray, DL, 1978:205 ISS
            Syn.: Hydroisotoma stachi Bagnall, 1949


             Syn.: Proisotoma (Ballistura) Schäfferi(sic) Heymons, R et Heymons, H in Brauer, A, 1909:5 ISS
Proisotoma Schäfferi(sic) from Germany
After Heymons R & Heymons H, 1909 Fig.6

             Syn.: Isotoma schäfferi(sic) Folsom, 1928 ISS
             Syn.: Proisotoma Schäfferi(sic) Denis, JR, 1931:116 ISS
             Syn.: Isotoma schäfferi(sic) Tarsia, 1943 ISS
             Syn.: Isotoma schäfferi(sic) Gisin, 1944 ISS
             Syn.: Hydroisotoma schäfferi(sic) Stach, J, 1947:250 ISS
Hydroisotoma schäfferi(sic) from Poland
After Stach, J., 1947 Plate XXIXa Fig.1

             Syn.: Isotoma (Isotoma) schäfferi(sic) Maynard, EA, 1951:108 ISS
             Syn.: Isotoma schäfferi(sic) Maynard, EA, 1951:108 ISS
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.198
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.199

             Syn.: Hydroisotoma schafferi(sic) Gisin, H, 1960:213 ISS
             Syn.: Hydroisotoma schafferi(sic) Snider, RJ, 1967:212 ISS
Hydroisotoma schaefferi ♂ from the USA
Habitus lateral
After Wray DL, 1978 Fig.5
Hydroisotoma schaefferi ♂ from the USA
Habitus dorsal
After Wray DL, 1978 Fig.6
Hydroisotoma schaefferi ♂ from the USA
Head with curled antennae
After Wray DL, 1978 Fig.2
Hydroisotoma schaefferi ♂ from the USA
Antenna with sense organs
After Wray DL, 1978 Fig.1
Hydroisotoma schaefferi from Germany
After Schulz, H.-J., 1989
Hydroisotoma schaefferi from Germany
2006 © Schulz, H.-J. & Schubert H.D.
Hydroisotoma schaefferi from the USA
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
2007.03.01 © Murray, T.
Hydroisotoma schaefferi from the USA
2008.01.13 © Murray, T.
Hydroisotoma schaefferi from France
2008.03.16 © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi from the USA
2011.02.20 © Husic, C.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2012.12.02 © Fleming, K.
Hydroisotoma schaefferi ♂ from the USA
Male with bent apical antennal segments and warts on 2nd antennal segment
2013.03.28 © Murray, T.

             Syn.: Hydroisotoma schafferi(sic) Glime, JM, 2015:3 ISS
Hydroisotoma schaefferi from Germany
2015.03.31 © Schulz, H.-J.
Hydroisotoma schaefferi from the USA
2015.04.16 © Mallory, C.
Hydroisotoma schaefferi from the USA
2017.01.19 © Fleming, K.
Hydroisotoma schaefferi from France
2018.03.03 © Tiky, S.







Genus Agrenia Börner, C, 1906:171 key

  Syn.: Isotomini Ågrenia(sic) Börner, C, 1906:171 IOS
Species-group bidenticulata
Remark by Fjellberg, A. & Bernard, E.C. 2009:19: Lateral mucronal seta present.



Species sp.1 Fjellberg, A, 1988:112

Species bidenticulata (Tullberg, T, 1877:35) Denis, J-R, 1924:231, t.t., B

            Syn.: Isotoma lanuginosa Carl, J, 1899:307
            Syn.: Agrenia bidenticulata var. elongata Carpenter, 1908
            Syn.: Agrenia bidenticulata var. nigra Linnaniemi, 1935


Agrenia bidenticulata from Belgium
After Jacquemart S, 1975 p.83
Agrenia bidenticulata from Russia
2006 © Boldsystems.org
Agrenia bidenticulata from the USA
2011.05.30 © Eiseman, C.
Agrenia bidenticulata from Norway
2015.mm.dd © Fjellberg, A.
Agrenia bidenticulata from Buryatia
With long claws
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.37
Agrenia bidenticulatus from France
2018.01.27 © Picard, J.
Agrenia bidenticulatus from France
Eyepatch with 6 ocelli
2018.01.27 © Picard, J.
Agrenia bidenticulatus juv. from France
2018.01.27 © Picard, J.
Agrenia bidenticulatus from France
Third foot with long narrow unguis(**),
and unguiculus with large exad lamella (*)
2018.01.27 © Picard, J.

Species lamellosa Fjellberg, A, 1988:110

Species pilosa Fjellberg, A, 1986:101

Species polymorpha Fjellberg, A, 1986:96


Species-group agilis
Remark by Fjellberg, A. & Bernard, E.C. 2009:20: Lateral mucronal seta absent.

Species agilis Fjellberg, A, 1986:101

Agrenia agilis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3A

Species atroviridis Fjellberg, A, 1986:104

Agrenia atroviridis from the USA
2008.02.09 © Murray, T.
Agrenia atroviridis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3B

Species cyanura Fjellberg, A, 1986:102

Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3C
Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3D

Species extrema Fjellberg, A et Bernard, EC, 2009:24

Species falcula Fjellberg, A et Bernard, EC, 2009:25

Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5A
Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5B

Species parkeri Fjellberg, A et Bernard, EC, 2009:20

Agrenia parkeri from the USA
2006 © Bernard, E.C.
Det. Bernard, E.C., 2010.01.25
Agrenia parkeri ♀ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri ♂ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4B

Species riparia Fjellberg, A, 1986:103

Species tarashchukae Fjellberg, A et Bernard, EC, 2009:27


Genus Pteronychella Börner, C, 1909:113

Species andina Yoshii, 1990

Species babai Yosii, 1939

Species ezoensis Yosii, 1965

Species perpulchra Börner, C, 1909:113, t.t.


Genus Isotomedia Salmon, JT, 1944:148

Species triseta Salmon, JT, 1944:148, t.t.


Genus Procerura Salmon, JT, 1941:326

Species aequaoculata Salmon, JT, 1941:328

Species dissimilis (Salmon, JT, 1944:146) Christiansen, K & Bellinger, P, 2000:107


Procerura fasciata from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.198.
Species fasciata Salmon, JT, 1941:329


Species fusca (Salmon, JT, 1941:330) Christiansen, K & Bellinger, P, 2000:107

  Subspecies pallida Salmon, JT, 1941:331

Species indica (Baijal, HN, 1958:355) Greenslade, P, 2003:1, s.i.

Procerura indica from India
After Yadav RS et al, 2018 Fig.1

Species magnifica (Salmon, JT, 1949:29) Greenslade, P, 2003:1, s.i.

Species montana Salmon, JT, 1941:328


Isotoma parva from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.a.
Species parva (Salmon, JT, 1937:352) Greenslade, P, 2003:1, s.i.

             Syn.: Papillomurus parva(sic) Salmon, JT, 1964:372 ISS

Species purpurea Salmon, JT, 1941:328

Species ochracea (Salmon, JT, 1949:27) Christiansen, K & Bellinger, P, 2000:107

Species serrata Salmon, JT, 1941:329

Species transequatoria (Salmon, 1969) Greenslade, P, 2003:1, s.i.

            Syn.: Papillomurus transequatoria(sic) Salmon, 1969 IOS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:24 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:74 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:78 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:116 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:126 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:128 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:178 ISS



Species violacea Salmon, JT, 1941:327, t.t.


Genus Spinocerura Salmon, 1941


Spinocerura capillata from New Zealand
After Salmon, J.T. 1941 Pl.52 Fig.212.
Species capillata Salmon, JT, 1941:332, t.t.


Species dreuxi (Delamare Deboutteville & Massoud, 1966)


Genus Setocerura Salmon, JT, 1949:33

Species georgiana (Schäffer, 1891) Wise, KAJ, 1970:198

Species maruiensis (Salmon, JT, 1941:326) Deharveng, L & de Oliveira, EP, 1994:442


Tomocerura rubenota from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.193.
Species rubenota (Salmon, JT, 1941:325) Salmon, JT, 1949:33, t.t.


Species sp. Greenslade, P in Dallai, R, 1989:112

Species striata (Schäffer, 1897)

Species wahlgreni (Womersley, H, 1939:321) Salmon, 1949


Genus Tomocerura Wahlgren, 1901

Species colonavia Salmon, JT, 1949:31

Species conjungens (Schäffer, 1897) Wahlgren, E, 1906:17

Species faguagae Izarra, 1970

Species georgiana (Schäffer, 1897) Massoud & Rapoport, 1968

Species picta Wahlgren, 1901, t.t.

Species swani (Womersley, 1934)

Isotoma swani from Australia
After Womersley H, 1939 Pl.1 Fig.2.


Genus Isotoma Bourlet, C, 1839:399 key

  Syn.: Desoria Deuxième division Nicolet H in Desor, E, 1841:385
  Syn.: Podurelles (A-tête-directe) Isotama(sic) Nicolet, H, 1847:374 ISS
Isotoma from Germany
Schematic sagittal section
After Anonymous Fig.E

  Syn.: Arthropleona Isootoma(sic) Richards, WR, 1968:15 ISS
Remark by ICZN. 2007:419: Placed on the Official List of Names in Zoology. Opinion 291.

Isotoma sp. from the USA
2009.01.29 © Justis, S.
Isotoma sp. from Tunesia
Predation by pseudoscorpion Garypus
2009.04.14 © Henderickx, H.
Isotoma sp. from Norway
3+3 macrosetae on abd.4 in one line
After Fjellberg A, 2010 p.2
Isotoma sp. from Norway
More elongate head
After Fjellberg A, 2010 p.2
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.
Isotoma nr viridis from the USA
2010.01.15 © Parker, H.S.
Isotoma anglicana from Belgium
Eyepatch with 6 large ocelli and 2 small (*)
2018.02.02 © Huskens, M.-L.
Isotoma viridis from the UK
Abdominal dorsal macrosetae in transverse row
2019.02.04 © Cann, A.J.



Species acrea Wray, 1953

Species anglicana Lubbock, 1862

Isotoma anglicana from the UK
After Lubbock J, 1873 Pl.38
Isotoma anglicana from the UK
2005 © Hopkin, S.P.
Isotoma anglicana from the UK
Manubrial thickening with 4 teeth
2005 © Hopkin, S.P.
Isotoma anglicana from Germany
Manubrial thickening with 4 teeth
Det. Schulze, H.J.
2005 © Mebes, H.
Isotoma anglicana from Sweden
2006 © Hall, K.
Isotoma anglicana juv. from Holland
On snow
2006.03.02 © Hamers., B
Isotoma anglicana from the UK
2007.03.25 © Barnes, C.
Isotoma anglicana from Holland
2008.01.27 © Baas, A.H.
Isotoma anglicana from Belgium
2008.02.07 © Hanssens., B
Isotoma anglicana from Holland
2008.02.08 © Baas, A.H.
Isotoma anglicana from the UK
2008.04.13 © Kilford., B
Isotoma anglicana from France
2008.11.13 © Lavoué,B.
 
Isotoma anglicana from the UK
2008.12.10 © Talbot, M.
Isotoma anglicana from the UK
2009.03.01 © Kilford., B
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana juv. from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from the UK
2009.03.21 © Talbot, M.
Isotoma anglicana from Holland
2009.03.20 © Baas, A.H.
Isotoma anglicana from the UK
2009.08.20 © Ransom, T.
Isotoma anglicana from Norway
After Fjellberg A, 2010 p.3
Isotoma anglicana from Norway
Manubrial dorsal macrosetae: A>B
After Fjellberg A, 2010 p.8
Isotoma anglicana from Norway
3 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
 
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana from Holland
2010.02.06 © van Duinen, J.
Isotoma anglicana from the UK
2010.05.02 © Robertson, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from the UK
2012.02.16 © Talbot, M.
Isotoma anglicana from Holland
Ovoid post-antennal organ (PAO)
2012.05.13 © van Duinen, J.
Isotoma anglicana from Holland
Abdominal pigmentation
2012.12.14 © Hamers., B
Isotoma anglicana from Belgium
On snow
2013.02.15 © Hamers., B
Isotoma anglicana from Holland
2014.02.21 © van Duinen, J.
Isotoma anglicana from Germany
2015.01.17 © Landgraf, A.
Isotoma anglicana from Holland
2015.02.04 © van Duinen, J.
Isotoma anglicana from Holland
2015.03.10 © van Duinen, J.
Isotoma anglicana from the UK
2015.08.16 © Boardman, P.
Isotoma anglicana from the UK
Manubrial thickening with 2+2 teeth (*)
2015.08.16 © Boardman, P.
Isotoma anglicana from Holland
2016.03.17 © van Duinen, J.
Isotoma anglicana from Holland
Distinctly spotted skin
2016.03.17 © van Duinen, J.
Isotoma anglicana from Belgium
Parasitic protonymph Acari
2018.02.02 © Huskens, M.-L.
Isotoma anglicana from the UK
Manubrial thickening with 2 teeth (*)
2019.01.15 © Cann, A.J.
Isotoma anglicana from the UK
With Vertagopus sp.
2019.02.24 © Jeffery, J.

Species arctica Schött, H, 1893:61

Isotoma viridis var. arctica from the American Arctics
After Schött H, 1893 Taf.V Fig.4
Isotoma viridis var. arctica from Alaska
After Folsom JW, 1902 Pl.IV Fig.5

             Syn.: Isotoma viridis var. artica(sic) Stach, J, 1924:114 ISS
Isotoma arctica from Alaska
After Fjellberg A, 2003 Fig.9
Isotoma arctica from the USA
Det. Christiansen, K.
2006 © Schimming, L.



Species aspera Bacon, 1914

Species burkhardti Winter, 1967

Species caeca Reuter, 1880

Species caerulea Bourlet, C, 1839:401, B

             Syn.: Isotoma coerulea(sic) Nicolet, H, 1847:374 ISS
             Syn.: Isotoma viridis f. coerulea(sic) Handschin, E in Dahl, F, 1929:67 ISS
             Syn.: Isotoma (Isotoma) viridis var. coerulea(sic) Palissa, A, 1960:378 ISS
             Syn.: Isotoma (Isotoma) viridis ssp. coerulea(sic) Palissa, A in Schwoerbel, J et Zwick, P, 2000:100 ISS
             Syn.: Isotoma coerulea(sic) Fjellberg, A, 2003:94 ISS
             Syn.: Isotoma cerulea(sic) Gudleiffsson, BE et Bjarnadottir, B, 2008:56 ISS
Isotoma caerulea from Belgium
2008.02.07 © Hanssens., B
Isotoma caerulea from the UK
2008.02.27 © Spears, G.
Isotoma caerulea from the UK
2009.03.25 © Ransom, T.
Isotoma caerulea from Norway
After Fjellberg A, 2010 p.4
Isotoma caerulea from Norway
Manubrial dorsal macrosetae: A<B
After Fjellberg A, 2010 p.8
Isotoma caerulea from Norway
2 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma caerulea from Denmark
2010.11.10 © Bildstrup, O.
Isotoma caerulea from the UK
2011.12.05 © Jones, A.
Isotoma caerulea from the UK
2012.11.08 © Jones, A.
Isotoma caerulea from the UK
2013.09.22 © Murray, A.
Isotoma caerulea from Germany
2015.12.18 © Kluck, A.
Isotoma caerulea from the UK
2015.12.17 © David W.
Isotoma caerulea from Holland
2016.11.15 © van Duinen, J.
Isotoma caerulea from France
2017.02.19 © Ramel, A.



Species crassicornis (Handschin, 1926) Weitschat, W & Wichard, W, 2002:86, -

Species decorata Brown, JM, 1926:205

Isotoma viridis var. decorata from Mesopotamia
After Brown JM, 1926 Fig.11

            Syn.: Isotoma viridis var. decorata f. obscura Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. obscura from Palestine
After Handschin E, 1942 Fig.1a

            Syn.: Isotoma viridis var. decorata f. pallida Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. pallida from Palestine
After Handschin E, 1942 Fig.1b

            Syn.: Isotoma viridis var. decorata f. principalis Handschin, E, 1942:405



             Syn.: Isotoma viridis-decorata(sic) Handschin, E, 1942:404 ISS

Species delta MacGillivray, 1896

            Syn.: Isotoma catena Guthrie, JE, 1903:69


Isotoma delta from the USA
2007.03.03 © Montgomery, G.
Isotoma delta from the USA
2007.12.29 © Bryant, P.
Isotoma delta from the USA
2008.03.08 © Justis, P.S.
Isotoma delta from the USA
2008.03.12 © Houston, S.
Isotoma delta from the USA
2008.05.09 © Bradford, A.
Isotoma delta from the USA
2008.11.08 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2008.11.26 © Justis, S.
Isotoma delta from the USA
2008.12.15 © of Baltimore City, T.
Isotoma delta from the USA
2009.01.16 © Montgomery, G.
Isotoma delta from the USA
2009.04.16 © of Baltimore City, T.
Isotoma delta from the USA
2009.04.27 © Kropiewnicki, T.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2012.01.04 © Kouri, J.
Isotoma delta juv. from the USA
2012.01.14 © Justis, S.
Isotoma delta from the USA
2015.03.10 © Himmler, K.
Isotoma delta from the USA
2015.12.31 © Schmidt, M.L.
Isotoma delta from the USA
2016.12.11 © McDonald, G.
Isotoma delta from the USA
2018.12.16 © Owl, Bo.

Species diverticula Yosii, 1966


Isotoma exiguadentata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.232.
Species exiguadentata Salmon, JT, 1941:336


Species fasciata Börner, C, 1909:112

            Syn.: Isotoma virgata Yosii, 1963 =?


Species finitima Scherbakov, 1899, s.d.

Species fulva Schäffer, 1897

Species galapagoensis Jacquemart, 1975 in Peck SB & Jacquemart J, 2011:3 u

Species gandhi Rapoport & Rubio, 1968

Species glauca Packard, AS, 1873:33

Species gracilliseta Börner, C, 1909:113

Species grana Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285

Species hibernica Carpenter, 1906


Podura himalayana from the Himalayas
After Baijal, H.N., 1955 Fig.1
Species himalayana (Baijal, HN, 1955:175)


            Syn.: Podura himmalayana(sic) Baijal, HN, 1955:175 IOS

Species incerta (Nicolet, H, 1847:370) Brook, G, 1884:280

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

             Syn.: Degeeria Incerta(sic) Kolenati, FA, 1858:132 ISS

Species innominata Yosii, 1963

Species iranica Arbea, JI et Kahrarian, M, 2015:77

Isotoma iranica from Iran
After Arbea JI & Kahrarian M, 2015 Fig.23

Species kosiana Bagnall, 1949

Species littoralis (Tanaka, 1981)

Species longispina (Mac Gillivray, 1893) Folsom, 1916:480

             Syn.: Isotoma longipenna(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS

Species mackenziana (Hammer, 1953) Fjellberg, A, 2003:96

Species malvinensis Wahlgren, E, 1906:13

Species manubriata Mac Gillivray, 1896

Species masatierrae Schött, 1921

Species melanocephala Börner, C, 1909:112

Species modesta Nicolet, 1845

Species nepalica (Yosii, 1966)

Species nishihirai Yosii, 1965

Species ornata Wahlgren, E, 1906:15

Species pallidafasciata Salmon, JT, 1941:336

Species perkinsi Carpenter, 1904

Species pinnata Börner, C, 1909:110

            Syn.: Isotoma (Isotoma) pinnata var. coracina Börner, C, 1909:112
            Syn.: Isotoma pinnata var. coracina Uchida, 1938
            Syn.: Isotoma pinnata var. coracina Uchida, 1957
            Syn.: Isotoma pinnata var. coracina Yosii, 1963
            Syn.: Isotoma pinnata ssp. coracina Salmon, JT, 1964:392
             Syn.: Isotoma pinnata ssp. voracina(sic) Salmon, JT, 1964:392 ISS
            Syn.: Isotoma pinnata var. coracina Yosii, R, 1977:157
            Syn.: Isotoma (Isotoma) pinnata var. v-album(sic) Börner, C, 1909:112 IOS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1938 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1957 ISS
             Syn.: Isotoma pinnata ssp. album(sic) Salmon, JT, 1964:392 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Yosii, R, 1977:157 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Potapov, M, 2001:282 ISS
            Syn.: Isotoma pinnata var. alba Uchida, 1943
            Syn.: Isotoma pinnata var. alba Uchida, 1957
            Syn.: Isotoma pinnata ssp. alba Salmon, JT, 1964:392
            Syn.: Isotoma pinnata var. alba Yosii, R, 1977:157




Species plumosa (Salmon, 1969) Lawrence, PN, 1978:269

             Syn.: Rhodanella plumose(sic) Mandal, GP et Hazra, AK, 2009:116 ISS

Species protocinerea Handschin, 1926 -

Species rhopalota Scherbakov, AM, 1898:9

Isotoma rhopalota from Russia
After Scherbakov AM, 1898 Table I Fig.9

Species riparia (Nicolet, H, 1842:62) Nicolet, H, 1847:373, B

Desoria riparia from Switzerland
After Nicolet, H., 1842 Pl.6 Fig.6

            Syn.: Podura aquatica grisea de Geer, 1740
             Syn.: Isotoma Aquaticum griseum(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Podura palustris Bourlet, 1843
             Syn.: Isotoma palustre(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Isotoma viridis var. aquatilis Schött, 1891



             Syn.: Isotoma Riparium(sic) Kolenati, FA, 1858:133 ISS
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.2
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.3
Isotoma viridis var. riparia from Germany
After Heymons R & Heymons H, 1909 Fig.10
Isotoma riparia from Norway
After Fjellberg A, 2003 Fig.8
Isotoma riparia from Estonia
2006.03.18 © Tartes, U.
Isotoma riparia from the UK
After Shaw P, 2009 ©
Isotoma riparia from France
Predation by pseudoscorpion
2009.11.16 © Lebeaux, P.
Isotoma riparia from Norway
After Fjellberg A, 2010 p.6
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.20 © Lind, J.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from the UK
2014.02.04 © Valentine., B
Isotoma nr riparia from Estonia
2015.02.07 © Tartes, U.
Isotoma riparia from Holland
2016.03.05 © de Wilde, A.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma riparia from Holland
2018.04.20 © van Duinen, J.
Isotoma riparia from Estonia
2019.02.09 © Tartes, U.



Species sarkundensis Baijal, HN, 1958:354

Species schalleri Winter, 1967

Species semenkevitshi Scherbakov, AM, 1898:8

Species silvatica Schäffer, 1897

Species sinensis Yue & Yin, 1999

Species sp. auct. anon., 1879

Species sp. Wise, 1970, s.i.

Species sp. Ellis, 1974:121, i.s.

Species sp. Maes & Palacios-Vargas, 1988:4

Species sp. Palacios-Vargas, 1990:21

Species sp. Palacios-Vargas & Gómez, 1991

Species sp. Palacios-Vargas & Gómez-Anaya, 1995:26

Species sp. Zeppelini & Castaño, 1995:10

Species sp. Palacios-Vargas, JG et Gamboa-Vargas, JA, 1997:90

Species sp. Brand, RH et Dunn, CP, 1998:238

Species sp. Brand, RH et Dunn, CP, 1998:238

Species sp. Rebek, EJ, Young, DK et Hogg, DB, 1999:60

Species spelaea Joseph, 1882

Species spinicauda (Bonet, 1930) Denis, JR, 1936:263

            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotoma decorata Martynova & Chikatunov, 1968 =?
            Syn.: Isotoma decorata Martynova, 1968 =?
            Syn.: Isotoma decorata Yosii, 1971 =?
            Syn.: Isotoma decorata Martynova & Vtorov, 1973 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1974 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1975 =?
            Syn.: Isotoma (Isotoma) decorata Tanaka & al, 1978=?
            Syn.: Isotoma (Isotoma) decorata Janetschek, 1990=?
            Syn.: Acanthisotoma quadridentata Huang & Liang, 1994 =?


Species subcaerulea Kos, 1952

Species subviridis Folsom, JW, 1937:115

Species tahitiensis Salmon, 1964

Species termitophila Womersley, 1934

Species tesselata Rapoport, 1962

Species tigrinella Wahlgren, E, 1906:14

            Syn.: Isotoma tigrina Schött, 1899 nec Nicolet, 1842

Species tridenticulata Schäffer, 1896

             Syn.: Isotoma 3-denticulata(sic) Absolon, K, 1901:32 ISS

Species tridentifera Schött, H, 1917:9

Isotoma tridentifera from Australia
After Schött H, 1917 Fig.12

  Subspecies edenticulata Womersley, 1934

Species tridentina Tarsia in Curia, 1941

Species turbotti (Salmon, JT, 1949:30) Greenslade P, 1990

Species vaillanti Murphy, 1958

Species variodentata Dunger, WG, 1982:60

  Forma marginata Dunger, 1982:62

  Forma principalis Dunger, 1982:63

  Forma unidentata Dunger, 1982:62

Species viridis Bourlet, C, 1839:401, B

            Syn.: Podura viatica Linnæus, 1746
            Syn.: Podura aquatilis Müller OF, 1776, sensu Gisin, 1960
            Syn.: Podura plumbea Müller, 1776
Isotoma plumbea from the USA
After Packard AS, 1872 Pl.10 Fig.6
Isotoma plumbea from the USA
After Packard AS, 1872 Pl.10 Fig.7

            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
            Syn.: Isotoma arborea Bourlet, C, 1839:94
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H in Desor, E, 1841:385
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H, 1842:60
Desoria cylindrica from Switzerland
After Nicolet H, 1842 Pl.6 Fig.1

             Syn.: Heterotoma Cylindricum(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Desoria (Deuxième division) annulata Nicolet, H, 1842:62
Isotoma viridis var. annulata from the USA
2007.03.27 © McClarin, J.
Isotoma viridis var. annulata from the USA
2007.04.30 © Cowen, R.
Isotoma viridis var. annulata from Holland
2008.10.10 © Baas, A.H.
Isotoma viridis var. annulata from the UK
2013.10.05 © Evans, J.
Isotoma viridis var. annulata from Germany
2015.01.01 © Beck, K.
Isotoma viridis var. annulata juv. from Germany
2015.01.16 © Landgraf, A.
Isotoma viridis var. annulata from Holland
2015.02.05 © van Duinen, J.
Isotoma viridis var. annulata from the USA
2015.03.22 © Abbott, J.C.
Isotoma viridis var. annulata juv. from Belgium
2016.03.28 © Huskens, M.-L.
Isotoma viridis var. annulata from the USA
2018.01.10 © Abbott, J.C.


            Syn.: Desoria (Deuxième division) ebriosa Nicolet, H, 1842:61
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
            Syn.: Desoria (Deuxième division) fusca var. var. Nicolet, H, 1842:63
            Syn.: Desoria (Deuxième division) pallida Nicolet, H, 1842:61
Isotoma viridis var. pallida from Germany
2005 © Menzel, E.
Isotoma viridis var. pallida from the UK
2006 © Barton, T.
Isotoma viridis var. pallida from Sweden
2006 © Hall, K.
Isotoma viridis var. pallida
2014.01.19 © Evans, J.


            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
            Syn.: Heterotoma chlorata Gervais in Walckenaër, 1844
Hétérotome verte from France
Heterotoma chlorata
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.6

            Syn.: Isotoma Desmarestii(sic) Gervais in Walckenaër, 1844 IOS
Isotome de Desmarest from France
Isotoma Desmarestii(sic)
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.11

             Syn.: Isotoma Desmarestii(sic) Nicolet, H, 1847:374,si ISS
            Syn.: Isotoma Nicolet(sic) Gervais in Walckenaër, 1844 IOS
Isotome Nicolet(sic) from France
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.12

             Syn.: Isotama(sic) Nicoletii(sic) Nicolet, H, 1847:374,si ISS
             Syn.: Isotoma Nicoletii(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Isotoma lineata Lubbock, 1862
            Syn.: Isotoma palustris Tullberg, 1871
            Syn.: Isotoma palustris var. unicolor Tullberg, 1871
            Syn.: Isotoma Belfragei(sic) Packard, AS, 1873:33 IOS
             Syn.: Isotoma Belfragei(sic) MacGillivray, 1891 ISS
            Syn.: Isotoma purpurascens Packard, AS, 1873:34
            Syn.: Isotoma tricolor Packard, 1873 partim
            Syn.: Isotoma tricolor MacGillivray, 1891 partim
            Syn.: Isotoma tricolor Schött, 1893
            Syn.: Isotoma tricolor von Dalla Torre, 1895
            Syn.: Isotoma tricolor MacGillivray, 1896 partim
            Syn.: Isotoma tricolor Folsom, 1903 partim
            Syn.: Isotoma tricolor Linnaniemi, 1912
            Syn.: Isotoma tricolor Bacon, 1914 partim
            Syn.: Isotoma tricolor Bartholin, 1916
            Syn.: Isotoma tricolor Folsom, JW, 1919:12A
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Mills, HB, 1934:63
            Syn.: Isotoma tricolor Stach, J, 1947:422 partim
            Syn.: Isotoma tricolor Maynard, EA, 1951:110
            Syn.: Isotoma tricolor Uchida, 1957
            Syn.: Isotoma tricolor Salmon, JT, 1964:403
            Syn.: Isotoma tricolor Christiansen, K et Bellinger, P, 1998:853
            Syn.: Isotoma cincta Tullberg, 1876
Isotoma viridis var. cincta from Novaja Selmja
After Schött H, 1893 Taf.V Fig.5

            Syn.: Isotoma viridis var. arctica Schött, H, 1893:61
            Syn.: Isotoma viridis f. principalis Schött, H, 1893:59
Isotoma viridis forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.1

            Syn.: Isotoma riparia Lameere, A, 1895:594
            Syn.: Isotoma capitola MacGillivray, 1896
            Syn.: Isotoma delta Mac Gillivray, 1896
            Syn.: Isotoma glauca MacGillivray, 1896
            Syn.: Isotoma montana MacGillivray, 1896
            Syn.: Isotoma tridentata MacGillivray, 1896
            Syn.: Isotoma viridis var. violacea Lie-Pettersen, 1898
Isotoma viridis var. violacea from Holland
2008.01.23 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2008.01.27 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2009.03.18 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2009.04.05 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2014.01.05 © van Duinen, J.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.
Isotoma viridis var. violacea from Belgium
2014.03.02 © Huskens, M.-L.
Isotoma viridis var. violacea from Belgium
2018.01.18 © Huskens, M.-L.
Isotoma viridis var. violacea from Belgium
2018.03.28 © Huskens, M.-L.
Isotoma viridis var. violacea from the UK
2019.02.10 © Cann, A.J.
Isotoma viridis var. violacea from the UK
Manubrial thickening with 1 tooth
2019.02.10 © Cann, A.J.


            Syn.: Isotoma paradoxa Carl, J, 1899:317
            Syn.: Isotoma viridis f. genuina Börner, C, 1901:50
            Syn.: Isotoma catena Guthrie, JE, 1903:69
            Syn.: Isotoma viridis f. micans Ågren, 1903
            Syn.: Isotoma hibernica Carpenter, 1906 =?
            Syn.: Isotoma aspera Bacon, 1914
            Syn.: Isotoma spinicauda Bonet, 1930
            Syn.: Isotoma (Isotoma) viridis var. pomona Folsom, JW, 1937:115
            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotomurus bemakus Wray, 1952
            Syn.: Orchesella rubra Scott, 1963 partim holotype
            Syn.: Rhodanella plumosa Salmon, 1969 =?
            Syn.: Tetracanthura mirabilis Martynova, 1971
            Syn.: Desoria riparia Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Desoria riparia Christiansen, K et Bellinger, P, 1998:853



Isotoma viridis from Alaska
After Folsom JW, 1902 Pl.IV Fig.4
Isotoma viridis f. typica from Germany
After Heymons R & Heymons H, 1909 Fig.9
             Syn.: Isotoma ciridis(sic) Salmon, JT, 1964:407 ISS
Isotoma viridis from the UK
2005 © Hopkin, S.P.
Isotoma viridis from the UK
2006 © Valentine., B
Isotoma viridis from Germany
2006 © Steiner, A.
Isotoma viridis from the UK
2006.04.08 © Read, D.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from Germany
2007 © Peters, J.
Isotoma viridis from the UK
2007.02.23 © Valentine., B
Isotoma viridis from the UK
2007.02.24 © Barton, T.
Isotoma viridis from the UK
2007.02.28 © Valentine., B
Isotoma viridis from the UK
2007.03.02 © Barton, T.
Isotoma viridis from the UK
2007.03.05 © Barton, T.
Isotoma viridis from the Netherlands
2007.03.21 © Baas, A.H.
Isotoma viridis from the USA
2007.10.21 © Moorehead, C.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from the Netherlands
2008.01.22 © Baas, A.H.
Isotoma viridis from the UK
2008.02.03 © Tonsbeek, M.
Isotoma viridis from Belgium
2008.03.23 © Oger, P.
Isotoma viridis from the UK
2008.04.07 © Robertson, A.
Isotoma viridis from the Netherlands
2008.10.08 © Baas, A.H.
Isotoma viridis from the Netherlands
2008.11.16 © Baas, A.H.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from the USA
2009.02.10 © Cowen, R.
Isotoma viridis from the USA
2009.03.05 © Cowen, R.
Isotoma viridis from the USA
2009.03.04 © Murray, T.
Isotoma viridis from Norway
After Fjellberg A, 2010 p.5
Isotoma viridis from Norway
Single ventroapical manubrial tooth
After Fjellberg A, 2010 p.7
Isotoma viridis from the UK
2010.02.06 © Barton, T.
Isotoma viridis from the UK
2010.02.14 © Robertson, A.
Isotoma viridis from the UK
2011.12.03 © Jones, A.
Isotoma viridis (bar=2µm)
Hexagonal-rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B'
Isotoma viridis (bar=500nm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B''
Isotoma viridis from Holland
2012.01.27 © van Duinen, J.
Isotoma viridis from the UK
2012.11.08 © Jones, A.
Isotoma viridis from Holland
2015.02.06 © van Duinen, J.
Isotoma viridis from Germany
2015.02.15 © Kluck, A.
Isotoma viridis from Holland
With parasitic larva of Acari:Erythraeoidea
2015.07.26 © van Duinen, J.
Isotoma viridis from Belgium
Collophore pressed against surfacewaterfilm
2016.01.14 © Huskens, M.-L.
Isotoma viridis from the UK
2016.02.15 © Horton, M.
Isotoma viridis from Belgium
Cephalic pigmentation
2016.11.27 © Huskens, M.-L.
Isotoma viridis from Belgium
Post antennal organ (PAO), half the size of nearest ocellus
2016.11.27 © Huskens, M.-L.
Isotoma viridis from Belgium
With phoretic Nematoda
2018.01.06 © Huskens, M.-L.
Isotoma viridis from the UK
Single ventroapical manubrial tooth
2019.02.04 © Cann, A.J.
Isotoma viridis from the UK
Variant with patched abd.5-6
2019.02.04 © Cann, A.J.
Isotoma viridis from the UK
Variant with patched abd.5-6
2019.02.04 © Cann, A.J.







Genus Pseudisotoma Handschin, 1924

  Syn.: Isotoma Pseudoisotoma(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
  Syn.: Collembola Psedisotoma(sic) Hornung-Leoni, CT et Palacios-Vargas, JG, 2012:67 ISS

Species agrana (Lee BH & Kim JT in Lee, B-H, Kim, B-J & Kim, J-T, 1993:288) Potapov, 2001


Pseudisotoma anamola from Lebanon
After Christiansen, K., 1958 Pl.10 Fig.29.
Species anamola Christiansen, 1958:76


Species himalayana Yosii, 1971

Species microchaeta Cassagnau, P, 1959:85

Species monochaeta (Kos, 1942) Jacquemart, S, 1967:4, B

Species ohtanii (Yosii, R, 1972:90) Potapov, 2001

Species sensibilis (Tullberg, T, 1877:36) Agrell, I, 1939:8, t.t., B

            Syn.: Isotoma turicensis Haller, 1880
            Syn.: Isotoma dubia Reuter, 1895
            Syn.: Isotoma determinata Mac Gillivray, 1896
            Syn.: Isotoma parva Mac Gillivray, 1896
            Syn.: Isotoma longidens Schäffer, 1896
            Syn.: Isotoma monstrosa Schäffer, 1896
            Syn.: Isotoma inopinata Axelson, 1902
            Syn.: Isotoma arborea Evans, 1908


Pseudisotoma sensibilis from Belgium
2006.10.12 © Schelfhout, L.
Pseudisotoma sensibilis from Belgium
2008.06.09 © Vuijlsteke, M.
Pseudisotoma sensibilis from Holland
2012.10.28 © van Duinen, J.
Pseudisotoma sensibilis from the USA
Abd.5+6 fused
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the UK
2017.07.19 © Byrne, J.

Species wisei Deharveng, 1980


Genus Semicerura Maynard, EA, 1951:99

Species bishopi Maynard, EA, 1951:99, t.t.

Semicerura nr bishopi
2012.01.25 © Walter, D.E.
Semicerura nr bishopi
2012.01.25 © Walter, D.E.

Species goryshini Martynova, 1969

Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org

Species multispinata (James, 1933) Christiansen, KA in Dindal, DL, 1990:987

             Syn.: Semicerura miltispinata(sic) Christiansen, KA in Dindal, DL, 1990:987 ISS



Genus Heteroisotoma Stach, 1947

  Syn.: Isotomidae Sibirisotoma Rusek, 1991

Species carpenteri (Börner, C, 1909:109), t.t.

            Syn.: Isotoma (Isotoma) andrei Mills, HB, 1934:58
            Syn.: Isotoma (Isotoma) difficilis Folsom, JW, 1937:89

Species dilatosetosa Martynova, 1980

Species heteronomica Jie, D, Potapov, M et Sokolova, E, 2011:55

            Syn.: Heteroisotoma sp.nov. Potapov & Polyakov, 2007

Species jacutica Martynova, 1980

Species sinorossica Jie, D, Potapov, M et Sokolova, E, 2011:51

Heteroisotoma sinorossica from China
After Jie & al, 2011 Fig.25

Species stebaevae (Rusek, 1991) Potapov & Stebaeva, 1992

            Syn.: Sibirisotoma stebajevae(sic) Rusek, 1991 IOS


             Syn.: Heteroisotoma stebajevae(sic) Jie, D, Potapov, M et Sokolova, E, 2011:55 ISS
             Syn.: Heteroisotoma stebaievae(sic) Stevens, MI et D'Haese, CA, 2016:4 ISS





Genus Folsomotoma Bagnall, 1949 key

  Syn.: Isotoma Sorensia Salmon, 1948


Species anomala (Salmon, 1948)

            Syn.: Folsomotoma minuta Fanciulli, PP, Leo, C, Convey, P, Frati, F et Carapelli, A, 2018:394 =?


Species bioculata (Womersley, 1934) Salmon, JT, 1964:412, t.t.

            Syn.: Isotoma bi-oculata(sic) Womersley, 1934,tt IOS

Species boerneri (Enderlein, 1903)

            Syn.: Isotoma börneri(sic) Enderlein, 1903 IOS
            Syn.: Folsomotoma subflava Fanciulli, PP, Leo, C, Convey, P, Frati, F et Carapelli, A, 2018:394 =?




Species kerguelensis (Enderlein, 1903)

             Syn.: Isotoma octo-oculata(sic) ssp. Kerguelensis(sic) Carpenter, GH, 1907:476 ISS

Species marionensis (Deharveng, 1981)

Species minuta (Salmon, JT, 1949:24)

Species octooculata (Willem, V, 1901:262) Potapov, 2001

            Syn.: Isotoma octo-oculata(sic) Willem, V, 1901:262 IOS
            Syn.: Isotoma sp. Carl, 1907
            Syn.: Isotoma octo-oculata(sic) var. gracilis Carpenter, GH, 1907:476
Isotoma octo-oculatus(sic) var. gracilis
from Antarctica
After Carpenter GH, 1907 Plate II Fig.9


            Syn.: Isotoma masatierrae Stach, 1947 =?
            Syn.: Isotoma octo-oculata(sic) ssp. kerguelensis Salmon, JT, 1964:411



             Syn.: Isotoma octo-oculata(sic) Willem, V, 1902:13 ISS
Isotoma octo-oculata from Antarctica
After Willem V, 1902 Pl.IV Fig.7

             Syn.: Isotoma octo-oculata(sic) Wahlgren, 1906 ISS
             Syn.: Isotoma octo-oculata(sic) Carl, 1907 ISS
             Syn.: Isotoma octo-oculata(sic) Carpenter, 1907 ISS
             Syn.: Isotoma octo-oculata(sic) Enderlein, 1909 ISS
             Syn.: Isotoma octo-oculata(sic) Ivanoff, 1913 ISS
             Syn.: Isotoma octo-oculata(sic) Womersley, 1937 ISS
             Syn.: Isotoma octo-oculata(sic) Denis, 1947 ISS
             Syn.: Parisotoma octo-oculata(sic) Salmon, 1949 ISS
             Syn.: Isotoma octo-oculata(sic) Gressitt & Weber, 1959 ISS
             Syn.: Parisotoma octo-oculata(sic) Brown, 1964 ISS
             Syn.: Parisotoma octo-oculata(sic) Salmon, JT, 1964:411 ISS
             Syn.: Isotoma 8-oculata(sic) Deharveng, 1981 ISS
Folsomotoma octooculata from Antarctica
After Fanciulli PP & al, 2018 Fig.1A

  Subspecies ovata Salmon, JT, 1949:39





Species punctata (Wahlgren, E, 1906:14)

            Syn.: Sorensia dreuxi Delamare Deboutteville & Massoud, 1966
            Syn.: Isotoma (Folsomotoma) subflava Greenslade, P in Dallai, R, 1986:56


Species subflava (Salmon, JT, 1949:22)

            Syn.: Sorensia dreuxi Wise, KAJ, 1970:194
            Syn.: Sorensia dreuxi Gressitt, 1970
            Syn.: Sorensia dreuxi Dreux, 1978 =?



Genus Desoria Agassiz M & Nicolet H in Desor E, 1841:384

  Syn.: Podures(sic) Desoria Nicolet H in Desor, E, 1841:126
  Syn.: Desoria Première division Nicolet H in Desor, E, 1841:385
  Syn.: ... Istoma(sic) Davenport, 1903 ISS
  Syn.: ... Isotomurs(sic) Fjellberg, A, 2007:99 ISS


Desoria sp. from France
2008.02.24 © Jacquemin, G.
Desoria sp. from the USA
2009.12.29 © Parker, H.S.

Remark by Duffin, C. 2009:60: Bearing in mind how much Cécile (wife of Agassiz) detested Desor as a social parasite blighting the Agassiz family it is interesting to speculate how she might have appreciated the irony of his lending his name to a 'flea'.




Desoria sp. from Singapore
2011.01.04 © Koh, J.
Desoria sp. from Tasmania
2014.04.16 © Murray, A.
Desoria olivacea from Belgium
Ocelli BDE in straight line
2018.02.18 © Huskens, M.-L.

Species sp.1 Fjellberg, A, 2010:40

Species sp.2 Fjellberg, A, 2010:40

Species ahiehie Christiansen, K et Bellinger, P, 1992:166

Species alaskensis Fjellberg, 1978

Species albella (Packard, AS, 1873:32)

            Syn.: Isotoma Walkerii(sic) Packard, 1871
             Syn.: Isotoma walkerii(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS


            Syn.: Isotoma leonina Packard, AS, 1873:32

Species albicornis Fjellberg, A, 2010:32

Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.115C
Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.111

Species arctica White A in Sutherland, PC, 1852:ccviii

Desoria arctica from the Arctic region
After White A in Sutherland PC, 1852 Fig.7.

Species ater Fjellberg, A, 2010:34

Desoria ater from the USA
After Fjellberg A, 2010 Fig.115D
Desoria ater from the USA
After Fjellberg A, 2010 Fig.113

Species atkasukiensis Fjellberg, 1978

Species banffensis Christiansen, K et Bellinger, P, 1998:794

Species bendixenae Christiansen, K et Bellinger, P, 1992:168

Species blekeni Leinaas, 1980

Species blufusata Fjellberg, 1978

Species breviseta Babenko, Potapov & Taskaeva, 2017

Species caeruleatra (Guthrie, JE, 1903:70)

Species canadensis Brown, 1932

            Syn.: Isotoma olivacea Stach, J, 1947:404


Isotoma (Desoria) canadensis from Canada
After Brown in Christiansen K & Bellinger P, 1980 Fig.583A
Desoria canadensis from Canada
2016.03.12 © Schmidt, H.
Desoria canadensis from Canada
2017.01.02 © Levac, G.

Species cancellarei (Christiansen & Bellinger, 1980) Fjellberg, A, 2010:33

Species capra Fjellberg, A, 2010:40

Species cera Christiansen, K et Bellinger, P, 1998:804

Species choi (Lee, B-H, 1977:166)

Species christianseni Fjellberg, 1978

Species creli Fjellberg, 1978

Species cryophila Fjellberg, A, 2010:38

Desoria cryophila from the USA
After Fjellberg A, 2010 Fig.115G

Species divergens (Axelson, 1900) Kaprus, IJ, Pomorski, RJ, Skarzynski, D & Potapov, MB, 2005:894

Species duodecimoculata Denis, 1927

             Syn.: Desoria duodecemaculata(sic) Fiera, C, 2007:23 ISS

Species elegans (Carl, J, 1899:319)

Species eureka Christiansen, K et Bellinger, P, 1998:806

Species fennica (Reuter, 1895)

             Syn.: Isotoma Formáneki(sic) Absolon, K, 1901:212 ISS

Species fideli (Rapoport & Izarra, 1962)

Species fjellbergi Najt, 1981

Species flora (Christiansen & Bellinger, 1980)

Desoria flora from the USA
2014.02.19 © Klein, T.

Species garibaldii Fjellberg, A, 2010:36

Desoria garibaldii from the USA
After Fjellberg A, 2010 Fig.115F

Species gelida Folsom, JW, 1937:106

Desoria gelida from the USA
2010.04.01 © Murray, T.
Desoria gelida from the USA
2010.04.30 © Hodge, K.

Species gorodkovi Martynova, 1969

Species grisea (Lubbock, J, 1870:278) Fjellberg, A, 2007:130

            Syn.: Isotoma grisea var. intricata Parfitt, 1891
            Syn.: Isotoma grisescens Schäffer, 1896
             Syn.: Isotoma olivacea var. griscescens(sic) Handschin, E in Dahl, F, 1929:70 ISS
            Syn.: Isotoma grisescens Denis, 1931 =?


Isotoma grisea from the UK
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Abdomen
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Head and thorax
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Eyepatch
Ocelli G & H not drawn
After Lubbock J, 1873 Pl.39

             Syn.: Isotoma giresea(sic) Dürkop, 1934 ISS



Species hiemalis (Schött, H, 1893:70) Bulimar, 1987

            Syn.: Isotoma Theobaldi(sic) Carl, J, 1899:315 IOS
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma nivicola Axelson, 1900


Desoria hiemalis from Germany
2006 © Kohl, F.
Desoria hiemalis from the USA
2007.02.18 © Murray, T.
Desoria hiemalis from Western Estonia
2009.02.26 © Tartes, U.
Desoria hiemalis from Russia
2013.01.01 © Serge, L.



Species hissarica Martynova & Chikatunov, 1968

Species hoodensis Christiansen, K et Bellinger, P, 1998:814

Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.03.12 © Dobbs, Br.
Desoria hoodensis? from the USA
2018.09.22 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.

Species husaini Yosii & Ashraf, 1965

Species imparidentata (Stach, J, 1964:13) Potapov, 2001

Species infuscata Murphy, 1959

Species intermedia (Schött, 1902) Fjellberg, A, 2007:125, B

            Syn.: Isotoma germanica Hüther & Winter, 1961


Remark by Potapov, M. 2001:120: Obscure species.

Desoria intermedia from Scandinavia
2008.04.04 © Fjellberg, A.

Species iuxta Dunger, WG, 1982:68

Species japonica (Yosii, 1939) Yosii, 1954

Species jayasrae Bhattacharjee, 1984

Species juberthiei Gers & Najt, 1983

Species kaszabi Dunger, WG, 1982:70

Species komarkovae Fjellberg, 1978

Species kuraica (Martynova, 1974:19)

Species lucama Wray, 1952

Species manitobae Fjellberg, 1978

Species marissa (Folsom, JW, 1937:89) Miranda Rangel & Palacios-Vargas, 1992

Species maxillosa Fjellberg, 1978

Species mazda Yosii, 1971

Species mossopi (Womersley, H, 1934:454)

Species multisetis (Carpenter & Phillips, 1922) Miranda Rangel & Palacios-Vargas, 1992

Species nahuelensis (Rapoport & Rubio, 1968)

Species negishina (Börner, C, 1909:108) Yosii, R, 1977:156, s.i.

            Syn.: Isotoma sp. Andreen, A, Malterre, J, Needham, J, Olivo, O, Pessoa, SB et Zhukov-Verezhnikov, NN, 1952:18


Species neglecta (Schäffer, 1900) Dányi, L & Traser, G, 2008:46

            Syn.: Isotoma affinis Axelson, 1900
            Syn.: Isotoma tigrina ab. coerulea Ågren, 1903


Desoria neglecta from Norway
2011 © Fjellberg, A.

Species nigra Mac Gillivray, 1896

Species nigrifrons Folsom JW in Mills, HB, 1934:60

Species nivalis Papon, 1856

Species nivalis (Carl, 1910) Miranda Rangel & Palacios-Vargas, 1992

Species nivea (Schäffer, 1896) Fjellberg, 1978

            Syn.: Isotoma albella? Skarzynski, D, Pomorski, RJ, Smolis, A, Weiner, WM, Szeptycki, A, Slawska, M et Sterzynska, M, 2002:35=?


Species nixoni Fjellberg, 1978

Species nympha Snider, RJ et Calandrino, FJ, 1987:5

Isotoma (Desoria) nympha from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.1

Species occulta (Börner, C, 1909:110) Yosii, R, 1977:156

Species olivacea (Tullberg, 1871) Yosii, R, 1977:156, B

            Syn.: Isotoma voraginum Uzel, 1891
            Syn.: Isotoma olivacea f. genuina Börner, C, 1901:56
            Syn.: Isotoma olivacea f. principalis Linnaniemi, 1912
            Syn.: Spinisotoma regina Stach, J, 1930:290
            Syn.: Isotoma grisescens Denis, 1931
            Syn.: Isotoma tigrina Denis, 1937
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Spinisotoma ephippiata Brian, 1945
            Syn.: Proisotoma subsegmentata Scott, 1959
            Syn.: Isotoma grisea Womersley, 1930 =?
            Syn.: Isotoma grisea Gisin, H, 1960:211 =?
            Syn.: Isotoma (Isotoma) grisea Goto & Lawrence, 1964 =?
            Syn.: Isotoma grisea Palissa, 1964 =?
            Syn.: Isotoma grisea Bolger, 1987 =?


Desoria olivacea from Holland
2009.04.05 © Baas, A.H.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Norway
2011 © Fjellberg, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Ovoid post-antennal organ (PAO)
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Abd.5 mesosetae 2/3rds length abd.5
2012.10.31 © Murray, A.
Desoria olivacea from Holland
2014.05.04 © Wieringa, P.
Desoria olivacea from Belgium
2017.04.20 © Huskens, M.-L.
Desoria olivacea from Belgium
2018.02.18 © Huskens, M.-L.

Species olympica Fjellberg, A, 2010:39

Species pallipes (Kos, 1952)

Species pamirensis Martynova, 1975

Species pilifrons Fjellberg, A, 2010:37

             Syn.: Desoria pilifons(sic) Fjellberg, A, 2010:28 ISS

Species pjasini Martynova, 1974

             Syn.: Desoria pjaisini(sic) Greenslade, P et Potapov, M, 2015:338 ISS

Species potapovi Fjellberg, A, 2007:126

Species propinqua (Axelson, 1902) Potapov, 2001

            Syn.: Isotoma poppiusi Axelson, 1903
            Syn.: Spinisotoma pectinata Stach, 1926
Spinisotoma pectinata from Poland
After Stach, J., 1947 Plate XXXVIII Fig.1.

            Syn.: ... propinqua var. hexaspina Agrell, 1936
            Syn.: Desoria sp. Yosii, R, 1977:155
            Syn.: Isotoma (Desoria) graeca Najt, 1981



Species pseudomaritima (Stach, J, 1947:436)

Isotoma pseudomaritima from Poland
After Stach, J., 1947 Plate XLVII Fig.1.

Species quadra Christiansen & Bellinger, 1980

Species randiella Fjellberg, 1978

Species rosea Fjellberg, A, 2010:38

Species saltans Agassiz M & Nicolet H in Desor E, 1841:385 tt

Desoria saltans from Switzerland
After Nicolet H in Desor E, 1841 Pl.2 Fig.1

            Syn.: ... puce des neiges rougeatre et velue Aristote 350 BC
            Syn.: Thysanoures(sic) puce du Mont-Rose Agassiz M in Desor, E, 1841:125
            Syn.: Thysanoures(sic) puce de glacier Desor, E, 1841:126
            Syn.: Desoria (Première division) glacialis Nicolet, H, 1842:58
Desoria glacialis from Switzerland
After Nicolet H, 1842 Pl.5 fig.10




Isotoma saltans from Austria
After Kopeszki H, 2000 Abb.3
Desoria saltans from Austria
2016.02.27 © van Leeuwen, J.

Species sandersoni Wray, 1952

Species spatiosa (Uchida, H & Tamura, H, 1968:10) Potapov, 2001

Pteronychella spatiosa from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo31

Species speciosa Mac Gillivray, 1896

Species subaequalis Folsom, JW, 1937:107

Species subantarctica (Izarra, 1972)

Species tadzhika Martynova & Chikatunov, 1968

Species taigicola Fjellberg, 1978

Species taimyrica (Martynova, 1974) Potapov, 2001

Species tianshanica Hao & Huang, 1995

Species tigrina Nicolet, H, 1842:59, B

Desoria tigrina from Switzerland
After Nicolet H, 1842 Pl.5 Fig.11

            Syn.: Isotoma grisea Gisin, 1944
            Syn.: Isotoma grisea Fjellberg, 1979
            Syn.: Isotoma grisea Christiansen & Bellinger, 1980
            Syn.: Isotoma grisea Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Isotoma grisea Christiansen & Bellinger, 1998
            Syn.: Isotoma affinis Axelson
            Syn.: Isotoma stachi Denis, 1929
            Syn.: Spinisotoma dimorpha Womersley, 1934
            Syn.: Isotoma olivacea Womersley, 1934
            Syn.: Isotoma olivacea Womersley, 1939
            Syn.: Isotoma klovstadi Womersley, 1937
            Syn.: Desoria grisescens Hopkin, 2007



Desoria tigrina from the UK
Setae on fifth abdominal segment
2005 © Hopkin, S.P.
Desoria tigrina from France
2008.02.24 © Jacquemin, G.
Desoria tigrina from Holland
2011.03.22 © van Duinen, J.
Desoria tigrina from the UK
2012.11.05 © Jones, A.
Desoria tigrina from the UK
2012.11.05 © Jones, A.
Desoria tigrina juv. from Iran
Ecomorphic form
After Kahrarian M & Arbea J, 2013 Fig.2
Desoria tigrina juv. from Iran
Ecomorphic abdominal spines
After Kahrarian M & Arbea J, 2013 Fig.3
Desoria tigrina from Iran
After Kahrarian M & Arbea J, 2013 Fig.4
Desoria tigrina from Holland
2015.02.06 © van Duinen, J.
Desoria tigrina from the UK
2016.01.07 © Barton, T.
Desoria tigrina from France
2016.12.06 © Garcelon, P.
Desoria tigrina from France
2017.02.05 © Picard, J.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.
Desoria tigrina from Belgium
2017.06.29 © Huskens, M.-L.
Desoria tigrina from Belgium
2017.11.15 © Huskens, M.-L.
Desoria tigrina from Belgium
2017.11.15 © Huskens, M.-L.
Desoria tigrina from France
2017.11.18 © Picard, J.
Desoria tigrina from France
2017.11.18 © Picard, J.
Desoria tigrina from France
2018.02.15 © Real, J.-A.
Desoria tigrina from France
2018.08.14 © Vol, F.
Desoria tigrina from the UK
2018.08.31 © Cann, A.J.
Desoria tigrina from the UK
Longest setae on abd.5 shorter than abd.5
2019.02.15 © Cann, A.J.

Species tolya Fjellberg, A, 2007:120

Species torildae Fjellberg, 1978

Species triangularis Fjellberg, A, 2010:33

Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.115E
Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.110

Species trispinata (Mac Gillivray, 1896) Thibaud, 1977

            Syn.: Isotoma maritima ssp. meridionalis Altner, H, 1963:47
             Syn.: Isotoma meridionale(sic) Poinsot, 1971 ISS
             Syn.: Isotoma meridionale(sic) Poinsot, N, 1972:686 ISS
            Syn.: Isotoma setinornata Lee, B-H, 1977:164
            Syn.: Isotoma setinornata Yi, 1976 nn


             Syn.: Isotoma tarspinata(sic) Lesshafft, 1977 ISS
Desoria trispinata from the Azores
2010.09.15 © E.D.E.N.
Desoria trispinata from the UK
2010.04.22 © Shaw, P.
Det.: Fjellberg, A.

             Syn.: Desoria trispina(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1666 ISS
Desoria trispinata from Japan
2013.03.12 © Hara, Y.
Desoria trispinata from Japan
Abd.4-6
2013.03.12 © Hara, Y.
Desoria trispinata from Germany
2014.09.27 © Beck, K.
Desoria trispinata from Germany
2014.10.05 © Jäschke, J.
Isotoma (Desoria) trispinata from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574
Desoria trispinata from Holland
2016.02.11 © van Duinen, J.
Desoria trispinata from Belgium
Eggs deposited in cluster
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Freshly deposited eggs are white and smooth
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
In orange eggs the chorion has hardened
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Chorion is hydrophobe
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Chorion bursts at equator and the halves move to the poles
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
The egg flattens laterally
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Eyepatches of embryo become visible
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar caps are formed
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar cap in polar aspect
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar cap in frontal aspect
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Hatchling
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
2017.12.24 © Huskens, M.-L.
Desoria trispinata from Belgium
2018.06.30 © Huskens, M.-L.



Species truncata Waltz & Hart, 1995

Desoria truncata from the USA
2006 © Bernard, E.C.
Desoria truncata from the USA
2007.09.05 © Barnd., B

Species tshernovi Martynova, 1974

            Syn.: Isotoma nanseni Fjellberg, 1978


             Syn.: Isotoma tschernovi(sic) Fjellberg, 1995 ISS
Desoria tshernovi from Norway
2011 © Fjellberg, A.



Species tuckeri Christiansen & Bellinger, 1980

Species tunica Christiansen & Bellinger, 1980

Species uniens (Christiansen, K et Bellinger, P, 1980:760)

Desoria uniens from the USA
After Fjellberg A, 2010 Fig.115B

Species violacea (Tullberg, T, 1877:36), B

            Syn.: Isotoma neglecta Schäffer
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma olivacea Tuxen, 1944 partim
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Isotoma inupikella Fjellberg, 1978
            Syn.: Desoria tolya Hågvar, S, 2010:283


Desoria violacea from the UK
Mucro with 4 teeth
2005.04.01 © Hopkin, S.P.
Desoria violacea (bar=2µm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.4E
Desoria violacea from the UK
2014.01.19 © Valentine., B

Species yukinomi (Yosii, 1939) Yosii, R, 1977:157

Species zlotini Martynova, 1968


Genus Gnathisotoma Cassagnau, P, 1957:393

Species bicolor Cassagnau, P, 1957:393, t.t.

Species deharvengi Najt, 1981

Species pallens Yoshii, 1990

Species patagonica Yoshii, 1990

Species spinolabris Fjellberg, A, 2010:42

Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.115G
Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.114


Genus Parisotoma Bagnall, RS, 1940:171

  Syn.: Collembola Isotoma Bagnall, RS, 1940:163 partim
  Syn.: Isotomidae Proisotoma(sic) Greenslade, 2002 lapsus


Species agrelli (Delamare Deboutteville, 1950), B

Species amurica (Potapov, MB, 1991:282) Potapov, 2001

Species appressopilosa Potapov, MB, 1991:279

Species atroculata Potapov, MB, 1991:285

Species coeca Yosii, 1966

Species confusoculata Salmon, JT, 1944:150

Species dichaeta (Yosii, 1969)

Species dividua Salmon, JT, 1944:150

Species ekmani (Fjellberg, 1977) Fjellberg, A, 2007:47

            Syn.: Isotoma notabilis f. pallida Agrell, I, 1939:2


             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:365 ISS
             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:366,af ISS
Parisotoma ekmani from the UK
2012.11.18 © Lewington, A.
Det.: Shaw, P.



Species greensladeae Rusek, J, 1984:364

Species hyonosenensis (Yosii, 1939) Potapov, 2001

            Syn.: Desoria pseudosetispinosa Chen, 1985
            Syn.: Isotoma setispinosa Lee, B-H, 1977:162


             Syn.: Isotoma hyonosensensis(sic) Yosii, R, 1977:156 ISS
             Syn.: Parisotoma hyonosonensis(sic) Gao, Y, Xiong, Y et Potapov, MB, 2009:59 ISS

Species insularis Deharveng, 1981

Parisotoma insularis from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.69.

Species longa Potapov, MB, 1991:283

Species notabilis (Schäffer, 1896) Bagnall, RS, 1940:171, t.t., B

            Syn.: Isotoma menotabilis Börner, 1903
            Syn.: Isotoma delicatula Brown, 1929
            Syn.: Isotoma (Isotoma) eunotabilis Folsom J in Mills, HB, 1934:58
            Syn.: Desoria monticola Hao & Huang, 1995


             Syn.: Parisotoma nitabilis(sic) Boitaud, L, Salmon, S Bourlette, C et Ponge J-F, 2006:452 ISS
Parisotoma notabilis from the UK
2010.03.14 © Barton, T.
Parisotoma notabilis from Norway
2011 © Fjellberg, A.
Parisotoma notabilis from the UK
2012.03.21 © Nurcombe, E.
Parisotoma notabilis from the UK
2012.04.03 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from Iran
After Kahrarian M & Arbea J, 2013 Fig.7
Parisotoma notabilis from the UK
With square eyepatch
2013.02.03 © Jones, A.
Parisotoma notabilis juv. from Canada
2013.10.22 © Hochhalter, R.
Parisotoma notabilis from Holland
With square eyepatch of 4 ocelli
2013.11.13 © van Duinen, J.
Parisotoma notabilis from the UK
With reticulate gray body pigmentation
2013.12.23 © Murray, A.
Parisotoma notabilis from Germany
2014.10.20 © Beck, K.
Parisotoma notabilis from the UK
2016.01.13 © Horton, M.
Parisotoma notabilis from Belgium
2017.04.20 © Huskens, M.-L.
Parisotoma notabilis from the UK
2018.02.17 © Lewis, M.
Parisotoma notabilis from Holland
2019.01.09 © Kamsteeg, G.
Parisotoma notabilis from Holland
With square eyepatch
2019.01.09 © Kamsteeg, G.



Species obscurocellata Potapov, Janion & Deharveng, 2011

Species picea Salmon, JT, 1949:36

Species postantennala Salmon, JT, 1949:38

Species quinquedentata Salmon, JT, 1943:382

             Syn.: Parisotoma quinquedendata(sic) Salmon, JT, 1943:382 ISS

Species raffi (Womersley, 1934)

Species reducta Rusek, J, 1984:360

Species removeopthalma Martynova, 1977

Species sexsetosa Potapov, Janion & Deharveng, 2011

Species tariva (Wray, 1953) Palacios-Vargas, JG, Mejía-Recamier, BE & Cutz-Pool, LQ, 2009:205

Species terricola Rusek, J, 1984:357

Species travei Deharveng, 1981

Species trichaetosa Martynova, 1977

Species vtorovi Martynova, 1977


Genus Misturasotoma Bernard, EC et Christiansen, KA, 2010:58

Species brucealla (Wray, 1953) Bernard, EC & Christiansen, KA, 2010:58, t.t.

            Syn.: Isotoma brucei Wray, 1953, nec Carpenter, 1907


Misturasotoma brucealla from the USA
After Bernard EC & Christiansen KA, 2010 Fig.1


Genus Vertagopus Börner, C, 1906:171 key

Vertagopus? sp. from the Arctic region
After White A in Sutherland PC, 1852 Fig.8
Vertagopus sp. from Germany
2006 © Kohl, F.
Vertagopus sp. from the USA
2006.12.13 © Cresswell, S.
Vertagopus sp. from the UK
2007.11.10 © Valentine., B
Vertagopus arboreus from the UK
Legs with clavate tenent setae
2015.12.29 © Cunningham, T.
Vertagopus arboreus from the UK
Legs with clavate tenent setae (*)
2018.03.10 © Cann, A.J.

Species sp.1 Fjellberg, A, 1996:100

Species sp.2 Fjellberg, A, 1996:100

Species sp.3 Janssens F & Duncan WL, 2019

Vertagopus sp. from the USA
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
Relative short antennae
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
Dense cover of short setae
Blue iridescent cuticular wax layer
2018.12.24 © Duncan, W.L.

Species abeloosi Poinsot, 1965

Species alpinus Haybach, 1972

Vertagopus alpinus from Austria
After Schulz H-J, 2010 Abb.4
Vertagopus alpinus from Austria
Left eyepatch : G and H smaller
Modified after Schulz H-J, 2010 Abb.3

Species alpus Christiansen & Bellinger, 1980

Species arborata Fusheng, Huifen & Yueling, 1993

Species arboreus (Linnæus, C, 1758:609) Ågren, 1903, B

            Syn.: Podura nemotrosa Linnæus, C, 1748:67 i
            Syn.: Podura nigra pedibus albis Hill, 1752 i
            Syn.: Podura annularis de Fourcroy, AF, 1785:524
            Syn.: Podura oblonga nigra de Geer, C, 1778:17
            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95
Podura fuliginosa from Ireland
After Templeton, R, 1836 Pl.XII Fig.3

            Syn.: Isotoma rubricauda Bourlet, C, 1839:402
            Syn.: Isotoma violacea Bourlet, C, 1839:404
            Syn.: Desoria cylindrica Lubbock, J, 1873:167
            Syn.: Desoria (Deuxième division) ebriosa Nicolet, 1842
            Syn.: Desoria pallida Nicolet, 1842
            Syn.: Isotoma viatica Lubbock, J, 1873:168
Isotoma viatica from the UK
After Lubbock, J., 1873 Pl.36.
 
 

            Syn.: Isotoma synonymica Mac Gillivray, 1896
            Syn.: Isotoma terminata Mac Gillivray, 1896
            Syn.: Isotoma denticulata Schäffer, 1896



"Kleine schwarze Insekten, welche unter den
Rinden alter Bäume sitzen, und mit dem Sterze
hüpfen" from Sweden
After de Geer, C., 1749 Tab.I Fig.1

             Syn.: Podura Arborea(sic) Scopoli, JA, 1763:380 ISS
Podura arborea nigra from Sweden
After de Geer, C., 1778 Tom.VII Pl.2 Fig.2


             Syn.: Vertagopus arborea(sic) Christiansen, 1958 ISS
             Syn.: Vertagopus arborea(sic) Salmon, JT, 1964:412 ISS
             Syn.: Vertagopus arborea(sic) Snider, RJ, 1967:216 ISS
             Syn.: Vertagopus arborea(sic) Massoud & Rapoport, 1968 ISS
             Syn.: Vertagopus arborea(sic) Raynal, G, 1974:189 ISS
             Syn.: Vertagopus arborea(sic) Rubio, 1974 ISS
             Syn.: Vertagopus arborea(sic) Rusek, 1977 ISS
             Syn.: Vertagopus arborea(sic) Bödvarsson, H, 1982:63 ISS
             Syn.: Vertagopus arborea(sic) André, HM, 1983:274,B ISS
             Syn.: Vertagopus arborea(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Vertagopus arborea(sic) Dallai & Malatesta in Calderini, 1995 ISS
             Syn.: Vertagopus arborea(sic) Camousseight, A, 1999:5 ISS
Vertagopus arboreus from the USA
2006 © Bernard, E.
Vertagopus arboreus from the UK
2006 © Valentine., B
Vertagopus arboreus from the UK
2006 © Barton, T.
Vertagopus arboreus from the UK
2007 © Valentine., B
Vertagopus arboreus from the UK
2007 © Barton, T.
Vertagopus arboreus from Germany
2007 © Peters, J.
Vertagopus arboreus from the UK
2007 © Barton, T.
Vertagopus arboreus from Germany
2007.02.15 © Peters, J.
Vertagopus arboreus from the USA
2007.03.10 © Murray, T.
Vertagopus arboreus from the USA, juvenile
2007.03.13 © Murray, T.
Vertagopus arboreus from the USA
2007.03.14 © Murray, T.
Vertagopus arboreus from the USA
2007.11.04 © Moorehead, C.
Vertagopus arboreus from the UK
2007.12.30 © Valentine., B
Vertagopus arboreus from the UK
2008.01.04 © Oakshott, A.
Vertagopus arboreus from the UK
2008.01.12 © Farrell, S.
Vertagopus arboreus from the UK
2008.01.12 © Tonsbeek, M.
Vertagopus arboreus from Germany
2008.01.14 © Peters, J.
Vertagopus arboreus from the UK
2008.02.20 © Robertson, A.
Vertagopus arboreus from the UK
2008.02.21 © Valentine., B
Vertagopus arboreus from the UK
2008.03.25 © Robertson, A.
Vertagopus arboreus from the UK
2008.04.01 © Valentine., B
Vertagopus arboreus from the UK
2009.01.12 © Valentine., B
Vertagopus arboreus from the USA
2009.02.15 © Murray, T.
Vertagopus arboreus from the UK
2009.12.10 © Valentine., B
Vertagopus arboreus from Holland
2011.05.01 © van Duinen, J.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus juv. from the UK
2012.11.01 © Murray, A.
Vertagopus arboreus from the UK
2012.12.13 © Valentine., B
Vertagopus arboreus from Holland
Dense cover of short setae
2012.12.16 © van Duinen, J.
Vertagopus arboreus juv. from the UK
2012.12.25 © Murray, A.
Vertagopus arboreus from the UK
Dense cover of short setae
2013.01.04 © Phillips, E.
Vertagopus arboreus juv. & adult from Germany
2014.02.05 © Beck, K.
Vertagopus arboreus subadult from the UK
2015.12.04 © Barton, T.
Vertagopus arboreus from the UK
Eyepatch with 6 ocelli with crystal + 2 ocelli without crystal
2015.12.29 © Cunningham, T.
Vertagopus arboreus from Belgium
2015.12.31 © Janssens, F.
Vertagopus arboreus from Belgium
Ready to jump
2016.11.01 © Huskens, M.-L.
Vertagopus arboreus from Belgium
Knobbed tenent setae (*)
2016.11.01 © Huskens, M.-L.
Vertagopus arboreus from Holland
Adult with early instar juvenile
2016.12.28 © van Duinen, J.
Vertagopus arboreus from Holland
Tibiotarsus with clavate tenent seta (*)
2016.12.28 © van Duinen, J.
Vertagopus arboreus from France
2018.01.04 © Garcelon, P.
Vertagopus arboreus from Belgium
Ovoid post-antennal organ (PAO) size about that of ocellus A
2019.01.04 © Huskens, M.-L.



Species arcticus Martynova, 1969

Species asiaticus Potapov, M, Gulgenova, A et Babykina, M, 2016:115

Vertagopus asiaticus from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.27

            Syn.: Vertagopus arboreus Uchida, 1969 =?
            Syn.: Vertagopus arboreus Martynova, 1979 =?


Species beta Christiansen & Bellinger, 1980

Species brevicaudus (Carpenter, 1900) Fjellberg, A, 1996:100

Species ceratus Potapov, M, Gulgenova, A et Babykina, M, 2016:113

Vertagopus ceratus from Buryatia Barguzin Range
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.25
Vertagopus ceratus from Buryatia Balan-Tamur Range
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.26

Species ciliatus Christiansen, K, 1958:75

Vertagopus ciliatus from Lebanon
After Christiansen K, 1958 Pl.9 Fig.22

Species cinereus (Nicolet, H, 1842:60) Marlier, GJ, 1942:9, t.t., B

            Syn.: Isotoma quadridenticulata Tullberg, 1871
            Syn.: Isotoma lateraria MacGillivray, 1896
             Syn.: Isotoma latraria(sic) Mills, HB, 1934:57 ISS
            Syn.: Isotoma clavigera Krausbauer, 1901 =?


             Syn.: Vertagopus cinerea(sic) Marlier, GJ, 1942:9,B ISS
             Syn.: Vertagopus cinerea(sic) Snider, RJ, 1967:216 ISS
             Syn.: Vertagopus cinerea(sic) Rusek, 1977 ISS
             Syn.: Vertagopus cinerea(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Vertagopus cinerea(sic) Dallai & Malatesta in Calderini, 1995 ISS
Vertagopus cinereus
After Potapov M, 2001 Fig.1
Vertagopus cinereus ♀
After Potapov M, 2001 Fig.2
Vertagopus cinereus from the UK
2008.09.21 © Robertson, A.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.12 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.18 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.21 © Baas, A.H.
Vertagopus cinereus from Holland
2009.04.21 © Baas, A.H.
Vertagopus cinereus from the UK
2010.11.14 © Robertson, A.
Vertagopus cinereus from France
2013.04.13 © Sandra.
Vertagopus cinereus from Holland
2013.12.11 © Baas, A.H.
Vertagopus cinereus juv. from Germany
2014.02.14 © Beck, K.
Vertagopus cinereus juv. from Holland
2014.11.07 © van Duinen, J.
Vertagopus cinereus juv. from Holland
2014.11.07 © van Duinen, J.
Vertagopus cinereus from the UK
2015.01.01 © Barton, T.
Vertagopus cinereus from the UK
2015.03.08 © Outen, A.
Vertagopus cinereus from Belgium
2017.10.28 © Huskens, M.-L.



Species haagvari Fjellberg, A, 1996:100

Species helveticus Haybach, 1980

Species laricis Martynova, 1975

Vertagopus laricis from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.23
Vertagopus laricis from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.28


Species monta (Christiansen & Bellinger, 1980) Fjellberg, A, 1996:100,cf

Species montanus Stach, 1947

Species obscurus Wahlgren, 1906

Species pallidus Martynova, 1974

Species persea Wray, 1952

Species pseudocinereus Fjellberg, 1975

            Syn.: Isotoma dilatata Mac Gillivray, 1896 =?
            Syn.: Isotoma inclinata Mac Gillivray, 1896 =?
            Syn.: Isotoma lateraria Mac Gillivray, 1896 =?
            Syn.: Isotoma unica Mac Gillivray, 1896 =?


Vertagopus pseudocinereus
Courtship behaviour
After Goloschapova & al, 2006 Fig.1
Vertagopus pseudocinereus from Holland
2014.01.07 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.22 © van Duinen, J.
Vertagopus pseudocinereus from Holland
With short dorsal macrosetae on each trunk segment
2014.02.22 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.26 © van Duinen, J.
Vertagopus pseudocinereus from Holland
2014.02.26 © van Duinen, J.
Vertagopus pseudocinereus ♂ from the USA
With bent 2nd antennal segments
2014.06.27 © Curtis, R.
Vertagopus pseudocinereus from Germany
2014.10.19 © Beck, K.
Vertagopus pseudocinereus from Germany
2014.10.20 © Beck, K.
Vertagopus pseudocinereus from Germany
2014.11.02 © Beck, K.
Vertagopus pseudocinereus from the USA
2014.12.23 © Montgomery, G.
Vertagopus pseudocinereus from the USA
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Head dorsal pattern
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Pale colour form
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from the USA
Elliptic postantennal organ (PAO)
2015.03.09 © Eiseman, C.
Vertagopus pseudocinereus from Canada
2015.11.09 © Hogue, A.
Vertagopus pseudocinereus ♂ from the USA
2017.09.07 © Wechsler, D.
Vertagopus pseudocinereus ♂ from the USA
Modified ant.2+3
2017.09.07 © Wechsler, D.
Vertagopus pseudocinereus ♀ from the USA
2018.04.18 © Murray, T.
Vertagopus pseudocinereus ♂ from the USA
Modified ant.2+3
2018.04.18 © Murray, T.
Vertagopus pseudocinereus ♂ from the USA
Modified ant.2+3
2018.04.18 © Murray, T.
Vertagopus ?pseudocinereus? from France
2019.01.26 © Garcelon, P.
Vertagopus ?pseudocinereus? from France
2019.01.26 © Garcelon, P.

Species reuteri (Schött, H, 1893:71) Fjellberg

Species sarekensis (Wahlgren, 1906) Fjellberg, A, 1996:100

Vertagopus sarekensis from Scandinavia
After Fjellberg, A, 2007 Fig.61O

Species tianschanicus Martynova, 1969

Species westerlundi (Reuter, 1898) Schubert, 1933

            Syn.: Isotoma brevicauda Carpenter, 1900
            Syn.: Isotoma sarekensis Wahlgren, 1906


Vertagopus westerlundi from Italy
2006 © Paolucci, P.
Vertagopus westerlundi from Buryatia
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.24


Genus Paracerura Deharveng, L et de Oliveira, EP, 1994:442 key

Species airesi de Mendonça, Abrantes & Fernandes, 2009

Species bella de Mendonça, MC et Queiroz, GC, 2017:388

Paracerura bella from Brazil
Dorsal aspect
After de Mendonça MC & Queiroz GC, 2017 Fig.17
Paracerura bella from Brazil
Lateral aspect
After de Mendonça MC & Queiroz GC, 2017 Fig.18
Paracerura bella from Brazil
Ventral aspect
After de Mendonça MC & Queiroz GC, 2017 Fig.19

Species cristinae Abrantes, EA et Duarte, M, 2013:1394

Paracerura cristinae from Brazil
After Abrantes EA & Duarte M, 2013 Fig.8

Species gandarela de Mendonça & da Silveira, 2013

Species itatiaiensis (Arlé, R, 1960:167) Deharveng, L & de Oliveira, EP, 1994:443

Species pallida Abrantes, EA et Duarte, M, 2013:1392

Paracerura pallida from Brazil
After Abrantes EA & Duarte M, 2013 Fig.1

Species paulista Abrantes, EA et Duarte, M, 2013:1397

Paracerura paulista from Brazil
After Abrantes EA & Duarte M, 2013 Fig.15

Species pindorama Queiroz & de Mendonça, 2010

Species serrana de Mendonça, Abrantes & Fernandes, 2009

Species virgata Deharveng, L et de Oliveira, EP, 1994:443, t.t.


Genus Marisotoma Fjellberg, A, 1997:26

Species canaliculata Fjellberg, A, 1997:26, t.t.

Species iremeli (Potapov, MB, 1991:278) Potapov, 2001

Species tenuicornis (Axelson, WM, 1903:10) Bulimar & Hutu, 1984, B


Genus Araucanocyrtus Massoud & Rapoport, 1968

Species fileri Massoud & Rapoport, 1968, t.t.


Genus Psammisotoma Greenslade, P et Deharveng, L, 1986:89 key

Species sp.1 Thibaud, J-M et Palacios-Vargas, JG, 1999:31

Species dispar (Christiansen, K et Bellinger, P, 1988:233)

Species galapagoensis Thibaud, J-M, Najt, J et Jaquemart (sic), S (+), 1994:202

Species kingae Greenslade, P et Deharveng, L, 1986:90, t.t.

Species mariagalanteae Thibaud, 1993

Species restingae Abrantes, EA et de Mendonça, MC, 2009:28

Psammisotoma restingae from Brazil
After Abrantes EA & de Mendonça MC, 2009 Fig.13


Genus Aackia Yosii, 1966

Species karakoramensis Yosii, 1966, t.t.


Genus Antarcticinella Salmon, 1965

Species monoculata Salmon, 1965


Genus Cheirotoma Bagnall, 1949

Species spatulata (Chamberlain, 1943) Bagnall, 1949


Genus Halisotoma Bagnall, 1949 key

Halisotoma sp. from the UK
2016.09.23 © Trewhella, S.
Det.: Fjellberg, A.

Species arenicola Fjellberg, A, 2009:135

Species boneti (Delamare Deboutteville, 1954)

Species kainui (Christiansen, K et Bellinger, P, 1992:164)

Species marisca (Christiansen & Bellinger, 1988:235)

Species maritima (Tullberg, 1871 tt) Bagnall, 1949

            Syn.: Isotoma grisea Schött, 1893 =?
            Syn.: Isotoma grisea Schäffer, 1896 =?
            Syn.: Isotoma grisea Carl, 1899
            Syn.: Isotoma grisea Houlbert, 1924
            Syn.: Isotoma pacifica Yosii, 1971
            Syn.: Isotoma pacifica Yosii, R, 1977:156


Halisotoma maritima from France
Ant.4 distinctly more pale
2018.08.29 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Ant.1 with apical spur (*)
2018.08.29 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Abd.4 dorsal spot pattern
2018.08.29 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Broad ovoid post-antennal organ (*)
2018.08.29 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Eyepatch with reduced G & H
2018.08.30 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
2018.08.31 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Lateral spot pattern
2018.08.31 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Longer setae at posterior segment margins
2018.08.31 © Huskens, M.-L.
Leg.: Janssens, F.
Halisotoma maritima from France
Unguiculus with inner basal bulge (*)
2018.08.31 © Huskens, M.-L.
Leg.: Janssens, F.

Species poseidonis (Bagnall, 1939)

             Syn.: Halisotoma poseidons(sic) Fjellberg, A, 2009:138 ISS

Species pritchardi (Womersley, H, 1936:478) Bagnall, 1949

            Syn.: Isotoma maritima Womersley, H, 1936:321


Species sindentata (Salmon, JT, 1943:382)

Parisotoma sindentata from New Zealand
After Salmon JT, 1943 Pl.41 Fig.38


Genus Metisotoma Maynard, EA, 1951:94 key

  Syn.: Isotomidae Cephalotoma Bagnall, 1949, nec Lesne, 1911
  Syn.: Isotominae Isotomedia Salmon, JT, 1964:381


Species bermani Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:80

            Syn.: Cephalotoma grandiceps Martynova & al, 1977 partim =?


Species grandiceps (Reuter, 1891 tt) Lippert & Butler, 1976

            Syn.: Isotoma macnamarai Mills, HB, 1934:60
            Syn.: Isotoma macnamarai Poole, RW in Poole, RW et Gentili, P, 1997:52
            Syn.: Metisotoma capitona Maynard, EA, 1951:95
Metisotoma capitona from the USA
After Maynard EA, 1951 PlateX Fig.179

            Syn.: Metisotoma capitona Hart, 1972 =?
            Syn.: Metisotoma capitona Christiansen, K et Bellinger, P, 1980:666
            Syn.: Metisotoma capitona Waltz & Hart, 1997
            Syn.: Metisotoma capitona Christiansen, K et Bellinger, P, 1998:733
            Syn.: Metisotoma spiniseta Maynard, EA, 1951:94
Metisotoma spiniseta from the USA
After Maynard EA, 1951 PlateI Fig.7

            Syn.: Metisotoma spiniseta Christiansen, K et Bellinger, P, 1980:666 =?
            Syn.: Metisotoma spiniseta Christiansen, K et Bellinger, P, 1998:733 =?



Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.24
Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.23
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the Canada
2016.03.12 © Schmidt, H.
Metisotoma grandiceps from the Canada
Wide head; short antennae
2016.03.12 © Schmidt, H.
Metisotoma grandiceps from FE Russia
Lateral aspect
After Potapov M & al, 2018 Fig.2
Metisotoma grandiceps from FE Russia
Ventral aspect
After Potapov M & al, 2018 Fig.1
Metisotoma grandiceps from FE Russia
Mouth frontal aspect
After Potapov M & al, 2018 Fig.5
Metisotoma grandiceps from FE Russia
Attack & kill of Tomocerus sp.
After Potapov M & al, 2018 Fig.44-47
             Syn.: Metisotoma grandipes(sic) Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:69 ISS


Species macnamarai (Folsom, 1918) Potapov, M, Babenko, A, Fjellberg, A & Schulz, H-J, 2018:74

Species stebaevae Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:78

            Syn.: Metisotoma grandiceps Stebaeva, 2003 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2012 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2013 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2015 =?


Species ursi (Yosii, R, 1972:90) Potapov, 2001


Genus Myopia Christiansen, K et Bellinger, P, 1980:768

Species alaskana (Christiansen, K & Bellinger, P, 1980:769) Fjellberg, A, 2010:28, t.t.

Myopia alaskana from Alaska
2007.06.12 © Takeuchi, N.
Myopia alaskana gr. from the USA
2008.12.23 © Fjellberg, A.
Myopia alaskana from the USA
After Fjellberg A, 2010 Fig.112
Myopia alaskana from the USA
After Fjellberg A, 2010 Fig.115A



Genus Najtia Arlé & Mendonça, 1986

Species vicaria (Arlé, R, 1960:166) Arlé & Mendonça, 1986


Genus Pseudosorensia de Izarra, DC, 1972:77

  Syn.: Isotoma Pseudosorencia(sic) Thibaud, J-M et Palacios-Vargas, JG, 1999:25 ISS

Species atlantica (Wise, KAJ, 1970:197)

Species fueguensis de Izarra, DC, 1972:77, t.t.

Species sp. Thibaud, J-M et Palacios-Vargas, JG, 1999:29

Species subcaeca Deharveng, 1981


Genus Azoritoma Greenslade, P et Potapov, M, 2008:89

Species macquariensis Greenslade, P et Potapov, M, 2008:90, t.t.


Genus Sericeotoma Potapov, MB, 1991:287

Species knissi Potapov, MB, 1991:288


Genus Mucronia Fjellberg, A, 2010:43

Species enigmatica Fjellberg, A, 2010:43, t.t.

Mucronia enigmatica from the USA
After Fjellberg A, 2010 Fig.93

Species fjellbergi Potapov, M et Babenko, A, 2014:1

Mucronia fjellbergi from Russia
After Potapov M & Babenko A, 2014 Fig.3B


Genus Skadisotoma Greenslade & Fjellberg, 2015

Species inpericulosa Greenslade & Fjellberg, 2015


Genus Rhyniella Hirst & Maulik, 1926, -

Species praecursor Hirst & Maulik, 1926 -

Rhyniella praecursor from the UK Rhynie Chert
After Hirst S & Maulik S, 1926 Fig.A
Holotype as fixed by Tillyard, 1928
Rhyniella praecursor from the UK Rhynie Chert
After Scourfield DJ, 1940



Genus Protoisotoma Christiansen, KA et Pike, E, 2002:171, -

Species burma Christiansen, KA et Nascimbene, P, 2006:343, -

Species micromucra Christiansen, KA et Pike, E, 2002:171, -


References