http://www.collembola.org/taxa/katiidae.htm - Last updated on 2017.10.31 by Frans Janssens
Checklist of the Collembola: Katiannidae

Familia Katiannidae Börner, 1913 sensu Bretfeld G, 1999:13 key
Genus Number
of species
Arborianna1
Betschurinus1
Cretokatianna1
Dalianus1
Gen.nov.12
Gen.nov.21
Gisinianus1
Katianna52
Katiannellina1
Katiannina2
Keratopygos1
Neokatianna1
Papirinus4
Parakatianna10
Polykatianna12
Pseudokatianna12
Rusekianna4
Sminthurinus95
Stenognathellus4
Vesicephalus7
Zebulonia1
Katiannidae214
Genera with number of species
Two new Katiannidae from the UK
2009.06.06 © Ardron, P.A.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.


Genus Cretokatianna Sánchez-García, A et Engel, MS, 2016:7, -
Species bucculenta Sánchez-García, A et Engel, MS, 2016:7, -
Cretokatianna bucculenta in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.4

            Syn.: Fasciosminthurus? sp. Simón-Benito, Ortuño & Espantaleón, 2002

Tribus Katiannini Börner, 1913

Genus Gen.nov.1 Greenslade P in Ardron, PA, 2009:16
Genus nov.1 sp. nov. from the UK
Third antennal segment with papilla and long fine setae
2009.04.01 © Ardron, P.
Det. Greenslade, P.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
Third tibiotarsus distally with spatulate setae (*)
2015.11.23 © Phillips, E.

Species sp.nov.1 Greenslade P in Ardron, PA, 2009:16
Genus nov.1 sp. nov. from the UK
After Ardron PA, 2009 p.16

  Syn.: Gen.nov.1 sp.nov.1 f. nigra Janssens F & Phillips E, 2016
Remark by Janssens, F. 2017.02.02: Dark form. Anterior middorsal pale narrow stripe. Dark 6th abdominal segment.

Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2016.12.14 © Phillips, E.
Genus nov.1 sp. nov. f. nigra from the UK
Form with dark abd.6
2017.02.01 © Barton, T.

"Sphyrotheca multifasciata" from the UK
2007.03.04 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.20 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.20 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Polykatianna" sp. ♀ from the UK
Sub-anal appendage
2009.03.27 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. ♂ from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Polykatianna" sp. from the UK
2009.04.01 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov. from the UK
Mucrones
2009.05.20 © Janssens, F.
Genus nov.1 sp. nov. ♀ from the UK
Female sub-anal appendage
2009.05.20 © Janssens, F.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.21 © Barton, T.
Genus nov.1 sp. nov. ♀ from the UK
Sub-anal appendage
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov. from the UK
Preparing for jump step 1 : lifting prolegs
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
Spermatophore
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
Preparing for a jump
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
Consumed spermatophore
2009.12.12 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.17 © Jones, A.
Genus nov.1 sp. nov. from the UK
2009.12.20 © Jones, A.
Genus nov.1 sp. nov. from the UK
Egg with polar caps
2009.12.20 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.24 © Barton, T.
Remark by Janssens, F. 2010.02.14: Adult form. Thoracic middorsal pale narrow stripe. Pale 6th abdominal tergum.

Genus nov.1 sp. nov. ♂ from the UK
Furca
2015.03.29 © Phillips, E.
Genus nov.1 sp. nov. from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov. from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
Abd.5 with 2 distinct bothriothricha
Abd.6 without pigment
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2012.12.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
Eyes
2012.02.16 © Nurcombe, E.
Genus nov.1 sp. nov. from the UK
2014.12.14 © Garlick, T.
Genus nov.1 sp. nov. from the UK
2014.12.18 © Jeffery, J.
Genus nov.1 sp. nov. from the UK
2014.12.21 © Robertson, A.
Genus nov.1 sp. nov. from the UK
2015.12.09 © Boardman, P.
Genus nov.1 sp. nov. from the UK
Footcomplex with knobbed tenent setae (*)
2016.12.01 © Phillips, E.
Genus nov.1 sp. nov. from the UK
Bothriotricha D (*)
2016.12.01 © Phillips, E.
Genus nov.1 sp. nov. ♀ from the UK
Sub-anal appendage (SAA)
2017.01.04 © Phillips, E.
Genus nov.1 sp. nov. ♀ from the UK
Curved sub-anal appendage (SAA) (*)
2017.01.29 © David W.
Genus nov.1 sp. nov. ♀ from the UK
Footcomplex with knobbed tenent setae
2017.02.04 © David W.
Genus nov.1 sp. nov. ♀ from the UK
Curved sub-anal appendage (SAA) (*)
2017.02.04 © David W.

Species sp.nov.2 Janssens F & Phillips E, 2017
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov. from the UK
Preparing for jump step 2 : everting collophore vesicles
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
Everting collophore vesicles before jumping
2016.01.14 © Horton, M.
Genus nov.1 sp. nov. from the UK
Backwards extended furca
2016.01.14 © Horton, M.

  Forma juvenile1 Janssens F & Barton T, 2009
"Parakatianna" sp.from the UK
2009.02.23 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov. from the UK
2009.12.11 © Barton, T.
Genus nov.1 sp. nov. from the UK
2010.03.09 © Barton, T.
Genus nov.1 sp. nov. from the UK
2013.10.13 © Nurcombe, E.

  Forma juvenile2 Janssens F & Cornwall N, 2007
"Sphyrotheca multifasciata" juv. from the UK
2007.03.18 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Katianna" sp. from the UK
2009.03.06 © Barton, T.
"Polykatianna" sp. from the UK
2009.04.07 © Shaw, P.
"Parakatianna" sp. from the UK
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov.2 from the UK
2009.10.25 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.11 © Barton, T.

  Forma subadult Janssens F & Cornwall N, 2007
"Sphyrotheca multifasciata" from the UK
2007.03.04 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.02.23 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
2009.03.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.02 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2014.01.21 © Horton, M.
Genus nov.1 sp. nov.2 from the UK
Papilla on 3rd antennal segment (*)
2014.01.21 © Horton, M.
Genus nov.1 sp. nov.2 from the UK
Feeding instar with dark gut contents
2014.01.21 © Horton, M.

  Forma adult
Remark by Janssens, F. 2010.02.14: Adult form. Thoracic and abdominal middorsal pale or orange broad area. Pigmented (dark or reddish) 6th abdominal tergum with 2 pale spots.

"Sphyrotheca multifasciata" from the UK
2007.03.18 © Cornwall, N.J.
"Sphyrotheca multifasciata" from the UK
2008.01.20 © Spears, G.
"Parakatianna" sp. from the UK
2009.03.06 © Barton, T.
"Parakatianna" sp. from the UK
Form with middorsal pale winged pattern
2009.04.21 © Ardron, P.
"Parakatianna" sp. from the UK
Form with middorsal pale winged pattern
2009.04.21 © Ardron, P.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.15 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.21 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov.2 pale & dark form from the UK
2009.11.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
Grooming bubble blowing
2009.12.11 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.12 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.15 © Jones, A.
Genus nov.1 sp. nov.2 from the UK
2009.12.20 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2009.12.28 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.01.24 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.06 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2010.02.13 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
With everted collophore vesicles, ready to take-off
2010.02.21 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2012.01.06 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2012.01.27 © Nurcombe, E.
Genus nov.1 sp. nov.2 ♂ from the UK
Raised genital papilla
2012.01.30 © Nurcombe, E.
Genus nov.1 sp. nov.2 from Ireland
2012.02.19 © Cluskey, D.
Genus nov.1 sp. nov.2 from Ireland
2013.02.02 © Cluskey, D.
Genus nov.1 sp. nov.2 from the UK
Composed eye with 8 ocelli: B,C,H large, A,E,F,G smaller, D smallest
2013.02.07 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2013.12.17 © Barton, T.
Genus nov.1 sp. nov.2 from the UK
2014.11.26 © Nurcombe, E.
Genus nov.1 sp. nov.2 from the UK
2014.12.14 © Garlick, T.
Genus nov.1 sp. nov.2 ♂ from the UK
2015.12.09 © Boardman, P.
Genus nov.1 sp. nov.2 ♂ from the UK
Raised genital papilla (*)
2016.03.11 © Phillips, E.
Genus nov.1 sp. nov.2 ♀ from the UK
Curved sub-anal appendage on basal papilla
2017.02.09 © Phillips, E.
Genus nov.1 sp. nov.2 ♀ from the UK
Broad sub-anal appendage on basal papilla
2017.02.09 © Phillips, E.


Genus Gen.nov.2 Greenslade P, 2012:il
Species sp.nov. Greenslade P, 2012:il
            Syn.: Katianna sp.3 Janssens F et Ardron PA in Ardron, PA, 2009:15
Katianna sp.3 from the UK
After Ardron PA, 2009 p.15

            Syn.: Sphyrotheca multifasciata Ardron, PA, 2009:15
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Spines on head

  Forma A
Remark by Janssens, F. 2017.02.08: Form with pale flanks and middorsal stripe.
Genus nov.2 sp. nov. from the UK
2007 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.12.05 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.10 © Barton, T.
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Genus nov.2 sp. nov. from the UK
2008.08.16 © Ardron, P.A.
Genus nov.2 sp. nov. from the UK
2009.04.21 © Ardron, P.A.
Genus nov.2 sp. nov. from Tasmania
2009.08.16 © Daley, A.
Genus nov.2 sp. nov. from the UK
2013.03.21 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. ♂ from the UK
2014.11.29 © Nurcombe, E.
Genus nov.2 sp. nov. from France
2017.02.05 © Picard, J.
Genus nov.2 sp. nov. from the UK
2017.02.02 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.02 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.17 © David W.
Genus nov.2 sp. nov. from the UK
2017.02.17 © David W.

  Forma B female
Remark by Janssens, F. 2017.02.08: Form with dark flanks and middorsal stripe.
Genus nov.2 sp. nov. from the UK
2006 © Barton, T.
Genus nov.2 sp. nov. from the UK
2006 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.04 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.03.18 © Cornwall, N.J.
Genus nov.2 sp. nov. from the UK
2007.12.05 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.16 © Barton, T.
Genus nov.2 sp. nov. from the UK
2007.12.16 © Barton, T.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from Tasmania
2009.10.24 © Daley, A.
Genus nov.2 sp. nov. from the UK
2011.12.07 © Kent, J.J.
Genus nov.2 sp. nov. from the UK
2013.12.10 © Nurcombe, E.
Genus nov.2 sp. nov. from the UK
Eyebrow short and anteriorly pointed
2016.12.17 © David W.
Genus nov.2 sp. nov. from France
2016.12.23 © Picard, J.
Genus nov.2 sp. nov. from France
2016.12.23 © Picard, J.
Genus nov.2 sp. nov. from France
2016.12.24 © Picard, J.
Genus nov.2 sp. nov. ♀ from the UK
2017.02.17 © David W.
Genus nov.2 sp. nov. ♀ from the UK
Sub-anal appendage (saa) on papilla
2017.02.17 © David W.
Genus nov.2 sp. nov. ♀ from the UK
Bothriotrichum D
2017.02.17 © David W.
Genus nov.2 sp. nov. from France
2017.03.08 © Picard, J.
Genus nov.2 sp. nov. ♀ from France
Sub-anal appendages (*) on papilla
2017.03.11 © Picard, J.

  Forma flammea Ardron PA, 2009:16
  Syn.: Katianna sp.3 f. flammea Ardron, PA, 2009:16
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.06.06 © Ardron, P.A.
Genus nov.2 sp. nov. f. flammea from the UK
2009.07.24 © Barton, T.
Genus nov.2 sp. nov. f . flammea from the UK
2014.11.12 © Nurcombe, E.
Genus nov.2 sp. nov. f . flammea from the UK
2014.11.12 © Nurcombe, E.
Genus nov.2 sp. nov. f . flammea from the UK
2015.01.07 © Jeffery, J.
Genus nov.2 sp. nov. f . flammea from the UK
2017.10.21 © Cunningham, T.

  Forma obscura Janssens F & Jeffery J, 2015
Genus nov.2 sp. nov. f. obscura from the UK
2015.11.05 © Jeffery, J.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura? from Australia
2016.07.10 © Steve & Alison.
Genus nov.2 sp. nov. f. obscura from the UK
2016.12.03 © David W.
Genus nov.2 sp. nov. f. obscura from the UK
2016.12.03 © David W.
Genus nov.2 sp. nov. f. obscura from the UK
At least 3 tenent setae (*)
2016.12.03 © David W.


Genus Keratopygos Christiansen, KA et Pike, E, 2002:180, -
Species megalos (Christiansen, KA & Pike, E, 2002:181, -) Christiansen, KA & Pike, E, 2002:181, -, t.t.

Genus Arborianna Bretfeld, 2002
Species cupiubae Bretfeld, 2002, t.t.

Genus Parakatianna Womersley, H, 1932:24
  Syn.: KatiannaGroup Longkingia Salmon, JT, 1946:40
Species albirubrafrons Salmon, JT, 1943:6
  Subspecies niveanota Salmon, JT, 1943:7
             Syn.: Parakatianna albirubrafrons ssp. niveanoa(sic) Salmon, JT, 1943:plate1 ISS
Species anomala Womersley, 1933
Species cortica Salmon, JT, 1943:8
Parakatianna cortica from New Zealand
After Salmon JT, 1943 Pl.1 Fig.9

Species diversitata Salmon, JT, 1943:7
Parakatianna diversitata from New Zealand
After Salmon JT, 1943 Pl.1 Fig.4

  Subspecies viridis Salmon, JT, 1943:8
Parakatianna diversitata
subsp. viridis from New Zealand
After Salmon JT, 1943 Pl.1 Fig.11

Species hexagona Salmon, JT, 1941:414
Parakatianna hexagona from New Zealand
After Salmon JT, 1941 Pl.71 Fig.510

Species homerica (Salmon, JT, 1946:44) Betsch, 1980
Species prospina (Salmon, JT, 1946:40) Betsch, 1980, t.t.
Species salmoni Wise, 1964
Species spinata Womersley, H, 1932:25, t.t.
Species superba (Salmon, JT, 1946:42) Betsch, 1980
Longkingia superba from New Zealand
After Salmon JT, 1946 Fig.28


Genus Katianna Börner, C, 1906:182
  Syn.: ... Metakatiannidae Stach, 1956 partim
  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim
Katianna nr schotti from the UK
Spinelike setae on head
2008.05.06 © Ardron, P.A.
Katianna nr schotti from the UK
Peglike papilla on 3rd antennal segment
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? & ♀? from the UK
2008.12.09 © Ardron, P.A.
Katianna sp. from Australia
2009.08.29 © Hort, J.
Katianna sp. from Peru
2013.03.17 © Cuadros Barrenechea.M., B
Katianna sp. from Tasmania
2014.04.18 © Murray, A.
Katianna sp. from Tasmania
2016.01.11 © Murray, A.

Species sp.1 Delamare Deboutteville, C et Massoud, Z, 1963:241
Species sp.2 Greenslade, P et Ireson, JE, 1986:282
Katianna sp. from Australia
After Greenslade P & Ireson JE, 1986 Fig.2

Species sp.21 Ireson, JE, 1990:147
Katianna sp.1 from Tasmania
Yellow-green colour form
After Ireson JE, 1990 Fig.51
Katianna sp.1 from Tasmania
Striped colour form
After Ireson JE, 1990 Fig.52

Species sp.3 Janssens F & Ardron PA, 2008
             Syn.: Katianna schoetti(sic) Ardron, PA, 2009:14 ISS
Katianna schoetti(sic) from the UK
After Ardron PA, 2009 p.14

            Syn.: Katianna australis Ardron, PA, 2009:15
Katianna australis from the UK
After Ardron PA, 2009 p.15
Remark by Greenslade, P. 2013.11.19 il: However carefully you examine specimens from England and compare with published descriptions from Australia, you cannot determine species of Katianna with accuracy. The only desciptions of K. australis that exist were by Womersley in the, 1930s. His descriptions are notoriously brief and totally inadequate. We probably have over 50 undescribed species of Katianna in Australia and New Zealand, maybe more. Womersley's description of K. australis could apply to many of them.

Katianna nr schotti ♂? from the UK
2008.04.30 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.04.30 © Ardron, P.A.
Katianna nr schotti from the UK, mucrones
2008.04.30 © Ardron, P.A.
Katianna nr schotti from the UK
Dorsal macrosetae
2008.04.30 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2008.05.06 © Ardron, P.A.
Katianna nr schotti from the UK
Foot complex with tenent hairs
2008.05.06 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2008.12.09 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2008.12.11 © Ardron, P.A.
 
Katianna nr schotti ♀? from the UK
2008.12.19 © Ardron, P.A.
 
Katianna nr schotti ♂? & ♀? from the UK
2008.12.19 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.02.01 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
 
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
Grooming using everted collophore vesicle
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.05.06 © Ardron, P.A.
Katianna nr schotti ♂? & ♀? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♀? & ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♀? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.06.06 © Ardron, P.A.
Katianna nr schotti ♂? from the UK
2009.12.20 © Jones, A.
Katianna nr schotti ϓ';? from the UK
2011.12.04 © Jones, A.
Katianna nr schotti ♂? from the UK
2011.12.05 © Jones, A.
Katianna nr schotti ♂? from the UK
2011.12.05 © Jones, A.
Katianna nr schotti ♂? from the UK
2013.03.21 © Nurcombe, E.
Katianna nr schotti juv. from the UK
2013.03.21 © Nurcombe, E.
Katianna nr schotti ♀? from the UK
2013.04.13 © Henton-Jones, D.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2013.12.11 © Nurcombe, E.
Katianna nr schotti ♂? from the UK
2014.11.09 © Evans, M.
Katianna nr schotti ♂? from the UK
2014.11.09 © Evans, M.
Katianna nr schotti ♀? from the UK
2014.12.21 © Horton, M.
Katianna nr schotti ♀? from the UK
2014.12.21 © Horton, M.
Katianna nr schotti ♀? from the UK
2015.03.03 © Cunningham, T.
Katianna nr schotti ♂? from Holland
2015.11.18 © van Duinen, J.
Katianna nr schotti ♀? from Holland
2015.12.18 © van Duinen, J.
Katianna nr schotti ♀? from Holland
2015.12.18 © van Duinen, J.

Species sp.4 Janssens F et Ardron PA in Ardron, PA, 2009:16
Katianna sp.4 from the UK
After Ardron PA, 2009 p.16
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
Female subanal appendage
2009.04.27 © Ardron, P.A.
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from the UK
2009.06.06 © Ardron, P.A.
Katianna sp. from the UK
Dorsal pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Lateral pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Face pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Posterior dorsal pigmentation
2012.11.24 © Bird, S.
Katianna sp. from the UK
Footcomplex of 3rd leg
2012.11.24 © Bird, S.
Katianna sp. from the UK
Mucro
2012.11.24 © Bird, S.

Species sp.5 Ardron, PA, 2009:19
Unknown sp.1 from the UK
After Ardron PA, 2009 p.19

Species sp.6 Ardron, PA, 2009:19
Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Katianna s.l. sp. from the UK
2008.mm.dd © Ardron, P.
Sminthurinus nr elegans var. dorsalis from the UK
After Ardron PA, 2009 p.18
Unknown sp.2 from the UK
After Ardron PA, 2009 p.19

Species sp.8 Janssens F & Bouffler E, 2014
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.
Katianna sp. from Australia
2014.07.03 © Bouffler, E.

Species sp.9 Janssens F & Ardron PA, 2015
Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
2015.04.26 © Ardron, P.A.
Katianna sp. from the UK
Caudal aspect
2015.04.26 © Ardron, P.A.

Species sp.10 Janssens F & Dobbs B, 2016
Katianna sp. from the USA
2016.01.02 © Justis, S.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
Antenna
2016.11.24 © Dobbs, Br.
Katianna sp. from the USA
Ant.3 with wart (*)
2016.11.25 © Dobbs, Br.
Katianna sp. from the USA
Dorsal pigmentation
2016.11.25 © Dobbs, Br.
Katianna sp. from the USA
2017.01.01 © Hart, J.
Katianna sp. from the USA
2017.01.01 © Hart, J.

Species antennalis Rapoport, 1962
Species antennapartitia Salmon, JT, 1941:412
             Syn.: Katianna antennapartita(sic) Salmon, JT, 1943:4 ISS
Species aurantiaca (Womersley, H, 1935:218) Greenslade, P in Houston, WWK, 1994:124
            Syn.: Arrhopalites aurantiaca(sic) Womersley, H, 1935:218 IOS
             Syn.: Arrhopalites aurantiaca(sic) Womersley, H, 1939:9225 key) ISS
             Syn.: Arrhopalites aurantiaca(sic) Stach, J, 1945:6,is ISS
             Syn.: Arrhopalites aurantiaca(sic) Stach, 1956 ISS
             Syn.: Arrhopalites aurantiaca(sic) Salmon, 1964 ISS
Species australis Womersley, H, 1931:5
Katianna australis from Tasmania
After Womersley H, 1931 Pl.I Fig.1
Katianna australis from Australia
After Womersley H, 1939 Pl.1 Fig.5
Katianna australis from Australia
After Greenslade P & Ireson JE, 1986 Fig.6
Katianna australis from Tasmania
After Ireson JE, 1990 Fig.44
Katianna australis from Tasmania
After Ireson JE, 1990 Fig.45

             Syn.: Katianna australia(sic) Ireson, JE, 1990:83 ISS
             Syn.: Katianna australia(sic) Ireson, JE, 1990:147 ISS
Katianna australis from Australia
After Thurmer J in Greenslade P, 1994 p.117

  Subspecies tillyardi Womersley, H, 1932:23
  Subspecies viridula Womersley, H, 1932:23
Species banzarei Salmon, JT, 1964:314
             Syn.: Katianna banzare(sic) Greenslade P, 1990 ISS
Katianna banzarei from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.81.
Katianna banzarei from Australia
2010 © Stevens, M.

Species cardoni (Delamare Deboutteville & Massoud, 1963:237) Rapoport, 1968
            Syn.: Katianna Cardoni Delamare Deboutteville & Massoud, 1963 IOS
Species cobold Börner, 1907
            Syn.: Sminthurus luteus Parona, 1895
            Syn.: Sminthurus viridis Parona, 1895
            Syn.: Katianna cobold var. decora Börner, 1907
            Syn.: Katianna cobold var. guttulata Börner, 1907
Species coeruleocephala Handschin, 1920
Species drummondi (Womersley, H, 1932:18) Greenslade, P in Houston, WWK, 1994:125
Species gloriosa Salmon, JT, 1946:38
Species houssayi Delamare Deboutteville & Massoud, 1963:225
            Syn.: Katianna Houssayi(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species jeanneli Delamare Deboutteville & Massoud, 1963:222
Species kerguelenensis Denis, 1947
Remark by Deharveng, L. 1981 (cfr Greenslade, P., 2007.10.09:in litt.): This species probably belongs in the genus Pseudokatianna because it lacks the generic character of spines on the head according to the figures.

             Syn.: Katianna kerguelensis(sic) Greenslade, P, 1997-:www0312 ISS
Remark by Greenslade, P. 2007.10.09:in litt.: Taking into consideration the form of the mucro, the chaetotaxy of the dens, the slight antennal annulation and other features, the species probably is not a Pseudokatianna but may belong in Polykatianna, especially as Denis noted it had conspicuous interocular bosses. However, neither of these two genera are well defined, so the correct generic placement for this species is not possible to determine at present.

Species kuscheli Delamare Deboutteville & Massoud, 1963:228
            Syn.: Katianna Kuscheli(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species liliputana (Nicolet, H, 1847:360) Stach, J, 1956:239, i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Smynthurus(sic) liliputanus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.9
             Syn.: Smynthurus Liliputanus(sic) Lubbock, J, 1873:116 ISS
Species maryae Bernard, EC, 2014:149
Katianna sp. from the USA
2006 © Bernard, E.C.
Katianna maryae from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1E
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1A
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1B
Katianna maryae ♀ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.1C
Katianna maryae ♂ from the USA
Scale bar = 500 µm
After Bernard EC, 2014 Fig.4A

Species mnemosyne Börner, C, 1906:182, t.t.
            Syn.: Sminthurus multifasciatus Parona, 1895
Species mucina Womersley, 1933
Katianna mucina from Australia
After Greenslade P & Ireson JE, 1986 Fig.3
Katianna mucina from Tasmania
After Ireson JE, 1990 Fig.46

Species nunezi (Najt, 1967) Betsch, 1977
Species obscura (Womersley, H, 1936:485) Greenslade, P & Ireson, JE, 1986:275
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7a
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7b
Parakatianna obscura from Australia
After Womersley H, 1936 Fig.7c
Katianna sp. cf. obscura from Australia
After Greenslade P & Ireson JE, 1986 Fig.5
Katianna cf. obscura from Tasmania
After Ireson JE, 1990 Fig.47

Species oceanica Schött, H, 1917:57
Katianna oceanica from Australia
After Schött H, 1917 Taf.4 Fig.6
Katianna oceanica from Australia
Abdomen posterior part
After Schött H, 1917 Taf.4 Fig.7

Species ornata Womersley, H, 1932:23
Katianna ornata from Tasmania
After Ireson JE, 1990 Fig.48

Species patagonica (Delamare Deboutteville, C & Massoud, Z, 1963:198) Betsch, 1977
Remark by Mari Mutt J.A. & Bellinger P.F. 1990:138: The genus Metakatianna houses a number of species based on first instars of species of Katianna, Sminthurinus, Pseudokatianna, etc. (Betch, 1980:12). We have listed the species of Metakatianna under Katianna but the position of many is still undetermined.
Species perplexa Salmon, JT, 1944:176
Species pescotti Womersley, H, 1935:218
Katianna cf. pescotti from Tasmania
After Ireson JE, 1990 Fig.50
Katianna pescotti from Tasmania
2014.04.28 © Murray, A.
Katianna pescotti from Tasmania
Body entirely yellow; strong macrosetae on head and body
2015.09.26 © Tony.
Katianna pescotti from Tasmania
Antennae bluish
2015.09.26 © Tony.
Katianna pescotti from Tasmania
Ant.3 with peg (*); ant.4 with 12 subdivisions
2015.09.26 © Tony.

Species poivrei Delamare Deboutteville, C et Massoud, Z, 1963:233
            Syn.: Katianna Poivrei(sic) Delamare Deboutteville, C et Massoud, Z, 1963:233 IOS
Species puella (Denis, 1933) Betsch, 1977
             Syn.: Metakatianna peulla(sic) Salmon, JT, 1964:589 ISS
             Syn.: Katianna paella(sic) Palacios-Vargas, 1992 ISS
Species purpuravirida Salmon, JT, 1941:413
Katianna purpuravirida from New Zealand
After Salmon JT, 1941 Pl.71 Fig.502
             Syn.: Katianna purpurvirida(sic) Salmon, JT, 1946:38 ISS
Species reducta Salmon, JT, 1944:176
Species richardsi Najt, 1967
Species ruberoculata Salmon, JT, 1944:175
Species schotti Womersley, 1933
            Syn.: Katianna oceanica var. schötti(sic) Womersley, 1933 IOS
            Syn.: Katianna oceanica King, Hutchinson & Greenslade P, 1976
            Syn.: Katianna oceanica Greenslade P, 1981
             Syn.: Katianna oceania(sic) var. schötti(sic) Womersley, H, 1935:218 ISS
             Syn.: Katianna oceanica var. schötti(sic) Womersley, H, 1939:235 ISS
             Syn.: Katianna schoetti(sic) King, Greenslade P & Hutchinson, 1985 ISS
             Syn.: Katianna oceanica var. schoetti(sic) Greenslade, P et Ireson, JE, 1986:275 ISS
Katianna oceanica var. schoetti(sic) from Australia
After Greenslade P & Ireson JE, 1986 Fig.4

             Syn.: Katianna oceanica var. schoetti(sic) Greenslade & Greenslade, 1987 ISS
             Syn.: Katianna oceanica var. schoetti(sic) Ireson, JE, 1990:83 ISS
Katianna oceanica var. schoetti(sic) from Tasmania
After Ireson JE, 1990 Fig.49

             Syn.: Katianna oceanica var. schoetti(sic) Ireson, 1993 ISS
             Syn.: Katianna schoetti(sic) Greenslade, P in Houston, WWK, 1994:126 ISS
Species serrae Najt, 1967
Species serrata (Schäffer, 1897) Stach, 1956
Species serrata Womersley, 1936, nec Schäffer, 1897
Species steparia (Najt, 1967) Mari Mutt & Bellinger, 1990:140
Species uschuaiensis (Schäffer, 1897) Stach, 1956
Species viretorum Najt, 1967
Species viridis Delamare Deboutteville & Massoud, 1963:237
Species willincki (Delamare Deboutteville & Massoud, 1963) Betsch, 1980
Species wygodzinskyi Delamare Deboutteville & Massoud, 1963:228
            Syn.: Katianna Wygodzinskyi(sic) Delamare Deboutteville & Massoud, 1963 IOS
Katianna nr wygodzinskyi from Peru
2009.01-03.dd © Contreras Valdez, A.

Species zebra (Womersley, H, 1942:30) Greenslade, P in Houston, WWK, 1994:126
Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7A
Parakatianna zebra from Australia
After Womersley H, 1942 Fig.7B


Genus Polykatianna Salmon, JT, 1946:54
  Syn.: .. Millsurus Betsch, 1977
Species aurea (Womersley, H, 1932:26) Bruzzese, 1980
Species cremea Salmon, JT, 1949:55
Species davidi (Tillyard, 1920) Greenslade P & Wise, 1986
            Syn.: Metakatianna gressitti Salmon, JT, 1964:317
            Syn.: Parakatianna litorea Salmon, JT, 1943:5
Polykatianna litorea ssp. litorea from New Zealand
2014 © Te Papa Museum

            Syn.: Parakatianna litorea ssp. luteaterga Salmon, JT, 1943:6
Parakatianna litorea ssp. luteaterga from New Zealand
After Salmon, J.T., 1943 Pl.1 Fig.6
Polykatianna litorea ssp. luteaterga from New Zealand
2014 © Te Papa Museum

Polykatianna davidi from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.78.

Species flammea Salmon, JT, 1946:55, t.t.
Species intermedia (Snider, 1978) Mendoza Arviso, S, Villalobos, FJ, Ruíz Montoya, L & Castro, AER, 1999:90
Species montana (Womersley, H, 1936:483) Greenslade P in Houston, 1994:128
Parakatianna montana from Australia
After Womersley H, 1936 Fig.6a

Species polygonia Snider, 1978
Species pulchella (Womersley, 1933) Greenslade P in Houston, 1994:128
Species quinseta (Salmon, 1954) Stach, 1956
Species radicula (Maynard, EA, 1951:224) Arnett, 1970
Sminthurinus radiculus from the USA
After Maynard EA, 1951 Plate XXIII Fig.378

            Syn.: Sminthurinus radiculus f. pallens Maynard, EA, 1951:225
Sminthurinus radiculus f. pallens from the USA
After Maynard EA, 1951 Plate XXIII Fig.380

            Syn.: Sminthurinus radiculus f. pictus Maynard, EA, 1951:224
Sminthurinus radiculus f. pictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.379

Polykatianna nr radicula from Canada
With aberrant reddish eyepatches
2013.01.17 © Kyron, T.F.
Polykatianna nr radicula from Canada
With aberrant reddish eyepatches
2013.01.17 © Kyron, T.F.
Polykatianna nr radicula ♀ from Canada
Facial chaetotaxy
2013.01.21 © Kyron, T.F.
Polykatianna nr radicula ♀ from Canada
Female palmate sub-anal appendages (s.a.a.)
2013.01.21 © Kyron, T.F.
Polykatianna nr radicula ♀ from Canada
Anogenital abdominal segments
2013.01.21 © Kyron, T.F.
Polykatianna nr radicula ♀ from Canada
Ocelli C & H subequal in size but distinctly different in colour
2013.01.21 © Kyron, T.F.
Polykatianna nr radicula ♀ from Canada
Fourth antennal segment subdivided
2013.01.21 © Kyron, T.F.

Species sminthurinus (Mills, HB, 1934:99) Richards, 1968
Polykatianna sminthurinus from the USA
2013.02.11 © Anderson, J.
Polykatianna sminthurinus from the USA
2013.02.11 © Anderson, J.
?Polykatianna sminthurinus? from Germany
Ant.4 subdivided; ant.3 with warth (*)
2014.10.26 © Landgraf, A.
Polykatianna sminthurinus from the USA
2014.12.23 © Montgomery, G.

Species utahensis (Snider, RJ, 1988:1)
Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.1
Sminthurinus (Polykatianna) utahensis from the USA
After Snider RJ, 1988 Fig.2


Genus Sminthurinus Börner, C, 1901:99
  Syn.: Sminthuridæ(sic) Sminthurus Schött, H, 1891:7 partim
  Syn.: Smynthuri(sic) Smynthurus(sic) Folsom, JW, 1896:448 partim ISS
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
  Syn.: Inartuculati Eusminthurus Börner partim
  Syn.: Sminthuridinae Smynthurella Houlbert, C, 1924:157
  Syn.: Sminthuridae Metakatianna Gisin, H, 1960:277
  Syn.: Sminthuridae Stenognathellus Gisin, H, 1960:277
  Syn.: Katiannini Stenognathellus Richards, WR, 1968:23
  Syn.: Katiannini Smithurinus(sic) Stach, J, 1956:96 ISS
Remark by ICZN. 2007:759: Placed on the Official List of Names in Zoology. Opinion 1027.


  Syn.: Collembola Simthurinus(sic) Chauvat, M, Trap, J, Perez, G, Delporte, P et Aubert, M, 2011:412 ISS
Sminthurinus sp. juv. from France
2011.08.29 © Lebeaux, P.
Sminthurinus henshawi from the USA
4th antennal segment not subdivided
2011.12.03 © Justis, S.
Sminthurinus sp. from the UK
Facial chaetotaxy
2012.11.09 © Murray, A.


  Syn.: Symphypleona Smynthurinus(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:73 ISS
Sminthurinus sp. from the UK
Ventral aspect: furca and everted vesicles of collophore (*)
2013.11.07 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
Footcomplex with clavate tenant setae (*)
2014.02.06 © Phillips, E.
Sminthurinus aureus from the UK
Ant.4 not subsegmented
2016.02.23 © Phillips, E.
Sminthurinus sp. from the USA
2016.03.09 © Abela, A.

Species sp.1 Bretfeld, 2000
Species sp.2 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species sp.3 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166
Species sp.4 Hornung-Leoni, CT et Palacios-Vargas, JG, 2012:67
Species sp.5 Janssens F & Phillips E, 2017
Sminthurinus sp. ♀ from the UK
2017.10.09 © Phillips, E.
Sminthurinus sp. ♀ from the UK
Caudal aspect
2017.10.09 © Phillips, E.
Sminthurinus sp. juv. from the UK
2017.10.09 © Phillips, E.
Sminthurinus sp. juv. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.25 © Phillips, E.
Sminthurinus sp. from the UK
2017.10.28 © Lewis, M.
Sminthurinus sp. from the UK
2017.10.28 © Lewis, M.

Species albiceps (Schött, 1896) Stach, 1956, s.i.
Species albipes (Schött, 1896) Stach, 1956, i.s.
Species alpinus Gisin, 1953
Sminthurinus alpinus from Corsica
After Schulz H-J Abb.7

  Subspecies bisetosus Ellis, 1976
Species antennalis (Rapoport, 1962) Delamare Deboutteville & Massoud, 1963:213
Species atrapallidus Snider, 1978
Sminthurinus atrapallidus from the USA
2006.02.25 © Schimming, L.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2009.02.07 © Justis, S.
Sminthurinus atrapallidus from the USA
2012.02.15 © Kouri, J.
Sminthurinus atrapallidus from the USA
2012.02.15 © Kouri, J.
Sminthurinus atrapallidus from the USA
Egg cluster
2012.04.09 © Zungri, D.
Sminthurinus atrapallidus from the USA
2016.12.23 © Dobbs, Br.
Sminthurinus atrapallidus from the USA
2017.01.25 © Vitanza, S.
Sminthurinus nr atrapallidus from the USA
2017.04.05 © Johnson, S.T.
Sminthurinus nr atrapallidus from the USA
Pale form
2017.04.05 © Johnson, S.T.

Species aureus (Lubbock, 1862) Ågren, 1903, B
            Syn.: Sminthurus ochraceus Koch, 1840
            Syn.: Smynthurus(sic) signifer Fitch, 1862
            Syn.: Sminthurus quadrilineatus Tullberg, 1871
            Syn.: Sminthurus quadrilineatus var. ochropus Reuter, 1890
Sminthurinus aureus f. ochropus from the UK
2009.04.05 © Ardron, P.A.


            Syn.: Sminthurinus aureus Ardron, PA, 2009:19
Sminthurinus near aureus from the UK
After Ardron PA, 2009 p.19

Sminthurinus aureus f. ochropus from the UK
2011.11.28 © Kent, J.J.
Sminthurinus aureus f. ochropus from the UK
2011.11.30 © Nurcombe, E.
Sminthurinus aureus f. ochropus from Holland
2012.03.02 © van Duinen, J.
Sminthurinus aureus f. ochropus from the UK
2012.04.06 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.04.06 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.06.18 © Murray, A.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.07.05 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2012.10.02 © Murray, A.
Sminthurinus aureus f. ochropus from the UK
2012.10.02 © Murray, A.
Sminthurinus aureus f. ochropus from the UK
2012.12.18 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2012.12.18 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2013.01.07 © Nurcombe, E.
Sminthurinus aureus f. ochropus from the UK
2013.10.29 © Phillips, E.
Sminthurinus aureus f. ochropus from the UK
2014.08.15 © Nurcombe, E.
Sminthurinus aureus f. ochropus from Germany
2014.10.05 © Jäschke, J.
Sminthurinus aureus f. ochropus from the UK
2014.11.22 © Phillips, E.
Sminthurinus aureus f. ochropus juv. from Germany
2015.01.08 © Mattew, M.
Sminthurinus aureus f. ochropus from the UK
2016.11.12 © Phillips, E.
Sminthurinus aureus f. ochropus from Belgium
2016.12.18 © Huskens, M.-L.


            Syn.: Sminthurus aureus var. fusca Krausbauer, 1898
            Syn.: Sminthurus albus Schäffer, 1900
            Syn.: Sminthurinus aureus var. atrata(sic) Börner, C, 1901:104 IOS
            Syn.: Sminthurus aureus var. maculata(sic) Krausbauer, 1901 IOS
             Syn.: Sminthurinus aureus var. maculata(sic) Börner, C, 1901:138 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Linnaniemi, 1907 ISS
             Syn.: Sminthurus aureus var. maculata(sic) Linnaniemi, WM, 1912:285 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Linnaniemi, WM, 1912:285 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Denis, 1922 ISS
             Syn.: Sminthurinus aureus f. maculata(sic) Handschin, E in Dahl, F, 1929:114 ISS
             Syn.: Sminthurinus aureus var. maculata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurus aureus ssp. maculata(sic) Salmon, JT, 1964:578 = atratus ISS
             Syn.: Sminthurinus aureus ssp. maculata(sic) Salmon, JT, 1964:578 = atratus ISS
             Syn.: Sminthurinus aureus f. maculata(sic) Stach, J, 1964:58 ISS
Sminthurinus aureus f. maculatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus aureus f. maculatus from France
2008.03.24 © Lebeaux, P.
Sminthurinus aureus f. maculatus from the UK
2012.07.22 © Murray, A.
Sminthurinus aureus f. maculatus from the UK
2012.07.22 © Murray, A.
Sminthurinus aureus f. maculatus from Holland
2012.11.12 © van Duinen, J.
Sminthurinus aureus f. maculatus from the UK
2012.12.21 © Phillips, E.
Sminthurinus aureus f. maculatus from Holland
2013.10.23 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
2013.10.23 © van Duinen, J.
Sminthurinus aureus f. maculatus from the UK
Freeing itself from a silk fiber
2014.01.17 © Phillips, E.
Sminthurinus aureus f. maculatus from Holland
Abdominal bothriotricha (A-D)
2014.10.09 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
2014.10.09 © van Duinen, J.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Germany
2014.11.01 © Kluck, A.
Sminthurinus aureus f. maculatus from Holland
2014.11.17 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
Colour form inbetween f. maculatus and f. reticulatus
2014.11.21 © van Duinen, J.
Sminthurinus aureus f. maculatus from Holland
Colour form inbetween f. maculatus and f. reticulatus
2014.11.21 © van Duinen, J.
Sminthurinus aureus f. maculatus from Belgium
2016.12.09 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
2017.02.04 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
2017.02.04 © Huskens, M.-L.
Sminthurinus aureus f. maculatus from Belgium
2017.02.04 © Huskens, M.-L.


            Syn.: Sminthurinus aureus var. punctata Börner, C, 1901:103
            Syn.: ... pustulata Krausbauer, 1902
            Syn.: Sminthurinus aureus var. punctiformis Latzel, R, 1917:249
            Syn.: ... anglicana Salmon, 1955
            Syn.: Sminthurinus niger Tuxen, 1944
            Syn.: Sminthurinus igniceps da Gama, 1959
            Syn.: Sminthurinus aureus f. reticulatus Janssens F & Phillips E, 2013
Sminthurinus aureus f. reticulatus from the UK
2012.11.19 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2012.12.03 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2012.12.04 © Phillips, E.
Sminthurinus aureus f. reticulatus from Switzerland
2012.12.06 © Murray, A.
Sminthurinus aureus f. reticulatus from the UK
2012.12.21 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2013.01.28 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2013.02.12 © Phillips, E.
Sminthurinus aureus f. reticulatus juv. from the UK
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus juv. from the UK
Mottled appearance: ready to moult
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus ♂ from the UK
Raised genital papilla (*)
2013.11.30 © Nurcombe, E.
Sminthurinus aureus f. reticulatus from the UK
2014.01.17 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
2014.01.25 © Phillips, E.
Sminthurinus aureus f. reticulatus from the UK
Feeding instar
2014.02.06 © Phillips, E.
Sminthurinus aureus f. reticulatus from Holland
2014.10.31 © van Duinen, J.
Sminthurinus aureus f. reticulatus from Holland
2014.11.23 © van Duinen, J.
Sminthurinus aureus f. reticulatus from Belgium
2016.12.09 © Huskens, M.-L.
Sminthurinus aureus f. reticulatus juv. from Belgium
2016.12.20 © Huskens, M.-L.


            Syn.: Sminthurinus aureus f. apigmentatus Janssens F & Nurcombe E, 2013
Sminthurinus aureus f. apigmentatus ♂ from the UK
Form without black pigment having red eyes
2013.11.24 © Nurcombe, E.
Sminthurinus aureus f. apigmentatus ♂ from the UK
Form without black pigment having red eyes
2013.11.24 © Nurcombe, E.

Smynthurus(sic) aureus from the UK
After Lubbock J, 1873 Pl.7
Sminthurus aureus from the UK
After Lubbock J in Collinge WE & Shoebotham JW, 1909 Pl.III Fig.2
Sminthurinus aureus from the UK
Ventral subapical dental setae absent
2004 © Hopkin, S.P.
Sminthurinus aureus
2006 © Boldsystems.org
 
Sminthurinus aureus from France
2007.03.17 © Lebeaux, P.
Sminthurinus aureus from the Netherlands
2009.01.26 © Heemskerk, R.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.dd © Lebeaux, P.
Sminthurinus aureus from France
2009.09.16 © Lebeaux, P.
Sminthurinus aureus from Czechia
2010.11.25 © Fiala, M.
Sminthurinus aureus from Germany
2011.05.04 © Marquard, F.
Sminthurinus aureus from France
2011.11.13 © Lebeaux, P.
Sminthurinus aureus from France
2011.11.13 © Lebeaux, P.
Sminthurinus aureus from France
2011.11.14 © Lebeaux, P.
Sminthurinus aureus (bar=2µm)
Hexagonal epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.6C
Sminthurinus aureus from the UK
2012.01.06 © Jones, A.
Sminthurinus aureus from the UK
2012.01.06 © Jones, A.
Sminthurinus aureus from the UK
2012.01.29 © Nurcombe, E.
Sminthurinus aureus juv. from the UK
2012.01.29 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.04.10 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.06.14 © Murray, A.
Sminthurinus aureus from the UK
2012.02.16 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.06.18 © Murray, A.
Sminthurinus aureus juv. from the UK
2012.08.01 © Murray, A.
Sminthurinus aureus from the UK
Lateral abdominal bothriotricha
2012.10.26 © Nurcombe, E.
Sminthurinus aureus from the UK
4th antennal segment not subdivided
2012.10.26 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.10.29 © Jones, A.
Sminthurinus aureus from the UK
2012.11.15 © Kent, J.J.
Sminthurinus aureus from the UK
2012.11.25 © Nurcombe, E.
Sminthurinus aureus from the UK
2012.11.26 © Kent, J.J.
Sminthurinus aureus from the UK
2012.11.26 © Phillips, E.
Sminthurinus aureus from Switzerland
2012.12.06 © Murray, A.
Sminthurinus aureus from the UK
2012.12.07 © Phillips, E.
Sminthurinus aureus-group s.l. from the UK
Forma aureus (1,2,4,6,8,10), maculatus (12), reticulatus (3,5,7,9,11)
2012.12.24 © Phillips, E.
Sminthurinus aureus from Holland
2013.02.16 © van Duinen, J.
Sminthurinus aureus from the UK
2013.04.30 © Murray, A.
Sminthurinus aureus from the UK
2013.04.30 © Murray, A.
Sminthurinus aureus from the UK
Dark spots mark positions of 4 abdominal bothriotricha (A-D)
2013.10.08 © Phillips, E.
Sminthurinus aureus from France
2014.01.04 © Sandra.
Sminthurinus aureus juv. from the UK
2014.01.08 © Phillips, E.
Sminthurinus aureus from Germany
2014.11.15 © Kluck, A.
Sminthurinus aureus from Germany
Walking in a row
2015.01.07 © Landgraf, A.
Sminthurinus aureus juv. from Belgium
Distinctly separate thoracic segments
2016.01.26 © Huskens, M.-L.
Sminthurinus aureus from the UK
Bothriotricha A-B-C-D
2016.03.10 © Phillips, E.

  Subspecies purpureus Salmon, JT, 1941:410
Species aureussetosus Bretfeld, 2002
Species australis Najt, 1973
Species bimaculatus Axelson, 1902
            Syn.: Sminthurinus aureus f. fenestratus Handschin, E in Dahl, F, 1929:114
            Syn.: Sminthurus biflavopunctatus Linnaniemi, 1912
            Syn.: Sminthurinus aureus f. biflavopunctata Handschin, E in Dahl, F, 1929:114
            Syn.: Sminthurinus biflavopunctatus Womersley, 1930
            Syn.: Sminthurinus biflavopunctatus Davies, 1934
            Syn.: Sminthurinus biflavopunctatus Kloet & Hincks, 1934
            Syn.: Sminthurinus aureus f. biflavopunctatus Uchida, 1957
            Syn.: Sminthurus biflavopunctatus Hopkin, 2000
            Syn.: Sminthurinus aureus var. circumfasciata Stach, 1922
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from France
2005 © Losacco, S.
Sminthurinus bimaculatus from the UK
2008.06.06 © Barton, T.
Sminthurinus bimaculatus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus bimaculatus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus bimaculatus from the UK
2012.07.15 © Murray, A.
Sminthurinus bimaculatus juv. from the UK
2012.07.15 © Murray, A.

Species bisetosus Ellis, 1976
Species bullai Najt, 1973
Species cantonensis Rusek, 1971
Species castagninoi Najt, 1969
Species cingula Bagnall, 1932, s.d.
Species cingulatus Bagnall, 1921
Species conchyliatus Snider, 1978
Species concolor (Meinert, 1896) Stach, 1956
Sminthurinus concolor from the UK
Two setae on retinaculum
2004 © Hopkin, S.P.
Sminthurinus concolor from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.5
Sminthurinus concolor from Norway
2011 © Fjellberg, A.

Species discordipes Salmon, JT, 1949:49
Species dispar Babenko, AB, 2007:375
Species distinctus Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:240
Sminthurinus distinctus from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.4A

Species domesticus Gisin, 1963
            Syn.: Sminthurinus niger da Gama, 1959 partim
            Syn.: Sminthurinus niger da Gama, 1961 partim
            Syn.: Sminthurinus niger Gisin, 1963
            Syn.: Sminthurinus domesticus f. apigmentata Janssens F & van Duinen J, 2012
Sminthurinus domesticus f. apigmentata from Holland
Form without black pigment having red eyes
2012.07.11 © van Duinen, J.
Sminthurinus domesticus f. apigmentata from Holland
Form without black pigment having red eyes
2013.08.18 © van Duinen, J.
Sminthurinus domesticus f. apigmentata from Holland
Form without black pigment having red eyes
2013.09.03 © van Duinen, J.
Sminthurinus domesticus f. apigmentata from Holland
Form without black pigment having red eyes
2013.09.03 © van Duinen, J.

Sminthurinus domesticus from the UK
Two subapical ventral dental setae
2004 © Hopkin, S.P.
Sminthurinus domesticus from the UK, juvenile
2007.09.29 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.13 © Valentine.
 , B
Sminthurinus domesticus from the UK
2007.10.13 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.18 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.24 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.25 © Valentine., B
Sminthurinus domesticus from the UK
2007.10.25 © Valentine., B
Sminthurinus domesticus from the UK
2007.11.15 © Valentine., B
Sminthurinus domesticus from the UK
2008.08.27 © Valentine., B
Sminthurinus domesticus from the UK
2008.10.16 © Valentine., B
Sminthurinus domesticus from the UK
2008.10.16 © Valentine., B
Sminthurinus domesticus from the UK
2008.11.18 © Valentine., B
Sminthurinus domesticus from the UK
2008.11.18 © Valentine., B
Sminthurinus domesticus from the UK
2008.12.05 © Valentine., B
Sminthurinus domesticus from the UK
2008.12.05 © Valentine., B
Sminthurinus domesticus from the UK
After Ardron PA, 2009 p.18
Sminthurinus domesticus from Portugal
2009.04.21 © Marcelino, J.
Sminthurinus domesticus from the UK
2009.11.07 © Valentine., B
Sminthurinus domesticus from the UK
2009.11.07 © Valentine., B
Sminthurinus domesticus from the UK
2010.01.19 © Jones, A.
Sminthurinus domesticus from Holland
2011.09.11 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus domesticus from Holland
2011.11.08 © van Duinen, J.
Sminthurinus domesticus from Holland
2012.07.11 © van Duinen, J.
Sminthurinus domesticus from Holland
Pale colour form
2012.07.11 © van Duinen, J.
Sminthurinus domesticus from the UK
2012.08.30 © Murray, A.
Sminthurinus domesticus from the UK
Face
2012.08.30 © Murray, A.
Sminthurinus domesticus from the UK
Pale colour form
2012.08.30 © Murray, A.
Sminthurinus domesticus early instars from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus juv. from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus from Portugal
2012.10.11 © Murray, A.
Sminthurinus domesticus from the UK
2013.01.04 © Murray, A.
Sminthurinus domesticus from the UK
Asymmetric abberation in frontocellar area
2013.01.04 © Murray, A.
Sminthurinus domesticus from Morocco
2014.04.21 © Beck, K.
Sminthurinus domesticus juv. from Morocco
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.28 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.29 © Beck, K.
Sminthurinus domesticus from Morocco
2014.04.30 © Beck, K.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus juv. from Germany
2016.02.06 © Gutekunst, V.
Sminthurinus domesticus from Germany
2017.01.28 © Melzer, L.

Species duplicatus Salmon, JT, 1941:411
Species elegans (Fitch, 1862) Linnaniemi, WM, 1912:283, B
            Syn.: Smynthurus(sic) signifer Fitch, 1862
            Syn.: Sminthurus quadrilineatus Tullberg, 1871, partim
Sminthurus quadrilineatus from Sweden
After Schött H, 1893 Taf.I Fig.18

            Syn.: Sminthurinus aureus var. ornata Börner, C, 1901:104
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.520
Sminthurinus elegans f. ornata from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornata from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornata from the UK
2008.12.09 © Valentine., B
Sminthurinus elegans f. ornata from the USA
2009.11.25 © Gledhill, L.
Sminthurinus elegans f. ornata juv. from the UK
2012.06.10 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.11 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.11 © Murray, A.
Sminthurinus elegans f. ornata juv. from the UK
2012.06.13 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.06.14 © Murray, A.
Sminthurinus elegans f. ornata from the UK
Grooming
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornata from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornata from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans f. ornata from the UK
2012.08.17 © Nurcombe, E.
Sminthurinus elegans f. ornata from Portugal
2012.10.16 © Murray, A.
Sminthurinus elegans f. ornata from Portugal
2012.10.16 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2012.10.29 © Jones, A.
Sminthurinus elegans f. ornata from the UK
Dorsal aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornata from the UK
Facial aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornata from the UK
Abdomen lateral aspect
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornata from the UK
6th abdominal segment
2012.11.06 © Murray, A.
Sminthurinus elegans f. ornata from the UK
2013.09.18 © Nurcombe, E.
Sminthurinus elegans f. ornata from the UK
2013.11.02 © Phillips, E.
Sminthurinus elegans f. ornata from the UK
Note spermatophore of Isotomidae
2013.11.12 © Phillips, E.
Sminthurinus elegans f. ornata from the UK
2014.08.21 © Phillips, E.
Sminthurinus elegans f. ornata from the UK
2014.08.21 © Phillips, E.
Sminthurinus elegans f. ornata from Holland
2014.11.23 © van Duinen, J.
Sminthurinus elegans f. ornata from the UK
2016.01.21 © Phillips, E.
Sminthurinus elegans f. ornata from the UK
2016.10.26 © Phillips, E.


            Syn.: Sminthurinus aureus var. dorsalis Axelson, WM, 1905:794
            Syn.: Sminthurinus elegans f. cancellus Maynard, EA, 1951:238
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.522
Sminthurinus elegans f. cancellus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.523

             Syn.: Sminthurinus cancellatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurinus elegans f. cancellus from the USA
2012.04.18 © Kouri, J.
Sminthurinus elegans f. cancellus from the USA
2012.04.18 © Kouri, J.


            Syn.: Sminthurinus elegans f. ochropus Maynard, EA, 1951:238
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.524
Sminthurinus elegans f. ochropus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.525
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.
Sminthurinus elegans f. ochropus from the USA
2009.02.08 © Justis, S.
Sminthurinus elegans f. ochropus from the USA
2017.01.28 © Abela, A.


            Syn.: Sminthurinus similitortus Maynard, EA, 1951:232
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.413
 

            Syn.: Sminthurinus aureus ssp. transversalis Salmon, JT, 1964:579
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXIV Fig.424
Sminthurinus elegans from Australia
After Greenslade P & Ireson JE, 1986 Fig.2
Sminthurinus elegans
Rare form with lateral white spots
2004 © Baquero, E.
Sminthurinus elegans from the UK
Female sub-anal appendages
2004 © Hopkin, S.P.
Sminthurinus elegans from the USA
2006 © McClarin, J.
Sminthurinus elegans from the USA
2006 © McClarin, J.
Sminthurinus elegans from Sweden
2006 © Fjellberg, A.
Sminthurinus elegans from the USA
2008.05.16 © Thombs, D.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.02.14 © Justis, S.
Sminthurinus elegans from the USA
2009.05.21 © Cowen, R.
Sminthurinus elegans from the USA
2010.01.05 © Justis, S.
Sminthurinus elegans from the UK
2011.12.31 © Jones, A.
Sminthurinus elegans from the USA
2012.01.29 © Kouri, J.
Sminthurinus elegans from the USA
2012.02.24 © Mallory, C.
Sminthurinus elegans from the USA
Very dark specimen
2012.02.24 © Mallory, C.
Sminthurinus elegans juv. from the UK
2012.06.10 © Murray, A.
Sminthurinus elegans from the UK
2012.08.08 © Nurcombe, E.
Sminthurinus elegans from the UK
2012.08.16 © Nurcombe, E.
Sminthurinus elegans from Iran
After Falahati Hossein Abad A & al, 2013 Fig.4
Sminthurinus elegans from the USA
2013.04.04 © Kouri, J.D.
Sminthurinus elegans from the UK
Ant.3 with wart (*)
2013.08.25 © Henton-Jones, D.
Sminthurinus elegans from the UK
2013.09.18 © Nurcombe, E.
Sminthurinus elegans from the UK
2013.10.06 © Phillips, E.
Sminthurinus elegans from Germany
2014.05.05 © Beck, K.
Sminthurinus elegans juv. from the UK
2014.08.28 © Phillips, E.
Sminthurinus elegans juv. from Holland
Ready to moult
2014.11.22 © van Duinen, J.
Sminthurinus elegans from the USA
2017.01.26 © Dobbs, Br.
Sminthurinus elegans from Belgium
Very dark colour form
2017.01.31 © Huskens, M.-L.
Sminthurinus elegans juv. from Belgium
2017.10.17 © Huskens, M.-L.
Sminthurinus elegans from Holland
2017.10.17 © Wieringa, P.

Species exiguus (Nicolet, H, 1847:360)
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species gamae Gisin, 1963
            Syn.: Sminthurinus igniceps da Gama, 1959 cf
Sminthurinus gamae from Portugal
After Gisin H, 1963 in Bretfeld, 1999 Fig.273

Species gisini da Gama, 1965
            Syn.: Sminthurinus carpathicus Rusek, 1966

Sminthurinus glaucus from New Zealand
After Salmon, J.T. 1943 Pl.1 Fig.8.
Species glaucus Salmon, JT, 1943:2

Species gloriosus (Womersley, H, 1937:157) Greenslade & Mayer, 1993 cf
Arrhopalites gloriosa from Australia
After Womersley H, 1937 Fig.1m

             Syn.: Sminthurinus gloriosa(sic) Greenslade & Mayer, 1993 cf ISS
             Syn.: Sminthurinus gloriosa(sic) Greenslade, P in Houston, WWK, 1994:131 ISS
Species granulatus Salmon, JT, 1946:34
Species granulosus Enderlein, G, 1908:409
Species handschini Folsom, 1934
Species henshawi (Folsom, JW, 1896:449) Maynard, EA, 1951:233
            Syn.: Smynthurus(sic) henshawii(sic) Folsom, JW, 1896:449 IOS
            Syn.: Sminthurinus aureus Folsom, 1898
            Syn.: Sminthurus aureus Guthrie, JE, 1903:43
            Syn.: Smynthurus(sic) aureus Mills, 1930
            Syn.: Sminthurus aureus Maynard, 1931
            Syn.: Sminthurinus aureus James, 1933
            Syn.: Sminthurinus aureus Mills, HB, 1934:95
            Syn.: Sminthurinus aureus Folsom, 1934
            Syn.: Sminthurinus aureus Brimley, 1938
            Syn.: Sminthurinus aureus Maynard, EA, 1951:233
Sminthurinus aureus from the USA
After Maynard EA, 1951 Plate XXIII Fig.410
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.03.10 © Cowen, R.
Sminthurinus henshawi f. aureus from the USA
2009.11.21 © Gledhill, L.
Sminthurinus henshawi f. aureus from the USA
2011.11.19 © Justis, S.
Sminthurinus henshawi f. aureus from the USA
2012.03.03 © Montgomery, G.
Sminthurinus henshawi f. aureus from the USA
2013.04.14 © Uhey, D.
Sminthurinus henshawi f. aureus? from Holland
2015.02.03 © van Duinen, J.
Sminthurinus henshawi f. aureus? from Holland
2015.02.03 © van Duinen, J.
Sminthurinus henshawi f. aureus from the USA
2015.04.17 © Dankowicz, E.
Sminthurinus henshawi f. aureus from the USA
Feeding instar with dark gut
2015.11.20 © Schrayer, G.


            Syn.: Sminthurinus aureus var. novus Mills, HB, 1934:96
            Syn.: Sminthurinus megoculatus Maynard, EA, 1951:233
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.542
Sminthurinus megoculatus from the USA
After Maynard EA, 1951 Plate XXIX Fig.544
Sminthurinus henshawi f. megoculatus from the USA
2007.03.26 © Reed, D.E.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.
Sminthurinus henshawi f. megoculatus from the USA
2008.01.26 © Montgomery, G.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. megoculatus from the USA
2012.01.29 © Kouri, J.


            Syn.: Sminthurinus similitortus Maynard, EA, 1951:232
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIV Fig.414
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.537
Sminthurinus similitortus from the USA
After Maynard EA, 1951 Plate XXIX Fig.538
Sminthurinus elegans from the USA
After Maynard EA, 1951 Plate XXVIII Fig.521
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.05 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.02.07 © Justis, S.
Det. Snider, R.J.
Sminthurinus henshawi f. similitortus from the USA
2009.04.08 © Justis, S.
Sminthurinus henshawi f. similitortus from the USA
2011.01.21 © Gerber, G.K.
Sminthurinus henshawi f. similitortus from the USA
2011.01.21 © Gerber, G.K.
Sminthurinus henshawi f. similitortus from the USA
2011.01.21 © Gerber, G.K.
Sminthurinus henshawi f. similitortus from the USA
2011.12.24 © Justis, S.
Sminthurinus henshawi f. similitortus from the USA
2011.12.24 © Justis, S.
Sminthurinus henshawi f. similitortus juv. from Canada
2012.01.13 © Hochhalter, R.
Sminthurinus henshawi f. similitortus juv. from Canada
2012.01.13 © Hochhalter, R.
Sminthurinus henshawi f. similitortus juv. from Canada
2012.01.13 © Hochhalter, R.
Sminthurinus henshawi f. similitortus juv. from Canada
2012.01.13 © Hochhalter, R.
Sminthurinus henshawi f. similitortus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. similitortus from the USA
2012.01.29 © Kouri, J.
Sminthurinus henshawi f. similitortus from the USA
2012.02.05 © Kouri, J.

             Syn.: Sminthurinus henshawii(sic) Stach, 1956 ISS
Sminthurinus (Sminthurinus) henshawi from the USA
After Snider RJ in Christiansen KA & Bellinger PF, 1998 Fig.914A
Sminthurinus henshawi from the USA
2006 © McClarin, J.
Sminthurinus henshawi from the USA
2009.11.25 © Gledhill, L.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2011.12.03 © Justis, S.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2011.12.03 © Justis, S.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2011.12.03 © Justis, S.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2012.01.07 © Justis, S.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2012.01.07 © Justis, S.
Sminthurinus nr henshawi from the USA
Forma "bimaculata"?
2012.01.07 © Justis, S.
Sminthurinus henshawi from Canada
Feeding instar; with phoretic nematode
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from Canada
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from Canada
2013.01.15 © Kyron, T.F.
Sminthurinus henshawi from the USA
2013.04.14 © Uhey, D.
Sminthurinus henshawi juv. from Canada
2014.01.23 © Hochhalter, R.
Sminthurinus henshawi from the USA
2014.04.20 © Dankowicz, E.

Species hygrophilus Bretfeld, 2000
             Syn.: Sminthurinus hydrophilus(sic) Babenko, AB, 2007:377 ISS
Species igniceps (Reuter, 1878) Börner, C, 1901:102
            Syn.: Sminthurus rex Uzel, 1891
Sminthurus igniceps from Sweden
After Schött H, 1893 Taf.II Fig.17
Sminthurinus igniceps ♀ from Poland
After Stach J, 1956 Plate XV Fig.1
Sminthurinus igniceps ♂ from Poland
After Stach J, 1956 Plate XV Fig.2
Sminthurinus igniceps from the UK
2012.06.13 © Murray, A.
Sminthurinus igniceps from the UK
2012.06.13 © Murray, A.
Sminthurinus igniceps ♀ from the UK
2013.09.12 © Phillips, E.
Sminthurinus igniceps ♀ from the UK
2013.09.12 © Phillips, E.
Sminthurinus igniceps from the UK
2013.10.04 © Phillips, E.
Sminthurinus igniceps from the UK
Pale form
2013.10.04 © Phillips, E.
Sminthurinus igniceps ♀ from the UK
2013.10.08 © Phillips, E.
Sminthurinus igniceps from the UK
2013.10.08 © Phillips, E.
Sminthurinus igniceps from the UK
2013.10.14 © Phillips, E.
Sminthurinus igniceps ♀ from the UK
2013.10.29 © Phillips, E.
Sminthurinus igniceps juv. from the UK
2013.11.25 © Phillips, E.
Sminthurinus igniceps from the UK
2013.12.19 © Barton, T.
Sminthurinus igniceps from the UK
2013.12.19 © Barton, T.
Sminthurinus igniceps from the UK
2014.08.14 © Phillips, E.
Sminthurinus igniceps from the UK
Head more pale than trunk
2014.08.22 © Phillips, E.
Sminthurinus igniceps from the UK
2014.11.04 © Phillips, E.
Sminthurinus igniceps from the UK
2014.11.22 © Phillips, E.
Sminthurinus igniceps from the UK
2014.11.22 © Phillips, E.
Sminthurinus igniceps from the UK
2014.11.29 © Phillips, E.
Sminthurinus igniceps ♀ from the UK
2015.07.27 © Boardman, P.

Species imperialis Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:60
Sminthurinus imperialis from Mexico
After Palacios-Vargas JG & al, 2012 Fig.1A
             Syn.: Sminthurinus Imperialis(sic) Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:60 ISS
Species inexcussus Najt & Rapoport, 1965
Species insularis Najt, 1973
Species jonesi Wise, KAJ, 1970:203
Species kaha Christiansen, K et Bellinger, P, 1992:349
Species kerguelensis Salmon, JT, 1964:314
Species latimaculosus Maynard, EA, 1951:231
Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.389
Sminthurinus latimaculosus from the USA
After Maynard EA, 1951 Plate XXIII Fig.390

             Syn.: Sminthurinus latimaculatus(sic) Zeppelini, 1996 ISS
             Syn.: Sminthurinus latimaculosos(sic) Palacios-Vargas, JG et Castaño-Meneses, G in Basset, Y, Novotny, V, Miller, SE et Kitching, RL, 2003:169 ISS
Sminthurinus latimaculosus from the USA
2012.06.15 © Anderson, J.

Species lawrencei Gisin, 1963
Species lichenatus Salmon, JT, 1943:3
Species lineatus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species maculosus Snider, 1978
             Syn.: Sminthurinus maculosos(sic) Palacios-Vargas, JG, Hornung-Leoni, CT et Garrido, I, 2012:61 ISS
Species mime Börner in Voeltzkow, 1907
            Syn.: Sminthurus niger Parona, 1895
            Syn.: Sminthurinus terrestris Womersley, 1931
Species minutus (MacGillivray, AD, 1894:109) Maynard, EA, 1951:234
Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXIII Fig.402
Sminthurinus minutus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.535
Sminthurinus minutus from the USA
2006.12.02 © Murray, T.
Sminthurinus minutus from the USA
2007.01.09 © Montgomery, G.
Sminthurinus minutus from the USA
2008.02.09 © Montgomery, G.
Sminthurinus minutus from the USA
2009.04.11 © Zungri, D.
Sminthurinus minutus from the USA
2009.04.11 © Zungri, D.
Sminthurinus minutus from the USA
2009.12.29 © Zungri, D.
Sminthurinus minutus from the USA
2009.12.31 © Zungri, D.
Sminthurinus minutus from the USA
2010.01.01 © Zungri, D.
Sminthurinus minutus juv. from the USA
2010.01.24 © Zungri, D.
Sminthurinus minutus from the USA
2010.02.07 © Zungri, D.
Sminthurinus minutus from the USA
2010.03.23 © Montgomery, G.
Sminthurinus minutus from the USA
2010.03.23 © Montgomery, G.
Sminthurinus minutus from the UK
2014.11.24 © Nurcombe, E.
Sminthurinus minutus from the UK
2014.11.24 © Nurcombe, E.
Sminthurinus minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus minutus from the UK
2014.11.27 © Nurcombe, E.
Sminthurinus minutus from the UK
2015.01.04 © Nurcombe, E.
Sminthurinus minutus juv. from the UK
2015.01.04 © Nurcombe, E.
Sminthurinus minutus from the UK
2015.02.08 © Phillips, E.

Species modestus Yosii, 1970
Sminthurinus modestus from Japan
2014.01.13 © Komatsu, T.

Species molinai Arlé, 1940
Species mountfordi (Womersley, H, 1939:242) Greenslade P in Houston, 1994:132
Species muscophilus Salmon, JT, 1946:32
Species niger (Lubbock, 1862) Börner, C, 1901:101, B
            Syn.: Sminthurus bimaculatus Tullberg, 1871
            Syn.: ... concolor Meinert, 1896
            Syn.: ... niger f. ochracea Axelson, 1902
            Syn.: ... niger f. alpina Gisin, 1953
            Syn.: Sminthurinus niger var. Stach, 1956
Smynthurus(sic) niger from the UK
After Lubbock J, 1870 Pl.XXI Fig.11
Smynthurus(sic) niger from the UK
Saltatory appendage
After Lubbock J, 1870 Pl.XXI Fig.12

Smynthurus(sic) niger from the UK
After Lubbock J, 1873 Pl.6
Sminthurus niger from the USA
After Schött H, 1891 Taf.II Fig.1
Sminthurinus niger from France
2005 © Locasso, S.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus niger from the UK
2007.03.04 © Cornwall, N.J.
 
Sminthurinus niger from the UK
2007.03.18 © Cornwall, N.J.
Sminthurinus niger from the UK
2007.03.18 © Cornwall, N.J.
Sminthurinus niger from the UK
2007.12.22 © Barton, T.
Sminthurinus niger from the UK
2007.12.22 © Barton, T.
Sminthurinus niger from the UK
2008.01.10 © Valentine., B
Sminthurinus cf. niger from the USA
2008.01.21 © Moorehead, C.
Sminthurinus niger from the UK, female
2008.04.08 © Valentine., B
Sminthurinus niger from the UK, male
2008.04.08 © Valentine., B
Sminthurinus niger from the UK
2008.11.28 © Valentine., B
Sminthurinus niger from the UK
2008.12.25 © Corley, P.
Sminthurinus niger from the UK
2009.01.04 © Corley, P.
Sminthurinus niger from the UK
2009.03.10 © Valentine., B
Sminthurinus niger from the UK
2009.03.10 © Valentine., B
Sminthurinus nr niger from the USA
2009.04.08 © Zungri, D.
Sminthurinus niger from the UK
2009.04.12 © Barton, T.
Sminthurinus niger from the UK
2009.12.04 © Barton, T.
Sminthurinus niger from the UK
2009.12.06 © Barton, T.
Sminthurinus niger from the UK
2009.12.28 © Jones, A.
Sminthurinus nr niger from the USA
2010.02.07 © Zungri, D.
Sminthurinus niger from France
2011.08.29 © Lebeaux, P.
Sminthurinus niger from France
2011.08.29 © Lebeaux, P.
Sminthurinus niger from the UK
2011.12.11 © Nurcombe, E.
Sminthurinus niger from the UK
2011.12.11 © Nurcombe, E.
Sminthurinus niger from France
2012.01.15 © Lebeaux, P.
Sminthurinus niger from the UK
2012.08.21 © Murray, A.
Det.: Shaw, P.
Sminthurinus niger from the UK
2012.08.21 © Murray, A.
Det.: Shaw, P.
Sminthurinus niger juv. from the UK
2012.08.22 © Murray, A.
Det.: Shaw, P.
Sminthurinus niger from France
2012.11.25 © Sandra.
Sminthurinus niger from the UK
2012.12.07 © Nurcombe, E.
Sminthurinus niger from the UK
Wart on 3rd antennal segment
2012.12.13 © Valentine., B
Sminthurinus niger from the UK
Speckled specimen: ready to moult
2013.01.04 © Murray, A.
Sminthurinus niger juv. from the UK
2013.01.12 © Nurcombe, E.
Sminthurinus niger from Canada
Ant.3 with papilla (*)
2013.01.23 © Kyron, T.F.
Sminthurinus niger from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus niger from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus niger from Canada
2013.01.23 © Kyron, T.F.
Sminthurinus niger from Canada
Dens with 1 subapical ventral seta
2013.01.24 © Kyron, T.F.
Sminthurinus niger from Canada
Hind foot with 4 tenent setae
2013.01.24 © Kyron, T.F.
Sminthurinus niger ♀ from Canada
Female palmate sub-anal appendage (s.a.a) on papilla (*)
2013.01.24 © Kyron, T.F.
Sminthurinus niger ♀ from Canada
Circumanal setae: dorsal median bifid seta (*), setae with expanded base (**), s.a.a. (***)
2013.01.24 © Kyron, T.F.
Sminthurinus niger juv. from the UK
Speckled: preparing to moult
2013.02.06 © Murray, A.
Sminthurinus niger from the UK
Epicuticular ultrastructure
2013.02.06 © Murray, A.
Sminthurinus niger from the UK
Footcomplex with unguiculus (u), unguis (U) and clavate tenent macrosetae (*)
2013.12.09 © Phillips, E.
Sminthurinus niger from the UK
2013.12.10 © Phillips, E.
Sminthurinus niger from Morocco
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
Circumanal setae with expanded base (*)
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
Circumanal median seta bifid (*)
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
Secreting a drop of saliva
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
Picking up drop of saliva with proclaw
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
Cleaning antenna with drop of saliva
2014.04.28 © Beck, K.
Sminthurinus niger from Morocco
2014.04.30 © Beck, K.
Sminthurinus niger from the UK
Trapped in sticky spider web
2015.02.05 © Garlick, T.
Sminthurinus niger from the UK
2015.02.05 © Garlick, T.
Sminthurinus niger from the UK
Speckled specimen: ready to moult
2015.02.05 © Garlick, T.
Sminthurinus niger from the UK
Short white "eyebrows"
2015.02.15 © Phillips, E.
Sminthurinus niger juv. from the UK
Speckled specimen: ready to moult
2015.11.10 © Jeffery, J.
Sminthurinus niger from the UK
Genital abdominal segment not fused with great abdomen
2016.01.06 © Phillips, E.
Sminthurinus niger from Belgium
2016.02.02 © Huskens, M.-L.
Sminthurinus niger juv. from Belgium
Preparing to moult
2016.03.15 © Huskens, M.-L.
Sminthurinus niger ♀ from the UK
Abd.5 not fused with large abdomen
Sub-anal appendage (*)
2017.01.14 © David W.
Sminthurinus niger from France
Ant.3 with wart (*)
2017.10.15 © Picard, J.

Species nigrafuscus Salmon, JT, 1941:409
Species nigrescens Schött, H, 1917:54
Species nunezi Rapoport in Delamare Deboutteville & Massoud, 1963:210
            Syn.: Sminthurinus Nunezi(sic) Rapoport in Delamare Deboutteville & Massoud, 1963 IOS
Sminthurinus nunezi from Peru
2009.01-03.dd © Contreras Valdez, A.

Species obscurus Salmon, JT, 1944:173
Species oculatus Schött, H, 1917:53
Sminthurinus oculatus typus I from Australia
After Schött H, 1917 Taf.4 Fig.1
Sminthurinus oculatus typus I from Australia
After Schött H, 1917 Taf.4 Fig.2
Sminthurinus oculatus typus II from Australia
After Schött H, 1917 Taf.4 Fig.3
Sminthurinus oculatus typus III from Australia
After Schött H, 1917 Taf.4 Fig.4

Species oiskiyensis Bretfeld, G, 2010:305
Sminthurinus oiskyensis from Russia
After Bretfeld G, 2010 Fig.1

Species operosus Najt & Rapoport, 1965
Species orientalis Stach, J, 1964:21
Sminthurinus orientalis from China
After Stach J, 1964 Plate XI Fig.1
Sminthurinus orientalis from China
After Stach J, 1964 Plate XI Fig.2

Species ovatulus (Koch, CL & Berendt, GC in Berendt, GC, 1854:121) -
Smynthurus(sic) ovatulus from Baltic amber
After Koch CL & Berendt GC, 1845 Tab.XV Fig.132

Species packardi Denis, 1933
Species pallescens Yosii, 1970
Species pallidus Womersley, 1931
Species patagonicus Najt, 1971
Species pekinensis Stach, J, 1964:22
Sminthurinus pekinensis from China
After Stach J, 1964 Plate XII Fig.1
Sminthurinus pekinensis from China
After Stach J, 1964 Plate XII Fig.2

Species planasiensis Dallai, 1969
Species proceraseta Salmon, JT, 1946:32
Species pullus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species quadratus Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species quadrimaculatus (Ryder, 1878) Richards, WR, 1968:23
            Syn.: Smynthurus(sic) quadrimaculata(sic) Ryder, 1878 IOS
            Syn.: Smynthurus(sic) 6-maculata(sic) Harvey, 1892 IOS
            Syn.: Smynthurus(sic) sexmaculatus MacGillivray, AD, 1894:109
            Syn.: Smynthurus(sic) minutus MacGillivray, 1894
            Syn.: Smynthurus(sic) remotus Folsom, JW, 1896:448
            Syn.: Smynthurus(sic) niger Nearcticrecords
            Syn.: Sminthurus concolor Nearcticrecords
            Syn.: Sminthurinus quadrimaculatus f. bimaculatus Maynard, EA, 1951:230
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.490
Sminthurinus quadrimaculatus f. bimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.491

             Syn.: Sminthurinus quadriomaculatus(sic) Richards, WR, 1968:23 ISS
Sminthurinus quadrimaculatus from the USA
Forma bimaculatus
2003 © Schoenherr, J.
Sminthurinus quadrimaculatus from Canada
Forma bimaculatus
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus from Canada
Forma bimaculatus
2007.05.17 © Pilon, C.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2007.12.16 © Moorehead, C.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2008.06.03 © Schoenherr, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2008.06.04 © Schoenherr, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
Spotted: ready to moult
2008.09.08 © Murray, T.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2013.07.13 © Israel, D.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
2014.11.05 © Rosenfeld, J.
Sminthurinus quadrimaculatus f. bimaculatus from the USA
Spotted: ready to moult
2016.04.02 © Justis, S.


            Syn.: Sminthurinus minutus f. bipunctatus Maynard, EA, 1951:235
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.403
Sminthurinus minutus f. bipunctatus from the USA
After Maynard EA, 1951 Plate XXVIII Fig.536
Sminthurinus quadrimaculatus from the USA
Forma bipunctatus
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus from the USA
Forma bipunctatus
2008.02.07 © Gross, J.
Sminthurinus quadrimaculatus from the USA
Forma bipunctatus
2008.02.07 © Gross, J.


            Syn.: Sminthurinus brunneus Maynard, EA, 1951:226
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.400
Sminthurinus brunneus from the USA
After Maynard EA, 1951 Plate XXIII Fig.401


            Syn.: Sminthurinus downsi Maynard, EA, 1951:225
Sminthurinus downsi from the USA
After Maynard EA, 1951 Plate XXII Fig.377

             Syn.: Sminthurinus dowsi(sic) Stach, J, 1956:102 ISS

            Syn.: Sminthurinus quadrimaculatus f. latipictus Maynard, EA, 1951:231
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.391
Sminthurinus quadrimaculatus f. latipictus from the USA
After Maynard EA, 1951 Plate XXIII Fig.392
Sminthurinus quadrimaculatus from the USA
Pale form
2006 © McClarin, J.


            Syn.: Sminthurinus quadrimaculatus ssp. maynardi Snider, RJ, 1967:223
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.393
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXIII Fig.394
Sminthurinus quadrimaculatus from the USA
After Maynard EA, 1951 Plate XXVI Fig.489
             Syn.: Sminthurines(sic) cuadrimaculatus(sic) Palacios-Vargas, 1979 ISS
             Syn.: Sminthurus quadrimaaculatus(sic) Vázquez & Palacios-Vargas, 1987 ISS
Sminthurinus quadrimaculatus from the USA
2007 © McClarin, J.
 
Sminthurinus quadrimaculatus from the USA
First instar
2007.04.12 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
Second instar
2007.04.15 © Murray, T.
Sminthurinus quadrimaculatus from the USA
2007.10.23 © McClarin, J.
Sminthurinus quadrimaculatus from the USA
2008.10.04 © Murray, T.
Sminthurinus quadrimaculatus juv. from Canada
2016.08.28 © Levac, G.
Sminthurinus quadrimaculatus from the USA
2016.09.20 © Rosenfeld, J.

Species reticulatus Cassagnau, P, 1964:481
Sminthurinus reticulatus from France
After Cassagnau P, 1964
            Syn.: Sminthurinus aureus var. transversalis Hüther, 1961
            Syn.: Sminthurinus aureus Ellis, 1976
Sminthurinus reticulatus from Sweden
After Fjellberg A, 2006 ©
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from the UK
2006.11.01 © Brocklehurst, K.
Sminthurinus reticulatus from France
2007.09.08 © Lebeaux, P.
Sminthurinus reticulatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus reticulatus from France
2011.11.13 © Lebeaux, P.
Sminthurinus reticulatus from France
2011.11.13 © Lebeaux, P.
Sminthurinus reticulatus from the UK
2012.03.30 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.03.30 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.04.01 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.04.11 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.07.20 © Murray, A.
Sminthurinus reticulatus from the UK
2012.07.22 © Murray, A.
Sminthurinus reticulatus juv. from the UK
2012.07.22 © Murray, A.
Sminthurinus reticulatus from the UK
2012.09.13 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.09.26 © Nurcombe, E.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.09 © Shaw, P.
Sminthurinus reticulatus from the UK
2012.12.23 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.01.04 © Barton, T.
Sminthurinus reticulatus from the UK
2013.02.12 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.02.15 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.07.01 © Phillips, E.
Sminthurinus reticulatus from the UK
2013.11.04 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.01.10 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.01.25 © Phillips, E.
Sminthurinus reticulatus from the UK
2014.08.20 © Phillips, E.
Sminthurinus reticulatus juv. from Germany
2014.11.01 © Kluck, A.
Sminthurinus reticulatus from Holland
2015.10.09 © van Duinen, J.
Sminthurinus reticulatus from Belgium
2016.12.02 © Huskens, M.-L.
Sminthurinus reticulatus from Belgium
Grooming upside-down
2016.12.02 © Huskens, M.-L.
Sminthurinus reticulatus juv. from Belgium
2017.10.17 © Huskens, M.-L.

Species salti Delamare Deboutteville, 1953
Species sayi Denis, 1933
             Syn.: Sminthurinus Sayi(sic) Delamare Deboutteville & Massoud, 1963 ISS
Species signatus (Krausbauer, 1898 partim) Börner, 1900
            Syn.: Sminthurus aureus var. signata(sic) Krausbauer, 1898 partim IOS
             Syn.: Sminthurinus aureus var. signata(sic) Axelson, WM, 1905:794 ISS
             Syn.: Sminthurinus aureus var. signata(sic) Linnaniemi, WM, 1912:283 ISS
             Syn.: Sminthurinus aureus f. signata(sic) Handschin, E in Dahl, F, 1929:113 ISS
             Syn.: Sminthurinus aureus var. signata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus quadrilineata ab. signata(sic) Stach, J, 1956:122 ISS
             Syn.: Sminthurinus elegans var. signata(sic) Stach, J, 1956:127 ISS
             Syn.: Sminthurinus aureus var. signata(sic) Gisin, H, 1960:279 ISS
Sminthurinus signatus from Denmark
After Fjellberg A, 2007 Fig.13.4
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.
Sminthurinus signatus from France
2016.12.07 © Legrand, T.

Species signifer (Fitch, 1862)
Species speciosus Yosii, 1970
Species splendidus (Womersley, 1933) Greenslade, P in Houston, WWK, 1994:133
Parakatianna splendida from Australia
After Womersley H, 1939 Pl.1 Fig.4.
             Syn.: Sminthurinus splendida(sic) Greenslade, P in Houston, WWK, 1994:133 ISS
Species subalpinus Itoh, 2000
Species suborientalis Stach, J, 1965:367
Species timorensis Yoshii, R et Suhardjono, YR, 1992:92
Species transversalis Axelson, WM, 1905:794
            Syn.: Sminthurinus reticulatus Fjellberg, A, 2007:197 =?
Sminthurinus transversalis from Finland
Det. Axelson, W.M., 1902
2006 © Fjellberg, A.
Sminthurinus transversalis ♀ from Denmark
After Fjellberg A, 2007 Fig.13.6.left
Sminthurinus transversalis ♂ from Denmark
After Fjellberg A, 2007 Fig.13.6.right

             Syn.: Sminthurinus transvernalis(sic) Falahati HosseinAbad, A, Kheirodin, A et Bretfeld, G, 2013:208 ISS
             Syn.: Smynthurinus(sic) transvernalis(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:73 ISS
Species tricolor Schött, H, 1917:54
Sminthurinus tricolor from Australia
After Schött H, 1917 Taf.4 Fig.5

Species trinotatus Axelson, WM, 1905:794, B
            Syn.: Sminthurus biflavopunctatus Collinge, WE et Shoebotham, JW, 1909:9
Sminthurus biflavopunctatus from the UK
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.1

             Syn.: Sminthurinus bifavopunctatus(sic) Salmon, JT, 1964:582 ISS
            Syn.: Sminthurinus aureus var. bimaculata Carpenter H in Collinge WE & Shoebotham JW, 1909:10
            Syn.: Sminthurinus fenestratus Börner, C, 1909:124
            Syn.: Sminthurinus fenestratus Uchida, 1952
            Syn.: Sminthurinus fenestratus Yosii, R, 1977:164
            Syn.: Sminthurinus trinotatus f. incompleta Stach, J, 1965:367
Sminthurinus trinotatus from Germany
2006.12.26 © Kohl, F.
Sminthurinus trinotatus from Germany
2006.12.29 © Kohl, F.
Sminthurinus trinotatus from Belgium
2008.06.09 © Vuijlsteke, M.
Sminthurinus trinotatus from the UK
2008.07.22 © Shaw, P.
Sminthurinus trinotatus juv. from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus juv. from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from France
2008.12.07 © Jacquemin, G.
Sminthurinus trinotatus from the UK
After Ardron PA, 2009 p.17
Sminthurinus trinotatus from the UK
2009.09.01 © Thomason, R.
Sminthurinus trinotatus from the UK
2009.09.01 © Thomason, R.
Sminthurinus trinotatus from the UK
2010.10.23 © Jones, A.
Sminthurinus trinotatus from Holland
Distinct abdominal segments 5 & 6
2011.07.08 © van Duinen, J.
Sminthurinus trinotatus from Germany
2012.04.14 © Schmitt, R.
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
 
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
Sminthurinus trinotatus from the UK
2012.10.03 © Murray, A.
Sminthurinus trinotatus from the UK
2012.10.31 © Jones, A.
Sminthurinus trinotatus from Holland
2013.08.19 © van Duinen, J.
Sminthurinus trinotatus from Holland
2013.08.19 © van Duinen, J.
Sminthurinus trinotatus from the UK
2013.11.15 © Horton, M.
Sminthurinus trinotatus juv. from the UK
2013.12.11 © Nurcombe, E.
Sminthurinus trinotatus juv. from the UK
2013.12.11 © Nurcombe, E.
Sminthurinus trinotatus from Germany
2014.05.06 © Beck, K.
Sminthurinus trinotatus juv. from Germany
First instar
2014.05.06 © Beck, K.
Sminthurinus trinotatus from Japan
2014.05.15 © Hara, Y.
Sminthurinus trinotatus from Germany
2014.10.28 © Beck, K.
Sminthurinus trinotatus from Germany
Dorsal aspect mimicry
2014.11.02 © Beck, K.
Sminthurinus trinotatus juv. from Germany
2014.11.02 © Beck, K.
Sminthurinus trinotatus from Germany
2014.11.02 © Beck, K.
Sminthurinus trinotatus from the UK
2015.07.27 © Boardman, P.
Sminthurinus trinotatus from Germany
2015.09.01 © Jäschke, J.
Sminthurinus trinotatus from the UK
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Abdominal bothriotricha in red cups
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Caudal aspect
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Broad white "eyebrows"
2015.11.16 © Horton, M.
Sminthurinus trinotatus from the UK
Facial aspect
2015.11.16 © Horton, M.
Sminthurinus trinotatus from Belgium
2016.01.06 © Huskens, M.-L.
Sminthurinus trinotatus from Japan
2016.07.21 © Hara, Y.
Sminthurinus trinotatus from the UK
Caudal aspect
2017.09.25 © Phillips, E.
Sminthurinus trinotatus from Holland
2017.10.26 © Wieringa, P.

Species tuberculatus Delamare Deboutteville & Massoud, 1963:201
Species tucumanensis Delamare Deboutteville & Massoud, 1963:205
Species tunicatus Salmon, JT, 1954:216
Sminthurinus tunicatus from New Zealand
2009 © Te Papa Museum

Species victorius Nguyen, 2001
Species wakhanicus Yosii, 1966
Species woringeri (Delamare Deboutteville & Massoud, 1963:207) Delamare Deboutteville & Massoud, 1963
            Syn.: Sminthurinus Woringeri Delamare Deboutteville & Massoud, 1963 IOS

Genus Pseudokatianna Salmon, JT, 1944:179
  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim
Species campbellensis Salmon, JT, 1949:52
            Syn.: Pseudokatianna cambellensis(sic) Salmon, JT, 1949:52 IOS
Species fagophila Salmon, JT, 1946:50
Species fasciata (Salmon, JT, 1944:178)
Species livida (Salmon, JT, 1943:2) Salmon, JT, 1944:181
Species lutea Salmon, JT, 1946:51
Species minuta Salmon, JT, 1946:53
Species nigretalba Salmon, JT, 1944:179, t.t.
  Subspecies aurea Salmon, JT, 1944:180
Species niveovata Salmon, JT, 1946:48
Pseudokatianna niveovata from New Zealand
After Salmon JT, 1946 Fig.42

  Subspecies nigra Salmon, JT, 1946:50
Species triclavata Salmon, JT, 1949:53
Species triverrucata Salmon, JT, 1944:181
Species umbrosalata Salmon, JT, 1946:52
Species zebra Salmon, JT, 1946:47
Pseudokatianna zebra from New Zealand
After Salmon JT, 1946 Fig.36


Genus Stenognathellus Stach, 1956
  Syn.: Sminthuridinae Stenognathellus Cassagnau, P, 1953:226
Remark by Ellis, W.N. & Bellinger, P.F. 1973:54: Stenognathellus Cassagnau, 1953 is an unavailable name (art. 13b): type species not designated.

Species cassagnaui Yosii, 1966
Stenognathellus cassagnaui from the UAE
After Schulz H-J & van Harten A, 2013 Pl.4

Species denisi Cassagnau, P, 1953:227, t.t.
            Syn.: Stenognathellus Denisi(sic) Cassagnau, P, 1953:227,tt IOS
Species polygonalis Hüther, 1967
Species stenognathus (Börner, 1907) Mari Mutt, JA & Bellinger, PF, 1990:144
            Syn.: Sminthurinus stenognathus ssp. similis Najt & Rapoport, 1965

Genus Katiannellina Delamare Deboutteville, C et Massoud, Z, 1963:217
Species patagonica Delamare Deboutteville, C et Massoud, Z, 1963:217, t.t.

Genus Betschurinus Dallai, R et Martinozzi, I, 1980:31
Species farmae Dallai, R et Martinozzi, I, 1980:33, t.t.

Genus Dalianus Cassagnau, 1969
Species galae Cassagnau, 1969

Genus Katiannina Maynard EA et Downs WL in Maynard, EA, 1951:242
  Syn.: ... Deuterosminthurus Wray, 1948
  Syn.: ... Polyclavella Stach, 1955
  Syn.: Katiannini Metakatianna Richards, WR, 1968:22 partim
  Syn.: ... Metakatianna Denis, 1933 u
  Syn.: Bourletiellini Katiannia(sic) Snider, RJ, 1967:225 ISS
Katiannina sp. from the USA
2014.01.13 © Philonatura, N.

Species macgillivrayi (Banks, 1897) Maynard, EA, 1951:242, t.t.
            Syn.: Deuterosminthurus altamontus Wray, 1948
            Syn.: Deuterosminthurus batrachos Wray, 1948
            Syn.: Polykatianna batchi Snider, 1969
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXII Fig.377a
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.552
Katiannina macgillivrayi from the USA
After Maynard EA Plate XXIX Fig.553
Katiannina macgillivrayi from the USA
2006 © Bernard, E.C.
Katiannina macgillivrayi from the USA
2007.05.15 © Murray, T.
Katiannina macgillivrayi from the USA
2008.12.14 © Fleming, K.
Katiannina macgillivrayi from the USA
2009.05.08 © Loser, I.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi ♀ from the USA
2009.11.26 © Justis, S.
Katiannina macgillivrayi from the USA
2010.04.17 © Skvarla, M.
Katiannina macgillivrayi juv. from the USA
2011.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2012.02.20 © Fleming, K.
Katiannina macgillivrayi juv. from the USA
2012.03.27 © Eiseman, C.
Katiannina macgillivrayi from the USA
Red eyepatch
2012.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2012.04.21 © Eiseman, C.
Katiannina macgillivrayi from the USA
2015.01.19 © Fleming, K.

Species subalpina Itoh, 2000

Genus Neokatianna Snider, RJ, 1989:541
Species acantholaema Snider, RJ, 1989:542, t.t.

Genus Rusekianna Betsch, J-M, 1977:61
Rusekianna from Norway
2009.06.03 © Fjellberg, A.
Pores at base of tibiotarsus

Species albifrons (Tullberg, 1871) Fjellberg, A, 2007:il
            Syn.: Rusekianna sibirica Bretfeld, 2002 =?
             Syn.: Sminthurus albifronti(sic) Reuter, OM, 1876:85 ISS
Sminthurinus albifrons from Norway
2006 © Fjellberg, A.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.
Rusekianna albifrons from France
2007.03.17 © Lebeaux, P.

Species bescidica Smolis, A et Skarzynski, D, 2006:317
Species mongolica Betsch, J-M, 1977:62, t.t.
Rusekianna mongolica from Mongolia
After Betsch J-M, 1977 Fig.2A

Species sibirica Bretfeld, 2002

Genus Zebulonia Betsch, J-M, 1970:69
Species massoudi Betsch, J-M, 1970:52

Genus Gisinianus Betsch, 1977
Species flammeolus (Gisin, 1957) Szeptycki & Weiner in Razowski, 1990:24, t.t.

Genus Papirinus Yosii, 1954
Species ankaratrensis Betsch, J-M, 1974:530
Species ieti Yosii, 1966
Species leleupi Delamare Deboutteville, C et Massoud, Z, 1963:227
            Syn.: Papirinus Leleupi(sic) Delamare Deboutteville, C et Massoud, Z, 1963:227 IOS
Species prodigiosus Yosii, 1954 tt
            Syn.: Papirinus prodigiosum(sic) Yosii, 1954 tt IOS
             Syn.: Papirinus prodigiosum(sic) Delamare Deboutteville, C et Massoud, Z, 1963:225 ISS

Tribus Vesicephalini Richards, 1968:28

Genus Vesicephalus Richards WR in Delamare Deboutteville C & Massoud Z, 1964:75 key
Remark by Delamare Deboutteville, C. & Massoud, Z. 1964:75: A French translation of the original generic diagnosis of Vesicephalus Richards, W.R. (1956:79, unpublished thesis) was published.
  Syn.: Vesicephalini Vesicephalus Richards, 1956 =i
Remark by Bretfeld, G. 1999:98: The new monophylum Katiannida Breteld, 1986 contains the Katiannidae Börner, 1913 sensu Betsch, 1980 and the genera incertae sedis sensu Betsch, 1980 and Bretfeld, 1986, including Vesicephalus Richards in Delamare Deboutteville & Massoud, 1964, based on two diagnostic apomorphies: 1. the presence of 2 additional outer setae Ka and Kp distally at the tibiotarsus; 2. the presence of spatulate setae distally at the tibiotarsus.
Remark by Fauna Europaea 2005: Family placement is uncertain. Vesicephalus is intermediate between Katiannidae (anal appendage in females) and Sminthuridae (dorsal chaetotaxy).
Vesicephalus sp. from the USA
Specimen caught in spider web
2007.12.02 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Fourth antennal segment not subdivided
2010.01.17 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Blunt dorsal macrosetae
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Clear swollen interocular vesicles
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Fourth antennal segment not subdivided
2016.11.06 © Johnson, S.T.

Species sp.1 Richards, 1968
Species sp.2 Snider & Janssens, 2008
Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.
Vesicephalus sp. from the USA
2008.03.04 © Fleming, K.R.
Vesicephalus sp. from the USA
Reddish eyepatch
2017.02.03 © Fleming, K.R.

Species bellingeri Bretfeld, 2002
Species crossleyi Snider, RJ, 1985:567
Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.1
Vesicephalus crossleyi from the USA
Female, type I
After Snider R, 1985 Fig.2
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.3
Vesicephalus crossleyi from the USA
Female, type II
After Snider R, 1985 Fig.4
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.5
Vesicephalus crossleyi from the USA
Male
After Snider R, 1985 Fig.6

Species europaeus Ardanaz & Pozo, 1985
             Syn.: Vesicephalus europeus(sic) Jordana, R, Baquero, E et Montuenga, LM, 2001:289 ISS
Vesicephalus europaeus
2002 © Baquero, E. & Jordana, R.
Vesicephalus europaeus from Portugal
2011.02.01 © Almeida, J.
Vesicephalus europaeus from Portugal
2011.02.02 © Almeida, J.
Vesicephalus europaeus male from Portugal
2012.02.02 © Almeida, J.

Species longisetis (Guthrie, JE, 1903:51) Delamare Deboutteville, C & Massoud, Z, 1964:73, t.t.
Species occidentalis (Mills, HB, 1935:138) Delamare Deboutteville, C & Massoud, Z, 1964:75
Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.48
Vesicephalus occidentalis from the USA
Female
After Snider R, 1985 Fig.47
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.
Vesicephalus occidentalis from the USA
2008.11.22 © Hall, K.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
2010.01.10 © Ten Eyck, C.
Vesicephalus occidentalis from the USA
Pale colour form
2016.10.28 © Dobbs, Br.
Vesicephalus occidentalis from the USA
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
2016.11.06 © Johnson, S.T.
Vesicephalus occidentalis from the USA
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Foot with many tenent setae
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Head with long curved blunt macrosetae
2016.12.03 © Johnson, S.T.
Vesicephalus occidentalis from the USA
Face
2016.12.03 © Johnson, S.T.




References