http://www.collembola.org/taxa/isotinae.htm - Last updated on 2023.04.02 by Frans Janssens
Checklist of the Collembola: Isotominae

Subfamilia Isotominae Schäffer, 1896

Genus Number
of species
Aackia1
Acanthomurus6
Agrenia13
Antarcticinella1
Araucanocyrtus1
Axelsonia8
Azoritoma1
Cheirotoma1
Chionobora1
Desoria103
Folsomotoma9
Gnathisotoma5
Halisotoma8
Heteroisotoma6
Hydroisotoma1
Isotoma67
Isotomedia1
Isotomurus76
Kaylathalia1
Marisotoma3
Metisotoma5
Misturasotoma1
Mucronia2
Myopia1
Najtia1
Paracerura10
Parisotoma28
Procerura13
Protodesoria1
Protoisotoma2
Psammisotoma7
Pseudisotoma8
Pseudosorensia4
Pteronychella4
Rhyniella1
Semicerura5
Sericeotoma1
Setocerura6
Skadisotoma1
Spinocerura2
Tibiolatra1
Tomocerura6
Vertagopus30
Isotominae453
 Genera with number of species
  Syn.: Entomobryidae Isotominini(sic) Börner, C, 1901:43 ISS
  Syn.: Collembola Isotomiens(sic) Denis, J-R, 1924:231 ISS
  Syn.: Isotomidæ(sic) Isotominæ(sic) Bagnall, RS, 1949:531 partim ISS
  Syn.: Isotomidæ(sic) Istominæ(sic) Bagnall, RS, 1949:532 ISS
Isotominae from Belgium
2006 © Vuijlsteke, M.
Isotoma viridis from the USA
2009.03.10 © Cowen, R.
Isotominae gen. nov. from Croatia
Close to Isotomurus
2019.03.dd © Bedek, J.





Genus Protodesoria Christiansen, KA et Nascimbene, P, 2006:346, -

Species granda Christiansen, KA et Nascimbene, P, 2006:346, -


Genus Tibiolatra Salmon, JT, 1941:320


Tibiolatra latronigra from New Zealand
After Salmon, J.T. 1941 Pl.49 Fig.165.
Species latronigra Salmon, JT, 1941:320, t.t.


Genus Acanthomurus Womersley, 1934

Acanthomurus sp. from Tasmania
2014.04.29 © Murray, A.

Species sp.nov. Greenslade, P in Dallai, R, 1989:116

            Syn.: Isotomurus chiltoni Womersley, 1934
            Syn.: Isotomurus chiltoni Womersley, 1937
            Syn.: Isotomurus chiltoni Norris, 1938
            Syn.: Isotomurus chiltoni Womersley, H, 1939:149
            Syn.: Isotomurus chiltoni Salmon, 1949 partim
            Syn.: Isotomurus chiltoni Stach, 1949 partim
            Syn.: Isotomurus chiltoni Salmon, JT, 1964:361 partim
            Syn.: Isotomurus chiltoni Wallace & Mackerras, 1970
            Syn.: Isotomurus chiltoni Greenslade, P in Houston, WWK, 1994:64
            Syn.: Isotoma chiltoni Greenslade, P in Houston, WWK, 1994:74


Acanthomurus sp. from Tasmania
2017.08.22 © Ellingsen, K.

Species alpinus Salmon, JT, 1941:317

Acanthomurus alpinus from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.169.

  Subspecies obscuratus Salmon, JT, 1943:381

Acanthomurus alpinus
subsp. obscuratus from New Zealand
After Salmon J.T., 1943 Pl.41 Fig.34.

Species plumbeus Womersley, 1934, t.t.

  Subspecies lineatus Womersley, 1934

Species rivalis Wise, 1964

Species setosus Salmon, JT, 1941:319

Acanthomurus setosus from New Zealand
After Salmon JT, 1941 Pl.49 Fig.162
Acanthomurus setosus? from New Zealand
2018.09.09 © Schulz, K.

  Subspecies violaceus Salmon, JT, 1941:320

Species womersleyi Salmon, JT, 1941:318

Acanthomurus womersleyi from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.176.


Genus Axelsonia Börner, 1906

  Syn.: ... Moniezina Denis, 1922


Species crassicornis (Schött, 1893) Yayuk, RS (sic), 1989:8

Species johnstoni Jordana, 1997

Species littoralis (Moniez, R, 1890:324) Denis, J-R, 1923:57

            Syn.: Isotoma nitida Handschin, E in Dahl, F, 1929:60 =?
            Syn.: Isotoma nitida Gisin, H, 1960:213
            Syn.: Axelsonia thalassophila Handschin, E in Dahl, F, 1929:60
            Syn.: Axelsonia australis Bagnall, 1939


             Syn.: Isotoma litoralis(sic) Moniez, 1894 ISS
             Syn.: Axelsonia litoralis(sic) Handschin, E in Dahl, F, 1929:60 ISS
             Syn.: Axelsonia litoralis(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
Axelsonia littoralis from Holland
2018.09.30 © van Leeuwen, J.
Axelsonia littoralis from Holland
2018.09.30 © van Leeuwen, J.
Axelsonia littoralis from the UK
With predatory mite Halotydeus hydrodromus
After Shaw P & Trewhella S, 2019 Fig.14
Axelsonia littoralis from Portugal
2019.01.06 © Almeida, J.
Axelsonia littoralis from Portugal
2019.01.06 © Almeida, J.
Axelsonia littoralis from Portugal
2019.01.09 © Almeida, J.
Axelsonia littoralis from France
2021.11.28 © Quintin, C.
Axelsonia littoralis from France
2021.12.18 © Quintin, C.
Axelsonia littoralis from France
2022.03.20 © Quintin, C.



Species nitida (Folsom, JW, 1899:264) Börner, 1906

Isotoma nitida from Japan
After Folsom JW, 1899 Pl.1 Fig.14

            Syn.: Axelsonia littoralis Yosii, R, 1955:385
            Syn.: Axelsonia littoralis Yosii, R, 1977:155
            Syn.: Isotoma (Isotoma) pteromucronata Uchida, H, 1965:88
            Syn.: Isotoma (Isotoma) pteromucronata Yosii, R, 1977:155



Species sarahae Christiansen, K et Bellinger, P, 1992:151

Species thalassophila Börner, 1906, t.t.

Species tubifera Strenzke, 1958

Species yinii Huang & Liang, 1992


Genus Metisotoma Maynard, EA, 1951:94 key

  Syn.: Isotomidae Cephalotoma Bagnall, 1949, nec Lesne, 1911
  Syn.: Isotominae Isotomedia Salmon, JT, 1964:381


Species bermani Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:80

            Syn.: Cephalotoma grandiceps Martynova & al, 1977 partim =?


Species grandiceps (Reuter, 1891 tt) Lippert & Butler, 1976

            Syn.: Isotoma macnamarai Mills, HB, 1934:60
            Syn.: Isotoma macnamarai Poole, RW in Poole, RW et Gentili, P, 1997:52
            Syn.: Metisotoma capitona Maynard, EA, 1951:95
Metisotoma capitona from the USA
After Maynard EA, 1951 PlateX Fig.179

            Syn.: Metisotoma capitona Hart, 1972 =?
            Syn.: Metisotoma capitona Christiansen, K et Bellinger, P, 1980:666
            Syn.: Metisotoma capitona Waltz & Hart, 1997
            Syn.: Metisotoma capitona Christiansen, K et Bellinger, P, 1998:733
            Syn.: Metisotoma spiniseta Maynard, EA, 1951:94
Metisotoma spiniseta from the USA
After Maynard EA, 1951 PlateI Fig.7

            Syn.: Metisotoma spiniseta Christiansen, K et Bellinger, P, 1980:666 =?
            Syn.: Metisotoma spiniseta Christiansen, K et Bellinger, P, 1998:733 =?



Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.24
Isotoma grandiceps from Siberia
After Schött H, 1893 Taf.VI Fig.23
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the USA
2007.02.22 © Bernard, E.C.
Metisotoma grandiceps from the Canada
2016.03.12 © Schmidt, H.
Metisotoma grandiceps from the Canada
Wide head; short antennae
2016.03.12 © Schmidt, H.
Metisotoma grandiceps from FE Russia
Lateral aspect
After Potapov M & al, 2018 Fig.2
Metisotoma grandiceps from FE Russia
Ventral aspect
After Potapov M & al, 2018 Fig.1
Metisotoma grandiceps from FE Russia
Mouth frontal aspect
After Potapov M & al, 2018 Fig.5
Metisotoma grandiceps from FE Russia
Attack & kill of Tomocerus sp.
After Potapov M & al, 2018 Fig.44-47
             Syn.: Metisotoma grandipes(sic) Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:69 ISS


Species macnamarai (Folsom, 1918) Potapov, M, Babenko, A, Fjellberg, A & Schulz, H-J, 2018:74

Species stebaevae Potapov, M, Babenko, A, Fjellberg, A et Schulz, H-J, 2018:78

            Syn.: Metisotoma grandiceps Stebaeva, 2003 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2012 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2013 =?
            Syn.: Metisotoma grandiceps Stebaeva, 2015 =?


Species ursi (Yosii, R, 1972:90) Potapov, 2001


Genus Isotomurus Börner, 1903 key  key  key

  Syn.: Podurides(sic) Podura Bourlet, C, 1845:117
  Syn.: Podurides(sic) Podura Bourlet, C, 1845:168 ISS
Podura sp. from France
Ocelli of left eye
Ocelli BDE from straight line
Modified after Bourlet C, 1845 Fig.18

  Syn.: Collembola Istoma(sic) Childs, GH, 1915:21 ISS
  Syn.: ... Isotama(sic) Stach, J, 1924:114 ISS

  Syn.: ... Proisotomurus Womersley, 1934
  Syn.: Isotomidae Chisotoma Gisin, H, 1960:212
  Syn.: Isotomidae Eosotoma Gisin, H, 1960:212
  Syn.: Isotoma riparia Alijani-Ardeshir, M, Shayanmehr, M et Amiri-Besheli, B, 2017:45
  Syn.: Isotoma riparia Alijani-Ardeshir, M, Shayanmehr, M et Amiri-Besheli, B, 2017:53,cf
  Syn.: Isotoma riparia Alijani-Ardeshir, M, Shayanmehr, M et Amiri-Besheli, B, 2017:55,cf



  Syn.: ... Isotomorus(sic) Stach, J, 1924:115 ISS
  Syn.: Isotomidae Isotomourus(sic) Baijal, HN, 1958:355 ISS
  Syn.: Isotominae Isotomuruus(sic) Salmon, JT, 1964:370 ISS
  Syn.: Isotomidae Isotomorus(sic) Poinsot, N, 1972:687 ISS
  Syn.: Isotomidae Isomurys(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS
Remark by ICZN. 2007:420: Placed on the Official List of Names in Zoology. Opinion 291.

Isotomurus sp. from Norway
Triangular head shape
After Fjellberg A, 2010 p.2
Isotomurus sp. from Norway
3+3 macrosetae on abd.4 in triangle
After Fjellberg A, 2010 p.2
Isotomurus palustris from the UK
Triangular head shape
2010.02.08 © Robertson, A.
Isotomurus graminis from Sweden
Difference between bothriotrichum (bot) and macroseta (mac)
2011.09.24 © Hall, K.
Isotomurus sp. from the UK
2011.12.20 © Kent, J.J.
Isotomurus sp. from the UK
Clypeus dark
2012.12.18 © Phillips, E.
Isotomurus maculatus from Prince Edward & Marion Island
Abd.3 distinctly larger then abd.4
2015 © Janion-Scheepers, C.

  Syn.: Isotomidae isotomurus(sic) Glime, JM, 2015:26 ISS
Isotomurus sp. juv. from Holland
2015.01.20 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.

  Syn.: Isotomidae Isotomorus(sic) Qazi, F et Shayanmehr, M, 2016:131 ISS
  Syn.: Isotomidae Isotomorus(sic) Qazi, F et Shayanmehr, M, 2016:133 ISS
Isotomurus palustris from Belgium
Macrosetae only present on posterior body segments
2016.01.04 © Huskens, M.-L.
Isotomurus palustris from Belgium
Macrosetae on abd.4 in triangular constellation
2016.01.04 © Huskens, M.-L.
Isotomurus graminis from the UK
Mucro with 4 teeth (outer aspect)
2016.01.26 © Shepherd, M.J.
Isotomurus maculatus? from Belgium
Ovoid post-antennal organ (PAO)
2016.02.25 © Huskens, M.-L.
Isotomurus sp. from Argentina
2016.06.05 © Chiavone, J.
Isotomurus sp. from Argentina
Lateral macrosetae on abd.4 form triangle
2016.06.05 © Chiavone, J.
Isotomurus sp. from Holland
Embryo in egg
2017.04.28 © van Duinen, J.
Isotomurus sp. from the UK
Ovoid post-antennal organ (PAO)
2017.12.23 © Cann, A.J.
Isotomurus sp. from Belgium
Ocelli B-D-E in a straight line
2018.02.13 © Huskens, M.-L.
Isotomurus sp. from France
Abd.4 with 3 dorsolateral macrosetae in triangular constellation
2018.11.25 © Tiky, S.
Isotomurus pseudopalustris f. aquatilis from the UK
Mucro with 3 teeth + lateral ridge (inner aspect)
2019.01.21 © Cann, A.J.
Isotomurus nr pseudopalustris from Japan
With Sminthurides sp. juv.
2019.01.20 © Hara, Y.
Isotomurus sp. from Belgium
Abd.3 distinctly larger then abd.4
As in Tomocerinae
2020.09.01 © Huskens, M.-L.
Isotomurus sp. from Belgium
Ocelli A-D in a square
As in Tomocerus
2020.09.01 © Huskens, M.-L.
Isotomurus palustris juv. from Belgium
Ocelli G and H far apart
2020.11.29 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
2020.12.29 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
Dark antennal basis and dark clypeus
Pale frons
2020.12.29 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
2020.12.29 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
2021.01.03 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
2021.01.03 © Huskens, M.-L.
Isotomurus sp. juv. from Belgium
Dark clypeus
2021.01.03 © Huskens, M.-L.
Isotomurus sp. ♂ from Belgium
2021.01.06 © Huskens, M.-L.
Isotomurus sp. ♂ from Belgium
Furca long with crenulated dentes
2021.01.06 © Huskens, M.-L.
Isotomurus sp. ♂ from Belgium
Male genital plate
2021.01.06 © Huskens, M.-L.
Isotomurus sp. ♂ from Belgium
Mouthcone
Dark clypeus
2021.01.06 © Huskens, M.-L.
Isotomurus sp. from Belgium
With pale intersegmental bands
2021.02.17 © Huskens, M.-L.
Isotomurus sp. from Belgium
Eye circumvented with pale ring
2021.02.17 © Huskens, M.-L.
Isotomurus sp. from Belgium
2021.02.18 © Huskens, M.-L.
Isotomurus sp. from Belgium
2021.02.18 © Huskens, M.-L.
Isotomurus sp. from Belgium
2021.02.18 © Huskens, M.-L.
Isotomurus sp. from India
3rd foot
SEM x3300
2021.11.12 © Angurana, R.
Isotomurus sp. from India
Mucro with 4 teeth (inner aspect)
SEM x1400
2021.11.12 © Angurana, R.
Isotomurus sp. from India
Mucro with 4 teeth (outer aspect)
SEM x3000
2021.11.12 © Angurana, R.
Isotomurus sp. from India
Epicutilar hexagonal ultrastructure
SEM x7500
2021.11.12 © Angurana, R.
Isotomurus sp. from Taiwan
2021.11.13 © Cheng, H.-J.
Isotomurus sp. from Taiwan
2021.12.09 © Cheng, H.-J.
Isotomurus sp. from Taiwan
Egg deposition
2022.01.15 © Cheng, H.-J.
Isotomurus sp. from Taiwan
Moulting
2022.01.16 © Cheng, H.-J.
Isotomurus sp. from Taiwan
Palp of outer maxillary lobe flipping back and forth
2022.01.15 © Cheng, H.-J.



Species sp.I Christiansen, K et Bellinger, P, 1998:743

            Syn.: Isotomurus (Isotomurus) retardatus Christiansen, K et Bellinger, P, 1980:685 partim
            Syn.: Isotomurus retardatus Christiansen, K et Bellinger, P, 1998:754


Species sp. Zoughailech, A, 2017:91

Isotomurus sp. from Algeria
After Zoughailech A, 2017 Fig.47 left
Isotomurus sp. from Algeria
After Zoughailech A, 2017 Fig.47 right

Species aetnensis Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:116

Isotomurus aetnensis
2001 © Carapelli, A. et al.
Isotomurus nr aetnensis from the Azores
2010.09.22 © E.D.E.N.

Species afghanicus Yosii, 1966

Isotomurus afghanicus from Afghanistan
After Potapov M, 2001 Fig.374

Species alticolus (Carl, J, 1899:313) Handschin, 1919

             Syn.: Isotomurus alticola(sic) Kloet, GS et Hincks, WD, 1945:7 ISS
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate L Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.1.
Isotomurus alticolus from Poland
After Stach, J., 1947 Plate LI Fig.3.

             Syn.: Isotomurus alticola(sic) Rusek, 1977 ISS
             Syn.: Isotomurus alticola(sic) Dallai & Malatesta in Calderini, 1995 ISS
Isotomurus alticolus from Italy
2001 © Carapelli, A. et al.



Species annectens Yosii, 1963

Species antennalis (Bagnall, 1940) Gisin, 1944, B

             Syn.: Isotoma antenalis(sic) Camousseight, A, 1999:6 ISS
Isotomurus antennalis from Denmark
After Fjellberg,A, 2007 Plate.1 Fig.1
Isotomurus antennalis from Denmark
Head pattern
After Fjellberg,A, 2007 Plate.1 Fig.1
Isotomurus antennalis from Denmark
After Fjellberg,A, 2007 Plate.2 Fig.1
Isotomurus antennalis from Denmark
Head pattern
After Fjellberg,A, 2007 Plate.2 Fig.1
Isotomurus antennalis from Belgium
Macrosetae on abd.4-6 indistinct
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
8 equally sized ocelli
Oval post-antennal organ (PAO) (*)
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Unguiculus with basal broad lamella
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Abd.5+6 separated
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Macrosetae on abd.4-6 short
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Maxillary outer lobe with distinctly long palp
2020.11.26 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Back of head distinctly more pale
2020.11.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Ocelli F+G close together
2020.11.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Cephalic pattern arrow shaped
2020.11.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
2020.12.29 © Huskens, M.-L.
Isotomurus antennalis from Belgium
2020.12.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Cephalic pattern
2020.12.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Abd.5+6 separated
2020.12.30 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Dark dorsal antennal basis
Interconnected by dark frontal spot
2021.01.03 © Huskens, M.-L.
Isotomurus antennalis juv. from Belgium
2021.01.03 © Huskens, M.-L.
Isotomurus antennalis from Belgium
2021.01.03 © Huskens, M.-L.
Isotomurus antennalis from Belgium
2021.01.06 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Furca with long crenulated dentes
2021.01.06 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Cephalic pattern
Dorsal aspect
2021.01.17 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Cephalic pattern
Lateral aspect
2021.01.17 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Cephalic pattern
Frontal aspect
2021.01.18 © Huskens, M.-L.
Isotomurus antennalis from Belgium
Habitus in ethanol appears banded
2021.01.18 © Janssens, F. Leg.: Huskens, M.-L.
Isotomurus antennalis from Belgium
Mucro with 4 teeth
Apical tooth (1) very small
2021.01.18 © Janssens, F. Leg.: Huskens, M.-L.
Isotomurus antennalis from Belgium
Unguiculus with basal broad lamella (*)
Unguis with indistinct inner tooth
2021.01.18 © Janssens, F. Leg.: Huskens, M.-L.



Species atreus Christiansen & Bellinger, 1980

Species balteatus (Reuter, OM, 1876:86) Handschin, E in Dahl, F, 1929:73

Isotoma palustris var. balteata from Sweden
After Schött H, 1893 Taf.V Fig.10
Isotomurus palustris f. balteata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.6
Isotomurus balteatus from Korea
After Yosii R & Lee C-E, 1963 Fig.2A
Isotomurus palustris f. balteata(sic) from Poland
After Potapov M, 2001 Fig.371
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus
2007 © Fjellberg, A.
Isotomurus balteatus from India
After Mandal GP & Hazra AK, 2009 Fig.17a
             Syn.: Isotomurus palustris var. balteata(sic) Pfliegler, WP et Janssens, F, 2016:202 ISS
             Syn.: Isotomurus balteaus(sic) Yadav, RS, 2017:11058 ISS
Isotomurus balteaus(sic) from India
After Yadav RS, 2017 Img.2



Species beskidensis Rusek, 1963

Species bimaculatus Ågren, 1903

Species bimus Christiansen, K et Bellinger, P, 1980:679

Isotomurus (Isotomurus) bimus from the USA
After Christiansen K & Bellinger P, 1980 Fig.541H
            Syn.: Isotomurus palustris var. balteatus Folsom, 1937
             Syn.: Isotomurus palustris ssp. balteata(sic) Brimley, 1938 ISS
             Syn.: Isotomurus palustris ssp. balteata(sic) Mills, 1938 ISS
            Syn.: Isotomurus palustris f. balteatus Maynard, EA, 1951:123
Isotomurus palustris f. balteatus from the USA
After Maynard EA PlateXXII Fig.222

            Syn.: Isotomurus palustris ssp. balteatus Snider, RJ, 1967:213
Isotomurus palustris ssp. balteatus from the USA
After Snider RJ, 1967 Backcover

            Syn.: Isotomurus tricolor Hart, 1974 =?



Isotomurus bimus from the USA
2007.03.31 © Babin, P.
Isotomurus bimus from the USA
2009.07.05 © Zungri, D.
Isotomurus bimus juv. from the USA
2009.07.06 © Zungri, D.
Isotomurus bimus from the USA
2010.08.03 © Zungri, D.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.07.20 © Dankowicz, E.
Isotomurus bimus from the USA
2014.11.01 © Emile.
Isotomurus bimus from the USA
2015.07.22 © Dankowicz, E.
Isotomurus bimus from the USA
2016.12.24 © Montgomery, G.
Isotomurus bimus from the USA
2017.08.13 © Lee, C.
Isotomurus nr bimus from Holland
2017.09.21 © Wieringa, P.
Isotomurus nr bimus from Holland
With incomplete pigmentation pattern
2017.09.21 © van Duinen, J.
Isotomurus nr bimus from Holland
With incomplete pigmentation pattern
2017.09.21 © van Duinen, J.
Isotomurus nr bimus from Holland
2017.10.01 © van Duinen, J.
Isotomurus nr bimus from Holland
2017.10.01 © van Duinen, J.
Isotomurus bimus from the USA
2017.12.26 © Abbott, J.

Species cassagnaui Deharveng & Lek, 1993

Isotomurus cassagnaui
2001 © Carapelli, A. et al.
Isotomurus ?cassagnaui? from France
2008.02.24 © Jacquemin, G.
Isotomurus ?cassagnaui? from France
2008.11.08 © Jacquemin, G.
Isotomurus ?cassagnaui? from France
2020.01.18 © Joachim, J.
Isotomurus ?cassagnaui? from France
2020.01.18 © Joachim, J.

Species chaos Potapov & Babenko, 2011

Species chiltoni (Carpenter, GH, 1925:9) Womersley, H, 1936:321

Species cibus Christiansen & Bellinger, 1980

Species denticulatus Kos, 1937

Species dhanbadensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:100

Isotomurus dhanbadensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.2 Fig.1

Species festus Potapov, M, Porco, D et Deharveng, L, 2018:385

Isotomurus festus from FE Russia
After Potapov M & al, 2018 Fig.1
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.18
Isotomurus festus from FE Russia
Dark form
After Potapov M & al, 2018 Fig.19
Isotomurus festus from FE Russia
Pale banded form
After Potapov M & al, 2018 Fig.17
Isotomurus festus from FE Russia
Dark banded form
After Potapov M & al, 2018 Fig.20
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.26low
Isotomurus festus from FE Russia
Pale form
After Potapov M & al, 2018 Fig.25low
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.25mid
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.26up
Isotomurus festus from FE Russia
Banded form
After Potapov M & al, 2018 Fig.25up

Species fucicolus (Schött, H, 1893:66) Axelson, 1905

Isotoma palustris var. fucicola from Sweden
After Schött H, 1893 Taf.V Fig.6
            Syn.: Isotoma fucicola Reuter, 1891 il
            Syn.: Isotoma janthina Vogler, 1896
            Syn.: Isotomurus italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:120
            Syn.: Isotomurus italicus Potapov, M, 2001:331



             Syn.: Isotomurus palustris ssp. fucicola(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. fucicola(sic) Linnaniemi, WM, 1912:190 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris f. fascicola(sic) Womersley, H, 1939:149 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Stach, J, 1947:460 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Yosii, R, 1956:65 ISS
             Syn.: Isotomurus palustris var. fucicola(sic) Gisin, H, 1960:214 ISS
Isotomurus fucicolus
2001 © Carapelli, A. et al.


             Syn.: Isotomurus fucicola(sic) Fanciulli, PP, Summa, D, Dallai, R et Frati, F, 2001:251 ISS
Isotomurus fucicola(sic) from Scandinavia
2007 © Fjellberg, A.
Isotomurus ?fucicolus? from the UK
2009.03.29 © Robertson, A.
Isotomurus ?fucicolus? from the UK
2009.09.30 © Robertson, A.
Isotomurus ?fucicolus? from the UK
2009.11.15 © Robertson, A.
Isotomurus ?fucicolus? from the UK
2009.11.22 © Robertson, A.
Isotomurus ?fucicolus? from the UK
Abd.3 with 4 bothriotricha
2009.11.26 © Robertson, A.
Isotomurus ?fucicolus? from Sweden
2011.09.24 © Hall, K.

             Syn.: Isotomurus fucicola(sic) Santeshwari, X, Raghuraman, M et Singh, J, 2015:572 ISS
             Syn.: Isotomurus fucicolous(sic) Shaw, P, 2017:147 ISS
Isotomurus fucicolus from Belgium
2017.06.06 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.06.06 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.06.12 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.06.29 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2017.07.07 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
Bluish antennae
2017.07.07 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
Facial pattern
2017.08.04 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
Cephalic pattern
2017.08.05 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
Abdominal dorsal pattern
2017.08.05 © Huskens, M.-L.
Isotomurus fucicolus juv. from Belgium
2017.08.10 © Huskens, M.-L.
Isotomurus fucicolus from France
2017.09.29 © Tiky, S.
Isotomurus fucicolus from France
2018.05.21 © Tiky, S.
Isotomurus fucicolus from France
2019.10.26 © Vol, F.
Isotomurus fucicolus from France
Abd.4 with 3 dorsolateral macrosetae in triangular constellation
2019.10.26 © Vol, F.
Isotomurus fucicolus from France
2019.10.26 © Vol, F.
Isotomurus fucicolus from Belgium
2020.01.31 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
2020.09.01 © Huskens, M.-L.
Isotomurus fucicolus from Belgium
With 2 pale interocular spots
2020.09.01 © Huskens, M.-L.



Species fucicolus2 da Gama, 1964

            Syn.: Isotomurus (gallicus-group) fucicola(sic) da Gama, 1964 IOS
             Syn.: Isotomorus(sic) fucicola(sic) Poinsot, N, 1972:688 ISS
             Syn.: Isotomurus fucicola(sic) Poinsot, N, 1972:689 ISS
Isotomorus(sic) fucicola(sic) from France
After Poinsot N, 1972 Fig.6D





Species fuscus (Salmon, JT, 1944:145) Greenslade, P, 2003:1, s.i.

Species gallicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:120

Isotomurus gallicus from France
2001 © Carapelli, A. et al.
             Syn.: Isotomurus palustris var. balteata(sic) Handschin, E, 1942:445 ISS
            Syn.: Isotomurus balteatus Poinsot, N, 1972:689
            Syn.: Isotomurus balteatus Ellis, WN, 1974:121
            Syn.: Isotomurus balteatus Poinsot-Balaguer & Ferard, 1983
            Syn.: Isotomurus balteatus Deharveng & Lek, 1993
            Syn.: Isotomurus balteatus Sousa & al., 1997



Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from France
2008.03.15 © Lebeaux, P.
Isotomurus gallicus from Norway
2010 © Fjellberg, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.15 © Murray, A.
Isotomurus gallicus from the UK
2012.07.18 © Murray, A.
Isotomurus gallicus from the UK
2013.06.12 © Murray, A.
Isotomurus gallicus from Holland
Diffuse pigmentation
2016.02.16 © Wieringa, P.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
2016.02.19 © van Duinen, J.
Isotomurus gallicus from Holland
Diffuse pigmentation
2016.02.19 © Wieringa, P.
Isotomurus gallicus from Holland
2017.10.12 © van Duinen, J.
Isotomurus gallicus from France
2019.05.09 © Tiky, S.
Isotomurus gallicus from France
2020.06.10 © Reinach, Br.
Isotomurus gallicus juv. from France
2020.06.10 © Reinach, Br.
Isotomurus gallicus from France
2021.03.10 © Tiky, S.
Isotomurus gallicus from France
2021.07.15 © Picard, J.
Isotomurus gallicus from France
2021.07.26 © Picard, J.

Species ghibellinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94

Isotomurus ghibellinus
2001 © Carapelli, A. et al.
Isotomurus ghibellinus from France
2019.12.21 © Joachim, J.
Isotomurus ghibellinus from France
2019.12.21 © Joachim, J.
Isotomurus ghibellinus from France
2019.12.21 © Joachim, J.
Isotomurus ghibellinus from France
2020.03.14 © Joachim, J.
Isotomurus ghibellinus from France
2020.03.14 © Reinach, Br.
Isotomurus ghibellinus from France
2021.02.13 © Joachim, J.
Isotomurus ghibellinus from France
2021.02.13 © Joachim, J.

Species graminis Fjellberg, A, 2007:96

            Syn.: Isotomurus palustris var. pallida Schäffer, 1896 =?
            Syn.: Isotomurus palustris var. prasinus Deharveng & Lek, 1993
            Syn.: Isotomurus prasinus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127
Isotomurus prasinus
2001 © Carapelli, A. et al.
Isotomurus prasinus from Sweden
2005 © Hall, K.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.
Isotomurus prasinus from the UK
2006 © Cornwall, N.J.




Isotomurus graminis
2007 © Fjellberg, A.
Isotomurus graminis from the UK
2007.03.18 © Cornwall, N.J.
Isotomurus graminis from the UK
2007.04.01 © Cornwall, N.J.
Isotomurus graminis from the UK
2009.03.29 © Michaelson, J.
Isotomurus graminis from the UK
2012.04.21 © Murray, A.
Isotomurus graminis from the UK
2012.11.21 © Murray, A.
Isotomurus graminis from the UK
2013.08.10 © Fenwick, D.
Isotomurus graminis from Holland
2014.01.14 © van Duinen, J.
Isotomurus graminis from Belgium
2014.01.18 © Wallays, H.
Isotomurus graminis ♂ from Holland
2014.02.24 © van Duinen, J.
Isotomurus graminis ♂ from Holland
Spiny setae of male
2014.02.24 © van Duinen, J.
Isotomurus graminis from Germany
2014.09.20 © Beck, K.
Isotomurus graminis from Germany
2014.11.09 © Landgraf, A.
Isotomurus graminis from the UK
2015.01.15 © Thomas, Br.
Isotomurus graminis from Holland
2015.02.06 © van Duinen, J.
Isotomurus graminis from Germany
Ovoid post-antennal organ (PAO)
2015.02.14 © Kluck, A.
Isotomurus graminis from Germany
Macrosetae on abd.4 in triangular constellation
2015.02.14 © Kluck, A.
Isotomurus graminis from Belgium
2016.01.10 © Huskens, M.-L.
Isotomurus graminis from the UK
2016.01.26 © Shepherd, M.J.
Isotomurus graminis from Belgium
2016.11.23 © Huskens, M.-L.
Isotomurus graminis from Belgium
2017.01.31 © Huskens, M.-L.
Isotomurus graminis from Belgium
2019.08.30 © Huskens, M.-L.
Isotomurus graminis from Belgium
Maxillary palp with long bristle
2019.08.30 © Huskens, M.-L.
Isotomurus graminis from Belgium
2020.01.14 © Huskens, M.-L.
Isotomurus graminis from Belgium
2020.01.19 © Huskens, M.-L.
Isotomurus graminis from Belgium
Intersegmental membrane (ism)
2020.01.19 © Huskens, M.-L.
Isotomurus graminis from France
2020.09.30 © Reinach, Br.
Isotomurus graminis from France
Ocelli BDE in a straight line
2020.09.30 © Reinach, Br.
Isotomurus graminis from France
Macrosetae on abd.4 in triangular constellation
2020.10.10 © Joachim, J.

Species griseus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:282

Species hadriaticus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:121

Isotomurus hadriaticus
2001 © Carapelli, A. et al.

Species hyrcanicus Lafooraki, EY, Hajizadeh, J et Shayanmehr, M, 2023:?

Species indicus Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:102

Isotomurus indicus from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.3 Fig.1

Species infuscatus Yosii, 1963

             Syn.: Isotomurus palustris f. fucicola(sic) Yosii, R, 1955:379 ISS
             Syn.: Isotomurus palustris f. fucicola(sic) Yosii, R, 1955:387 ISS
            Syn.: Isotomurus palustris ssp. fucicolus Yosii, R, 1977:156


Species innominatus Denis, J-R, 1933:243

            Syn.: Isotomurus innominata(sic) Denis, J-R, 1933:243 IOS

Species italicus Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94

Isotomurus italicus
2007 © Fjellberg, A.


Species jharkhandensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:103

Isotomurus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.4 Fig.1
Isotomurus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.4 Fig.2


Species katule Lafooraki, EY, Hajizadeh, J et Shayanmehr, M, 2023:?

Isotomurus katule from Iran
After Lafooraki EY & al, 2023 cover

Species lapidosus (Salmon, JT, 1949:25) Greenslade, P, 2003:1, s.i.

Species lateclavus Kos, 1937

Species lineatus (Salmon, JT, 1941:322) Christiansen, K & Bellinger, P, 2000:114

  Subspecies violaceus Salmon, JT, 1941:323

Species louisiana (Scott, 1962) Waltz, RD, 1998:195

Species maculatus (Schäffer, 1896) Ågren, 1903, B

            Syn.: Isotoma bifasciata Bourlet, C, 1839:401
            Syn.: Desoria (Première division) fulvomaculata Nicolet, H, 1842:59
             Syn.: Isotoma fulvo-maculata(sic) Nicolet, 1847 ISS
            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
Desoria virescens from Switzerland
After Nicolet, H, 1842 Pl.5 Fig.12.

            Syn.: Desoria tigrina Bourlet, C, 1845:67
            Syn.: Desoria tigrina Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura viridis Bourlet, 1843 partim Schött, 1891
            Syn.: Isotoma palustris f. principalis Tullberg, 1872, partim Schött, 1891
            Syn.: Desoria riparia Lubbock, J, 1873:170
            Syn.: Isotomurus palustris var. texensis Stach, J, 1947:459
            Syn.: Isotomurus palustris ssp. texensis Salmon, JT, 1964:368
            Syn.: Isotomurus palustris f. hortorum Cassagnau, 1987
            Syn.: Isotomurus palustris Greenslade, 2002



             Syn.: Isotomurus palustris var. maculata(sic) Stach, J, 1924:113 ISS
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.9
Isotomurus palustris var. maculata(sic) from Malta
After Stach J, 1924 Taf.II Fig.10
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.11
Isotomurus palustris var. maculata(sic) from Tunis
After Stach J, 1924 Taf.II Fig.12

             Syn.: Isotomurus palustris f. maculata(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Kseneman, M, 1932:20 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Stach, J, 1947:459 partim ISS
Isotomurus palustris f. maculata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.1
Isotomurus palustris f. maculata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.4
Isotomurus palustris f. maculata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.5

             Syn.: Isotomurus palustris var. maculata(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Stach, J, 1967:401 ISS
Isotomurus palustris ssp. maculatus from France
After Poinsot N, 1972 Fig.6C

             Syn.: Isotomurus palustris f. maculata(sic) Verlinden, A, 1977:114,B ISS
             Syn.: Isotomurus maculata(sic) Deharveng, 1981 gr ISS
             Syn.: Isotomurus maculata(sic) Gabriel & al, 1990 gr ISS
Isotomurus maculatus
2001 © Carapelli, A. et al.
Isotomurus maculatus from Italy
After Potapov M, 2001 Fig.368-369
Isotomurus maculatus from the UK
2005 © Hopkin, S.P.
Isotomurus maculatus from the UK
2005 © Brocklehurst, K.
Isotomurus maculatus from Scandinavia
After Fjellberg, A, 2006 Fig.4
Isotomurus maculatus from the UK
2006.04.08 © Read, D.
Isotomurus maculatus from the UK
2007 © Cornwall, N.J.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus palustris f. maculatus from the USA
2007 © Moorehead, C.
Isotomurus maculatus
2007 © Fjellberg, A.
Isotomurus maculatus from Italy
2007 © Bartolini, L.
Isotomurus maculatus from the UK
2007.12.01 © Warner, P.
Isotomurus palustris f. maculatus from the USA
2008.05.09 © Zungri, D.
Isotomurus maculatus from Hungary
2008.11.08 © Keresztes, G.
Isotomurus maculatus from Holland
2008.11.14 © Baas, A.H.
Isotomurus maculatus from Holland
2008.11.19 © Baas, A.H.
Isotomurus maculatus from Holland
2008.11.19 © Baas, A.H.
Isotomurus maculatus from Holland
2008.11.19 © Baas, A.H.
Isotomurus maculatus from Holland
2008.11.19 © Baas, A.H.
Isotomurus maculatus from France
2009.02.25 © Lefebvre, V.
Isotomurus maculatus from Belgium
2009.03.08 © De Groot, S.
Isotomurus maculatus from the UK
2009.04.06 © Robertson, A.
Isotomurus maculatus from Tasmania
2009.08.16 © Daley, A.

             Syn.: Isotomurus maculata(sic) Greenslade, 2010 gr ISS
Isotomurus maculatus from Belgium
2010.01.31 © Wallays, H.

             Syn.: Isotomurus maculata(sic) Greenslade, P et Convey, P, 2012:411,gr ISS
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2012.11.07 © Jones, A.
Isotomurus maculatus from the UK
2013.12.01 © van Duinen, J.
Isotomurus maculatus from the UK
Clypeus and labrum dark
2015.02.05 © Horton, M.
Isotomurus maculatus from the UK
Pre-jump posture
2015.02.05 © Horton, M.
Isotomurus maculatus from Prince Edward & Marion Island
2015 © Janion-Scheepers, C.
Isotomurus maculatus from Prince Edward & Marion Island
2015 © Janion-Scheepers, C.

             Syn.: Isotomurus palustris var. maculata(sic) Pfliegler, WP et Janssens, F, 2016:202 ISS
             Syn.: Isotomurus palustris f. maculata(sic) Pfliegler, WP et Janssens, F, 2016:204 ISS
Isotomurus maculatus from Belgium
2016.01.11 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Unguis with basal lateral teeth (*)
2016.03.08 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.04.24 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2016.09.04 © Huskens, M.-L.
Isotomurus palustris f. maculatus from the USA
2017.01.22 © Dobbs, Br.
Isotomurus maculatus from Belgium
2017.02.07 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Clypeus dark
2017.02.08 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Ventral aspect; with open valves of collophore
2017.12.08 © Huskens, M.-L.
Isotomurus maculatus from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus maculatus from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomururus palustris f. maculatus from the USA
2017.12.27 © Dobbs, Br.
Isotomururus palustris f. maculatus from the USA
2017.12.27 © Dobbs, Br.
Isotomurus maculatus ♂ from Belgium
Ventral aspect; roundish genital plate
2018.01.11 © Huskens, M.-L.
Isotomurus maculatus ♂ from Belgium
Latero-ventral aspect; circumgenital setae
2018.01.11 © Huskens, M.-L.
Isotomururus palustris f. maculatus from the USA
2018.12.16 © Duncan, W.L.
Isotomururus maculatus juv. from France
Moulting
Bulging thorax
2020.10.19 © Joachim, J.
Isotomururus maculatus juv. from France
Ecdysial cleavage line
2020.10.19 © Joachim, J.
Isotomurus maculatus juv. from the UK
2021.01.06 © Uncle, F.
Isotomurus maculatus juv. from the UK
2021.01.06 © Uncle, F.
Isotomurus maculatus from Belgium
2021.09.10 © Huskens, M.-L.
Isotomurus maculatus from Belgium
Lateral abdominal pattern
2021.09.10 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2021.10.22 © Huskens, M.-L.
Isotomurus maculatus from Belgium
2021.10.22 © Huskens, M.-L.



Species mankyungensis Kim, JT et Lee, B-H, 2000:181

Species martiniquensis Stach, J, 1947:449

Isotomurus tricuspis ssp. martiniquensis from Martinique
After Stach, J., 1947 Plate XLIX Fig.8.

Species mauretanicus (Handschin, 1926, s.d.) Handschin, 1928

Species myohyangsanus Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285

Species nebulosus Lek, S et Carapelli, A, 1998:125

Isotomurus nebulosus from France
After Lek & Carapelli, 1998 Fig.1
Isotomurus nebulosus
2001 © Carapelli, A. et al.
Isotomurus ?nebulosus? from France
Dark form
2018.04.05 © Garcelon, P.
Isotomurus ?nebulosus? from France
Pale form
2018.04.06 © Garcelon, P.

Species nicolae Stomp, 1983

Species novaezealandiae (Salmon, JT, 1941:321) Christiansen, K & Bellinger, P, 2000:114

            Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1941:321 IOS
Proisotomurus novae-zealandiae(sic) from New Zealand
After Salmon J.T., 1941 Pl.50 Fig.179




             Syn.: Proisotomurus novae-zealandiae(sic) Salmon, JT, 1964:359 ISS



Species obscurus Carpenter, GH, 1916:34

Species opala Christiansen, K et Bellinger, P, 1992:155

Isotomurus opala from Hawaii
Type 1
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.C
Isotomurus opala from Hawaii
Type 2
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.D
Isotomurus opala from Hawaii
Intermediate between type 3 & 4
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.A
Isotomurus opala from Hawaii
Type 4
After Christiansen K & Bellinger P, 1992 Plate 64 Fig.B

            Syn.: Isotomurus palustris var. balteatus Folsom, 1932



Isotomurus opala? from Hawaii
2017.10.03 © Nelson, S.
Isotomurus opala? from Hawaii
2017.10.06 © Nelson, S.
Isotomurus opala? from Hawaii
2017.10.10 © Nelson, S.

Species palliceps (Uzel, 1891) Stach, 1929

Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.1.
Isotomurus palliceps from Poland
After Stach, J., 1947 Plate LII Fig.2.

Species palustris (Müller OF, 1776) Börner, 1903, t.t., B

            Syn.: Podura aquatica cinerea de Geer, C, 1740:63
"Graue Insekten die unter den vorigen schwarzen [Podura aquatica] zusammen auf den Wasser sitzen und etwas grösser sind" from Sweden
After de Geer, C., 1749 Tab.IV Fig.1.

            Syn.: Podura aquatica secunda Lin
            Syn.: Podura aquatica secunda Bourlet, C, 1845:112
             Syn.: Podura Aquatica(sic) secunda Bourlet, C, 1845:153 ISS
            Syn.: Podura aquatilis Müller OF, 1776
            Syn.: Podura aquatica grisea de Geer, C, 1778:28
Podura aquatica grisea from Sweden
After de Geer, C., 1778 Tom.VII Pl.2 Fig.19.

             Syn.: Podura Grisea(sic) Herbst, JFW, 1787:350 ISS
Podura Grisea(sic) from Germany
After Herbst JFW, 1787 Taf.LXXV Fig.4ab

            Syn.: Podura Stagnorum(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:93 IOS
Podura Stagnorum(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.4

             Syn.: Podura Stagnorum(sic) Bourlet, C, 1845:160 ISS
Isotomurus palustris f. stagnorum from Belgium
2021.09.03 © Huskens, M.-L.

            Syn.: Podura fuliginosa Templeton, R in Templeton, R et Westwood, JO, 1836:95 =?
            Syn.: Podura armata Nicolet, 1842 lapsus
            Syn.: Aetheocerus aquaticus Bourlet, 1842
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
            Syn.: Isotoma spilosoma Gervais in Walckenaër, 1844
Isotome spilosome from France
Isotoma spilosoma
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.9


             Syn.: Isotoma Spilosoma(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Isotoma palustris var. spilosoma Parfitt, 1891
             Syn.: Isotomurus spilosoma(sic) Houlbert, C, 1924:106 ISS
            Syn.: Podura Aquatica(sic) Bourlet, C, 1845:151
            Syn.: Desoria riparia Bourlet, C, 1845:67
            Syn.: Desoria riparia Bourlet, C in Erichson, WP, 1846:303
            Syn.: Podura psi Herklots, 1857
            Syn.: Degeeria pi Herklots, 1858
            Syn.: Isotoma tricolor MacGillivray, 1891
            Syn.: Isotoma tricolor MacGillivray, 1896
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Womersley, 1934
            Syn.: Isotomurus tricolor Womersley, 1934
            Syn.: Isotoma tricolor Stach, J, 1947:422
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotoma tricolor Cavalloro, 1956
            Syn.: Isotomurus tricolor Hart, 1974 =?
            Syn.: Isotomurus tricolor Fjellberg, 1986
            Syn.: Isotoma balteata Reuter, 1876

            Syn.: Isotoma Stuxbergii(sic) Tullberg, 1876 IOS
            Syn.: Isotoma Tullbergi(sic) Moniez, 1889
            Syn.: Isotoma aequalis MacGillivray, 1896
            Syn.: Isotoma palustris f. pallida Schäffer, 1896
             Syn.: Isotomurus palustris ssp. pallida(sic) Salmon, JT, 1964:368 ISS
            Syn.: Isotoma maculata Börner, 1901
            Syn.: Isotoma palustris f. cincta Krausbauer, 1902
            Syn.: Isotoma palustris ab. bimaculata Agren, 1903
            Syn.: Isotomurus palustris ssp. tricuspis Börner, C, 1906:173
            Syn.: Proisotomurus papillatus Womersley, 1934
            Syn.: Isotoma lineata Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:8
            Syn.: Isotomurus lineatus Hopkin, SP, 2007:26
            Syn.: Isotomurus antennalis Poinsot, N, 1972:687
            Syn.: Isotomurus palustris f. catalana Cassagnau, 1987
Isotomurus palustris f. catalana from France
After Potapov M, 2001 Fig.370

            Syn.: Isotomurus independente Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:94
Isotomurus independente
2001 © Carapelli, A. et al.

            Syn.: Isotomurus sp. Dethier, M et Hubart, J-M, 2000:29,B
            Syn.: Isotomurus palustris f. poinsotae Janssens F & Joachim J, 2019
            Syn.: Isotomurus palustris ssp. palustris Poinsot, N, 1972:687
Isotomurus palustris ssp. palustris from France
After Poinsot N, 1972 Fig.6A
Isotomurus palustris from the UK
Aberrant abdominal pigmentation
2015.12.04 © Barton, T.
Isotomurus palustris f. poinsotae juv. from the UK
2019.10.20 © Marquis, A.
Isotomurus palustris f. poinsotae from Spain
2019.12.31 © Joachim, J.
Isotomurus palustris f. poinsotae from Spain
2019.12.31 © Joachim, J.
Isotomurus palustris f. poinsotae from Spain
2019.12.31 © Joachim, J.




             Syn.: Podura Palustris(sic) Bourlet, C, 1845:153 ISS
Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.9

             Syn.: Isotoma pulustris(sic) Stach, J, 1924:114 ISS
Isotomurus palustris from the USA
After Folsom JW, 1937 Plate25 Fig.281

Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.1
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.2
Isotomurus palustris from China
After Stach J, 1964 PlateVIII Fig.3
Isotomurus palustris ssp. palustris from the USA
After Snider RJ, 1967 Cover


             Syn.: Isotomurus palustis(sic) Heungens, A et Van Daele, E, 1973:141,B ISS
             Syn.: Isotomurus pallustris(sic) Greenslade, P in Dallai, R, 1989:111 ISS
Isotomurus palustris from Hawaii
After Christiansen K & Bellinger P, 1992 Plate 65 Fig.A

Isotomurus palustris
2001 © Carapelli, A. et al.
Isotomurus palustris from Italy
After Potapov M, 2001 Fig.372

Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2005 © Hopkin, S.P.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from the UK
2006 © Barton, T.
Isotomurus palustris from Belgium
2006 © Vuijlsteke, M.
Isotomurus palustris from the UK
2007 © Valentine., B
Isotomurus palustris from the USA
2007.02.20 © Moorehead, C.
Isotomurus palustris
2007 © Fjellberg, A.
Isotomurus palustris from the UK
2007.10.16 © Campbell, A.
Isotomurus palustris from the UK
2007.10.19 © Campbell, A.
Isotomurus palustris from Holland
2007.11.08 © Baas, A.H.
Isotomurus palustris from the UK
2007.11.24 © Campbell, A.
Isotomurus palustris from the USA
2008.01.12 © Moorehead, C.
Isotomurus palustris from the UK
2008.01.12 © Farrell, S.
Isotomurus palustris from Holland
2008.01.26 © Baas, A.H.
Isotomurus palustris? from France
2008.02.24 © Jacquemin, G.
Isotomurus palustris from Germany
2008.04.17 © Peters, J.
Isotomurus palustris from Holland
2008.11.14 © Baas, A.H.
Isotomurus palustris
2008.12.25 © onefivenine.


             Syn.: Isotomurus palustres(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Cutz-Pool, LQ, 2009:205 ISS
Isotomurus palustris from the USA
2009.01.31 © Justis, S.
Isotomurus palustris from the USA
Clypeus dark
2009.02.01 © Justis, S.
Isotomurus palustris from France
2009.02.25 © Lefebvre, V.
Isotomurus palustris from the USA
2009.03.03 © Justis, S.
Isotomurus palustris from the UK
2009.03.07 © Barton, T.
Isotomurus palustris from Holland
2009.03.27 © Baas, A.H.
Isotomurus palustris from Holland
2009.03.27 © Baas, A.H.
Isotomurus palustris from the UK
2009.12.26 © Barton, T.
Isotomurus palustris from the UK
Clypeus dark
2009.12.26 © Barton, T.
Isotomurus palustris? from Holland
2011.03.22 © van Duinen, J.
Isotomurus palustris from the UK
Post-antennal organ
2012.01.03 © Kent, J.J.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus cf palustris from the UK
2012.08.30 © Murray, A.
Isotomurus palustris? from the UK
2012.11.05 © Jones, A.
Isotomurus palustris? from the UK
2012.11.05 © Jones, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from the UK
2012.11.07 © Jones, A.
Isotomurus palustris from La Palma
2012.12.01 © Barton, T.
Isotomurus palustris from Holland
2014.11.07 © van Duinen, J.
Isotomurus palustris from the UK
Phoretic curled up nematode
2015.02.24 © Thomas., B
Isotomurus palustris from the UK
2015.03.23 © Barton, T.
Isotomurus palustris from the UK
Clypeus dark
2015.03.23 © Barton, T.
Isotomurus palustris from Belgium
2016.01.04 © Huskens, M.-L.
Isotomurus palustris? from the UK
2016.01.07 © Barton, T.
Isotomurus palustris from the UK
2016.07.22 © David W.
Isotomurus palustris from the USA
2016.11.28 © Dobbs, Br.
Isotomurus palustris? from France
2016.12.06 © Garcelon, P.

             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:101,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:103,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:104,gr ISS
             Syn.: Isotomurus Palustris(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:105,gr ISS
Isotomurus palustris from Belgium
2017.11.15 © Huskens, M.-L.
Isotomurus palustris from Belgium
2017.11.15 © Huskens, M.-L.
Isotomurus palustris juv. from the UK
2018.02.01 © Horton, M.
Isotomurus palustris juv. from Belgium
2020.11.28 © Huskens, M.-L.
Isotomurus palustris juv. from Belgium
2020.11.29 © Huskens, M.-L.
Isotomurus palustris juv. from Belgium
Ventral anal valve dark
2020.12.09 © Huskens, M.-L.
Isotomurus palustris juv. from Belgium
Ventral anal valves dark
2020.12.09 © Huskens, M.-L.





Species palustroides Folsom, JW, 1937:77

             Syn.: Isotomurus paulustroides(sic) Weber, 1950 ISS
             Syn.: Isotomurus pallstroides(sic) Huang & Liang, 1991 ISS

Species paraciliatus Cardoso, MA, 1970:109

Species philfrancisi Bernard, 2006

Isotomurus philfrancisi
2006 © Bernard, E.

Species plumosus Bagnall, 1940, B

            Syn.: Isotoma lineata Lubbock, J, 1862:597
Isotoma lineata from the UK
After Lubbock J, 1862 Pl.LIX Fig.9
Isotomurus plumosus f. lineatus from the UK
2019.10.20 © Marquis, A.
Isotomurus plumosus f. lineatus from France
2020.11.21 © Joachim, J.
Isotomurus plumosus f. lineatus from France
Eyepatch extended with fine stripe at ocellus F
2020.11.21 © Joachim, J.
Isotomurus plumosus f. lineatus from Belgium
Head pattern
2021.06.02 © Huskens, M.-L.
Isotomurus plumosus f. lineatus from Belgium
2021.06.03 © Huskens, M.-L.
Isotomurus plumosus f. lineatus from Belgium
Head pattern
2021.06.03 © Huskens, M.-L.
Isotomurus plumosus f. lineatus from Belgium
2021.06.03 © Huskens, M.-L.
Isotomurus plumosus f. lineatus from Belgium
Clypeus dark
2021.06.03 © Huskens, M.-L.
Isotomurus plumosus f. lineatus juv. from Belgium
2023.01.03 © Huskens, M.-L.

             Syn.: Isotoma Nicoletii(sic) Lubbock, J, 1862:597 =? ISS
            Syn.: Isotomurus palustroides ssp. subciliatus Stach, J, 1947:461
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.1
Isotomurus palustroides subciliatus from Poland
After Stach, J., 1947 Plate L Fig.2

             Syn.: Isotomurus palustroides ssp. subcialiatus(sic) Rusek, J in Dlabola, J, 1977:15 ISS
            Syn.: Isotomurus plumosus f. bimis Janssens F & Huskens ML, 2019
Isotomurus plumosus f. bimis from Belgium
Dark banded bimus-like form
2019.07.23 © Huskens, M.-L.
Isotomurus plumosus f. bimis from Belgium
Dark banded bimus-like form
2019.07.23 © Huskens, M.-L.
Isotomurus plumosus f. bimis from Belgium
Dark banded bimus-like form
2019.07.27 © Huskens, M.-L.
Isotomurus plumosus f. bimis from Belgium
Dark banded bimus-like form
2019.07.27 © Huskens, M.-L.
Isotomurus plumosus f. bimis from Belgium
Almost black form
2019.08.04 © Huskens, M.-L.




Isotomurus plumosus from Belgium
2001 © Hopkin, S.P.
Isotomurus plumosus from Poland
After Potapov M, 2001 Fig.373
Isotomurus plumosus from Sweden
2006 © Hall, K.
Isotomurus plumosus
2007 © Fjellberg, A.
Isotomurus plumosus from the UK
2007.04.09 © Cornwall, N.J.
Isotomurus plumosus from Belgium
2007.03.28 © Lindsley, J.
Isotomurus plumosus juv. from France
2008.03.02 © Lebeaux, P.
Isotomurus plumosus from Denmark
2012.03.19 © Bildstrup, O.
Isotomurus plumosus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus plumosus from the UK
2013.06.24 © Fenwick, D.
Isotomurus plumosus juv. from the UK
2014.09.16 © Garlick, T.
Isotomurus plumosus juv. from the UK
2014.09.16 © Garlick, T.
Isotomurus plumosus juv. from the UK
2014.10.24 © Garlick, T.
Isotomurus plumosus from Czechia
With Sminthurinus elegans f. ornata
2015.03.05 © Krásenský, P.
Isotomurus plumosus? juv. from Holland
2015.07.02 © van Leeuwen, J.
Isotomurus plumosus from Holland
2015.12.04 © van Duinen, J.
Isotomurus plumosus from Belgium
2016.03.13 © Janssens, F.
Isotomurus plumosus juv. from Belgium
2016.04.17 © Huskens, M.-L.
Isotomurus plumosus from Belgium
2016.07.10 © Janssens, F.
Isotomurus plumosus from Belgium
Postantennal organ (PAO) elliptical with central constriction
2016.08.10 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Frontal facial pattern
2016.08.11 © Huskens, M.-L.
Isotomurus plumosus? from the UK
2017.02.01 © David W.
Isotomurus plumosus juv. from Belgium
2017.09.08 © Huskens, M.-L.
Isotomurus plumosus from France
Head without interocular stripe
2017.09.10 © Picard, J.
Isotomurus plumosus juv. from Belgium
Different juvenile stages
+ Sminthurides malmgreni ♀
2019.02.15 © Janssens, F.
Isotomurus plumosus juv. from Belgium
+ Podura aquatica
2019.02.15 © Janssens, F.
Isotomurus plumosus from the UK
2019.02.23 © Smart, J.
Isotomurus plumosus from Belgium
'F-patch': elongate patch near ocellus F
2019.08.30 © Huskens, M.-L.
Isotomurus plumosus from the UK
2019.10.20 © Marquis, A.
Isotomurus plumosus juv. from Belgium
2019.10.24 © Huskens, M.-L.
Isotomurus plumosus from the UK
2020.02.05 © Cann, A.J.
Isotomurus plumosus from the UK
Cephalic pattern
2020.02.05 © Cann, A.J.
Isotomurus plumosus from Belgium
'F-patch': elongate patch near ocellus F
2020.02.18 © Huskens, M.-L.
Isotomurus plumosus juv. from Belgium
2020.02.29 © Huskens, M.-L.
Isotomurus plumosus juv. from Belgium
Extra ocular patch (EOP)
2020.05.16 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Dark form near to f. bimis
2020.07.22 © Huskens, M.-L.
Isotomurus plumosus from Belgium
2021.06.03 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Eye with post-antennal organ (PAO)
Distinct extra ocellar patch at ocellus F
2021.11.05 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Unguiculus (u) with basal lamella
2021.11.05 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Eye with distinct extra ocellar patch at ocellus F
2021.11.06 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Eye with post-antennal organ (PAO)
Distinct extra ocellar patch at ocellus F
2021.11.06 © Huskens, M.-L.
Isotomurus plumosus from Belgium
Post-antennal organ (PAO)
Extra ocular patch (EOP)
2021.11.16 © Huskens, M.-L.

Species potapovi Lafooraki, EY, Hajizadeh, J et Shayanmehr, M, 2023:?

Species pseudopalustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127

            Syn.: Podura palustris ssp. aquatilis Müller OF, 1776
Isotoma aquatilis from the UK
After Lubbock, J., 1873 Pl.37.

             Syn.: Isotoma aquatelis(sic) Parfitt, 1891 ISS
Isotoma palustris var. aquatilis from Sweden
After Schött H, 1893 Taf.V Fig.7
Isotomurus palustris f. aquatilis from the USA
2010.01.23 © Justis, S.
Isotomurus pseudopalustris nr f. aquatilis from France
2013.01.13 © Quintin, C.
Isotomurus pseudopalustris f. aquatilis from the UK
2013.01.25 © Cobby, P.
Isotomurus palustris f. aquatilis from the USA
2017.02.06 © Dobbs, Br.
Isotomurus pseudopalustris nr f. aquatilis from France
2018.02.24 © Tiky, S.
Isotomurus pseudopalustris f. aquatilis from the Canary Islands
2018.03.09 © Huskens, M.-L.
Isotomurus pseudopalustris f. aquatilis from the UK
2018.12.28 © Marquis, A.
Isotomurus pseudopalustris f. aquatilis from the UK
Abd.4 pattern
2019.01.21 © Cann, A.J.
Isotomurus palustris f. aquatilis from the USA
2019.11.15 © Christopher, Br.
Isotomurus pseudopalustris f. aquatilis from Holland
2022.01.13 © Kamsteeg, G.


            Syn.: Isotoma trifasciata Bourlet, C, 1839:402
Isotoma trifasciata from the UK
After McIntire SJ, 1867 Fig.37
After McIntire SJ, 1868
Isotoma trifasciata from the UK
After Lubbock J, 1870 Pl.XXII Fig.20
Isotoma trifasciata from the UK
Foot of anterior leg
After Lubbock J, 1870 Pl.XXII Fig.21

            Syn.: Isotomurus palustris f. trimaculata(sic) Womersley, H, 1939:149 lapsus
            Syn.: Isotoma Gervaisii(sic) Nicolet, H, 1847:374 IOS
Isotoma Gervaisii(sic) from Switzerland
After Nicolet,, 1847 Pl.6 Fig.9


             Syn.: Isotoma Gervaisii(sic) Brauer, 1855 ISS
             Syn.: Isotoma Gervaisii(sic) Kolenati, FA, 1858:133 ISS
             Syn.: Isotoma Gervaisii(sic) Lubbock, J, 1873:176 ISS
             Syn.: Isotoma Gervaisii(sic) Stach, J, 1947:367 ISS
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2004 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2005.06.02 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from Belgium
2008.12.11 © Vuijlsteke, M.
Isotomurus pseudopalustris f. gervaisi from the UK
2011.08.26 © Barton, T.
Isotomurus pseudopalustris f. gervaisi from Holland
2014.02.11 © van Duinen, J.
Isotomurus pseudopalustris f. gervaisi from the UK
2014.12.20 © Evans, K.
Isotomurus pseudopalustris f. gervaisi from Belgium
2016.05.27 © Huskens, M.-L.
Isotomurus pseudopalustris f. gervaisi from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus pseudopalustris f. gervaisi from Belgium
2018.12.27 © Huskens, M.-L.
Isotomurus pseudopalustris f. gervaisi from France
Very faint dorsal pigmentation
2019.01.06 © Picard, J.
Isotomurus pseudopalustris f. gervaisi from France
Dark clypeus
2019.01.06 © Picard, J.
Isotomurus pseudopalustris f. gervaisi from Belgium
2019.10.03 © Huskens, M.-L.
Isotomurus pseudopalustris f. gervaisi from Holland
2019.11.24 © Kamsteeg, G.


            Syn.: Isotoma tricolor Schött, H, 1891:22
            Syn.: Isotoma tricolor MacGillivray, AD, 1894:105
            Syn.: Isotoma lineata Lubbock, J, 1873:170
            Syn.: Isotoma palustris f. principalis Schött, H, 1893:63 partim
Isotoma palustris forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.8
Isotomurus palustris forma principalis from Germany
After Heymons R & Heymons H, 1909 Fig.11


            Syn.: Isotomurus palustris f. maculata(sic) Stach, J, 1947:459 partim
Isotomurus palustris f. maculata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.2
Isotomurus palustris f. maculata(sic) from Poland
After Stach, J., 1947 Plate XLVIII Fig.3


            Syn.: Isotomurus palustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 1995:87
            Syn.: Isotomurus pseudopalustris f. fuscescens Janssens F & Huskens M-L, 2021 B
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish colour form
2021.06.24 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish colour form
2021.06.24 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish colour form
2021.06.24 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Pale brownish colour form
2021.06.25 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Pale brownish colour form
Head with arrow pattern
2021.06.25 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Pale brownish colour form
Clypeus dark
2021.06.25 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish banded colour form
2021.07.18 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish banded colour form
2021.07.18 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish banded colour form
Dark clypeus + labrum
2021.07.18 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish banded colour form
2021.07.18 © Huskens, M.-L.
Isotomurus pseudopalustris f. fuscescens from Belgium
Brownish pale colour form
2021.07.18 © Huskens, M.-L.




Isotomurus pseudopalustris from Italy
After Potapov M, 2001 Fig.366-367
Isotomurus pseudopalustris from the UK
2006 © Brocklehurst, K.
Isotomurus pseudopalustris from the UK
2006.06.24 © Botting, J.
Isotomurus palustris f. pseudopalustris from the USA
2007 © Moorehead, C.
Isotomurus pseudopalustris from Germany
2008.04.17 © Peters, J.
Isotomurus pseudopalustris from the UK
2008.09.24 © Robertson, A.
Isotomurus palustris f. pseudopalustris from the USA
2010.02.07 © Zungri, D.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.10.21 © Nurcombe, E.
Isotomurus pseudopalustris from the UK
2012.11.03 © Nurcombe, E.
Isotomurus pseudopalustris from France
2012.11.05 © Quintin, C.
Isotomurus pseudopalustris from Germany
2015.01.20 © Landgraf, A.
Isotomurus pseudopalustris? from Holland
2015.02.06 © van Duinen, J.
Isotomurus pseudopalustris from Holland
2015.02.24 © van Duinen, J.
Isotomurus pseudopalustris from Holland
Two pale dots on abd.4
2015.10.19 © van Duinen, J.
Isotomurus pseudopalustris from the UK
2015.11.25 © Evans, K.
Isotomurus pseudopalustris from the UK
2015.12.08 © Horton, M.
Isotomurus pseudopalustris from the UK
2015.12.28 © Evans, K.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
2016.02.07 © David W.
Isotomurus pseudopalustris from the UK
Unguiculus with distinct inner tooth
2016.02.13 © Phillips, E.
Isotomurus pseudopalustris from Belgium
2016.04.09 © Huskens, M.-L.
Isotomurus palustris f. pseudopalustris from the USA
2016.04.17 © Dobbs, Br.
Isotomurus palustris f. pseudopalustris from the USA
2016.04.21 © Dobbs, Br.
Isotomurus pseudopalustris from Belgium
Dorsal anal valve pigmentation
2016.05.05 © Huskens, M.-L.
Isotomurus pseudopalustris from Belgium
Clypeus midfacial darker
2016.05.05 © Huskens, M.-L.
Isotomurus pseudopalustris from Belgium
2016.05.08 © Huskens, M.-L.
Isotomurus pseudopalustris? from France
2017.02.05 © Picard, J.
Isotomurus pseudopalustris? from Belgium
2017.06.29 © Huskens, M.-L.
Isotomurus pseudopalustris? from France
2017.11.18 © Picard, J.
Isotomurus pseudopalustris from France
Clypeus midfacial darker
2017.11.18 © Picard, J.
Isotomurus pseudopalustris from Holland
Abd.4 pigmentation pattern
2017.12.24 © Kamsteeg, G.
Isotomurus pseudopalustris? from France
2018.02.15 © Real, J.-A.
Isotomurus pseudopalustris? from France
2018.08.14 © Vol, F.
Isotomurus pseudopalustris? from the UK
2018.08.31 © Cann, A.J.
Isotomurus pseudopalustris? from Australia
2018.09.28 © Buck, D.
Isotomurus pseudopalustris from Holland
2018.11.23 © Kamsteeg, G.
Isotomurus palustris f. pseudopalustris from the USA
2019.01.04 © Pearson, R.R.
Isotomurus pseudopalustris juv. from the UK
2019.02.20 © Jeffery, J.
Isotomurus palustris f. pseudopalustris from the USA
2021.01.23 © Schneider, K.
Isotomurus palustris f. pseudopalustris from the USA
2021.01.23 © Schneider, K.
Isotomurus palustris f. pseudopalustris from the USA
2021.01.23 © Schneider, K.
Isotomurus palustris f. pseudopalustris from the USA
2021.01.23 © Schneider, K.
Isotomurus pseudopalustris from Belgium
2021.03.30 © Huskens, M.-L.
Isotomurus pseudopalustris juv. from Belgium
Abd.6 with bilateral spots
2021.03.30 © Huskens, M.-L.
Isotomurus pseudopalustris juv. from Belgium
2021.03.30 © Huskens, M.-L.



Species pseudosensillatus de Mendonça, 1990

Species punctiferus Yosii, 1963

             Syn.: Isotomurus punctatus(sic) Yosii, R, 1967:2 lapsus ISS
             Syn.: Isotomurus punctiferous(sic) Gupta & Mukharji, 1977 ISS
Isotomurus punctiferus from Japan
After Potapov M, 2001 Fig.375
Isotomurus nr punctiferus from Norway
2007.03.29 © Fauskanger, I.
Isotomurus nr punctiferus from France
2008.11.08 © Jacquemin, G.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.09.25 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.
Isotomurus nr punctiferus juv. from the UK
2012.10.02 © Murray, A.

             Syn.: Isotomurus punctiferous(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:104 ISS
Isotomurus punctiferus from Taiwan
2022.05.29 © Cheng, H.-J.
Isotomurus punctiferus from Taiwan
2022.05.29 © Cheng, H.-J.



Species qinpuensis Chen, 1985

Isotomurus cf. qinpuensis from Japan
After Sumida C in Sakai S & Nakamori T, 2020 Fig.1
Isotomurus ?qinpuensis? from Taiwan
2021.08.12 © Cheng, H.-J.
Isotomurus ?qinpuensis? from Taiwan
2022.01.27 © Cheng, H.-J.
Isotomurus ?qinpuensis? from Taiwan
2022.03.15 © Cheng, H.-J.

Species quadrisetosus Rusek, 1971

Species rabili Deharveng & Lek, 1993

Isotomurus rabili
2001 © Carapelli, A. et al.

Species retardatus Folsom, JW, 1937:71

            Syn.: Isotomurus retardatus Mills & Rolfs, 1933 ???
             Syn.: Isotomurus ratardatus(sic) Keilbach, 1982 ISS
Isotomurus retardatus from the USA
2021.06.15 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Head lateral pattern
2021.06.15 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
2021.06.15 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Head dorsal pattern
2021.06.15 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Eyepatch with 8 ocelli
Ocelli BDE in straight line
2021.06.17 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Dens crenulated
2021.06.21 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Mucro with basal lobe (*)
Teeth lamellate
2021.06.21 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Unguiculus with basal lamella no tooth
Unguis slender without inner teeth
2021.06.22 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Mucro with 4 teeth in inner aspect
Tooth 2 with lamella; basal rounded lobe (*)
2021.06.25 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Mucro with 4 teeth in posterior aspect
Outer basal tooth (1)
2021.06.25 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Trunk with longitudinal thin median line
2021.06.25 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Explosive take-offs when disturbed
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Take-off from water surface
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Stabilising body in mid-air after jump
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Using collophore as anchor during landing on water to prevent bouncing
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Everting collophore vesicles just before impact to adhere to watersurface
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Traject in mid-air after jump; sticky collophore used at landing to prevent bouncing
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Spring of furca reloaded during traject in air
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Collophore releases substrate to free energy stored in spring of furca
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Collophore picks up drop of water during take-off to be used at landing
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Almost vertical take-off
Collophore released from watersurface
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Sliding take-off
Uplifted abdomen; collophore not released
2022.08.31 © Ortega-Jiminez, V.M.
Isotomurus retardatus from the USA
Take-off
Down pushed abdomen
2022.08.31 © Ortega-Jiminez, V.M.



Species riparius de Mendonça, 1990

Species sahebganjensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:105

Isotomurus sahebganjensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.5 Fig.1

Species sensillatus Winter, 1967

Species stepposus Potapov, MB et Starostenko, EV, 2002:331

Isotomurus stepposus from Ukraine
After Potapov MB & Starostenko EV, 2002 Fig.1

Species stuxbergi (Tullberg, T, 1877:35) Babenko & Bulavintsev, 1993

            Syn.: Isotoma Stuxbergii(sic) Tullberg, T, 1877:35 IOS
            Syn.: Isotoma Stuxbergi(sic) var. prasina Reuter, 1891
            Syn.: Isotoma Stuxbergii(sic) var. prasina Reuter, OM, 1895:26 IOS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1905 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Wahlgren, 1907 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Collinge & Shoebotham, 1910 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1911 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Linnaniemi, 1912 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Shoebotham, 1914 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Brown, 1918 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Axelson, 1906 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1920 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Denis, 1921 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1923 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Thompson, 1924 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stirnimann, 1926 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Jackson, 1928 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Handschin, E in Dahl, F, 1929:73 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Stach, 1929 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1930 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1930 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Kseneman, M, 1932:20 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Turk, 1933 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills, 1934 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Schubert, 1935 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Drenowski, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Folsom, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1937 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Womersley, 1939 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Szent-Ivany, 1940 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Buitendijk, 1941 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Stach, J, 1947:459 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Frenzel, 1949 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Mills & Richards, 1953 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, 1954 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Goto, 1955 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, R, 1955:387 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Grinbergs, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) von Torne, 1958 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Gisin, H, 1960:214 ISS
             Syn.: Isotomurus (Isotomurus) palustris var. prasina(sic) Grinbergs, 1960 ISS
             Syn.: Isotomurus palustris ssp. prasina(sic) Salmon, JT, 1964:368 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Poinsot-Balaguer, 1976 ISS
             Syn.: Isotomurus palustris f. prasina(sic) Yosii, R, 1977:156 ISS
             Syn.: Isotomurus palustris var. prasina(sic) Fjellberg, A, 2007:98=? ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:3 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:4 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:5 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:6 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:7 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:9 ISS
             Syn.: Isotomurus prasis(sic) Gundersen, H, Leinaas, HP, et Thaulow, C, 2014:10 ISS
            Syn.: Isotomurus palustris ssp. palustris Brimley, 1942
            Syn.: Isotomurus ciliatus Stach, 1947
            Syn.: Isotomurus tricolor Babenko & Bulavintsev, 1993 =?


             Syn.: Isotoma Stuxbergi(sic) Moniez, R, 1890:325 ISS
             Syn.: Isotoma Stuxbergi(sic) Moniez, R, 1890:326 ISS
Isotomurus stuxbergi
2007 © Fjellberg, A.





Species subterraneus Stach, 1946

Species takahashii (Yosii, R, 1940:115) Yosii, 1963

Isotoma takhaashii(sic) from Formosa
After Yosii R, 1940 Fig.7
            Syn.: Isotomurus alticulus ssp. japonicus Yosii, R, 1956:64
            Syn.: Isotomurus alticolus ssp. japonicus Yosii, R, 1977:156



             Syn.: Isotoma takhaashii(sic) Yosii, R, 1940:116 ISS

Species texensis Folsom, JW, 1937:76

Isotomurus palustris texensis from the USA
After Folsom JW, 1937 Plate25 Fig.282
Isotomurus palustris texensis from the USA
After Folsom JW, 1937 Plate25 Fig.283

Species tricolor (Packard, AS, 1873:34) Lippert & Butler, 1976

            Syn.: Isotoma palustris Folsom, JW, 1901:159
            Syn.: Isotoma palustris Guthrie, JE, 1903:70
            Syn.: Isotoma palustris f. principalis Guthrie, JE, 1903:70
            Syn.: Isotoma palustris Folsom, JW, 1919:11A
            Syn.: Isotomurus palustris Mills, HB, 1934:54
            Syn.: Isotomurus palustris Maynard, EA, 1951:122
            Syn.: Isotomurus palustris Salmon, JT, 1964:362 partim
            Syn.: Isotomurus palustris Salmon, JT, 1964:363 partim
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1980:687
            Syn.: Isotomurus palustris Christiansen, K et Bellinger, P, 1998:756


Isotomurus (Isotomurus) tricolor from the USA
After Folsom JW in Christiansen K & Bellinger P, 1980 Fig.546A
Isotomurus tricolor from the USA
2006 © Murray, T.
Isotomurus tricolor from the USA
2007.05.18 © Cowen, R.
Isotomurus tricolor from the USA
2008.01.06 © Moorehead, C.
Isotomurus tricolor from the USA
2009.01.07 © Fleming, K.
Isotomurus tricolor from the USA
2009.03.04 © Murray, T.
Isotomurus tricolor from the USA
2009.08.19 © Zungri, D.
Isotomurus tricolor from the USA
2010.02.08 © Zungri, D.
Isotomurus tricolor from the USA
2010.01.21 © Murray, T.
Isotomurus tricolor from the USA
2012 © Steffen, J. & al.
Isotomurus tricolor from the USA
2012.01.14 © Justis, S.
Isotomurus tricolor from the USA
2012.09.22 © Murray, T.
Isotomurus tricolor from the USA
2015.04.16 © Mallory, C.
Isotomurus tricolor from the USA
2017.01.20 © Fleming, K.
Isotomurus tricolor from the USA
2020.04.10 © Alloway, L.
Isotomurus tricolor from the USA
2021.01.12 © Pavy, R.
Isotomururus tricolor from the USA
2021.02.04 © Duncan, W.L.
Isotomururus tricolor from the USA
Abd.4 with 2 macrosetae (m) and bothriotrichum (b) in a triangle
2021.02.04 © Duncan, W.L.
Isotomururus tricolor from the USA
2021.02.04 © Duncan, W.L.
Isotomururus tricolor from the USA
2022.12.30 © Hendrix, S.
Isotomururus tricolor from the USA
Cephalic pattern
2022.12.30 © Hendrix, S.

Species tricuspis Börner, C, 1906:173

Species unifasciatus (Börner, C, 1901:50) Stach, J, 1947:459, B

            Syn.: Isotoma palustris f. principalis Schött partim


Isotomurus unifasciatus
2001 © Carapelli, A. et al.
Isotomurus unifasciatus from the UK
2005 © Hopkin, S.P.
Isotomurus unifasciatus from the UK
2007 © Barton, T.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus from the UK
2007 © Cornwall, N.J.
Isotomurus unifasciatus
2007 © Fjellberg, A.
Isotomurus unifasciatus from the UK
2007 © Spears, G.
Isotomurus unifasciatus from Norway
2009.11.29 © Haraldseide, H.
Isotomurus unifasciatus from the UK
2012.01.26 © Kent, J.J.
Isotomurus unifasciatus from the UK
2012.11.02 © Murray, A.
Isotomurus unifasciatus juv. from the UK
2013.01.18 © Nurcombe, E.
Isotomurus unifasciatus from Holland
2013.02.07 © van Duinen, J.
Isotomurus nr unifasciatus from New Zealand
2014.06.01 © Murray, A.
Isotomurus unifasciatus from Germany
2015.01.05 © Beck, K.
Isotomurus unifasciatus juv. from Germany
2015.03.28 © Kluck, A.
Isotomurus unifasciatus from the UK
2018.12.04 © Smart, J.
Isotomurus unifasciatus from Belgium
2020.02.25 © Huskens, M.-L.
Isotomurus unifasciatus from Belgium
2020.02.25 © Huskens, M.-L.
Isotomurus unifasciatus from Belgium
2020.02.25 © Huskens, M.-L.
Isotomurus unifasciatus juv. from Belgium
2020.07.12 © Huskens, M.-L.
Isotomurus unifasciatus from France
2021.03.22 © Vol, F.

Species venustus Barra, J-A, 2006:67

Species viridipalustris Kos, 1937

Species yamaquizuensis Winter, 1967

            Syn.: Isotomurus yamaquizuensis Winter, 1963 nn


Isotomurus cf. yamaquizuensis from Colombia
After Mari Mutt JA, 1987 Fig.11


Genus Chionobora Greenslade, P et Potapov, M, 2015:334

Species amila Greenslade, P et Potapov, M, 2015:334, t.t.

Chionobora amila from Tasmania
After Greenslade P & Potapov M, 2015 Fig.1
Chionobora amila ♂ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1A
Chionobora amila ♀ from Tasmania
After Stevens MI & D'Haese CA, 2016 Fig.1B



Genus Kaylathalia Stevens, MI et D'Haese, CA, 2016:3

Species klovstadi (Carpenter, 1902) Stevens, MI & D'Haese, CA, 2016:3, t.t.

            Syn.: Isotoma Klovstadi(sic) Carpenter, 1902 IOS


             Syn.: Isotoma Klovstadi(sic) Enderlein, 1909 ISS
             Syn.: Isotoma Klovstadi(sic) McNamara, 1919 ISS
             Syn.: Isotoma Klovstadi(sic) Gressitt & Weber, 1959 ISS
Desoria klovstadi from Antarctica
2008.05.13 © O'Brien., B

             Syn.: Chionobora klovsatdi(sic) Stevens, MI et D'Haese, CA, 2016:3 ISS
             Syn.: Kaylathalia klovsadi(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:7 ISS






Genus Hydroisotoma Stach, J, 1947:249

  Syn.: Proisotominæ(sic) Chisotoma Bagnall, RS, 1949:533
  Syn.: Isotomidæ(sic) Eosotoma Bagnall, RS, 1949:532
  Syn.: Collembola Eosotoma Ellis, WN et Bellinger, PF, 1973:19


Species schaefferi (Krausbauer, 1898) Stach, J, 1947:250, t.t.

            Syn.: Isotoma schäfferi(sic) Krausbauer, 1898,tt IOS
            Syn.: Isotoma rivularis Latzel, 1921
            Syn.: Archisotoma dimorpha Denis, 1922
            Syn.: Proisotoma Schäfferi(sic) ssp. aculeata Stach, J, 1930:289
             Syn.: Proisotoma schaefferi ssp. aculata(sic) Wray, DL, 1978:205 ISS
            Syn.: Hydroisotoma stachi Bagnall, 1949


             Syn.: Proisotoma (Ballistura) Schäfferi(sic) Heymons, R et Heymons, H in Brauer, A, 1909:5 ISS
Proisotoma Schäfferi(sic) from Germany
After Heymons R & Heymons H, 1909 Fig.6

             Syn.: Isotoma schäfferi(sic) Folsom, 1928 ISS
             Syn.: Proisotoma Schäfferi(sic) Denis, J-R, 1931:116 ISS
             Syn.: Isotoma schäfferi(sic) Tarsia, 1943 ISS
             Syn.: Isotoma schäfferi(sic) Gisin, 1944 ISS
             Syn.: Hydroisotoma schäfferi(sic) Stach, J, 1947:250 ISS
Hydroisotoma schäfferi(sic) from Poland
After Stach, J., 1947 Plate XXIXa Fig.1

             Syn.: Isotoma schäfferi(sic) Bagnall, RS, 1949:534 ISS
             Syn.: Proisotoma (Proisotoma) schäfferi(sic) Bagnall, RS, 1949:534 ISS
             Syn.: Eosotoma schäfferi(sic) Bagnall, RS, 1949:534 ISS
             Syn.: Isotoma (Isotoma) schäfferi(sic) Maynard, EA, 1951:108 ISS
             Syn.: Isotoma schäfferi(sic) Maynard, EA, 1951:108 ISS
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.198
Isotoma schäfferi(sic) from the USA
After Maynard EA PlateXII Fig.199

             Syn.: Hydroisotoma schafferi(sic) Gisin, H, 1960:213 ISS
             Syn.: Hydroisotoma schafferi(sic) Snider, RJ, 1967:212 ISS
Hydroisotoma schaefferi ♂ from the USA
Habitus lateral
After Wray DL, 1978 Fig.5
Hydroisotoma schaefferi ♂ from the USA
Habitus dorsal
After Wray DL, 1978 Fig.6
Hydroisotoma schaefferi ♂ from the USA
Head with curled antennae
After Wray DL, 1978 Fig.2
Hydroisotoma schaefferi ♂ from the USA
Antenna with sense organs
After Wray DL, 1978 Fig.1
Hydroisotoma schaefferi from Germany
After Schulz, H.-J., 1989
Hydroisotoma schaefferi from Germany
2006 © Schulz, H.-J. & Schubert H.D.
Hydroisotoma schaefferi from the USA
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2007 © Murray, T.
Hydroisotoma schaefferi from the USA
2007.03.01 © Murray, T.
Hydroisotoma schaefferi from the USA
2008.01.13 © Murray, T.
Hydroisotoma schaefferi from France
2008.03.16 © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi juv. from France
2009.09.dd © Lebeaux, P.
Hydroisotoma schaefferi from the USA
2011.02.20 © Husic, C.
Hydroisotoma schaefferi from the USA
Furca with subcylindrical dentes
2012.12.02 © Fleming, K.
Hydroisotoma schaefferi ♂ from the USA
Male with bent apical antennal segments and warts on 2nd antennal segment
2013.03.28 © Murray, T.

             Syn.: Hydroisotoma schafferi(sic) Glime, JM, 2015:3 ISS
Hydroisotoma schaefferi from Germany
2015.03.31 © Schulz, H.-J.
Hydroisotoma schaefferi from the USA
2017.01.19 © Fleming, K.
Hydroisotoma schaefferi from France
2018.03.03 © Tiky, S.
Hydroisotoma schaefferi from France
2018.03.03 © Tiky, S.
Hydroisotoma schaefferi from France
2019.02.25 © Tiky, S.
Hydroisotoma schaefferi from France
2019.02.25 © Tiky, S.
Hydroisotoma schaefferi from Canada
2020.03.27 © Armstrong, Be.
Hydroisotoma schaefferi ♀ from the USA
2021.11.16 © Shultz, E.P.
Hydroisotoma schaefferi ♀ from the USA
2021.11.16 © Shultz, E.P.
Hydroisotoma schaefferi from the USA
2022.02.11 © Karighattam, A.






Genus Agrenia Börner, C, 1906:171 key

  Syn.: Isotomini Ågrenia(sic) Börner, C, 1906:171 IOS
  Syn.: Isotomidae Ågrenia(sic) Folsom, JW, 1937:68 IOS
Agrenia sp. from the USA
2022.03.05 © Feldman, A.


Species-group bidenticulata
Remark by Fjellberg, A. & Bernard, E.C. 2009:19: Lateral mucronal seta present.




Species sp.1 Fjellberg, A, 1988:112

Species bidenticulata (Tullberg, T, 1877:35) Denis, J-R, 1924:231, t.t., B

            Syn.: Isotoma lanuginosa Carl, J, 1899:307
            Syn.: Agrenia bidenticulata var. elongata Carpenter, 1908
            Syn.: Agrenia bidenticulata var. nigra Linnaniemi, 1935


Agrenia bidenticulata from Belgium
After Jacquemart S, 1975 p.83
Agrenia bidenticulata from Russia
2006 © Boldsystems.org
Agrenia bidenticulata from Norway
2015.mm.dd © Fjellberg, A.
Agrenia bidenticulata from Buryatia
With long claws
After Potapov M, Gulgenova A & Babykina M, 2016 Fig.37

Species lamellosa Fjellberg, A, 1988:110

Species pilosa Fjellberg, A, 1986:101

Species polymorpha Fjellberg, A, 1986:96

Agrenia ?polymorpha? from the USA
2011.05.30 © Eiseman, C.
Agrenia polymorpha from Canada
2020.02.23 © Majka, C.
Agrenia polymorpha from Canada
Eyepatches interconnected with transverse band
2020.02.23 © Majka, C.
Agrenia polymorpha from Canada
Uniform body colour
2020.02.25 © Majka, C.
Agrenia polymorpha from Canada
With post-ocular median spot
2020.02.25 © Majka, C.

Species-group agilis
Remark by Fjellberg, A. & Bernard, E.C. 2009:20: Lateral mucronal seta absent.


Species agilis Fjellberg, A, 1986:101

Agrenia agilis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3A

Species atroviridis Fjellberg, A, 1986:104

Agrenia atroviridis from the USA
2008.02.09 © Murray, T.
Agrenia atroviridis from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3B

Species cyanura Fjellberg, A, 1986:102

Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3C
Agrenia cyanura from the USA
After Fjellberg A & Bernard EC, 2009 Fig.3D

Species extrema Fjellberg, A et Bernard, EC, 2009:24

Species falcula Fjellberg, A et Bernard, EC, 2009:25

Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5A
Agrenia falcula from the USA
After Fjellberg A & Bernard EC, 2009 Fig.5B

Species parkeri Fjellberg, A et Bernard, EC, 2009:20

Agrenia parkeri from the USA
2006 © Bernard, E.C.
Det. Bernard, E.C., 2010.01.25
Agrenia parkeri ♀ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri ♂ from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4A
Agrenia parkeri from the USA
After Fjellberg A & Bernard EC, 2009 Fig.4B

Species riparia Fjellberg, A, 1986:103

Agrenia riparia from Norway
After Fjellberg A, 2007 Fig.58A

Species tarashchukae Fjellberg, A et Bernard, EC, 2009:27


Genus Pteronychella Börner, C, 1909:113

  Syn.: Collembola Pteronychlla(sic) Yosii, R, 1961:14 ISS
  Syn.: Collembola Pteronvchella(sic) Yosii, R, 1961:18 ISS

Species andina Yoshii, 1990

Species babai Yosii, 1939

Species ezoensis Yosii, R, 1965:30

Species perpulchra Börner, C, 1909:113, t.t.

             Syn.: Pteronychella perpulchura(sic) Yosii, R, 1961:19 ISS


Genus Isotomedia Salmon, JT, 1944:148

Species triseta Salmon, JT, 1944:148, t.t.


Genus Procerura Salmon, JT, 1941:326

  Syn.: Isotomini Papillomurus Salmon, JT, 1941:330
  Syn.: ... Salmonides Bagnall, 1949


Species aequaoculata Salmon, JT, 1941:328

Species dissimilis (Salmon, JT, 1944:146) Christiansen, K & Bellinger, P, 2000:107


Procerura fasciata from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.198.
Species fasciata Salmon, JT, 1941:329


Species fusca (Salmon, JT, 1941:330) Christiansen, K & Bellinger, P, 2000:107

  Subspecies pallida Salmon, JT, 1941:331

Species indica (Baijal, HN, 1958:355) Greenslade, P, 2003:1, s.i.

Procerura indica from India
After Yadav RS et al, 2018 Fig.1

Species magnifica (Salmon, JT, 1949:29) Greenslade, P, 2003:1, s.i.

Species montana Salmon, JT, 1941:328


Isotoma parva from New Zealand
After Salmon, J.T. 1937 Pl.49 Fig.a.
Species parva (Salmon, JT, 1937:352) Greenslade, P, 2003:1, s.i.

             Syn.: Papillomurus parva(sic) Salmon, JT, 1964:372 ISS

Species purpurea Salmon, JT, 1941:328

Species ochracea (Salmon, JT, 1949:27) Christiansen, K & Bellinger, P, 2000:107

Species serrata Salmon, JT, 1941:329

Species transequatoria (Salmon, 1969) Greenslade, P, 2003:1, s.i.

            Syn.: Papillomurus transequatoria(sic) Salmon, 1969 IOS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:24 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:74 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:78 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:116 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:126 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:128 ISS
             Syn.: Papillomurus transequatoria(sic) Mandal, GP et Hazra, AK, 2009:178 ISS



Species violacea Salmon, JT, 1941:327, t.t.


Genus Spinocerura Salmon, 1941


Spinocerura capillata from New Zealand
After Salmon, J.T. 1941 Pl.52 Fig.212.
Species capillata Salmon, JT, 1941:332, t.t.


Species dreuxi (Delamare Deboutteville & Massoud, 1966)


Genus Setocerura Salmon, JT, 1949:33

Species georgiana (Schäffer, 1891) Wise, KAJ, 1970:198

Species maruiensis (Salmon, JT, 1941:326) Deharveng, L & de Oliveira, EP, 1994:442


Tomocerura rubenota from New Zealand
After Salmon, J.T. 1941 Pl.51 Fig.193.
Species rubenota (Salmon, JT, 1941:325) Salmon, JT, 1949:33, t.t.


Species sp. Greenslade, P in Dallai, R, 1989:112

Species striata (Schäffer, 1897)

Species wahlgreni (Womersley, H, 1939:321) Salmon, 1949


Genus Tomocerura Wahlgren, 1901

Species colonavia Salmon, JT, 1949:31

Species conjungens (Schäffer, 1897) Wahlgren, E, 1906:17

Species faguagae Izarra, 1970

Species georgiana (Schäffer, 1897) Massoud & Rapoport, 1968

Species picta Wahlgren, 1901, t.t.

Species swani (Womersley, 1934)

Isotoma swani from Australia
After Womersley H, 1939 Pl.1 Fig.2.


Genus Isotoma Bourlet, C, 1839:399 key

  Syn.: Podurides(sic) Podura Bourlet, C, 1845:114
  Syn.: Isotominae Panurotoma Bagnall, RS, 1949:540
  Syn.: Collembola Panurotoma Poole, RW in Poole, RW et Gentili, P, 1997:51
  Syn.: Isotominae Pathisotoma Bagnall, RS, 1949:540
  Syn.: Collembola Pathisotoma Poole, RW in Poole, RW et Gentili, P, 1997:51


  Syn.: Desoria Deuxième division Nicolet H in Desor, E, 1841:385
  Syn.: Podurides(sic) Podura Elditt, HL, 1854:42 partim
  Syn.: Podurelles (A-tête-directe) Isotama(sic) Nicolet, H, 1847:374 ISS
  Syn.: Podurides(sic) Isotoma Elditt, HL, 1854:42
Isotoma from Germany
Schematic sagittal section
After Anonymous Fig.E

  Syn.: Arthropleona Isootoma(sic) Richards, WR, 1968:15 ISS
Remark by ICZN. 2007:419: Placed on the Official List of Names in Zoology. Opinion 291.

Isotoma sp. from the USA
2009.01.29 © Justis, S.
Isotoma sp. from Tunesia
Predation by pseudoscorpion Garypus
2009.04.14 © Henderickx, H.
Isotoma sp. from Norway
3+3 macrosetae on abd.4 in one line
After Fjellberg A, 2010 p.2
Isotoma sp. from Norway
More elongate head
After Fjellberg A, 2010 p.2
Isotoma anglicana from Belgium
Eyepatch with 6 large ocelli and 2 small (*)
And ocelli CDE in a straight line
2018.02.02 © Huskens, M.-L.
Isotoma viridis from the UK
Abdominal dorsal macrosetae in transverse row
2019.02.04 © Cann, A.J.
Isotoma viridis from the UK
Eyepath with ocelli CDE in straight line
2019.07.13 © Cann, A.J.
Isotoma sp. from Belgium
Eyepatch with ocelli CDE in straight line
2021.04.22 © Huskens, M.-L.
Isotoma sp. from Taiwan
2022.02.07 © Lee, R.
Isotoma sp. from Taiwan
2022.02.07 © Lee, R.
Isotoma sp. from Taiwan
2023.02.23 © Lin, T.-P.



Species sp.1 Janssens F sensu auct. anon., 1879

            Syn.: Isotoma sp. auct. anon., 1879
            Syn.: Isotoma sp. auct. anon., 1889
            Syn.: Isotoma spec. Willem, V, 1902:15


Species sp.2 Janssens F sensu Wise, 1970

            Syn.: Isotoma (Isotoma) sp. Wise, 1970 si
            Syn.: Isotoma sp.indet.nov.comb. Greenslade, P in Dallai, R, 1989:112
            Syn.: Isotoma sp. Greenslade, P in Houston, WWK, 1994:64,si
            Syn.: Isotoma georgiana Womersley, 1934
            Syn.: Isotoma georgiana Womersley, 1939

Species sp.3 Janssens F sensu Brand & Dunn, 1998

            Syn.: Isotoma sp.2 Brand, RH et Dunn, CP, 1998:238


Species sp.4 Janssens F sensu Brand & Dunn, 1998

            Syn.: Isotoma sp.3 Brand, RH et Dunn, CP, 1998:238


Species sp.5 Janssens F sensu Rebek, Young & Hogg, 1999

            Syn.: Isotoma sp.1 Rebek, EJ, Young, DK et Hogg, DB, 1999:60


Species acrea Wray, 1953

Species anglicana Lubbock, J, 1862:596

            Syn.: Isotoma Anglicana(sic) Lubbock, J, 1862:596 IOS
Isotoma Anglicana(sic) from the UK
After Lubbock J, 1862 Pl.LIX Fig.8
Isotoma anglicana from the UK
After Lubbock J, 1873 Pl.38
Isotoma anglicana from the UK
2005 © Hopkin, S.P.
Isotoma anglicana from the UK
Manubrial thickening with 4 teeth
2005 © Hopkin, S.P.
Isotoma anglicana from Germany
Manubrial thickening with 4 teeth
Det. Schulze, H.J.
2005 © Mebes, H.
Isotoma anglicana from Sweden
2006 © Hall, K.
Isotoma anglicana juv. from Holland
On snow
2006.03.02 © Hamers., B
Isotoma anglicana from the UK
2007.03.25 © Barnes, C.
Isotoma anglicana from Belgium
2008.02.07 © Hanssens., B
Isotoma anglicana from Holland
2008.02.08 © Baas, A.H.
Isotoma anglicana from the UK
2008.04.13 © Kilford., B
Isotoma anglicana from France
2008.11.13 © Lavoué,B.
 
Isotoma anglicana from the UK
2008.12.10 © Talbot, M.
Isotoma anglicana from the UK
2009.03.01 © Kilford., B
Isotoma anglicana from Holland
2009.03.18 © Baas, A.H.
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from Holland
Neck pattern of bilateral spots
2009.03.19 © Baas, A.H.
Isotoma anglicana from Holland
2009.03.19 © Baas, A.H.
Isotoma anglicana from Holland
2009.03.20 © Baas, A.H.
Isotoma anglicana from the UK
2009.03.21 © Talbot, M.
Isotoma anglicana from the UK
2009.08.20 © Ransom, T.
Isotoma anglicana from Norway
After Fjellberg A, 2010 p.3
Isotoma anglicana from Norway
Manubrial dorsal macrosetae: A>B
After Fjellberg A, 2010 p.8
Isotoma anglicana from Norway
3 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma anglicana juv. from the UK
Neck pattern of bilateral spots
2010.02.01 © Robertson, A.
 
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana juv. from the UK
2010.02.01 © Robertson, A.
Isotoma anglicana from Holland
2010.02.06 © van Duinen, J.
Isotoma anglicana from the UK
2010.05.02 © Robertson, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from the UK
2011.12.01 © Jones, A.
Isotoma anglicana from Holland
Ovoid post-antennal organ (PAO)
2012.05.13 © van Duinen, J.
Isotoma anglicana from the UK
Neck pattern of bilateral spots
2012.08.13 © Murray, A.
Isotoma anglicana from Holland
Abdominal pigmentation
2012.12.14 © Hamers., B
Isotoma anglicana from Belgium
On snow
2013.02.15 © Hamers., B
Isotoma anglicana from Holland
2014.02.21 © van Duinen, J.
Isotoma anglicana from Germany
2015.01.17 © Landgraf, A.
Isotoma anglicana from Holland
2015.02.04 © van Duinen, J.
Isotoma anglicana juv. from the UK
Neck pattern of bilateral spots
2015.02.23 © Cunningham, T.
Isotoma anglicana from Holland
2015.03.10 © van Duinen, J.
Isotoma anglicana from the UK
2015.08.16 © Boardman, P.
Isotoma anglicana from the UK
Manubrial thickening with 2+2 teeth (*)
2015.08.16 © Boardman, P.
Isotoma ?anglicana? juv. from Sweden
2016.03.03 © Neergaard, R.
Isotoma ?anglicana? juv. from Sweden
2016.03.03 © Neergaard, R.
Isotoma anglicana from Holland
2016.03.17 © van Duinen, J.
Isotoma anglicana from Holland
Distinctly spotted skin
2016.03.17 © van Duinen, J.
Isotoma anglicana from Belgium
Extra interantennal spots
2017.06.09 © Huskens, M.-L.
Isotoma anglicana from Belgium
Neck with spotted pattern
2017.06.09 © Huskens, M.-L.
Isotoma anglicana from Belgium
Neck with spotted pattern
2017.06.09 © Huskens, M.-L.
Isotoma anglicana from Belgium
2018.01.18 © Huskens, M.-L.
Isotoma anglicana from Belgium
Extra interantennal spots
2018.01.18 © Huskens, M.-L.
Isotoma anglicana from Belgium
Parasitic protonymph Acari
2018.02.02 © Huskens, M.-L.
Isotoma anglicana from the UK
With dark spotted neck pattern
2018.03.21 © David W.
Isotoma anglicana from the UK
Manubrial thickening with 2 teeth (*)
2019.01.15 © Cann, A.J.
Isotoma anglicana from the UK
With Vertagopus sp.
2019.02.24 © Jeffery, J.
Isotoma anglicana from the UK
2019.08.13 © Smart, J.
Isotoma anglicana from the UK
2019.08.13 © Smart, J.
Isotoma anglicana from the UK
2019.08.13 © Smart, J.
Isotoma anglicana juv. from Belgium
Neck pattern
2019.09.21 © Huskens, M.-L.
Isotoma anglicana from Belgium
2020.01.14 © Huskens, M.-L.
Isotoma anglicana juv. from Belgium
2020.01.14 © Huskens, M.-L.
Isotoma anglicana from Belgium
Extra interantennal spots
2020.12.23 © Huskens, M.-L.
Isotoma anglicana juv. from Belgium
Extra interantennal spots
2020.12.23 © Huskens, M.-L.
Isotoma anglicana from Belgium
2020.12.30 © Huskens, M.-L.
Isotoma anglicana from Belgium
Ocelli CDE in straight line
2020.12.30 © Huskens, M.-L.
Isotoma anglicana from Belgium
Extra interantennal spots
2020.12.30 © Huskens, M.-L.
Isotoma anglicana from Belgium
Furca with crenulated dentes
2021.01.07 © Huskens, M.-L.
Isotoma anglicana from Belgium
Collophore with open apical valves
2021.01.07 © Huskens, M.-L.
Isotoma anglicana from Belgium
Linea ventralis
2021.01.07 © Huskens, M.-L.
Isotoma anglicana from Belgium
Extra interantennal spots
2021.01.07 © Huskens, M.-L.
Isotoma anglicana from Belgium
2021.01.15 © Huskens, M.-L.
Isotoma anglicana from Belgium
With interantennal spots
2021.01.15 © Huskens, M.-L.
Isotoma anglicana ♂ from Belgium
2021.05.24 © Huskens, M.-L.
Isotoma anglicana ♂ from Belgium
Inter-antennal spots
2021.05.24 © Huskens, M.-L.
Isotoma anglicana ♂ from Belgium
Manubrial macrosetae
A distinctly larger than D (B+C not in focus)
2021.05.24 © Huskens, M.-L.
Isotoma anglicana ♂ from Belgium
Genital plate roundish
2021.05.24 © Huskens, M.-L.
Isotoma anglicana from the UK
Furca flexed under ventrum
2022.10.12 © Barnett, D.
Isotoma anglicana from the UK
Furca extended backwards
2022.10.12 © Barnett, D.
Isotoma anglicana from the UK
Composed eye
2022.10.12 © Barnett, D.



Species arctica Schött, H, 1893:61

Isotoma viridis var. arctica from the American Arctics
After Schött H, 1893 Taf.V Fig.4
Isotoma viridis var. arctica from Alaska
After Folsom JW, 1902 Pl.IV Fig.5

             Syn.: Isotoma viridis var. artica(sic) Stach, J, 1924:114 ISS
Isotoma arctica from Alaska
After Fjellberg A, 2003 Fig.9
Isotoma arctica from the USA
Det. Christiansen, K.
2006 © Schimming, L.
Isotoma arctica? from the USA
2021.01.02 © Kneidel, K.



Species aspera Bacon, 1914

Species burkhardti Winter, 1967

Species caeca Reuter, 1880

Species caerulea Bourlet, C, 1839:401, B

             Syn.: Isotoma coerulea(sic) Nicolet, H, 1847:374 ISS
             Syn.: Isotoma coerulea(sic) Elditt, HL, 1854:37 ISS
             Syn.: Isotoma viridis f. coerulea(sic) Handschin, E in Dahl, F, 1929:67 ISS
             Syn.: Isotoma (Isotoma) viridis var. coerulea(sic) Palissa, A, 1960:378 ISS
             Syn.: Isotoma (Isotoma) viridis ssp. coerulea(sic) Palissa, A in Schwoerbel, J et Zwick, P, 2000:100 ISS
             Syn.: Isotoma coerulea(sic) Fjellberg, A, 2003:94 ISS
             Syn.: Isotoma cerulea(sic) Gudleiffsson, BE et Bjarnadottir, B, 2008:56 ISS
Isotoma caerulea from Belgium
2008.02.07 © Hanssens., B
Isotoma caerulea from the UK
2008.02.27 © Spears, G.
Isotoma caerulea from the UK
2009.03.25 © Ransom, T.
Isotoma caerulea from Norway
After Fjellberg A, 2010 p.4
Isotoma caerulea from Norway
Manubrial dorsal macrosetae: A<B
After Fjellberg A, 2010 p.8
Isotoma caerulea from Norway
2 dorsal basal dental macrosetae
After Fjellberg A, 2010 p.9
Isotoma caerulea from Denmark
2010.11.10 © Bildstrup, O.
Isotoma caerulea from the UK
2011.12.05 © Jones, A.
Isotoma caerulea from the UK
2012.11.08 © Jones, A.
Isotoma caerulea from the UK
2013.09.22 © Murray, A.
Isotoma caerulea from Germany
2015.12.18 © Kluck, A.
Isotoma caerulea from the UK
2015.12.17 © David W.
Isotoma caerulea from Holland
2016.11.15 © van Duinen, J.
Isotoma caerulea from France
2017.02.19 © Ramel, A.
Isotoma caerulea from Belgium
2017.06.13 © Huskens, M.-L.
Isotoma caerulea from the UK
Manubrial thickening with 2+2 teeth
2019.12.06 © Cann, A.J.
Isotoma caerulea from Belgium
2021.02.25 © Huskens, M.-L.
Isotoma caerulea from Belgium
Interantennal spots triangular
2021.02.25 © Huskens, M.-L.
Isotoma caerulea from Belgium
2021.02.25 © Huskens, M.-L.
Isotoma caerulea from Belgium
Left eye with aberration
Ocelli CDF in straight line
2021.02.25 © Huskens, M.-L.

             Syn.: Isotoma coerulea(sic) Bonfanti, J, Krogh, PH, Hedde, M et Cortet, J, 2022:4 ISS


Species crassicornis (Handschin, 1926 nec Schött, 1893 -) Weitschat, W & Wichard, W, 2002:86, -

Species decorata Brown, JM, 1926:205

Isotoma viridis var. decorata from Mesopotamia
After Brown JM, 1926 Fig.11

            Syn.: Isotoma viridis var. decorata f. obscura Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. obscura from Palestine
After Handschin E, 1942 Fig.1a

            Syn.: Isotoma viridis var. decorata f. pallida Handschin, E, 1942:405
Isotoma viridis-decorata(sic) f. pallida from Palestine
After Handschin E, 1942 Fig.1b

            Syn.: Isotoma viridis var. decorata f. principalis Handschin, E, 1942:405



             Syn.: Isotoma viridis-decorata(sic) Handschin, E, 1942:404 ISS

Species delta MacGillivray, 1896

            Syn.: Isotoma catena Guthrie, JE, 1903:69


Isotoma (Isotoma) viridis delta from the USA
After Folsom JW, 1937 Plate37 Fig.435
Isotoma delta from the USA
2007.12.29 © Bryant, P.
Isotoma delta juv. from the USA
2008.03.12 © Houston, S.
Isotoma delta juv. from the USA
2008.05.09 © Bradford, A.
Isotoma delta from the USA
2008.11.08 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2008.11.15 © Justis, S.
Isotoma delta from the USA
2009.01.16 © Montgomery, G.
Isotoma delta from the USA
2009.04.27 © Kropiewnicki, T.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2011.03.18 © Hoyer, R.
Isotoma delta from the USA
2012.01.04 © Kouri, J.
Isotoma delta juv. from the USA
2012.01.14 © Justis, S.
Isotoma delta from the USA
2015.12.31 © Schmidt, M.L.
Isotoma delta from the USA
2018.12.16 © Owl, Bo.
Isotoma delta from the USA
2 interantennal spots
2020.01.20 © Scholnick, S.
Isotoma delta from the USA
2021.02.16 © Martin, D.
Isotoma delta juv. from the USA
2021.05.06 © Pavy, R.
Isotoma delta juv. from the USA
2021.11.09 © Cordes, F.L.
Isotoma delta juv. from the USA
2021.11.09 © Cordes, F.L.
Isotoma delta juv. from the USA
2021.11.09 © Cordes, F.L.
Isotoma delta from the USA
2022.01.24 © Moore, J.
Isotoma delta from the USA
2022.12.21 © Shahan, T.
Isotoma delta from the USA
2022.12.21 © Shahan, T.
Isotoma delta from the USA
Dark colour form
2023.01.10 © Fleming, K.
Isotoma delta from the USA
Dark colour form
2023.01.10 © Fleming, K.

Species diverticula Yosii, 1966


Isotoma exiguadentata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.232.
Species exiguadentata Salmon, JT, 1941:336


Species finitima Scherbakov, 1899, s.d.

Species fulva Schäffer, 1897

Species galapagoensis Jacquemart, 1975 in Peck SB & Jacquemart J, 2011:3 u

Species gandhi Rapoport & Rubio, 1968

Species glauca Packard, AS, 1873:33

Species gracilliseta Börner, C, 1909:113

Species grana Lee BH et Kim JT in Lee, B-H, Kim, B-J et Kim, J-T, 1993:285

Species hibernica Carpenter, 1906


Podura himalayana from the Himalayas
After Baijal, H.N., 1955 Fig.1
Species himalayana (Baijal, HN, 1955:175)


            Syn.: Podura himmalayana(sic) Baijal, HN, 1955:175 IOS

Species incerta (Nicolet, H, 1847:370) Brook, G, 1884:280

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

             Syn.: Degeeria Incerta(sic) Kolenati, FA, 1858:132 ISS

Species innominata Yosii, 1963

Species iranica Arbea, JI et Kahrarian, M, 2015:77

Isotoma iranica from Iran
After Arbea JI & Kahrarian M, 2015 Fig.23

Species kosiana Bagnall, 1949

Species littoralis (Tanaka, 1981)

Species longispina (Mac Gillivray, 1893) Folsom, 1916:480

             Syn.: Isotoma longipenna(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS

Species mackenziana (Hammer, 1953) Fjellberg, A, 2003:96

Species malvinensis Wahlgren, E, 1906:13

Species masatierrae Schött, 1921

Species melanocephala Börner, C, 1909:112

Species modesta Nicolet, 1845

Species nepalica (Yosii, 1966)

Species nishihirai Yosii, R, 1965:31

Isotoma nishihirai from Japan
After Yosii R, 1965 Fig.22A

Species ornata Wahlgren, E, 1906:15

Species pallidafasciata Salmon, JT, 1941:336

Species perkinsi Carpenter, GH, 1904:302

Isotoma perkensi from Hawaii
After Carpenter GH, 1904 Pl.IX Fig.17

Species pinnata Börner, C, 1909:110

            Syn.: Isotoma (Isotoma) pinnata var. coracina Börner, C, 1909:112
            Syn.: Isotoma pinnata var. coracina Uchida, 1938
            Syn.: Isotoma pinnata var. coracina Uchida, 1957
            Syn.: Isotoma pinnata var. coracina Yosii, 1963
            Syn.: Isotoma pinnata ssp. coracina Salmon, JT, 1964:392
             Syn.: Isotoma pinnata ssp. voracina(sic) Salmon, JT, 1964:392 ISS
            Syn.: Isotoma pinnata var. coracina Yosii, R, 1977:157
            Syn.: Isotoma (Isotoma) pinnata var. v-album(sic) Börner, C, 1909:112 IOS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1938 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Uchida, 1957 ISS
             Syn.: Isotoma pinnata ssp. album(sic) Salmon, JT, 1964:392 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Yosii, R, 1977:157 ISS
             Syn.: Isotoma pinnata var. v-album(sic) Potapov, M, 2001:282 ISS
            Syn.: Isotoma pinnata var. alba Uchida, 1943
            Syn.: Isotoma pinnata var. alba Uchida, 1957
            Syn.: Isotoma pinnata ssp. alba Salmon, JT, 1964:392
            Syn.: Isotoma pinnata var. alba Yosii, R, 1977:157


  Subspecies fasciata Börner, C, 1909:112


            Syn.: Isotoma virgata Yosii, 1963 =?


Isotoma pinnata ssp. fasciata from Taiwan
2021.11.01 © Cheng, H.-J.



Species plumosa (Salmon, 1969) Lawrence, PN, 1978:269

             Syn.: Rhodanella plumose(sic) Mandal, GP et Hazra, AK, 2009:116 ISS

Species protocinerea Handschin, 1926 -

Species rhopalota Scherbakov, AM, 1898:9

Isotoma rhopalota from Russia
After Scherbakov AM, 1898 Table I Fig.9

Species riparia (Nicolet, H, 1842:62) Nicolet, H, 1847:373, B

Desoria riparia from Switzerland
After Nicolet, H., 1842 Pl.6 Fig.6

            Syn.: Podura aquatica grisea de Geer, 1740
             Syn.: Isotoma Aquaticum griseum(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Podura palustris Bourlet, 1843
             Syn.: Isotoma palustre(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Isotoma viridis var. aquatilis Schött, 1891



             Syn.: Isotoma Riparium(sic) Kolenati, FA, 1858:133 ISS
Desoria riparia from Europe
After Houttuyn M, 1881-1883 Pl.99 Fig.4
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.2
Isotoma viridis var. riparia from Sweden
After Schött H, 1893 Taf.V Fig.3
Isotoma viridis var. riparia from Germany
After Heymons R & Heymons H, 1909 Fig.10
Isotoma (Isotoma) viridis riparia from the USA
After Folsom JW, 1937 Plate37 Fig.434
Isotoma riparia from Norway
After Fjellberg A, 2003 Fig.8
Isotoma riparia from Estonia
2006.03.18 © Tartes, U.
Isotoma riparia from the UK
After Shaw P, 2009 ©
Isotoma riparia from France
Predation by pseudoscorpion
2009.11.16 © Lebeaux, P.
Isotoma riparia from Norway
After Fjellberg A, 2010 p.6
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.07 © Lind, J.
Isotoma riparia from Sweden
2013.04.20 © Lind, J.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from Holland
2014.01.22 © Wieringa, P.
Isotoma riparia from the UK
2014.02.04 © Valentine., B
Isotoma nr riparia from Estonia
2015.02.07 © Tartes, U.
Isotoma riparia from Holland
2016.03.05 © de Wilde, A.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma nr riparia from Canada
2016.03.13 © Schmidt, H.
Isotoma riparia from Sweden
2017.02.17 © Neergaard, R.
Isotoma riparia from Sweden
2017.11.15 © Neergaard, R.
Isotoma riparia from Holland
2018.04.20 © van Duinen, J.
Isotoma riparia from Estonia
2019.02.09 © Tartes, U.
Isotoma riparia from Belgium
2019.04.17 © Huskens, M.-L.
Isotoma riparia juv. from Belgium
2019.06.30 © Huskens, M.-L.
Isotoma riparia juv. from Belgium
2020.11.18 © Huskens, M.-L.
Isotoma riparia from Belgium
2021.03.05 © Huskens, M.-L.
Isotoma riparia from Holland
2021.06.30 © Kamsteeg, G.



Species sarkundensis Baijal, HN, 1958:354

Species schalleri Winter, 1967

Species semenkevitshi Scherbakov, AM, 1898:8

Species silvatica Schäffer, 1897

Species sinensis Yue & Yin, 1999

Species spelaea Joseph, 1882

Species spinicauda Bonet, F, 1930:249

            Syn.: Isotoma spinicauda f. principalis Bonet, F, 1930:251
Isotoma spinicauda from India
After Bonet F, 1930 Fig.1A

            Syn.: Isotoma spinicauda var. decolor Bonet, F, 1930:251
            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotoma decorata Martynova & Chikatunov, 1968 =?
            Syn.: Isotoma decorata Martynova, 1968 =?
            Syn.: Isotoma decorata Yosii, 1971 =?
            Syn.: Isotoma decorata Martynova & Vtorov, 1973 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1974 =?
            Syn.: Isotoma decorata Vtorov & Martynova, 1975 =?
            Syn.: Isotoma (Isotoma) decorata Tanaka & al, 1978=?
            Syn.: Isotoma (Isotoma) decorata Janetschek, 1990=?
            Syn.: Acanthisotoma quadridentata Huang & Liang, 1994 =?



Species subcaerulea Kos, 1952

Species subviridis Folsom, JW, 1937:115

Isotoma subviridis from Canada
2010.03.08 © Doucet, D.A.
Isotoma subviridis from the USA
2012 © Steffen, J. & al.
Isotoma ?subviridis juv.? from the USA
2013.07.13 © Nanz, S.
Isotoma subviridis from Canada
2020.02.26 © Doucet, D.A.

Species tahitiensis Salmon, 1964

Species termitophila Womersley, 1934

Species tesselata Rapoport, 1962

Species tigrinella Wahlgren, E, 1906:14

            Syn.: Isotoma tigrina Schött, 1899 nec Nicolet, 1842

Species tridenticulata Schäffer, 1896

             Syn.: Isotoma 3-denticulata(sic) Absolon, K, 1901:32 ISS

Species tridentifera Schött, H, 1917:9

Isotoma tridentifera from Australia
After Schött H, 1917 Fig.12
             Syn.: Isotoma tridentifiera(sic) Greenslade, P et Kitching, RL, 2011:337 ISS
  Subspecies edenticulata Womersley, 1934



Species tridentina Tarsia in Curia, 1941

Species turbotti (Salmon, JT, 1949:30) Greenslade P, 1990

Species vaillanti Murphy, 1958

Species variodentata Dunger, WG, 1982:60

            Syn.: Isotoma (Isotoma) variodentata f. marginata Dunger, WG, 1982:62
            Syn.: Isotoma (Isotoma) variodentata f. principalis Dunger, WG, 1982:63
            Syn.: Isotoma (Isotoma) variodentata f. unidentata Dunger, WG, 1982:62


Species viridis Bourlet, C, 1839:401, B

            Syn.: Podura viatica Linnæus, 1746
            Syn.: Podura aquatilis Müller OF, 1776, sensu Gisin, 1960
            Syn.: Podura plumbea Müller, 1776
Isotoma plumbea from the USA
After Packard AS, 1872 Pl.10 Fig.6
Isotoma plumbea from the USA
After Packard AS, 1872 Pl.10 Fig.7

            Syn.: Podura fuliginosa Nicolet, H, 1847:373
            Syn.: Isotoma arborea Bourlet, C, 1839:94
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H in Desor, E, 1841:385
            Syn.: Desoria (Deuxième division) cylindrica Nicolet, H, 1842:60
Desoria cylindrica from Switzerland
After Nicolet H, 1842 Pl.6 Fig.1

             Syn.: Heterotoma Cylindricum(sic) Kolenati, FA, 1858:133 ISS
            Syn.: Desoria (Deuxième division) annulata Nicolet, H, 1842:62
Isotoma viridis var. annulata from the USA
2007.03.27 © McClarin, J.
Isotoma viridis var. annulata from the USA
2007.04.30 © Cowen, R.
Isotoma viridis var. annulata from Holland
2008.10.10 © Baas, A.H.
Isotoma viridis var. annulata from the UK
2013.10.05 © Evans, J.
Isotoma viridis var. annulata from Germany
2015.01.01 © Beck, K.
Isotoma viridis var. annulata juv. from Germany
2015.01.16 © Landgraf, A.
Isotoma viridis var. annulata from Holland
2015.02.05 © van Duinen, J.
Isotoma viridis var. annulata from the USA
2015.03.22 © Abbott, J.C.
Isotoma viridis var. annulata juv. from Belgium
2016.03.28 © Huskens, M.-L.
Isotoma viridis var. annulata from the USA
2018.01.10 © Abbott, J.C.


            Syn.: Desoria (Deuxième division) ebriosa Nicolet, H, 1842:61
            Syn.: Desoria (Deuxième division) fusca Nicolet, H, 1842:63
             Syn.: Desoria fusia(sic) Elditt, HL, 1854:40 ISS
            Syn.: Desoria (Deuxième division) fusca var. var. Nicolet, H, 1842:63
            Syn.: Desoria (Deuxième division) pallida Nicolet, H, 1842:61
Isotoma viridis var. pallida from Germany
2005 © Menzel, E.
Isotoma viridis var. pallida from the UK
2006 © Barton, T.
Isotoma viridis var. pallida from Sweden
2006 © Hall, K.
Isotoma viridis var. pallida
2014.01.19 © Evans, J.


            Syn.: Desoria (Première division) virescens Nicolet, H, 1842:59
            Syn.: Heterotoma chlorata Gervais in Walckenaër, 1844
Hétérotome verte from France
Heterotoma chlorata
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.6

            Syn.: Isotoma Desmarestii(sic) Gervais in Walckenaër, 1844 IOS
Isotome de Desmarest from France
Isotoma Desmarestii(sic)
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.11

             Syn.: Isotoma Desmarestii(sic) Nicolet, H, 1847:374,si ISS
            Syn.: Isotoma Nicolet(sic) Gervais in Walckenaër, 1844 IOS
Isotome Nicolet(sic) from France
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.12

             Syn.: Isotama(sic) Nicoletii(sic) Nicolet, H, 1847:374,si ISS
             Syn.: Isotoma Nicoletii(sic) Kolenati, FA, 1858:133 ISS
             Syn.: Podura Fuliginosa(sic) Bourlet, C, 1845:160 ISS
             Syn.: Podura Grisea(sic) Bourlet, C, 1845:160 ISS
            Syn.: Podura viridis var.1 Bourlet, C, 1845:114
            Syn.: Podura viridis var.2 Bourlet, C, 1845:114
            Syn.: Podura viridis var.3 Bourlet, C, 1845:114
            Syn.: Podura viridis var.4 Bourlet, C, 1845:114
            Syn.: Isotoma palustris Tullberg, 1871
            Syn.: Isotoma palustris var. unicolor Tullberg, 1871
            Syn.: Isotoma Belfragei(sic) Packard, AS, 1873:33 IOS
             Syn.: Isotoma Belfragei(sic) MacGillivray, 1891 ISS
            Syn.: Isotoma purpurascens Packard, AS, 1873:34
            Syn.: Isotoma tricolor Packard, 1873 partim
            Syn.: Isotoma tricolor MacGillivray, 1891 partim
            Syn.: Isotoma tricolor Schött, 1893
            Syn.: Isotoma tricolor von Dalla Torre, 1895
            Syn.: Isotoma tricolor MacGillivray, 1896 partim
            Syn.: Isotoma tricolor Folsom, 1903 partim
            Syn.: Isotoma tricolor Linnaniemi, 1912
            Syn.: Isotoma tricolor Bacon, 1914 partim
            Syn.: Isotoma tricolor Bartholin, 1916
            Syn.: Isotoma tricolor Folsom, JW, 1919:12A
            Syn.: Isotoma tricolor Handschin, E in Dahl, F, 1929:67
            Syn.: Isotoma tricolor Mills, HB, 1934:63
            Syn.: Isotoma tricolor Stach, J, 1947:422 partim
            Syn.: Isotoma tricolor Maynard, EA, 1951:110
            Syn.: Isotoma tricolor Uchida, 1957
            Syn.: Isotoma tricolor Salmon, JT, 1964:403
            Syn.: Isotoma tricolor Christiansen, K et Bellinger, P, 1998:853
            Syn.: Isotoma cincta Tullberg, 1876
Isotoma viridis var. cincta from Novaja Selmja
After Schött H, 1893 Taf.V Fig.5
Isotoma viridis var. cincta from the UK
2010.04.04 © Botting, J.

            Syn.: Isotoma lineata Schött, 1893
            Syn.: Isotoma viridis var. arctica Schött, H, 1893:61
            Syn.: Isotoma viridis f. principalis Schött, H, 1893:59
Isotoma viridis forma principalis from Sweden
After Schött H, 1893 Taf.V Fig.1

            Syn.: Isotoma riparia Lameere, A, 1895:594
            Syn.: Isotoma capitola MacGillivray, 1896
            Syn.: Isotoma delta Mac Gillivray, 1896
            Syn.: Isotoma glauca MacGillivray, 1896
            Syn.: Isotoma montana MacGillivray, 1896
            Syn.: Isotoma tridentata MacGillivray, 1896
            Syn.: Isotoma viridis var. violacea Lie-Pettersen, 1898
Isotoma viridis var. violacea from Holland
2008.01.23 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2008.01.27 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2009.04.05 © Baas, A.H.
Isotoma viridis var. violacea from Holland
2014.01.05 © van Duinen, J.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.
Isotoma viridis var. violacea from France
2014.01.17 © Sandra.
Isotoma viridis var. violacea from Belgium
2014.03.02 © Huskens, M.-L.
Isotoma viridis var. violacea from Belgium
2018.03.28 © Huskens, M.-L.
Isotoma viridis var. violacea from the UK
2019.02.10 © Cann, A.J.
Isotoma viridis var. violacea from the UK
Manubrial thickening with 1 tooth
2019.02.10 © Cann, A.J.
Isotoma viridis var. violacea from Belgium
2021.02.18 © Huskens, M.-L.


            Syn.: Isotoma paradoxa Carl, J, 1899:317
            Syn.: Isotoma viridis f. genuina Börner, C, 1901:50
            Syn.: Isotoma catena Guthrie, JE, 1903:69
            Syn.: Isotoma viridis f. micans Ågren, 1903
            Syn.: Isotoma viridis f. typica Heymons, R et Heymons, H in Brauer, A, 1909:7
Isotoma viridis f. typica from Germany
After Heymons R & Heymons H, 1909 Fig.9

            Syn.: Isotoma aspera Bacon, 1914
            Syn.: Isotoma turkestanica Stach, 1947
            Syn.: Isotomurus bemakus Wray, 1952
            Syn.: Isotoma hibernica Gisin, H, 1960:206 =?
            Syn.: Isotoma spinicauda Gisin, H, 1960:206
            Syn.: Orchesella rubra Scott, 1963 partim holotype
            Syn.: Rhodanella plumosa Salmon, 1969 =?
            Syn.: Tetracanthura mirabilis Martynova, 1971
            Syn.: Isotoma viridis var. pomona Christiansen, K et Bellinger, P, 1980:710
            Syn.: Isotoma viridis var. pomona Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Isotoma viridis var. pomona Christiansen, K et Bellinger, P, 1998:785
            Syn.: Desoria riparia Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Desoria riparia Christiansen, K et Bellinger, P, 1998:853



             Syn.: Podura Viridis(sic) Bourlet, C, 1845:125 ISS
             Syn.: Podura Viridis(sic) Bourlet, C, 1845:153 ISS
             Syn.: Podura Viridis(sic) Bourlet, C, 1845:160 ISS
             Syn.: Podura Viridis(sic) Bourlet, C, 1845:168 ISS
Podura Viridis(sic) from France
After Bourlet C, 1845 Fig.9
Isotoma viridis from Alaska
After Folsom JW, 1902 Pl.IV Fig.4
Isotoma (Isotoma) viridis from the USA
After Folsom JW, 1937 Plate37 Fig.428


             Syn.: Isotoma ciridis(sic) Salmon, JT, 1964:407 ISS
Isotoma viridis from the UK
2005 © Hopkin, S.P.
Isotoma viridis from the UK
2006 © Valentine., B
Isotoma viridis from Germany
2006 © Steiner, A.
Isotoma viridis from the UK
2006.04.08 © Read, D.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from the USA
2007 © Murray, T.
Isotoma viridis from Germany
2007 © Peters, J.
Isotoma viridis from the UK
2007.02.23 © Valentine., B
Isotoma viridis from the UK
2007.02.24 © Barton, T.
Isotoma viridis from the UK
2007.02.28 © Valentine., B
Isotoma viridis from the UK
2007.03.02 © Barton, T.
Isotoma viridis from the UK
2007.03.05 © Barton, T.
Isotoma viridis from Holland
2007.03.21 © Baas, A.H.
Isotoma viridis from the USA
2007.10.21 © Moorehead, C.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from the UK
2007.12.29 © Barton, T.
Isotoma viridis from Holland
2008.01.22 © Baas, A.H.
Isotoma viridis from the UK
2008.02.03 © Tonsbeek, M.
Isotoma viridis from Belgium
2008.03.23 © Oger, P.
Isotoma viridis from the UK
2008.04.07 © Robertson, A.
Isotoma viridis from Holland
2008.10.08 © Baas, A.H.
Isotoma viridis from Holland
2008.11.16 © Baas, A.H.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from Germany
2009.01.12 © Peters, J.
Isotoma viridis from the USA
2009.02.10 © Cowen, R.
Isotoma viridis from the USA
2009.03.05 © Cowen, R.
Isotoma viridis from the USA
2009.03.04 © Murray, T.
Isotoma viridis from Norway
After Fjellberg A, 2010 p.5
Isotoma viridis from Norway
Single ventroapical manubrial tooth
After Fjellberg A, 2010 p.7
Isotoma viridis from the UK
2010.02.06 © Barton, T.
Isotoma viridis from the UK
2010.02.14 © Robertson, A.
Isotoma viridis from the UK
2011.12.03 © Jones, A.
Isotoma viridis (bar=2µm)
Hexagonal-rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B'
Isotoma viridis (bar=500nm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2B''
Isotoma viridis from Holland
2012.01.27 © van Duinen, J.
Isotoma viridis from the UK
2012.11.08 © Jones, A.
Isotoma viridis from Holland
2015.02.06 © van Duinen, J.
Isotoma viridis from Germany
2015.02.15 © Kluck, A.
Isotoma viridis from Holland
With parasitic larva of Acari:Erythraeoidea
2015.07.26 © van Duinen, J.
Isotoma viridis from Belgium
Collophore pressed against surfacewaterfilm
2016.01.14 © Huskens, M.-L.
Isotoma viridis from the UK
2016.02.15 © Horton, M.
Isotoma viridis from Belgium
Cephalic pigmentation
2016.11.27 © Huskens, M.-L.
Isotoma viridis from Belgium
Post antennal organ (PAO), half the size of nearest ocellus
2016.11.27 © Huskens, M.-L.
Isotoma viridis from Belgium
With phoretic Nematoda
2018.01.06 © Huskens, M.-L.
Isotoma viridis from the UK
Single ventroapical manubrial tooth
2019.02.04 © Cann, A.J.
Isotoma viridis from the UK
Variant with patched abd.5-6
2019.02.04 © Cann, A.J.
Isotoma viridis from the UK
Variant with patched abd.5-6
2019.02.04 © Cann, A.J.
Isotoma viridis juv. from Belgium
2019.06.30 © Huskens, M.-L.
Isotoma viridis from the UK
Abd.4 reduced in size
to free space for the extended furca
2019.11.11 © Cann, A.J.
Isotoma viridis from the UK
Flexible cuticula at base of manubrium (*)
to support folding of furca
2019.11.11 © Cann, A.J.
Isotoma viridis from Belgium
2019.11.21 © Huskens, M.-L.
Isotoma viridis from Belgium
Footcomplex with inner unguiculus (u) and outer unguis (U)
2020.01.05 © Huskens, M.-L.
Isotoma viridis from Belgium
Neck (= protergum) pattern
2020.03.07 © Huskens, M.-L.
Isotoma viridis ♂ from Belgium
Grooming in tripod posture
2020.04.06 © Huskens, M.-L.
Isotoma viridis ♂ from Belgium
Male genital papilla roundish
2020.04.06 © Huskens, M.-L.
Isotoma viridis ♂ from Belgium
"Furcal subcoxae"
2020.04.06 © Huskens, M.-L.
Isotoma viridis from Belgium
Ocelli G & H far apart
2020.10.11 © Huskens, M.-L.
Isotoma viridis from Belgium
Ocelli CDE in a straight line
2020.12.09 © Huskens, M.-L.
Isotoma viridis from Belgium
2020.12.17 © Huskens, M.-L.
Isotoma viridis from Belgium
Dorsal thoracic ecdysial line
At which the cuticula split when moulting
2020.12.17 © Huskens, M.-L.
Isotoma viridis juv. from Belgium
Frontal eyes
2021.01.06 © Huskens, M.-L.
Isotoma viridis from Belgium
Egg cluster
Ready to hatch
2021.01.22 © Huskens, M.-L.
Isotoma viridis from Belgium
Eyepatch with 8 ocelli
Ocelli A+B largests
2021.01.22 © Huskens, M.-L.
Isotoma viridis from Belgium
Ant.4 distinctly longer then ant.3
2021.01.22 © Huskens, M.-L.
Isotoma viridis juv. from Belgium
First instar
2021.01.24 © Huskens, M.-L.

  Subspecies pomona Folsom, JW, 1937:115

            Syn.: Isotoma (Isotoma) viridis var. pomona Folsom, JW, 1937:115
Isotoma (Isotoma) viridis pomona from the USA
After Folsom JW, 1937 Plate37 Fig.436
Isotoma viridis ssp. pomona from the USA
2010.01.15 © Parker, H.S.
Isotoma viridis ssp. pomona from the USA
2010.01.15 © Parker, H.S.
Isotoma viridis ssp. pomona from the USA
2010.01.15 © Parker, H.S.
Isotoma viridis ssp. pomona from the USA
2010.01.15 © Parker, H.S.
Isotoma viridis ssp. pomona from the USA
2010.01.15 © Parker, H.S.
Isotoma viridis ssp. pomona from the USA
2016.12.11 © McDonald, G.
Isotoma viridis ssp. pomona from the USA
2017.01.29 © Gross, J.
Isotoma viridis ssp. pomona from the USA
2019.02.22 © Schneider, K.
Isotoma viridis ssp. pomona from the USA
Thoracic pattern
2019.02.22 © Schneider, K.
Isotoma viridis ssp. pomona from the USA
2019.03.07 © Flores, J.
Isotoma viridis ssp. pomona from the USA
2019.03.07 © Flores, J.
Isotoma viridis ssp. pomona from the USA
2021.11.19 © Loarie, D.
Isotoma viridis ssp. pomona from the USA
Cephalic pattern
2021.11.19 © Loarie, D.
Isotoma viridis ssp. pomona from the USA
Abd.3-6 pattern
2021.11.19 © Loarie, D.
Isotoma viridis ssp. pomona from the USA
2021.12.28 © Wildwings.
Isotoma viridis ssp. pomona from the USA
2022.02.25 © Wildwings.
Isotoma viridis ssp. pomona from the USA
2023.01.23 © Wang, G.
Isotoma viridis ssp. pomona from the USA
2023.01.23 © Wang, G.
Isotoma viridis ssp. pomona from the USA
2023.01.25 © Wang, G.
Isotoma viridis ssp. pomona from the USA
2023.01.25 © Wang, G.
Isotoma viridis ssp. pomona from the USA
Egg cluster
2023.02.05 © Wang, G.







Genus Pseudisotoma Handschin, 1924

  Syn.: Isotoma Pseudoisotoma(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
  Syn.: Collembola Psedisotoma(sic) Hornung-Leoni, CT et Palacios-Vargas, JG, 2012:67 ISS

Species agrana (Lee BH & Kim JT in Lee, B-H, Kim, B-J & Kim, J-T, 1993:288) Potapov, 2001


Pseudisotoma anamola from Lebanon
After Christiansen, K., 1958 Pl.10 Fig.29.
Species anamola Christiansen, 1958:76


Species himalayana Yosii, 1971

Species microchaeta Cassagnau, P, 1959:85

Species monochaeta (Kos, 1942) Cassagnau, P, 1954:623, B

            Syn.: Isotoma unipila Stach, 1947


             Syn.: Pseudisotoma monochæta(sic) Cassagnau, P, 1954:623 ISS

Species ohtanii (Yosii, R, 1972:90) Potapov, 2001

Species sensibilis (Tullberg, T, 1877:36) Agrell, I, 1939:8, t.t., B

            Syn.: Isotoma turicensis Haller, 1880
            Syn.: Isotoma dubia Reuter, OM, 1895:27
            Syn.: Isotoma determinata Mac Gillivray, 1896
            Syn.: Isotoma parva Mac Gillivray, 1896
            Syn.: Isotoma longidens Schäffer, 1896
            Syn.: Isotoma monstrosa Schäffer, 1896
            Syn.: Isotoma inopinata Axelson, 1902
            Syn.: Isotoma arborea Evans, 1908 =?


Pseudisotoma sensibilis from Belgium
2006.10.12 © Schelfhout, L.
Pseudisotoma sensibilis from Belgium
2008.06.09 © Vuijlsteke, M.
Pseudisotoma sensibilis from Holland
2012.10.28 © van Duinen, J.
Pseudisotoma sensibilis from the USA
Abd.5+6 fused
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the USA
2016.12.21 © Dobbs, Br.
Pseudisotoma sensibilis from the UK
2017.07.19 © Byrne, J.
Pseudisotoma sensibilis from Belgium
2020.11.25 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Abd.5+6 fused
2020.11.25 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Alternating short and long setae at abdominal posterior segment margins
2020.11.25 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Unguis with 2 distinct basal lateral teeth (*)
2020.11.25 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
2020.11.28 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Clavate tenent macroseta of footcomplex (*)
2020.12.09 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Iridisation of epicuticular wax layer
2020.12.09 © Huskens, M.-L.
Pseudisotoma sensibilis from the USA
2021.01.09 © Hendrix, S.
Pseudisotoma sensibilis from Belgium
2021.01.11 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Leg2(+3) with 3 tenent macrosetae (*)
2021.01.11 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Footcomplex with clavate tenent macrosetae (*)
2021.01.11 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
2022.01.19 © Huskens, M.-L.
Pseudisotoma sensibilis from Belgium
Posterior segment margin with alternating long and short setae
2022.01.19 © Huskens, M.-L.

Species wisei Deharveng, 1980


Genus Semicerura Maynard, EA, 1951:99

  Syn.: Isotominae Spinocerura Salmon, JT, 1964:382
  Syn.: Isotomidae Tomocerura Martynova, 1969
  Syn.: Isotomidae Setocerura Greenslade, P in Dallai, R, 1989:108


Species bishopi Maynard, EA, 1951:99, t.t.

Semicerura bishopi from the USA
After Maynard EA, 1951 PlateXI Fig.185
Semicerura nr bishopi from Canada
2012.01.25 © Walter, D.E.
Semicerura nr bishopi from Canada
2012.01.25 © Walter, D.E.

Species bryophila Potapov M et Sun X in Potapov, M, Xie, Z-J, Kuprin, A et Sun, X, 2020:106

Semicerura bryophila from China
After Potapov M & al, 2020 Fig.1
Semicerura bryophila from China
After Potapov M & al, 2020 Fig.38

            Syn.: Semicerura bishopi Martynova, 1969
            Syn.: Semicerura bishopi Stebaeva, 1976
            Syn.: Semicerura bishopi Christiansen, K et Bellinger, P, 1980:694
            Syn.: Semicerura bishopi Christiansen, K et Bellinger, P, 1998:766



Species draconis Potapov M et Sun X in Potapov, M, Xie, Z-J, Kuprin, A et Sun, X, 2020:111

Semicerura draconis from Far East Russia
After Potapov M & al, 2020 Fig.2
Semicerura draconis from Far East Russia
After Potapov M & al, 2020 Fig.39

Species goryshini (Martynova, 1969) Jie, D, Potapov, M & Sokolova, E, 2011:50

Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org
Semicerura goryshini
2006 © Boldsystems.org

Species multispinata (James, 1933) Christiansen, KA in Dindal, DL, 1990:987

             Syn.: Semicerura miltispinata(sic) Christiansen, KA in Dindal, DL, 1990:987 ISS



Genus Heteroisotoma Stach, 1947

  Syn.: Isotominae Istoma(sic) Bagnall, RS, 1949:540 ISS
  Syn.: Isotomidae Sibirisotoma Rusek, 1991


Species carpenteri (Börner, C, 1909:109), t.t.

            Syn.: Isotoma (Isotoma) andrei Mills, HB, 1934:58
            Syn.: Isotoma (Isotoma) difficilis Folsom, JW, 1937:89

Species dilatosetosa Martynova, 1980

Species heteronomica Jie, D, Potapov, M et Sokolova, E, 2011:55

            Syn.: Heteroisotoma sp.nov. Potapov & Polyakov, 2007

Species jacutica Martynova, 1980

Species sinorossica Jie, D, Potapov, M et Sokolova, E, 2011:51

Heteroisotoma sinorossica from China
After Jie & al, 2011 Fig.25

Species stebaevae (Rusek, 1991) Potapov & Stebaeva, 1992

            Syn.: Sibirisotoma stebajevae(sic) Rusek, 1991 IOS


             Syn.: Heteroisotoma stebajevae(sic) Jie, D, Potapov, M et Sokolova, E, 2011:55 ISS
             Syn.: Heteroisotoma stebaievae(sic) Stevens, MI et D'Haese, CA, 2016:4 ISS





Genus Folsomotoma Bagnall, 1949 key

  Syn.: Isotoma Sorensia Salmon, 1948


Remark by Xie, Z.-J. & al. 2020:507: In the mitochondrial maximum likelihood phylogenetic tree (Fig.1) Folsomotoma clusters with Folsomia and Cryptopygus. Reduced s-chaetotaxy of the body and the intermediate number of setae on manubrium suggest Anurophorinae (versus Isotominae).
Species anomala (Salmon, 1948)

            Syn.: Folsomotoma minuta Fanciulli, PP, Leo, C, Convey, P, Frati, F et Carapelli, A, 2018:394 =?


Species bioculata (Womersley, 1934) Salmon, JT, 1964:412, t.t.

            Syn.: Isotoma bi-oculata(sic) Womersley, 1934,tt IOS

Species boerneri (Enderlein, 1903)

            Syn.: Isotoma börneri(sic) Enderlein, 1903 IOS
            Syn.: Folsomotoma subflava Fanciulli, PP, Leo, C, Convey, P, Frati, F et Carapelli, A, 2018:394 =?




Species kerguelensis (Enderlein, 1903)

             Syn.: Isotoma octo-oculata(sic) ssp. Kerguelensis(sic) Carpenter, GH, 1907:476 ISS

Species marionensis (Deharveng, 1981)

Folsomotoma marionensis from Marion-Island
2015 © Janion-Scheepers, C.

Species minuta (Salmon, JT, 1949:24)

Species octooculata (Willem, V, 1901:262) Potapov, 2001

            Syn.: Isotoma octo-oculata(sic) Willem, V, 1901:262 IOS
            Syn.: Isotoma sp. Carl, 1907
            Syn.: Isotoma octo-oculata(sic) var. gracilis Carpenter, GH, 1907:476
Isotoma octo-oculatus(sic) var. gracilis
from Antarctica
After Carpenter GH, 1907 Plate II Fig.9


            Syn.: Isotoma masatierrae Stach, 1947 =?
            Syn.: Isotoma octo-oculata(sic) ssp. kerguelensis Salmon, JT, 1964:411




            Syn.: Isotoma octo-oculata(sic) Willem, V, 1902:13 IOS
Isotoma octo-oculata(sic) from Antarctica
After Willem V, 1902 Pl.IV Fig.7

            Syn.: Isotoma octo-oculata(sic) Wahlgren, 1906 IOS
            Syn.: Isotoma octo-oculata(sic) Carl, 1907 IOS
            Syn.: Isotoma octo-oculata(sic) Carpenter, 1907 IOS
            Syn.: Isotoma octo-oculata(sic) Enderlein, 1909 IOS
            Syn.: Isotoma octo-oculata(sic) Ivanoff, 1913 IOS
            Syn.: Isotoma octo-oculata(sic) Womersley, 1937 IOS
            Syn.: Isotoma octo-oculata(sic) Denis, 1947 IOS
            Syn.: Parisotoma octo-oculata(sic) Salmon, 1949 IOS
            Syn.: Isotoma octo-oculata(sic) Gressitt & Weber, 1959 IOS
            Syn.: Parisotoma octo-oculata(sic) Brown, 1964 IOS
            Syn.: Parisotoma octo-oculata(sic) Salmon, JT, 1964:411 IOS
             Syn.: Isotoma 8-oculata(sic) Deharveng, 1981 ISS
Folsomotoma octooculata from Antarctica
After Fanciulli PP & al, 2018 Fig.1A

  Subspecies ovata Salmon, JT, 1949:39





Species punctata (Wahlgren, E, 1906:14)

            Syn.: Sorensia dreuxi Delamare Deboutteville & Massoud, 1966
            Syn.: Isotoma (Folsomotoma) subflava Greenslade, P in Dallai, R, 1986:56


Species subflava (Salmon, JT, 1949:22)

            Syn.: Sorensia dreuxi Wise, KAJ, 1970:194
            Syn.: Sorensia dreuxi Gressitt, 1970
            Syn.: Sorensia dreuxi Dreux, 1978 =?



Genus Desoria Agassiz M & Nicolet H in Desor E, 1841:384

  Syn.: Podures(sic) Desoria Nicolet H in Desor, E, 1841:126
  Syn.: Desoria Première division Nicolet H in Desor, E, 1841:385
  Syn.: Podurides(sic) Podura Elditt, HL, 1854:42 partim
  Syn.: ... Istoma(sic) Davenport, 1903 ISS

  Syn.: Spinisotoma Stach, 1926 Spinisotoma Stach, 1926 Spinisotoma Stach, 1926
  Syn.: Collembola Isotoma? Ellis, WN, 1974:121 sp
Isotoma? spec. juv. from Rhodos
After Ellis WN, 1974 Fig.3b
Isotoma? spec. juv. from Rhodos
Ocelli BDE in straight line
Modified after Ellis WN, 1974 Fig.3a

  Syn.: ... Isotomurs(sic) Fjellberg, A, 2007:99 ISS




Remark by Duffin, C. 2009:60: Bearing in mind how much Cécile (wife of Agassiz) detested Desor as a social parasite blighting the Agassiz family it is interesting to speculate how she might have appreciated the irony of his lending his name to a 'flea'.



Desoria sp. from Singapore
2011.01.04 © Koh, J.
Desoria sp. from Tasmania
2014.04.16 © Murray, A.
Desoria olivacea from Belgium
Ocelli BDE in straight line
2018.02.18 © Huskens, M.-L.
Desoria sp. from the USA
2020.01.19 © Hendrix, S.

Species sp.1 Fjellberg, A, 2010:40

Species sp.2 Fjellberg, A, 2010:40

Species ahiehie Christiansen, K et Bellinger, P, 1992:166

Species alaskensis (Fjellberg, 1978)

Species albella (Packard, AS, 1873:32)

            Syn.: Isotoma Walkerii(sic) Packard, 1871
             Syn.: Isotoma walkerii(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS


            Syn.: Isotoma leonina Packard, AS, 1873:32

Species albicornis Fjellberg, A, 2010:32

Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.115C
Desoria albicornis from the USA
After Fjellberg A, 2010 Fig.111

Species arctica White A in Sutherland, PC, 1852:ccviii

Desoria arctica from the Arctic region
After White A in Sutherland PC, 1852 Fig.7.

Species ater Fjellberg, A, 2010:34

Desoria ater from the USA
After Fjellberg A, 2010 Fig.115D
Desoria ater from the USA
After Fjellberg A, 2010 Fig.113

Species atkasukiensis Fjellberg, 1978

Species banffensis (Christiansen, K et Bellinger, P, 1998:794)

Species bendixenae Christiansen, K et Bellinger, P, 1992:168

Species blekeni Leinaas, 1980

Desoria blekeni from Scandinavia
2015 © Boldsystems.org.

Species blufusata Fjellberg, 1978

Species breviseta Babenko, Potapov & Taskaeva, 2017

Species caeruleatra (Guthrie, JE, 1903:70)

Species calderonis Valle B in Valle, B, Cucini, C, Nardi, F, Caccianiga, M, Gobbi, M, Di Musciano, M, Carapelli, A, Ficetola, GF, Guerrieri, A et Fanciulli, PP, 2021:36

Desoria calderonis from Italy
After Valle B & al, 2021 Fig.1
Desoria calderonis from Italy
After Valle B & al, 2021 Fig.2A

Species canadensis Brown, 1932

            Syn.: Isotoma olivacea Stach, J, 1947:404


Isotoma (Desoria) canadensis from Canada
After Brown in Christiansen K & Bellinger P, 1980 Fig.583A
Desoria canadensis from Canada
2016.03.12 © Schmidt, H.
Desoria canadensis from Canada
2017.01.02 © Levac, G.

Species cancellarei (Christiansen & Bellinger, 1980) Fjellberg, A, 2010:33

Species capra Fjellberg, A, 2010:40

Species cera Christiansen, K et Bellinger, P, 1998:804

Species choi (Lee, B-H, 1977:166)

Species christianseni Fjellberg, 1978

Species creli Fjellberg, 1978

Species cryophila Fjellberg, A, 2010:38

Desoria cryophila from the USA
After Fjellberg A, 2010 Fig.115G

Species divergens (Axelson, 1900) Kaprus, IJ, Pomorski, RJ, Skarzynski, D & Potapov, MB, 2005:894

Species duodecimoculata Denis, 1927

             Syn.: Desoria duodecemaculata(sic) Fiera, C, 2007:23 ISS

Species elegans (Carl, J, 1899:319)

Species eureka Christiansen, K et Bellinger, P, 1998:806

Species fennica (Reuter, OM, 1895:5)

            Syn.: Isotoma Formáneki(sic) Absolon, K, 1901:32 IOS
            Syn.: Isotoma Formáneki(sic) Absolon, K, 1901:212 IOS
            Syn.: Isotoma formaneki Gisin, H, 1960:209 =?
            Syn.: Isotoma formaneki Rusek, J in Dlabola, J, 1977:14 =?
            Syn.: Isotoma ruseki Fjellberg, 1979
            Syn.: Isotoma (Desoria) gersi Najt, 1981
            Syn.: Isotoma (Desoria) huetheri Dunger, WG, 1982:66




Species fideli (Rapoport & Izarra, 1962)

Species fjellbergi Najt, 1981

Species flora (Christiansen & Bellinger, 1980)

Desoria flora from the USA
2012 © Steffen, J. & al.
Desoria flora from the USA
2014.02.19 © Klein, T.

Species garibaldii Fjellberg, A, 2010:36

Desoria garibaldii from the USA
After Fjellberg A, 2010 Fig.115F

Species gelida Folsom, JW, 1937:106

Desoria gelida from the USA
2010.04.01 © Murray, T.
Desoria gelida from the USA
2010.04.30 © Hodge, K.
Desoria gelida from the USA
2022.01.07 © Feldman, A.
Desoria gelida from the USA
2022.02.21 © Feldman, A.
Desoria gelida from the USA
2023.01.28 © Geisert, K.

Species gorodkovi Martynova, 1969

Species grisea (Lubbock, J, 1870:278) Fjellberg, A, 2007:130

            Syn.: Isotoma grisea var. intricata Parfitt, 1891
            Syn.: Isotoma grisescens Schäffer, 1896
            Syn.: Isotoma grisescens Collinge & Shoebotham, 1910
             Syn.: Isotoma olivacea var. griscescens(sic) Handschin, E in Dahl, F, 1929:70 ISS
            Syn.: Isotoma grisescens Denis, 1931 =?
            Syn.: Isotoma grisescens Stach, J, 1947:401
            Syn.: Isotoma tigrina Stach, J, 1947:403


Isotoma grisea from the UK
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Abdomen
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Head and thorax
After Lubbock J, 1873 Pl.39
Isotoma grisea from the UK
Eyepatch
Ocelli G & H absent
After Lubbock J, 1873 Pl.39

             Syn.: Isotoma giresea(sic) Dürkop, 1934 ISS
Desoria grisea? from the UK
2008.09.21 © Robertson, A.
Desoria grisea? from the UK
2010.11.14 © Robertson, A.
Desoria grisea? from the UK
2015.03.08 © Outen, A.
Desoria grisea from Denmark
2019 © Poul, U.
Det.: Fjellberg, A., 2019.03.07
Desoria ?grisea? from Belgium
2022.08.23 © Huskens, M.-L.
Desoria ?grisea? from Belgium
2022.08.23 © Huskens, M.-L.



Species hiemalis (Schött, H, 1893:70) Bulimar, 1987

            Syn.: Isotoma Theobaldi(sic) Carl, J, 1899:315 IOS
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma nivicola Axelson, 1900


Desoria hiemalis from Scandinavia
2015 © Boldsystems.org.



Species hissarica Martynova & Chikatunov, 1968

Species hoodensis Christiansen, K et Bellinger, P, 1998:814

Desoria hoodensis? from the USA
2009.12.29 © Parker, H.S.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.01.28 © Dobbs, Br.
Desoria hoodensis? from the USA
2017.03.12 © Dobbs, Br.
Desoria hoodensis? from the USA
2018.09.22 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.
Desoria hoodensis? from the USA
2018.12.27 © Owl, Bo.
Desoria hoodensis? from the USA
2019.11.24 © Dobbs, Br.
Desoria hoodensis? from the USA
2019.11.24 © Dobbs, Br.
Desoria hoodensis? from the USA
2019.11.24 © Dobbs, Br.
Desoria hoodensis? from the USA
Eyepatch
2019.11.24 © Dobbs, Br.
Desoria hoodensis? from the USA
2020.12.16 © Pearson, R.R.
Desoria hoodensis? from Canada
2021.04.16 © Barbin, T.
Desoria hoodensis? from Canada
Cephalic dorsal pattern
2021.04.16 © Barbin, T.

Species husaini Yosii & Ashraf, 1965

Species imparidentata (Stach, J, 1964:13) Potapov, 2001

Species infuscata Murphy, 1959

Species intermedia (Schött, 1902) Yosii, 1963, B

            Syn.: Isotoma germanica Hüther & Winter, 1961
            Syn.: Isotoma germanica Fjellberg, A, 2007:125


             Syn.: Isotoma media(sic) Schött, 1902 ISS
Remark by Potapov, M. 2001:120: Obscure species.

Desoria intermedia from Scandinavia
2008.04.04 © Fjellberg, A.
Desoria intermedia from Scandinavia
2015 © Boldsystems.org
Desoria intermedia from Scandinavia
2015 © Boldsystems.org
Desoria intermedia from Scandinavia
Cephalic pattern
2015 © Boldsystems.org
Desoria intermedia from Scandinavia
2015 © Boldsystems.org
Desoria intermedia from Scandinavia
Head pale laterally
2015 © Boldsystems.org
Desoria intermedia from Denmark
2017.02.02 © Krone, F.
Det.: Fjellberg, A.
Desoria intermedia from Denmark
2017.02.02 © Krone, F.
Det.: Fjellberg, A.
Desoria intermedia from Denmark
Abd.5+6 fused
2017.02.07 © Fjellberg, A.
Leg.: Krone, F.
Desoria intermedia from Denmark
Abd.5+6 fused
2017.02.07 © Fjellberg, A.
Leg.: Krone, F.
Desoria intermedia from Denmark
Mucro
2017.02.07 © Fjellberg, A.
Leg.: Krone, F.
Desoria intermedia from Denmark
2019.02.13 © Poul, U.
Det.: Fjellberg, A.
Desoria intermedia from Denmark
2019.02.13 © Poul, U.
Det.: Fjellberg, A.



Species iuxta Dunger, WG, 1982:68

Species japonica (Yosii, 1939) Yosii, 1954

Species jayasrae Bhattacharjee, 1984

Species juberthiei Gers & Najt, 1983

Species kaszabi Dunger, WG, 1982:70

Species komarkovae Fjellberg, 1978

Species kuraica (Martynova, 1974:19)

Species lucama Wray, 1952

Species manitobae Fjellberg, 1978

Species marissa (Folsom, JW, 1937:89) Miranda Rangel & Palacios-Vargas, 1992

Species maxillosa Fjellberg, 1978

Species mazda Yosii, 1971

Species mossopi (Womersley, H, 1934:454)

             Syn.: Collembola Isotomamossopi(sic) Thibaud, J-M, 2013:241 lapsus ISS

Species multisetis (Carpenter & Phillips, 1922) Miranda Rangel & Palacios-Vargas, 1992

Species nahuelensis (Rapoport & Rubio, 1968)

Species negishina (Börner, C, 1909:108) Yosii, R, 1977:156, s.i.

            Syn.: Isotoma sp. Andreen, A, Malterre, J, Needham, J, Olivo, O, Pessoa, SB et Zhukov-Verezhnikov, NN, 1952:18


Species neglecta (Schäffer, 1900) Dányi, L & Traser, G, 2008:46

            Syn.: Isotoma affinis Axelson, 1900
            Syn.: Isotoma tigrina ab. coerulea Ågren, 1903


Desoria neglecta from Norway
2011 © Fjellberg, A.

Species nigra Mac Gillivray, 1896

Species nigrifrons (Folsom JW in Mills, HB, 1934:60)

Desoria nigrifrons from the USA
2012 © Steffen, J. & al.

Species nivalis Papon, 1856

Species nivalis (Carl, 1910) Miranda Rangel & Palacios-Vargas, 1992

Species nivea (Schäffer, 1896) Fjellberg, 1978

            Syn.: Isotoma albella? Skarzynski, D, Pomorski, RJ, Smolis, A, Weiner, WM, Szeptycki, A, Slawska, M et Sterzynska, M, 2002:35=?


Species nixoni Fjellberg, 1978

Species nympha Snider, RJ et Calandrino, FJ, 1987:5

Isotoma (Desoria) nympha from the USA
After Snider RJ & Calandrino FJ, 1987 Fig.1

Species occulta (Börner, C, 1909:110) Yosii, R, 1977:156

Species olivacea (Tullberg, 1871) Yosii, R, 1977:156, B

            Syn.: Isotoma tigrina Linnaniemi, 1912
            Syn.: Isotoma tigrina Denis, 1937
            Syn.: Desoria tigrina Stach, J, 1947:396 =?
            Syn.: Isotoma tigrina Stach, J, 1947:396
            Syn.: Isotoma voraginum Uzel, 1891
             Syn.: Isotoma vorraginum(sic) Stach, J, 1947:396 ISS
            Syn.: Isotoma olivacea f. genuina Börner, C, 1901:56
            Syn.: Isotoma olivacea f. principalis Linnaniemi, 1912
            Syn.: Spinisotoma regina Stach, J, 1930:290
            Syn.: Isotoma olivacea f. grisescens Börner, C, 1901:56
            Syn.: Isotoma grisescens Linnaniemi, 1912
            Syn.: Isotoma grisescens Denis, 1931
            Syn.: Isotoma grisescens Stach, J, 1947:396
            Syn.: Isotoma grisea Womersley, 1930 =?
            Syn.: Isotoma grisea Stach, J, 1947:396 =?
            Syn.: Isotoma grisea Gisin, H, 1960:211 =?
            Syn.: Isotoma (Isotoma) grisea Goto & Lawrence, 1964 =?
            Syn.: Isotoma grisea Palissa, 1964 =?
            Syn.: Isotoma grisea Bolger, 1987 =?
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Spinisotoma ephippiata Brian, 1945
            Syn.: Isotoma affinis Stach, J, 1947:396
            Syn.: Isotoma neglecta Stach, J, 1947:396
            Syn.: Isotoma olivacea f. principalis Stach, J, 1947:402
            Syn.: Proisotoma subsegmentata Scott, 1959


Desoria olivacea from Holland
2009.04.05 © Baas, A.H.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Holland
2010.04.06 © van Duinen, J.
Desoria olivacea from Norway
2011 © Fjellberg, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Ovoid post-antennal organ (PAO)
2012.10.31 © Murray, A.
Desoria olivacea from the UK
Abd.5 mesosetae 2/3rds length abd.5
2012.10.31 © Murray, A.
Desoria olivacea from Holland
2014.05.04 © Wieringa, P.
Desoria olivacea from Belgium
2017.04.20 © Huskens, M.-L.
Desoria olivacea from Belgium
2018.02.18 © Huskens, M.-L.

Species olympica Fjellberg, A, 2010:39

Species pallipes (Kos, 1952)

Species pamirensis Martynova, 1975

Species pilifrons Fjellberg, A, 2010:37

             Syn.: Desoria pilifons(sic) Fjellberg, A, 2010:28 ISS

Species pjasini Martynova, 1974

             Syn.: Desoria pjaisini(sic) Greenslade, P et Potapov, M, 2015:338 ISS

Species potapovi Fjellberg, A, 2007:126

Species propinqua (Axelson, 1902) Potapov, 2001

            Syn.: Isotoma poppiusi Axelson, 1903
            Syn.: Spinisotoma pectinata Stach, 1926
Spinisotoma pectinata from Poland
After Stach, J., 1947 Plate XXXVIII Fig.1.

            Syn.: ... propinqua var. hexaspina Agrell, 1936
            Syn.: Desoria sp. Yosii, R, 1977:155
            Syn.: Isotoma (Desoria) graeca Najt, 1981



Species pseudomaritima (Stach, J, 1947:436)

Isotoma pseudomaritima from Poland
After Stach, J., 1947 Plate XLVII Fig.1.

Species quadra Christiansen & Bellinger, 1980

Species randiella Fjellberg, 1978

Species rosea Fjellberg, A, 2010:38

Species saltans Agassiz M & Nicolet H in Desor E, 1841:385 tt

Desoria saltans from Switzerland
After Nicolet H in Desor E, 1841 Pl.2 Fig.1

            Syn.: ... puce des neiges rougeatre et velue Aristote 350 BC
            Syn.: Thysanoures(sic) puce du Mont-Rose Agassiz M in Desor, E, 1841:125
            Syn.: Thysanoures(sic) puce de glacier Desor, E, 1841:126
            Syn.: Desoria (Première division) glacialis Nicolet, H, 1842:58
Desoria glacialis from Switzerland
After Nicolet H, 1842 Pl.5 fig.10




Isotoma saltans from Austria
After Schaller F & Kopeszki H, 1991 Abb.2
Isotoma saltans from Austria
After Kopeszki H, 2000 Abb.3
Desoria saltans? from Germany
2006 © Kohl, F.
Desoria saltans? from the USA
2007.02.18 © Murray, T.
Desoria saltans? from Western Estonia
2009.02.26 © Tartes, U.
Desoria saltans? from Russia
2013.01.01 © Serge, L.
Desoria saltans from Austria
2016.02.27 © van Leeuwen, J.

Species sandersoni Wray, 1952

Species spatiosa (Uchida, H & Tamura, H, 1968:10) Potapov, 2001

Pteronychella spatiosa from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo31

Species speciosa Mac Gillivray, 1896

Species subaequalis Folsom, JW, 1937:107

Species subantarctica (Izarra, 1972)

Species tadzhika Martynova & Chikatunov, 1968

Species taigicola Fjellberg, 1978

Species taimyrica (Martynova, 1974) Potapov, 2001

Species tianshanica Hao & Huang, 1995

Species tigrina (Tullberg, 1871) Parona, 1875, B

            Syn.: Isotoma affinis Axelson
            Syn.: Isotoma stachi Denis, 1929
            Syn.: Spinisotoma dimorpha Womersley, 1934
            Syn.: Isotoma tigrina var. olivacea Agren, 1903
            Syn.: Isotoma olivacea Womersley, 1934
            Syn.: Isotoma olivacea Womersley, 1939
            Syn.: Isotoma klovstadi Womersley, 1937
            Syn.: Isotoma grisea Gisin, 1944
            Syn.: Isotoma grisea Fjellberg, 1979
            Syn.: Isotoma grisea Christiansen & Bellinger, 1980
            Syn.: Isotoma grisea Poole, RW in Poole, RW et Gentili, P, 1997:51
            Syn.: Isotoma grisea Christiansen & Bellinger, 1998
            Syn.: Isotoma grisescens Agren, 1903
            Syn.: Desoria grisescens Hopkin, 2007


Desoria tigrina from the UK
Setae on fifth abdominal segment
2005 © Hopkin, S.P.
Desoria tigrina-complex from Holland
2009.04.12 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.12 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.12 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.12 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.18 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.21 © Baas, A.H.
Desoria tigrina-complex from Holland
2009.04.21 © Baas, A.H.
Desoria tigrina juv. from Iran
Ecomorphic form
After Kahrarian M & Arbea J, 2013 Fig.2
Desoria tigrina juv. from Iran
Ecomorphic abdominal spines
After Kahrarian M & Arbea J, 2013 Fig.3
Desoria tigrina from Iran
After Kahrarian M & Arbea J, 2013 Fig.4
Desoria tigrina-complex from Holland
2013.12.11 © Baas, A.H.
Desoria tigrina-complex juv. from Germany
2014.09.27 © Beck, K.
Desoria tigrina-complex juv. from Holland
2014.11.07 © van Duinen, J.
Desoria tigrina-complex juv. from Holland
2014.11.07 © van Duinen, J.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.
Desoria tigrina juv. ecomorph from Belgium
Abd.5 with transverse row of spines
2017.04.26 © Huskens, M.-L.
Desoria tigrina-complex from Holland
2018.02.21 © Kamsteeg, G.
Desoria tigrina-complex from Belgium
2019.03.03 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Abd.5 & 6 separate
Largest setae on abd.5 < length abd.5
2019.03.03 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Cephalic pattern
2019.03.03 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Eyepatch with 8 ocelli, G & H small
2019.03.07 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Eyepatch of adult with 8 ocelli, G & H small
2019.03.07 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Eyepatch of juvenile with 6 ocelli, G & H not visible
2019.03.07 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
2019.03.08 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Eyepatch with H & G far apart
2019.03.08 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Pinkish juvenile
2019.03.08 © Huskens, M.-L.
Desoria tigrina-complex from Holland
2019.04.06 © Wieringa, P.
Desoria tigrina-complex from Belgium
2019.11.06 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Abd.4 with star-shaped mark
2019.11.06 © Huskens, M.-L.
Desoria tigrina-complex from Holland
2020.01.25 © Kamsteeg, G.
Desoria tigrina-complex from Belgium
2020.02.15 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
2020.06.26 © Huskens, M.-L.
Desoria tigrina-complex from Belgium
Abd.3-6 pattern
2020.06.26 © Huskens, M.-L.
Desoria tigrina from the USA
2023.03.26 © Wang, G.
Desoria tigrina from the USA
Third foot
2023.03.26 © Wang, G.
Desoria tigrina from the USA
Eye with 8 ocelli (A-H)
2023.03.27 © Wang, G.
Desoria tigrina from the USA
Mucro with 4 teeth
Mucronal outer seta absent
2023.03.30 © Wang, G.

Species tolya Fjellberg, A, 2007:120

            Syn.: Isotoma violacea Fjellberg, 1979
            Syn.: Desoria violacea Potapov, M, 2001:157
            Syn.: ... violacea Babenko & Flellberg, 2006


Desoria tolya from Denmark
2019.02.16 © Poul, U.
Det.: Fjellberg, A.
Desoria tolya from Denmark
2019.02.16 © Poul, U.
Det.: Fjellberg, A.
Desoria tolya from Denmark
2019.02.16 © Poul, U.
Det.: Fjellberg, A.

Species torildae Fjellberg, 1978

Species triangularis Fjellberg, A, 2010:33

Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.115E
Desoria triangularis from the USA
After Fjellberg A, 2010 Fig.110

Species trispinata (Mac Gillivray, 1896) Thibaud, 1977

            Syn.: Isotoma maritima ssp. meridionalis Altner, H, 1963:47
             Syn.: Isotoma meridionale(sic) Poinsot, 1971 ISS
             Syn.: Isotoma meridionale(sic) Poinsot, N, 1972:686 ISS
            Syn.: Isotoma setinornata Lee, B-H, 1977:164
            Syn.: Isotoma setinornata Yi, 1976 nn


Isotoma trispinata from Tokara Island
Mucro
After Yosii R, 1955 Abb.4F

             Syn.: Isotoma tarspinata(sic) Lesshafft, 1977 ISS
Desoria trispinata from the Azores
2010.09.15 © E.D.E.N.
Desoria trispinata from the UK
2010.04.22 © Shaw, P.
Det.: Fjellberg, A.

             Syn.: Desoria trispina(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1666 ISS
Desoria trispinata from Japan
2013.03.12 © Hara, Y.
Desoria trispinata from Japan
Abd.4-6
2013.03.12 © Hara, Y.
Desoria trispinata from Germany
2014.10.05 © Jäschke, J.
Isotoma (Desoria) trispinata from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.574
Desoria trispinata from Holland
2016.02.11 © van Duinen, J.
Desoria trispinata from Belgium
Eggs deposited in cluster
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Freshly deposited eggs are white and smooth
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
In orange eggs the chorion has hardened
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Chorion is hydrophobe
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Chorion bursts at equator and the halves move to the poles
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
The egg flattens laterally
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Eyepatches of embryo become visible
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar caps are formed
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar cap in polar aspect
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Polar cap in frontal aspect
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
Hatchling
2017.12.14 © Huskens, M.-L.
Desoria trispinata from Belgium
2017.12.24 © Huskens, M.-L.
Desoria trispinata from Belgium
2018.06.30 © Huskens, M.-L.
Desoria trispinata from the UK
After Shaw P & Trewhella S, 2019 Fig.7
Desoria trispinata from Belgium
2019.08.04 © Huskens, M.-L.
Desoria trispinata from Belgium
2019.10.03 © Huskens, M.-L.
Desoria trispinata from the USA
In normal light
2021.12.21 © Pavy, R.
Desoria trispinata from the USA
Fluorescence under 365nm UV light
2022.12.17 © Pavy, R.
Desoria trispinata from the USA
Fluorescence under 365nm UV light
2022.12.19 © Pavy, R.
Desoria trispinata from the USA
Mucro with 3rd basal lateral tooth (*)
2022.12.19 © Pavy, R.



Species truncata Waltz & Hart, 1995

Desoria truncata from the USA
2006 © Bernard, E.C.
Desoria truncata from the USA
2007.09.05 © Barnd., B

Species tshernovi Martynova, 1974

            Syn.: Isotoma nanseni Fjellberg, 1978


             Syn.: Isotoma tschernovi(sic) Fjellberg, 1995 ISS
Desoria tshernovi from Norway
2011 © Fjellberg, A.



Species tuckeri Christiansen & Bellinger, 1980

Species tunica Christiansen & Bellinger, 1980

Species uniens (Christiansen, K et Bellinger, P, 1980:760)

Desoria uniens from the USA
After Fjellberg A, 2010 Fig.115B
Desoria ?uniens? from the USA
2012 © Steffen, J. & al.

Species violacea (Tullberg, T, 1877:36), B

            Syn.: Isotoma neglecta Schäffer
            Syn.: Isotoma violacea var. mucronata Axelson, 1900
            Syn.: Isotoma olivacea Tuxen, 1944 partim
            Syn.: Isotoma trispinata Tuxen, 1944 partim
            Syn.: Isotoma inupikella Fjellberg, 1978
            Syn.: Desoria tolya Hågvar, S, 2010:283


             Syn.: Isotoma violacaea(sic) Kopeszki, H et Meyer, E, 1996:230 ISS
Desoria violacea from the UK
Mucro with 4 teeth
2005.04.01 © Hopkin, S.P.
Desoria violacea (bar=2µm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.4E
Desoria violacea from the UK
2014.01.19 © Valentine., B



Species yukinomi (Yosii, 1939) Yosii, R, 1977:157

Species zlotini Martynova, 1968


Genus Gnathisotoma Cassagnau, P, 1957:393

Species bicolor Cassagnau, P, 1957:393, t.t.

Species deharvengi Najt, 1981

Species pallens Yoshii, 1990

Species patagonica Yoshii, 1990

Species spinolabris Fjellberg, A, 2010:42

Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.115G
Gnathisotoma spinolabris from the USA
After Fjellberg A, 2010 Fig.114


Genus Parisotoma Bagnall, RS, 1940:171

  Syn.: Collembola Isotoma Bagnall, RS, 1940:163 partim
  Syn.: Isotomidae Proisotoma(sic) Greenslade, 2002 lapsus


Species agrelli (Delamare Deboutteville, 1950), B

Species amurica (Potapov, MB, 1991:282) Potapov, 2001

Species appressopilosa (Potapov, MB, 1991:279) Potapov, M, Huang, C-W, Gulgenova, A & Luan, Y-X, 2020:19

Species atroculata Potapov, MB, 1991:285

Species baicalica Potapov M et Gulgenova A in Potapov, M, Huang, C-W, Gulgenova, A et Luan, Y-X, 2020:16

Parisotoma baicalica from Central Asia
After Potapov M & al, 2020 Fig.43

Species coeca Yosii, 1966

Species confusoculata Salmon, JT, 1944:150

Species dichaeta (Yosii, 1969)

Species dividua Salmon, JT, 1944:150

Species ekmani (Fjellberg, 1977) Fjellberg, A, 2007:47

            Syn.: Isotoma notabilis f. pallida Agrell, I, 1939:2
            Syn.: Isotoma sphagneticola Fjellberg, 2007=?
            Syn.: Isotomina sphagneticola Berg, MP, van Leeuwen, J et van Bezouw, R, 2020:49 =?


             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:365 ISS
             Syn.: Isotoma ekmany(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:366,af ISS
Parisotoma ekmani from the UK
2012.11.18 © Lewington, A.
Det.: Shaw, P.



Species greensladeae Rusek, J, 1984:364

Species hyonosenensis (Yosii, 1939) Potapov, 2001

            Syn.: Desoria pseudosetispinosa Chen, 1985
            Syn.: Isotoma setispinosa Lee, B-H, 1977:162


             Syn.: Isotoma hyonosensensis(sic) Yosii, R, 1977:156 ISS
             Syn.: Parisotoma hyonosonensis(sic) Gao, Y, Xiong, Y et Potapov, MB, 2009:59 ISS

Species insularis Deharveng, 1981

Parisotoma insularis from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.69.

Species longa Potapov, MB, 1991:283

Species notabilis (Schäffer, 1896) Bagnall, RS, 1940:171, t.t., B

            Syn.: Isotoma menotabilis Börner, 1903
            Syn.: Isotoma menotabilis Brown, JM, 1929:426 =?
            Syn.: Isotoma delicatula Brown, JM, 1929:425
            Syn.: Isotoma (Isotoma) eunotabilis Folsom J in Mills, HB, 1934:58
            Syn.: Desoria monticola Hao & Huang, 1995


             Syn.: Parisotoma nitabilis(sic) Boitaud, L, Salmon, S Bourlette, C et Ponge J-F, 2006:452 ISS
Parisotoma notabilis from the UK
2010.03.14 © Barton, T.
Parisotoma notabilis from Norway
2011 © Fjellberg, A.
Parisotoma notabilis from the UK
2012.03.21 © Nurcombe, E.
Parisotoma notabilis from the UK
2012.04.03 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from the UK
2012.08.23 © Murray, A.
Parisotoma notabilis from Iran
After Kahrarian M & Arbea J, 2013 Fig.7
Parisotoma notabilis from the UK
With square eyepatch
2013.02.03 © Jones, A.
Parisotoma notabilis juv. from Canada
2013.10.22 © Hochhalter, R.
Parisotoma notabilis from Holland
With square eyepatch of 4 ocelli
2013.11.13 © van Duinen, J.
Parisotoma notabilis from the UK
With reticulate gray body pigmentation
2013.12.23 © Murray, A.
Parisotoma notabilis from Germany
2014.10.20 © Beck, K.
Parisotoma notabilis from the UK
2016.01.13 © Horton, M.
Parisotoma notabilis from Belgium
2017.04.20 © Huskens, M.-L.
Parisotoma notabilis from the UK
2018.02.17 © Lewis, M.
Parisotoma notabilis from Holland
2019.01.09 © Kamsteeg, G.
Parisotoma notabilis from Holland
With square eyepatch
2019.01.09 © Kamsteeg, G.
Parisotoma notabilis from Holland
2019.11.27 © Kamsteeg, G.
Parisotoma notabilis from Belgium
2020.12.17 © Huskens, M.-L.
Parisotoma notabilis from Belgium
Eyepatch short square
3 or 4 ocelli
2020.12.17 © Huskens, M.-L.



Species obscurocellata Potapov, Janion & Deharveng, 2011

             Syn.: Parisotoma obscurocellatus(sic) Thibaud, J-M, 2013:240 ISS

Species picea Salmon, JT, 1949:36

Species postantennala Salmon, JT, 1949:38

Species quinquedentata Salmon, JT, 1943:382

             Syn.: Parisotoma quinquedendata(sic) Salmon, JT, 1943:382 ISS

Species raffi (Womersley, 1934)

Species reducta (Rusek, J, 1984:360) Potapov, M, Huang, C-W, Gulgenova, A & Luan, Y-X, 2020:19

Species removeopthalma Martynova, 1977

Species sexsetosa Potapov, Janion & Deharveng, 2011

Species tariva (Wray, 1953) Potapov, 1991

             Syn.: Isotoma (Desoria) tarvia(sic) Stevenson & Dindal, 1982 ISS
?Parisotoma tariva? from the USA
2020.01.09 © Fleming, K.
?Parisotoma tariva? from the USA
Eyes absent
2020.01.09 © Fleming, K.



Species terricola (Rusek, J, 1984:357) Potapov, M, Huang, C-W, Gulgenova, A & Luan, Y-X, 2020:19

Species travei Deharveng, 1981

Species trichaetosa Martynova, 1977

Species vtorovi Martynova, 1977


Genus Misturasotoma Bernard, EC et Christiansen, KA, 2010:58

Species brucealla (Wray, 1953) Bernard, EC & Christiansen, KA, 2010:58, t.t.

            Syn.: Isotoma brucei Wray, 1953, nec Carpenter, 1907


Misturasotoma brucealla from the USA
After Bernard EC & Christiansen KA, 2010 Fig.1


Genus Vertagopus Börner, C, 1906:171 key

  Syn.: Podurides(sic) Podura Bourlet, C, 1845:115
  Syn.: ... Parafolsomia auctt. partim nec Salmon, 1949


Vertagopus? sp. from the Arctic region
After White A in Sutherland PC, 1852 Fig.8
Vertagopus sp. from Germany
2006 © Kohl, F.
Vertagopus sp. from the USA
2006.12.13 © Cresswell, S.
Vertagopus sp. from the UK
2007.11.10 © Valentine., B
Vertagopus arboreus from the UK
Legs with clavate tenent setae
2015.12.29 © Cunningham, T.
Vertagopus arboreus from the UK
Legs with clavate tenent setae (*)
2018.03.10 © Cann, A.J.
Vertagopus arboreus from Belgium
Knobbed tenent macrosetae (*)
2020.03.05 © Huskens, M.-L.

Species sp.1 Fjellberg, A, 1996:100

Species sp.2 Fjellberg, A, 1996:100

Species sp.3 Janssens F & Duncan WL, 2019

Vertagopus sp. from the USA
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
Relative short antennae
2018.12.24 © Duncan, W.L.
Vertagopus sp. from the USA
Dense cover of short setae
Blue iridescent cuticular wax layer
2018.12.24 © Duncan, W.L.

Species abeloosi Poinsot, 1965

Species alpinus Haybach, 1972

Vertagopus alpinus from Austria
After Schulz H-J, 2010 Abb.4
Vertagopus alpinus from Austria
Left eyepatch : G and H smaller
Modified after Schulz H-J, 2010 Abb.3

Species alpus Christiansen & Bellinger, 1980

Species arboratus Huang, 1991

             Syn.: Vertagopus arborata(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:128 ISS
             Syn.: Vertagopus arborata(sic) Gao, Y, Xiong, Y et Potapov, MB, 2009:59 ISS
             Syn.: Vertagopus arborata(sic) Potapov, M, Gulgenova, A et Babykina, M, 2016:119 ISS

Species arboreus (Linnæus, C, 1758:609) Ågren, 1903, B

            Syn.: Podura nemotrosa Linnæus, C, 1748:67 i
            Syn.: Podura nigra pedibus albis Hill, 1752 i
            Syn.: Podura annularis de Fourcroy, AF, 1785:524
            Syn.: Podura oblonga nigra de Geer, C, 1778:17
            Syn.: Po