http://www.collembola.org/taxa/neaninae.htm - Last updated on 2017.05.04 by Frans Janssens
Checklist of the Collembola: Neanurinae

Subfamilia Neanurinae Börner C, 1901:33, sensu Cassagnau, 1989 key
Genus Number
of species
Adbiloba6
Afrobella9
Albanura1
Americanura22
Australonura22
Balkanura3
Bilobella14
Blasconura10
Blasconurella5
Caledonimeria1
Caledonura1
Calvinura2
Camerounura1
Cansilianura1
Cassagnaua1
Catalanura3
Caucasanura2
Chaetobella9
Chirolavia2
Christobella1
Coecoloba7
Coreanura1
Crossodonthina12
Cryptonura7
Deuterobella4
Deutonura56
Digitanura1
Echinanura2
Ectonura17
Edoughnura1
Elgonura1
Endonura42
Galanura1
Ghirkanura1
Gnatholonche25
Graniloba2
Hazaranura1
Hemilobella5
Himalmeria (Himalmeria)3
Himalmeria (Yetimeria)17
Himalmeria20
Hyperlobella1
Imparitubercula1
Inameria2
Itanura1
Kalanura4
Lathriopyga11
Lobella (Lobella)14
Lobella (Protolobella)1
Lobella15
Lobellina19
Metanura7
Monobella10
Morulodes4
Nahuanura5
Neanura8
Neanuratobeassignedtopropergenus33
Neanurella1
Nepalanura1
Nepalimeria6
Nilgirella5
Oregonanura1
Paleonura50
Palmanura17
Paralobella12
Paramanura2
Paranura36
Parectonura1
Parvatinura5
Penelopella1
Persanura1
Phradmon6
Phylliomeria4
Pronura55
Propeanura14
Protanura12
Pseudadbiloba1
Pseudobiloba1
Pseudoxenylla1
Pumilinura7
Rambutanura4
Riozura1
Sensillanura8
Siamanura15
Singalimeria1
Speleonura1
Sphaeronura2
Stenomeria1
Sulobella1
Synameria3
Tabasconura1
Tamulmeria2
Telobella2
Tetraloba1
Thaianura1
Thaumanura5
Travura3
Vietnura1
Vitronura20
Womersleya3
Xylanura1
Yosialina1
Yuukianura9
Zelandanura1
Neanurinae771
Genera with number of species
Remark by ICZN. 2007:543: Placed on the Official List of Names in Zoology. Opinion 435.

Neanurinae from Singapore
2007 © Anker, A.
Neanurinae from Croatia
2008.05.25 © Keresztes, G.
Neanurinae from China
2012.10.18 © Deart, Y.
  Syn.: Neanuroidea) Neanirinae(sic) Shveenkova, YB, 2017:76 ISS

Tribus Incerta sedis

Genus Pseudoxenylla Christiansen, KA et Pike, E, 2002:175, -
Species fovealis (Christiansen, KA & Pike, E, 2002:176, -) Christiansen, KA & Pike, E, 2002:176, -, t.t.

Genus Echinanura Carpenter, 1935
Species elegans Carpenter, 1935
Species pacifica (Carpenter, 1934) Yosii, R, 1967:268

Genus Pseudadbiloba Massoud, 1963
Species africana Massoud, 1963

Genus Pseudobiloba Stach, 1951
Species tenebrarum (Absolon, K, 1901:575) Stach, 1951, t.t.

Genus Caledonimeria Delamare Deboutteville, C et Massoud, Z, 1962:331
Species mirabilis Delamare Deboutteville, C et Massoud, Z, 1962:332, t.t.
Caledonimeria mirabilis from New Caledonia
2006 Thibaud, J.-M. © MNHN.


Tribus Neanurini Börner, 1901, sensu Cassagnau P, 1983:6 key
Neanurini from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Neanurini from Malaysia
2008.08.22 © Chin, H.-S.
Neanurini from Indonesia
2013.06.18 © Koh, J.
Neanurini from Tasmania
2014.04.04 © Murray, A.


Genus Edoughnura Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:58
Species rara Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:58, t.t.
Edoughnura rara from Algeria
After Deharveng, L & Hamra-Kroua, S, 2008 ©


Genus Pumilinura Cassagnau, 1979 key
Species albanica Peja & Deharveng, 1995
Species croatica Smolis, A et Skarzynski, D, 2004:9
Species gneiweri (da Gama, 1963) Barra, 1993:76
Species hidalgoi Cassagnau, P et Palacios-Vargas, JG, 1983:3
             Syn.: Pumilinura hildagoi(sic) Barra, J-A, 1993:77 ISS
Species loksai (Dunger, 1974) Barra, J-A, 1993:77
Species travei Cassagnau, P et Peja, N, 1979:220, t.t.
Species vickae Barra, 1993:74

Genus Protanura Börner, C, 1906:167 key
  Syn.: ... Yuukianura Yosii, 1953
Species aleo (Christiansen, K et Bellinger, P, 1992:72)
Species burti Fernando, 1957
Species capitata Folsom, JW, 1932:56
Species carpenteri Mukerji, 1932
             Syn.: Protanura carpenter(sic) Mandal, GP et Hazra, AK, 2009:2 ISS
Species citronella (Carpenter, 1904) Bellinger, PF & Christiansen, KA, 1974:32
Species deharvengi Thibaud & Massoud, 1980
Species lutea Cassagnau, P et Peja, N, 1979:219
Species mediterranea Stach, J, 1967:399
Protanura mediterranea from Malta
After Stach J, 1967 Pl.XXIX Fig.1

            Syn.: Protanura pseudomuscorum Pfliegler, WP et Janssens, F, 2016:204
Species monticellii Caroli, 1912
Species papillata Cassagnau, P et Delamare Deboutteville, C, 1955:374
Species pseudomuscorum (Börner, 1903 tt) Caroli, 1912
            Syn.: Protanura mediterranea Dallai, R, 1973:500
            Syn.: Protanura monticellii Dallai, R, 1973:501
            Syn.: Protanura papillata Dallai, R, 1973:501
Protanura pseudomuscorum from Algeria
After Boldsystems.org, 2006 ©
Remark by Deharveng, L., Hamra-Kroua, S. & Bedos, A. 2007:58: This species and P. monticellii Caroli, 1912 requires redescription from topotypic material.


Species quadrioculata (Börner, 1901) Börner, C, 1906:167

Genus Lathriopyga Caroli, 1912 key
  Syn.: Neanuridae Beanura(sic) Lawrence, PN, 1970:151 lapsus ISS
  Syn.: Achorutes Lathriopyga Caroli, 1910 nn
  Syn.: Neanurinae Lathiopyga(sic) Palacios-Vargas, JG et Najt, J, 1986:24 ISS
Lathriopyga longiseta from the UK
Abd.6 hidden underneath abdomen
2009.04.08 © Crone, S.H.
Lathriopyga longiseta from the UK
Tertiary cuticular granules
2013.01.07 © Murray, A.
Lathriopyga longiseta from the UK
"Missing" 6th abdominal segment
2015.01.28 © Murray, A.

Species belisaria Cassagnau, P et Peja, N, 1979:211
Species bulgarica Smolis, A, Skarzynski, D et Pomorski, RJ, 2004:158
            Syn.: Lathriopyga hellenica Pomorski & Skarzynski, 1999
Species delmasi Massoud, Z, 1963:73
Species hellenica Ellis, WN, 1974:112
Species krizevacensis Barra, J-A, 1993:72
Species longiseta (Caroli, 1910 nn) Stach, J, 1951:71
            Syn.: Achorutes (Lathriopyga) longisetus var. flava Caroli, 1912
            Syn.: Neanura longiseta f. flava Caroli, 1910 nn
Remark by Busmachiu, G., Deharveng, L. & Weiner, W.M. 2010:57: Lathriopyga longiseta flava Caroli, 1912 is a species inquirenda because of an inadequate original description. It could be an albino form of L. longiseta or a good species.


            Syn.: Neanura hystrix Bagnall, RS, 1940:170
            Syn.: Achorutes longisetus Caroli, 1910 nn
             Syn.: Achorutes longiseta(sic) Tarsia, 1936 ISS
Neanura longiseta from Madeira
Dorsal chaetotaxy
Modified after da Gama MM, 1964 Fig.31


             Syn.: Achorutes longisetis(sic) Massoud, Z, 1967:322 ISS
Neanura longiseta from the UK
After Lawrence PN, 1970 Fig.1
Neanura longiseta from the UK
After Lawrence PN, 1970 Fig.15
Beanura(sic) longiseta from the UK
After Lawrence PN, 1970 Fig.2
Lathriopyga longiseta from Greece
Cephalic and prothoracic dorsal chaetotaxy
After Ellis WN, 1976 Fig.30
Lathriopyga longiseta from the UK
After Hopkin, S.P., 1997 ©
Lathriopyga longiseta from the UK
After Hopkin, S.P., 2004 ©
Lathriopyga longiseta from the UK
After Hopkin, S.P., 2004 ©
Lathriopyga longiseta from the UK
After Hopkin SP, 2007 Fig.7d
NOT Lathriopyga longiseta...
Lathriopyga longiseta from the UK
2007.03.04 © Cornwall, N.J.
Lathriopyga longiseta from the UK
2009.04.08 © Crone, S.H.
Lathriopyga longiseta from the UK
2013.01.04 © Murray, A.
Lathriopyga longiseta from the UK
Pointed buccal cone
2013.01.04 © Murray, A.
Lathriopyga longiseta from the UK
Prothoracic dorso-internal-tubercles divergent
2013.01.07 © Murray, A.
Lathriopyga longiseta from the UK
Cephalic posterior dorso-internal tubercles fused with dorso-external (**)
2015.01.24 © Murray, A.
Lathriopyga longiseta from the UK
Cephalic anterior tubercles fused but with granulated fields present
2015.01.24 © Murray, A.
Lathriopyga longiseta from the UK
Abd.6 (partly) hidden under abd.5
2015.01.25 © Murray, A.
Lathriopyga longiseta from the UK
Abdominal macrosetae long and curved
2015.01.25 © Murray, A.
Lathriopyga longiseta from the UK
Dorso-internal tubercles on abd.4 fused
2015.01.28 © Murray, A.

Species microchaeta Cassagnau, P et Peja, N, 1979:212
Species montana Deharveng, L, 1985:213
Lathriopyga montana from France
After Deharveng, L., 1985 Fig.1A

Species nistru Busmachiu, G, Deharveng, L et Weiner, WM, 2010:54
Lathriopyga nistru from Moldova
After Busmachiu G & al, 2010 Fig.1

            Syn.: Lathriopyga bulgarica Busmachiu & Deharveng, 2008

Species primigenia Cassagnau, P et Peja, N, 1979:213
Species willemi Deharveng, L, 1985:216
Lathriopyga willemi from Greece
After Deharveng, L., 1985 Fig.2A


Genus Balkanura Cassagnau, 1979 key
Species cainadasi Cassagnau, P et Peja, N, 1979:207
             Syn.: Balkanura caindasi(sic) Smolis, A, 2003:313 ISS
Species jugoslawica (Palissa & Zivadinovic, 1974) Cassagnau, 1979, t.t.
             Syn.: Balkanura jugoslavica(sic) Smolis, A, 2003:313 ISS
Species wiktori Smolis, A, 2003:315

Genus Monobella Cassagnau, 1979 key
Species calva Bedos, A et Deharveng, L, 1998:292
Species cassagnaui Deharveng, L, 1981:98
Species dubia Deharveng, 1986
Species edaphica Bedos, A et Deharveng, L, 1998:294
Monobella edaphica from France
After Bedos A & Deharveng L, 1998 Fig.4B

Species gersi (Deharveng, 1979) Deharveng, 1986
Species grassei (Denis, 1923) Cassagnau, 1979, t.t.
             Syn.: Neanura graccei(sic) Bagnall, RS, 1940:163 ISS
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
2007 © Cornwall, N.J.
Monobella grassei from the UK
2007 © Cornwall, N.J.
Monobella grassei from the UK
2008.04.30 © Farrell, S.
Monobella grassei from the UK
Pointed mouthcone
2009.02.20 © Michaelson, J.
Monobella grassei from the UK
2011.12.09 © Nurcombe, E.
Monobella grassei from the UK
Ocular tubercle with 2 separate ocelli
2012.01.24 © Jones, A.
Monobella grassei from the UK
Pointed mouthcone
2012.01.24 © Jones, A.
Monobella grassei from the UK
Pointed mouthcone
2012.03.22 © Nurcombe, E.
Monobella grassei from the UK
Note curled-up phoretic nematode
2012.10.20 © Murray, A.
Monobella grassei from the UK
6th abdominal segment hidden under 5th
2012.10.20 © Murray, A.
Monobella grassei from France
2012.11.02 © Quintin, C.
Monobella grassei from France
Two distinctly separated ocelli
2012.11.05 © Quintin, C.
Monobella grassei from the UK
Macrosetae on 1st abdominal segment about the same length as the segment
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Head
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Antenna with curved blunt sensilla
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Two distinctly separated ocelli
2014.01.16 © Phillips, E.
Monobella grassei from the UK
Pointed mouthcone
2014.01.16 © Phillips, E.
Monobella grassei from the UK
Two distinctly separated ocelli
2015.12.13 © Horton, M.
Monobella grassei from the UK
Pointed mouthcone
2015.12.13 © Horton, M.
Monobella grassei from Holland
2016.01.15 © Krediet, A.
Monobella grassei from France
Abd.6 hidden under abd.5
2016.12.03 © Garcelon, P.
Monobella grassei from France
Distinctly pointed mouthcone
2017.01.13 © Garcelon, P.

  Subspecies banyulensis Deharveng, L, 1981:100
  Subspecies distincta Deharveng, 1979
  Subspecies minor Bedos, A et Deharveng, L, 1998:297
  Subspecies prodistincta Deharveng, 1986
Species jau Bedos, A et Deharveng, L, 1998:293
Monobella jau from France
After Bedos A & Deharveng L, 1998 Fig.4A

Species pyrenaica (Deharveng, 1979) Deharveng, 1986
Monobella pyrenaica from France
After Boldsystems.org, 2006 ©

Species rostrata Deharveng, 1986
Species rousseti Bedos, A et Deharveng, L, 1998:294
Monobella rousseti from France
After Bedos A & Deharveng L, 1998 Fig.5A


Genus Endonura Cassagnau, 1979 key
Species sp.1 Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178
Species alavensis Pozo & Simón, 1981
Species arbasensis Deharveng, 1979
Species asiatica Smolis, Deharveng & Kaprus, 2011
Species baculifer Deharveng, 1979
Species caeca (Gisin, 1963) Smolis, A, Kahrarian, M, Piwnik, A, & Skarzynski, D, 2016:66
Species cantabrica Deharveng, 1979
Endonura cantabrica from Romania
After Boldsystems.org, 2006 ©

Species carpatica Smolis, 2006
Species centaurea (Cassagnau, P & Peja, N, 1979:216) Pomorski, RJ & Skarzynski, D, 1999:118
Species ceratolabralis Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:59
Species colorata (da Gama, 1964)
            Syn.: Neanura stachi Gisin, 1952 partim
Species corsica (Denis, 1948)
Species cretensis (Ellis, WN, 1976:277)
Endonura cretensis from Israel
After Smolis A & Kaprus' IJ, 2009 Fig.1

Species dalensi (Deharveng, 1979) Dallai, 1980
Endonura dalensi from Albania
After Traser G & Kontschán J, 2004 Fig.2

  Subspecies sardoa Dallai, R, [1984]:431
             Syn.: Endonura dalensi ssp. sardosa(sic) Traser, G et Kontschán, J, 2004:74 ISS
Species deharvengi (Cassagnau, P & Peja, N, 1979:217) Smolis, A, Kahrarian, M, Piwnik, A, & Skarzynski, D, 2016:67
Species dentifera Smolis, Skarzynski, Pomorski & Kaprus, 2007
Species dichaeta Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:55
Species dudichi (Loksa, I, 1967:289) Smolis, A, 2008:?
Neanura dudichi from Hungary
After Loksa I, 1967 Abb.1

Species gladiirostra Smolis, A et Kaprus, IJ, 2003:327
             Syn.: Endonura gladirostra(sic) Smolis, A et Kaprus, IJ, 2003:327 ISS
             Syn.: Endonura gladirostris(sic) Fanciulli, PP et Dallai, R, 2008:23 ISS
Species gladiolifer (Cassagnau, P, 1954:29)
Species gracilirostris Smolis, Skarzynski, Pomorski & Kaprus, 2007
Species granulata (Cassagnau, P & Delamare Deboutteville, C, 1955:376) Pomorski & Skarzynski, 2000
Species ichnusae Dallai, R, [1984]:435
Species immaculata Deharveng, 1980
Species incolorata (Stach, J, 1951:56) Smolis, A, 2008:?
Species levantica Smolis, Deharveng & Kaprus, 2011
Species ludovicae (Denis, 1948)
Species lusatica (Dunger, 1966) Smolis, A, 2008:?
Species occidentalis Deharveng, 1979
Species pejai Deharveng, 1980
Species persica Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:62
Species poinsotae Deharveng, 1980
Species quadriseta (Cassagnau, P & Peja, N, 1979:218) Fanciulli, PP & Dallai, R, 2008:23
Species reticulata (Axelson, WM, 1905:790) Babenko, 2002
            Syn.: Protanura quadrioculata Linnaniemi, 1912 =?
            Syn.: Achorutes tetrophthalmus Stach, 1929 =?
            Syn.: Neanura (Endonura) tundricola Fjellberg, 1985
Species saleri Fanciulli, PP et Dallai, R, 2008:19
Species tartaginensis (Deharveng, 1980) Dallai, 1983
             Syn.: Endonura tartaginenis(sic) Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:58 ISS
Species tatricola (Stach, J, 1951:53) Gruia, 1975
            Syn.: Neanurella szeptyckii Weiner, 1973
Species taurica (Stach, J, 1951:34) Kaprus, IJ, Pomorski, RJ, Skarzynski, D & Potapov, MB, 2005:893
Species tetrophthalma (Stach, J, 1930:282) Cassagnau, 1979, t.t.
            Syn.: Biloba tetrophthalma f. principalis Stach, J, 1951:52
             Syn.: Achorutes tetrophtalmus(sic) Smolis, A et Kaprus, IJ, 2009:1 ISS
             Syn.: Endonura tetrophtalma(sic) Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:67 ISS
Species transcaucasica (Stach, J, 1951:34) Fanciulli, PP & Dallai, R, 2008:22
Endonura transcaucasica
After Boldsystems.org, 2006 ©

Species tyrrhenica Dallai, R, [1984]:437
Species urotuberculata Pomorski, RJ et Skarzynski, D, 1999:118

Genus Cryptonura Cassagnau, 1979 key
Species anthrenoidea (Ellis, WN, 1976:278)
Species dirfysensis (Cassagnau, P & Peja, N, 1979:214) Smolis, A, 2002:325
Species franzi (Stach, J, 1951:70) Cassagnau, 1979, t.t.
            Syn.: Lathriopyga Franzi(sic) Stach, J, 1951:70,tt IOS
Species jubilaria Smolis, A, 2002:326
Species kuehnelti (Gisin, 1954)
             Syn.: Cryptonura kühnelti(sic) Smolis, A, Falahati, A et Skarzynski, D, 2012:58 ISS
Species maxima Smolis, A, Falahati, A et Skarzynski, D, 2012:55
Species persica Smolis, A, Falahati, A et Skarzynski, D, 2012:52

Genus Neanurella Cassagnau, 1968
Species microphthalma Cassagnau, 1968, t.t.

Genus Deutonura Cassagnau, 1979
Species sp.1 Thibaud & Massoud, 1980
             Syn.: Deutonura conjoncta(sic) Thibaud & Massoud, 1980 ISS
            Syn.: Deutonura conjuncta? Deharveng, L, Zoughailech, A, Hamra-Kroua, S et Porco, D, 2015:281
Species abietis (Yosii, 1969)
Species albella (Stach, 1921) Szeptycki, A & Weiner, WM in Razowski J, 1990:20
Species alduidensis Arbea JI, 1987, n.n.
Remark by Arbea, J.I. 2007.05.07:in litt.: Unavailable manuscript name. Nomen nudum.

Species alticola (Stach, 1951:36)
Species anophthalma (Massoud & Thibaud, 1968)
Species arbeai Lucianez & Simon, 1996
Species atlantica Deharveng, 1981
Deutonura atlantica from France
After Boldsystems.org, 2006 ©

Species balsainensis (Simón, 1978)
Species benzi Traser, G, Thibaud, J-M et Najt, J, 1993:221
Species betica (Deharveng, 1979) Dallai, R, [1984]:444
            Syn.: Neanura (Lathriopyga) stachi Dallai, 1969
            Syn.: Lathriopyga stachi Dallai, 1976
Species binatuber (Lee, 1974)
Species caerulescens Deharveng, 1982
Species caprai Dallai, R, [1984]:456
Species carinthiaca Deharveng, 1982
Species centralis (da Gama, 1964)
Species coiffaiti Deharveng, 1979
Species conjuncta (Stach, 1926) Gruia, 1969
            Syn.: Neanura comata Börner in litt.
            Syn.: Achorutes monoculatus Denis, 1931
Lathriopyga conjuncta from Poland
After Stach J, 1951 Pl.1 Fig.2

             Syn.: Neanura conjucta(sic) Farkas, J, 1995:46 ISS
Deutonura conjuncta from Italy
After Paolucci, P., 2006 ©
Deutonura conjuncta from Hungary
2012.05.10 © Lásló, D.
Deutonura conjuncta from Slovenia
2017.03.04 © Pisec, N.
Deutonura conjuncta from Slovenia
2017.03.04 © Pisec, N.
Deutonura conjuncta from Slovenia
Abd.6 under abd.5
2017.03.06 © Pisec, N.
Deutonura conjuncta from Slovenia
Abd.6 (*) under abd.5
2017.03.07 © Pisec, N.
Deutonura conjuncta from Slovenia
2017.03.10 © Pisec, N.

Species czarnohorensis Deharveng, 1982
Species decolorata (da Gama & Gisin in Gisin, 1964:391):391
             Syn.: Neanura decolarata(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:259 ISS
Species deficiens Deharveng, 1979
Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©

  Subspecies meridionalis Deharveng, 1979
  Subspecies sylvatica Deharveng, 1982
Species deharvengi Arbea, JI et Jordana, R, 1991:100
Species dextra (Gisin, 1954)
Species ezomontana (Yosii, R, 1972:82)
Species fodinarum (Yosii, R, 1956:45)
Neanura fodinarum from Japan
After Yosii R, 1956 Pl.XXII Fig.134

Species frigida (Yosii, 1969)
Deutonura frigida from Russia
After Boldsystems.org, 2006 ©
Deutonura frigida from Russia
After Boldsystems.org, 2006 ©
Deutonura frigida from Russia
After Boldsystems.org, 2006 ©

Species gibbosa Porco, Bedos & Deharveng, 2010
Species gisini Deharveng, 1982
Species granatuberis (Lee, 1974)
Species gurreae Lucianez & Simon, 1996
Species ibicensis (Ellis, WN, 1974:128)
Species igilica Dallai, R, [1984]:451
Species ilvatica Dallai, R, [1984]:449
Species inopinata Deharveng, 1979
Species insularis Deharveng, 1982
Species leei Deharveng, L et Weiner, WM, 1984:19
Species luberonensis Deharveng, 1982
Species mirabilis Deharveng, 1987
Species monticola (Cassagnau, P, 1954:33)
Deutonura monticola from France
After Boldsystems.org, 2006 ©

Species oglasicola Dallai, R, [1984]:453
Species phlegraea (Caroli, 1912) Cassagnau, 1979, t.t.
             Syn.: Lathriopyga plegraea(sic) Stach, J, 1951:Pl.1 Fig.3 ISS
Lathriopyga plegraea(sic) from Poland
After Stach J, 1951 Pl.1 Fig.3

  Subspecies implena Kseneman, M, 1938:467
Species picea (Yosii, 1969)
Species plena (Stach, J, 1951:68) Szeptycki, A & Weiner, WM in Razowski J, 1990:20
Species portucalensis (da Gama, 1964) Deharveng, 1982
Species provincialis Deharveng, 1979
Deutonura provincialis from France
After Boldsystems.org, 2006 ©
Deutonura provincialis from France
After Boldsystems.org, 2006 ©

Species quinquesetosa Deharveng, 1982
Species selgae Deharveng, 1979
Species similis Deharveng, 1979
Species sinistra (Denis, 1935) Dallai & Malatesta in Calderini, 1995:6
Species stachi (Gisin, 1952) Szeptycki, A & Weiner, WM in Razowski J, 1990:20
            Syn.: Lathriopyga phlegraea Stach, 1951, nec Caroli, 1912
            Syn.: Achorutes phlegraeus f. albella Stach, 1921
Deutonura stachi from Romania
After Dányi, L. & Traser G, 2008 Fig.1,2

Species sylviae Dallai, R, [1984]:446
Species urbionensis Deharveng, 1979
Species vallespirensis Deharveng, 1982
Species weinerae Deharveng, 1982
Species yoshiiana Deharveng, L et Weiner, WM, 1984:21
Species zana Deharveng, L, Zoughailech, A, Hamra-Kroua, S et Porco, D, 2015:282
            Syn.: Deutonura sp. Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:57
Deutonura zana from Algeria
After Deharveng L et al, 2015 Fig.1
Deutonura zana from Algeria
After Deharveng L et al, 2015 Fig.2


Genus Albanura Deharveng, 1982
Species nana (Cassagnau, P & Peja, N, 1979:215) Deharveng, 1982

Genus Catalanura Deharveng, 1982
Species catalana (Deharveng, 1979) Cassagnau, P, 1983:15, t.t.
Species florae Arbea, JI et Jordana, R, 1991:100
Species najtae (Deharveng, 1979)

Genus Metanura Yosii, 1954
Species cassagnaui Deharveng, L et Weiner, WM, 1984:24
Species iriomotensis (Yosii, 1965)
Species kitayamana Yoshii, 1995
Species nagatonis Yoshii, 1995
Species okinawana (Yosii, 1965)
Species sanctisebastiani Yosii, 1954, t.t.
            Syn.: Metanura st-sebastiani(sic) Yosii, 1954,tt IOS
             Syn.: Metanura sebastiani(sic) Yosii, R, 1956:44 ISS
Neanura (Metanura) sebastiani(sic) from Japan
After Yosii R, 1956 Pl.XXIV Fig.141


Species yamashironis Yoshii, 1995

Genus Nahuanura Palacios-Vargas, JG et Najt, J, 1986:18 key
Species ce Palacios-Vargas, JG et Najt, J, 1986:19, t.t.
Species ome Palacios-Vargas, JG et Simón-Benito JC, 2007:170
Species suenoi (Yosii, 1955) Palacios-Vargas, JG & Simón-Benito JC, 2007:175
            Syn.: Paranura s-uenoi(sic) Yosii, 1955 IOS
             Syn.: Paranura s-uenoi(sic) Massoud, Z, 1967:238 ISS
             Syn.: Paranura s-uenoi(sic) Yosii, R, 1977:146 ISS
             Syn.: Paranura s-uenoi(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Paranura s-uenoi(sic) Deharveng, L, 1989:104 ISS
Remark by Nicolson, D. 2007.mm.dd:in litt.: Based on art.NN the hyphen in the original spelling should be removed.


             Syn.: Nahuanura? s-uenoi(sic) Palacios-Vargas, JG et Simón-Benito JC, 2007:175 ISS
             Syn.: Paranura? s-uenoi(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:683 ISS
Species trioculata Palacios-Vargas, JG et Simón-Benito JC, 2007:172
Species yei Palacios-Vargas, JG et Simón-Benito JC, 2007:172

Genus Neanuratobeassignedtopropergenus Janssens F, 2017
Species aethiopica (Tarsia in Curia, 1939)
Species albipes (Nicolet, H, 1847:389), i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species alpestris (Latzel, R, 1917:236)
Species atra (Nicolet, H, 1847:389), i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species azrouensis (Thibaud & Massoud, 1980)
Species bakeri (Handschin, 1926), s.i.
Species casabianca (Fernando, 1959), i.s.
Species caucasica (Stach, 1951:35)
Species chilensis (Nicolet, H, 1847:389), i.s.
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Anoura chilensis from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.3.
             Syn.: Anoura Chilensis(sic) Lubbock, J, 1873:200 ISS
Species coronifera (Axelson, WM, 1905:789)
Species eburnea (Gisin, 1963)
Species florida (Latzel, R, 1917:236)
Species folsomi (Stach, 1951:26), n.n.
Species growae (Christiansen, K et Bellinger, P, 1980:377)
Neanura growae from the USA
Tubercles only on head and posterior body segments
2015.01.01 © Fleming, K.

Species hirta (Joseph, 1881)
Species infernalis (Joseph, 1881)
Species japonica (Börner, C, 1909:107)
Species kentingensis (Lee & Kim, 1990)
Species laricis (Martynova, 1977)
Species loricata (Börner in Schille, 1912), n.n.
Species magna (MacGillivray, AD, 1893:128)
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2009.03.23 © Leppin, M.
Neanura magna from the USA
2009.03.24 © Leppin, M.
Neanura magna from the USA
Tubercles on abd.4 in triangular constellation
2016.11.28 © Dobbs, Br.
Neanura magna ♂ from the USA
Bilobed abd.6 hidden under abd.5
2016.11.28 © Dobbs, Br.
Neanura magna from the USA
Eye with 3 ocelli in triangle
2016.11.28 © Dobbs, Br.
Neanura magna from the USA
Cephalic tubercles: (1)+2+4+6
2016.11.28 © Dobbs, Br.
Neanura magna from the USA
Abdominal dorsal tubercles: 6+6+6+6+2
2016.12.07 © Dobbs, Br.
Neanura magna ♀ from the USA
Rudiment of furca
2016.12.07 © Dobbs, Br.

Species mirabilis (Massoud, 1963), i.s.
Species multituberculata (Coleman, 1941), s.i.
Species pallida (Willem, V, 1902:10)
            Syn.: Biclavella patagonica Wahlgren, 1907
Species patagonica (Wahlgren, 1901)
            Syn.: Biclavella pallida Börner, 1906
Species pini (Becker, 1948), s.i.
Species pohli (Barra, J-A, 2006:63)
Species pseudornata (Scott, 1961)
Species sexoculata (Carpenter, GH, 1916:29), i.s.
Species takoensis (Kinoshita, 1916)
Neanura takoensis from China
2006.03.14 © Botting, J.
Neanura takoensis from China
After www.yellowman.cn., 2007 ©
Neanura takoensis from China
2011.02.11 © Botting, J.
Neanura takoensis from China
2011.02.11 © Botting, J.

Species tigrina (Harvey, 1900), s.i.
Species tumulosa (Li, 1991)
Species weigneri (Stach, 1922)

Genus Neanura MacGillivray, AD, 1893:314 key
  Syn.: ... Achorutes Templeton, 1835,nr ICZN
  Syn.: ... Achoreutes(sic) Koch, 1840 nr ISS
  Syn.: ... Achoreutes(sic) Templeton, 1843 nr ISS
  Syn.: ... Acharutes(sic) Harvey, 1893 ISS
  Syn.: ... Blax Koch, 1840 =?
  Syn.: ... Anoura(sic) Gervais, 1843 nec Gray, 1838 IOS
  Syn.: ... Anura Agassiz, 1846
  Syn.: Lipurelas Anoura(sic) Nicolet, H in Gay, C, 1849:95 ISS
  Syn.: Thysanura Anoura(sic) MacGillivray, AD, 1893:128 partim ISS
  Syn.: Aphoruridæ(sic) Anoura(sic) MacGillivray, AD, 1893:314 ISS
  Syn.: Thysanura Anoura(sic) Cockerell, TDA, 1894:116 ISS
  Syn.: ... Rathumoutes Templeton, 1844 nr
  Syn.: ... Biclavella Willem, 1902
  Syn.: ... Biloba Stach, 1949,nr Opinion 435, 1956
  Syn.: ... Imparitubercula Stach, 1951
  Syn.: Bilobidae Kodiakia Bödvarsson, H, 1960:43
Remark by ICZN. 2007:543: Placed on the Official List of Names in Zoology. Opinion 435.

Species alba von Törne, 1957
            Syn.: Neanura parva Dunger, 1975 partim
Remark by Dányi, L. & Traser, G. 2008:16: The data of Neanura parva (Stach) reported by Dunger (1975) included Neanura alba as well.


Species judithae Smolis, A et Deharveng, L, 2017:39
Neanura judithae from Poland
After Smolis A & Deharveng L, 2017 Fig.1A

Species minuta Gisin, 1963
Species moldavica Busmachiu, G et Deharveng, L, 2008:?
Species muscorum (Templeton, R in Templeton, R & Westwood, JO, 1836:97) MacGillivray, AD, 1893:314, t.t., B
            Syn.: Blax ater Koch, 1840
            Syn.: Achorutes tuberculatus Nicolet, H, 1842:51
Achorutes tuberculatus from Switzerland
After Nicolet, H., 1842 Pl.5 Fig.1.
Anoure tuberculée from Europe
Anoura tuberculata
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.1

             Syn.: Achorutes Tuberculatus(sic) Bourlet, C, 1845:67 ISS
            Syn.: Anoura crassicornis Müller J, 1859
            Syn.: Anoura nigra Wankel, 1860
            Syn.: Anura gibbosa Packard, AS, 1873:27
            Syn.: Anoura sextuberculata Harvey, 1896
            Syn.: Neanura muscorum var. purpurascens Axelson, WM, 1905:790
            Syn.: Neanura muscorum ssp. alpestris Latzel, 1917
            Syn.: Neanura persimilis Mills, HB, 1934:30 =?
Neanura persimilis from the USA
2015.09.15 © Wechsler, D.
Neanura persimilis from Canada
2016.04.29 © de Repentigny, L.-G.

            Syn.: Neanura betulae Selga, 1959
Achorutes Muscorum(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.6
Anoura muscorum from the UK
After Lubbock, J., 1873 Pl.48 Fig.1.
Neanura muscorum from New Zealand
After Salmon, J.T., 1941 Pl.46 Fig.104.
Remark by Deharveng, L. & Bedos, A. 2000:211: The Vietnamese record possibly corresponds to Vietnura caerulea, new species, which is the most common Neanurini of Vietnam. Neanura muscorum, widespread in temperate zones, has never been met in the numerous samples we collected in the region, nor in any country of Southeast Asia.

Neanura muscorum from the UK
After Hopkin, S.P., 2002 ©
Neanura muscorum from Sweden
2005 © Hall, K.
Neanura muscorum
After Boldsystems.org, 2006 ©
Neanura muscorum from Sweden
2006 © Hall, K.
Neanura muscorum from Norway
2006.07.29 © Fauskanger, I.
Neanura muscorum from the UK
2007 © Barton, T.
Neanura muscorum from the USA
2007 © McClarin, J.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from France, juvenile
2007.03.15 © Lebeaux, P.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from the UK
2007.04.01 © Cornwall, N.J.
Neanura muscorum from the UK
2008.01.06 © Campbell, A.
Neanura muscorum from the Netherlands
2008.01.15 © van Erkelens, J.A.
Neanura muscorum from France
2008.01.19 © Hugues, C.
Neanura muscorum from the UK
2008.04.18 © Farrell, S.
Neanura muscorum from the UK
2008.04.18 © Farrell, S.
Neanura muscorum from Algeria
After Hamra-Kroua, S., 2009.04.02 ©
Det. Deharveng, L.
Neanura muscorum with egg deposition
After Thibaud J-M & D'Haese C, 2010 Fig.50
2009 © D'Haese, C.
Neanura muscorum from Belgium
2011.09.09 © Henderickx, H.
Neanura muscorum from the UK
With phoretic nematodes
2012.02.29 © Nurcombe, E.
Neanura muscorum from Denmark
2012.03.21 © Bildstrup, O.
Neanura muscorum from the USA
With parasitic mite
2012.05.04 © Murray, T.
Neanura muscorum from the UK
Ocular tubercles not united with frontal tubercle
2012.06.27 © Murray, A.
Neanura muscorum from the UK
Ocular tubercle with 2+1 ocelli
2012.09.07 © Murray, A.
Neanura muscorum from the UK
Flower-like cuticular ornamentation on tubercles
2012.12.11 © Murray, A.
Neanura muscorum from the UK
Sharply pointed mouth cone
2012.12.11 © Murray, A.
Neanura muscorum from the UK
Bilobed 6th abdominal segment
2013.01.06 © Nurcombe, E.
Neanura muscorum from France
Ocular tubercle with 2 anterior ocelli and 1 posterior ocellus
2013.03.24 © Sandra.
Neanura muscorum from France
Cephalic tuberculae and pointed mouthcone
2013.03.24 © Sandra.
Neanura muscorum from the UK
Tuberculate hydrophobic cuticula
2013.12.04 © Murray, A.
Neanura muscorum from the UK
2+2 anterior ocelli
Cephalic tubercles not fused
2014.08.22 © Murray, A.
Neanura muscorum from the USA
2014.11.09 © Silsbee, D.
Neanura muscorum from the UK
Bilobed 6th abdominal segment
2015.01.28 © Murray, A.
Neanura muscorum from the UK
With uplifted abdomen
2016.01.08 © Jeffery, J.

Species pallida Deharveng, 1979
Species parva (Stach, J, 1951:48) Gisin, 1960:99
Species pseudoparva Rusek, 1963

Genus Itanura Queiroz, GC et Deharveng, L, 2015:140
Itanura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.16

Species brasiliensis (Arlé, R, 1960:164) Queiroz, GC & Deharveng, L, 2015:142
Itanura brasiliensis from Brazil
From type locality
After Queiroz GC et Deharveng L, 2015 Fig.17
Itanura brasiliensis from Brazil
From Rio de Janeiro: Itatiatia
After Queiroz GC et Deharveng L, 2015 Fig.22
Itanura brasiliensis from Brazil
From Rio de Janeiro: Serra dos Órgãos
After Queiroz GC et Deharveng L, 2015 Fig.28


Genus Thaumanura Börner, 1932
  Syn.: Bilobidae Thamanura(sic) Stach, J, 1951:Pl.1 Fig.4 ISS
Species carolii (Stach, 1920) Börner, 1932, t.t.
            Syn.: Achorutes Carolii(sic) Stach, 1920,tt IOS
Thamanura(sic) carolii from Poland
After Stach J, 1951 Pl.1 Fig.4

             Syn.: Thaumanura caroli(sic) Dallai, R, 1969:377 ISS
Thaumanura carolii from Romania
After Boldsystems.org, 2006 ©
Thaumanura carolii from Czechia
2007.07.05 © Krásenský, P.
Thaumanura carolii from Slovenia
2013.06.09 © Miran, C.
Thaumanura carolii from Slovenia
Posterior abdominal segments
2013.06.09 © Miran, C.
Thaumanura carolii from Slovenia
Ventral aspect
2013.06.09 © Miran, C.

Species dallaii Cassagnau, P et Peja, N, 1979:221
Species echinata (Kos, 1940)
            Syn.: Achorutes bornensis Schött, 1925 =?
            Syn.: Achorutes japonicus Börner, 1909 =?
            Syn.: Achorutes pterothrix Börner, 1909 =?
Species oniscoides (Latzel, R, 1917:237) Stach, J, 1951:65, s.i.
             Syn.: Achorutes Carolii(sic) Stach, 1919 =? ISS
             Syn.: Thaumanura oniscoïdes(sic) Massoud, Z, 1967:329 ISS
Thaumanura oniscoides from Hungary
2012.05.10 © Lásló, D.

Species ruffoi Dallai, R, 1969:377
Thaumanura ruffoi from Italy
2011.04.09 © Piredda, C.


Genus Kalanura Smolis, A, 2007:?
Species babenkoi Smolis, A, 2007:?
Species compacta Smolis, A, 2007:?
Species spectabilis Smolis, A, 2007:?
Species stebaevae Smolis, A, 2007:?

Genus Vietnura Deharveng, L et Bedos, A, 2000:211
Species caerulea Deharveng, L et Bedos, A, 2000:212, t.t.
Vietnura caerulea from Vietnam
After Deharveng, P. & Bedos, A., 2000 Fig.1.


Genus Cansilianura Dallai, R et Fanciulli, PP, 1983:235
Species malatestai Dallai, R et Fanciulli, PP, 1983:237, t.t.
Cansilianura malatestai from Italy
After Boldsystems.org, 2006 ©


Genus Caucasanura Kuznetsova & Potapov, 1988
Species besucheti Deharveng, L, 1989:817
Species stebaevae Kuznetsova & Potapov, 1988

Genus Ghirkanura Kuznetsova & Potapov, 1988
  Syn.: Neanuridae (Neanurinae) Girkanura(sic) Hopkin, SP, 1997:205 ISS
Species chernovae Kuznetsova & Potapov, 1988

Genus Imparitubercula Stach, 1951
Species villosa (Kos, 1940) Stach, 1951:11
             Syn.: Imparitubercula villosia(sic) Cassagnau, P, 1983:21,si ISS

Genus Tetraloba Lee, 1983
Species seolagensis Lee, 1983

Genus Xylanura Smolis, 2011
Species oregonensis Smolis, 2011

Genus Persanura Mehrafroz Mayvan, M, Shayanmehr, M, Smolis, A et Skarzynski, D, 2015:553
Species hyrcanica Mehrafroz Mayvan, M, Shayanmehr, M, Smolis, A et Skarzynski, D, 2015:553, t.t.

Genus Christobella Fjellberg, 1985
  Syn.: ... Cristobella Skidmore, 1995 ISS
Species ornata (Folsom, JW, 1902:89) Fjellberg, 1985
Neanura ornata from Alaska
After Folsom JW, 1902 Pl.IV Fig.2


Tribus Morulodini Cassagnau P, 1983:5

Genus Morulodes Cassagnau, P, 1955:160
Species ambiguus (Christiansen & Bellinger, 1980:368) Cassagnau, P, 1983:6
Species rishirianus Tanaka & Ischisawa, 2002
Species serratus (Folsom, 1916) Massoud, Z, 1967:287, t.t.
            Syn.: Morulodes millsi Cassagnau, P, 1955:160
            Syn.: Kodiakia aleuta Bödvarsson, H, 1960:46
            Syn.: Kodiakia prima Bödvarsson, H, 1960:43
Morulodes serratus from the USA
Setae very heavily serrate
2016.11.26 © Dobbs, Br.
Morulodes serratus from the USA
Bilobed 6th abdominal segment hidden under 5th
2016.11.26 © Dobbs, Br.

Species setosus (Canby, 1926) Cassagnau, P, 1983:6
Morulodes setosus from Canada
2013.04.13 © Hochhalter, R.
Morulodes setosus from Canada
Eyepatch with 3 anterior and 1 posterior ocelli
2013.04.13 © Hochhalter, R.
Morulodes setosus from Canada
Bilobed 6th abdominal segment hidden under 5th
2013.04.13 © Hochhalter, R.


Tribus Lobellini Cassagnau P, 1983:8
            Syn.: Neanurinae Crossodonthini(sic) Massoud, Z, 1967:342 partim IOS              Syn.: ... Crossodonthini(sic) Deharveng, L, 2004:424 partim ISS             Syn.: ... Neanurini Deharveng, L, 2004:424 partim
            Syn.: ... Protanurini Deharveng, L, 2004:424 partim

Genus Coreanura Deharveng, L et Weiner, WM, 1984:27
Species filiae Deharveng, L et Weiner, WM, 1984:27, t.t.

Genus Propeanura Yosii, R, 1956:46
Species garuda Yoshii, 1976
Species hygrophila (Cassagnau, P et Deharveng, L, 1981:158)
Species hypostoma (Denis, 1929)
Species kymgangensis Deharveng, L et Weiner, WM, 1984:44
Species labukae Yoshii, 1981
Species lapidicola Cassagnau, P et Deharveng, L, 1981:155
Species leei Deharveng, L et Weiner, WM, 1984:46
Species megalops Deharveng, L et Weiner, WM, 1984:48
Species nomurai (Yosii, R, 1956:43, i.s.) Deharveng & Weiner, 1984
Yuukianura nomurai from Japan
After Yosii R, 1956 Pl.XXII Fig.131

Species parvituberculata (Yosii, 1965) Cassagnau, P, 1983:22
             Syn.: Neanura parvitubercula(sic) Massoud, 1967 ISS
Species pterothrix (Börner, C, 1909:107) Yosii, 1959, t.t.
             Syn.: Neanura pterothryx(sic) Yosii, 1954 ISS
             Syn.: Neanura (Propeanura) pterothryx(sic) Yosii, R, 1956:47,tt ISS
             Syn.: Neanura pterothryx(sic) Massoud, 1967 ISS
             Syn.: Propeanura pterothryx(sic) Cassagnau, P, 1983:15,tt ISS
Species sandakanensis Yoshii, 1981
Species soemarwotoi Yosii, 1976
Species zehntneri (Handschin, 1920)

Genus Coecoloba Yosii, R, 1956:33
Species hidana Yosii, R, 1956:36
Coecoloba hidana from Japan
After Yosii R, 1956 Pl.XIX Fig.115
             Syn.: Coecoloba hidani(sic) Massoud, 1967 ISS
Species kurasawana Yosii, R, 1956:35
Coecoloba kurasawana from Japan
After Yosii R, 1956 Pl.XVII Fig.96

Species lobella (Yosii, R, 1954:65) Yosii, R, 1956:34, t.t.
Coecoloba lobella from Japan
After Yosii R, 1956 Pl.XVII Fig.95

Species odai Yosii, R, 1956:36
Coecoloba odai from Japan
After Yosii R, 1956 Pl.XIX Fig.116

Species onychiuriformis (Yosii, 1976) Deharveng, L, 1983:59
Species plumleyi Deharveng, L, 1983:61
Species wakasana Yosii, R, 1956:37
Coecoloba wakasana from Japan
After Yosii R, 1956 Pl.XX Fig.117


Genus Lobella Börner, C, 1906:168
  Syn.: ... Biocularia Stach, J, 1951:6 partim
Lobella sp. from Japan
Probably self-luminous
After Oba Y & al, 2011 Fig.7E
Lobella sp. from Japan
Emits a continuous weak green light from abdominal tubercles by stimulation
After Oba Y & al, 2011 Fig.7F

Subgenus Protolobella Cassagnau, P, 1983:18
Species assamensis Yosii, 1966, t.t.
Lobella (Lobella) assamensis from India
After Mandal GP & Hazra AK, 2009 Fig.10a+b

Subgenus Lobella Cassagnau, P, 1983:18 ss
Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Indonesia
2013.10.26 © Bertner, P.

Species bicincta Cassagnau, P et Deharveng, L, 1984:3
Species bicolor Cassagnau, P et Deharveng, L, 1984:4
Species daeana Lee, 1982
Species fortis (Oudemans, JT in Weber, M, 1890:91) Yayuk, 1989:7
Species malabarica Yosii, 1966
Species maxillaris Yosii, 1966
Species monocincta Cassagnau, P et Deharveng, L, 1984:4
Species nana Lee & Kim, 1990
Species palmeri (Wray, 1967) Palacios-Vargas & Najt, 1986:25
Lobella palmeri from the USA
After Bernard, E.C., 2006 ©
Lobella palmeri from the USA
2015.05.04 © Anderson, J.
Lobella palmeri from the USA
2015.05.04 © Anderson, J.

Species punctata Cassagnau, P et Deharveng, L, 1984:7
Species sauteri (Börner, C, 1906:168) Stach, J, 1951:12, t.t.

             Syn.: Neanura (Lobella) Sauteri(sic) Carpenter, GH, 1916:30 ISS
Species semilunaris (Schött, 1925) Yayuk, 1989:7
Species sokamensis Deharveng, L et Weiner, WM, 1984:31
Species uozumii Yosii, 1956

Genus Telobella Cassagnau, P, 1983:18
Species kemiri Suhardjono, YR et Deharveng, L, 2001:?
Species wayang Yosii, 1976, t.t.

Genus Lobellina Yosii, 1956
Species sp. Yosii, R, 1977:149
            Syn.: Lobella sauteri Yosii, 1956 nec Börner
Species cavicola (Yosii, 1956), i.s.
Lobellina? cavicola from Japan
Abd.1-4 dorso-internal tubercles absent
2014.08.23 © Hara, Y.
Lobellina? cavicola from Japan
Ocular tubercle with 2 anterior +1 posterior ocelli
2014.08.23 © Hara, Y.

Species chosonica Deharveng, L et Weiner, WM, 1984:35
Species decipiens (Yosii, 1965)
Species gedehensis (Yosii, 1976), i.s.
Species ionescui (Massoud & Gruia, 1974) Cassagnau, P, 1983:10
Species ipohensis (Yosii, 1976) Deharveng, L & Weiner, WM, 1984:34
Species kitazawai (Yosii, 1969) Deharveng, L & Weiner, WM, 1984:34
Species minuta (Lee, 1980) Deharveng, L & Weiner, WM, 1984:34
Species mizunasiana (Yosii, 1956)
             Syn.: Lobella (Lobellina) mizunashiana(sic) Yosii, R, 1977:149 ISS
             Syn.: Lobella (Lobella) mizunashiana(sic) Yosii, R, 1977:149 ISS
Species montana Deharveng, L et Weiner, WM, 1984:37
Species musangensis (Yosii, 1976) Deharveng, L & Weiner, WM, 1984:34
Species nanjingensis Ma, Z-C et Chen, J-X, 2008:62
Species paraminuta Deharveng, L et Weiner, WM, 1984:39
Species perfusionides (Stach, J, 1965:346) Deharveng, L & Weiner, WM, 1984:34
             Syn.: Lobella perfusoïdes(sic) Massoud, Z, 1967:333 ISS
Species proxima Deharveng, L et Weiner, WM, 1984:41
Species roseola (Yosii, 1954) Cassagnau, P, 1983:15, t.t.
Species setapauca (Gapud, 1970), i.s.
Species stachi (Yosii, 1951)

Genus Cassagnaua Özdikmen, H, 2009:293
  Syn.: Lobellienne Pectinura Cassagnau, P, 1983:18 nec Forbes, 1843
Species hongkongensis (Yosii, 1976) Özdikmen, H, 2009:294, t.t.

Genus Crossodonthina Yosii, 1954
Species alatoserrata Yosii, 1965
Species altamontana Yoshii, 1981
             Syn.: Crossodenthina altamontana Yayuk, 1989 ISS
Species appendiculata Yosii, R, 1956:41
Species bidentata Luo & Chen, 2009
Species formosana Yosii, 1955
Species hainana Xiong, Chen & Yin, 2005
Species koreana Yosii, R et Lee, C-E, 1963:1
Crosodonthina koreana from Korea
After Yosii R & Lee C-E, 1963 Fig.1A

            Syn.: Crossodonthina delamarei Lee BH , 1973

Species montana Lee & Kim, 1990
Species nipponica Yosii, 1954, t.t.
            Syn.: Crossodonthina japonica Yosii, 1959 lapsus
            Syn.: Crossodonthina japonica Yosii, R et Lee, C-E, 1963:2 lapsus
Crossodonthina nipponica from Japan
After Yosii R, 1956 Pl.XX Fig.118

Species radiata (Salmon, JT, 1941:308) Massoud, Z, 1967:343
Neanura radiata from New Zealand
After Salmon J.T., 1941 Pl.47 Fig.117

Species tiantongshana Xiong, Chen & Yin, 2005
Species tridentiens Yue, Q-Y et Yin, W-Y, 1999:222

Genus Sphaeronura Cassagnau, P, 1983:19
Species bornensis (Schött, 1925) Deharveng, L & Bedos, A, 2000:210
Remark by Deharveng, L. & Bedos, A. 2000:210: It is likely that the Borneo and the Vietnam forms belong to different species.

Species chaotica (Yosii, 1976) Cassagnau, P, 1983:15, t.t.

Genus Riozura Cassagnau, P, 1983:18
Species yoshibai (Yosii, 1963) Cassagnau, P, 1983:15, t.t.

Genus Yuukianura Yosii, 1955
  Syn.: Lobellini Yukianura(sic) Smolis, A, Skazynski, D, Kahrarian, M et Kaprus, IJ, 2016:299 ISS
Species aphoruroides (Yosii, 1953) Yosii, 1955, t.t.
Yuukianura aphoruroides from Japan
After Yosii R, 1956 Pl.XXI Fig.119
Lobella (Yuukianura) aphoruroides from China
After Stach J, 1964 Plate III Fig.1

Species halophila Yosii, 1955
Species hawaiiensis (Bellinger, PF & Christiansen, KA, 1974:31) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58
Species judithae Deharveng, L, Palacios-Vargas, JG et Bedos, A, 2017:58
Yuukianura judithae from Vanuata
After Deharveng L, Palacios-Vargas JG & Bedos A, 2017 Fig.1B

Species pacifica (Yosii, 1971) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58
Species rosea (Kim, JT & Lee, B-H, 2000:180) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58
Species szeptyckii Deharveng, L et Weiner, WM, 1984:50
Species tongana (Yosii, R, 1964:9) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:57
Species yasudai (Yosii, 1966:479) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:57

Genus Deuterobella Yoshii, R et Suhardjono, YR, 1992:5
Species imadatei (Yosii, 1976), t.t.
Species murphyi (Yosii, 1976) Deharveng, L & Bedos, A, 2000:210
Remark by Deharveng, L. & Bedos, A. 2000:210: It is likely that the Malaya and the Vietnam forms belong in fact to different species.

Species tawauensis (Yoshii, 1981)
Species ternatensis (Yoshii, R et Suhardjono, YR, 1992:6)

Genus Yosialina Salmon, 1964
Species similis (Yosii, R, 1954:66) Salmon, 1964

Genus Hemilobella Deharveng & Greenslade P, 1992
Species ferrumequinum Deharveng, Agolin & D'Haese, 2008
Species newmani (Womersley, 1933) Deharveng & Greenslade P, 1992:734
Species perakensis (Yosii, 1976) Deharveng & Greenslade P, 1992:736
Species pluvia Deharveng & Greenslade P, 1992:732
Species rounsevelli Deharveng & Greenslade P, 1992:730, t.t.

Genus Hyperlobella Cassagnau, P, 1988:24
Species kraepelini (Börner, C, 1906:169) Cassagnau, P, 1988:24, t.t.
            Syn.: Neanura dubiosa Ritter, 1910
            Syn.: Neanura dubiosa Mandal, GP et Hazra, AK, 2009:13
            Syn.: Neanura pudibunda Imms, 1912
            Syn.: Neanura pudibunda Mandal, GP et Hazra, AK, 2009:13
            Syn.: Protanura spinifera Carpenter, 1917
            Syn.: Protanura spinifera Mandal, GP et Hazra, AK, 2009:13
            Syn.: Protanura sp. Goto, 1955 =?
            Syn.: Lobella cassagnaui Prabhoo, 1971
Lobella kraepelini from Singapore
After Yosii R, 1959 Fig.6A
Lobella kraepelini from India
After Mandal GP & Hazra AK, 2009 Fig.9a


Genus Paralobella Cassagnau, P et Deharveng, L, 1984:9 key
Species apsala (Yosii, 1976) Jiang & al, 2012
Species breviseta Luo, Y-Z et Palacios-Vargas, JG, 2016:344
Paralobella breviseta from China
Modified after Luo Y-Z & Palacios-Vargas JG, 2016 Fig.1

Species erawan (Yosii, 1976) Jiang & al, 2012
Species khaochongensis (Yosii, 1976) Bedos, 1994
             Syn.: Lobella? (Lobellina) Khaochonaensis(sic) Deharveng, 1987 ISS
Species kinabaluensis (Yoshii, 1981) Jiang & al, 2012
Species orousseti Cassagnau, P et Deharveng, L, 1984:9, t.t.
             Syn.: Paralobella orousetti(sic) Hopkin, SP, 2002:1 ISS
             Syn.: Paralobella orousetii(sic) Hopkin, SP, 2002:1 ISS
Paralobella orousseti
After Deharveng, L., 2002 ©

Species palustris Jiang, Luan & Yin, 2012
Species paraperfusa (Gapud, 1968) Jiang & al, 2012
Species penangensis (Yosii, 1976) Jiang & al, 2012
Species perfusa (Denis, 1934) Deharveng, L & Bedos, A, 2000:210
Species sabahna (Yosii, 1981) Jiang & al, 2012
Species selangorica (Yosii, 1976) Jiang & al, 2012

Genus Sulobella Deharveng & Suhardjono, 2000
Species yoshii Deharveng & Suhardjono, 2000, t.t.

Tribus Paleonurini Cassagnau, 1989

Genus Graniloba Cassagnau, 2000
Species imitator Cassagnau, 2000, t.t.
Species verrucosa Cassagnau, 2000

Genus Parectonura Deharveng in Tillier, 1988
Species ciliata Deharveng in Tillier, 1988:46, t.t.

Genus Caledonura Deharveng in Tillier, 1988
Species tillierae Deharveng in Tillier, 1988:49

Genus Penelopella Cassagnau, 1986
  Syn.: Paleonurini Penellopella(sic) Smolis, A, 2000:412 ISS
Species pacifica Cassagnau, 1986

Genus Siamanura Deharveng, L, 1987:2
Species bisetosa Deharveng, L, 1987:18
Species breviseta Deharveng, L, 1987:15
Species burkhardti Deharveng, L, 1987:24
Species clavata Deharveng, L, 1987:15
Species gouzei Deharveng, L, 1987:20
Species hanuman (Yoshii, 1976)
Species inthanonensis Deharveng, L, 1987:6
Species leksawasdii Deharveng, L, 1987:11
Species lisu Deharveng, L, 1987:20
Species loebli Deharveng, L, 1987:27
            Syn.: Siamanura löbli(sic) Deharveng, L, 1987:27 IOS
Species maffrigali Deharveng, L, 1987:14
Species media Deharveng, L, 1987:6
Species phyllioseta Deharveng, L, 1987:23
Species primadinae Suhardjono, YR et Deharveng, L, 1992:61
             Syn.: Siamanura primadina(sic) Suhardjono, YR et Deharveng, L, 1992:61 ISS
Species trisetosa Deharveng, L, 1987:9

Genus Chirolavia Deharveng, L, 1991:54
Species gabaudei Deharveng, L, 1991:55
Species murphyi Deharveng, L, 1991:54, t.t.

Genus Pronura Delamare Deboutteville, 1953
Species aciculifer Cassagnau, 1996
Species aequalis Cassagnau, 1991
Species alata Cassagnau, 1996
Species amazonica Cassagnau & Oliveira, 1990
Species anjavicola Cassagnau, 1996
Species annapurnensis Cassagnau, 1991
Species australica (Yosii, 1966) Greenslade P & Deharveng, 1991:852
            Syn.: Neanura cirrata Yosii, 1966 =?
            Syn.: Achorutes rosaceus Womersley, 1933 =:
Species bidoup Deharveng, L et Smolis, A, 2002:2
Species bipartita Cassagnau, 1991
Species bouakensis Cassagnau, 1996
Species caeca Cassagnau, 1996
Species camerounensis Cassagnau, 1996
Species capitata Cassagnau, 1991
Species celisi Cassagnau, 1996
Species clavisetis (Yoshii, 1981)
Species curviseta Cassagnau, 1996
Species dandicola Cassagnau, 1991
Species diplochaeta Cassagnau, 1991
Species dorsolateralis Deharveng, L et Bedos, A, 1993:191
Species edaphica Cassagnau, 1991
Species elegans Cassagnau, 1996
Species erythrina Cassagnau, 1991
Species fluctuans Cassagnau, 1996
             Syn.: Pronura fluctans(sic) Smolis, A, 2000:412 ISS
Species gaucheri Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:198
Pronura gaucheri from French Guyana
After Palacios-Vargas JG et al, 2011 Fig.1
2009 © D'Haese, C.

Species gosainkundi Cassagnau, 1991
Species indiana Salmon, 1969
Species kilimanjarica Delamare Deboutteville, 1953, t.t.
             Syn.: Pronura kilimandjarica(sic) Massoud, Z, 1967:336 ISS
             Syn.: Pronura kilimandjarica(sic) Cassagnau, P, 1983:16 ISS
             Syn.: Pronura kilimandjarica(sic) Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:198 ISS
Species kyandolirensis Cassagnau, 1996
Species laminata Cassagnau, 1996
Species lemurica Cassagnau, 1996
Species longiseta Cassagnau, 1991
Species madagascariensis Cassagnau, 1996
Species major Cassagnau, 1996
Species mignardi Massoud, 1963
Species nana Cassagnau, 1996
Species nepalica Cassagnau, 1991
Species ornata Deharveng, L et Bedos, A, 1993:190
Species paraguayana Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:202
Species parvati Cassagnau, 1991
Species plumosa Cassagnau, 1991
Species pomorskii Smolis & Deharveng, 2006
Species prima Cassagnau, 1996
Species prodoni Cassagnau, 1991
Species pumilio Cassagnau, 1996
Species regularis Cassagnau, 1996
Species reticulata Cassagnau, 1991
Species ruwenzoriensis Cassagnau, 1996
Species seticola Cassagnau, 1991
Species setimigrans Cassagnau, 1996
Species shimicola Cassagnau, 1991
Species sphaeroculata Cassagnau, 1996
Species straturata Cassagnau, 1996
Species travei Cassagnau, 1991
Species trisetosa Cassagnau, 1996
Species trisulensis Cassagnau, 1991

Genus Galanura Smolis, A, 2000:412
Species agnieskae Smolis, A, 2000:414, t.t.
             Syn.: Galanura agnieszkae(sic) Smolis, A, 2000:414 ISS

Genus Paleonura Cassagnau, P, 1982:6
Species aberdarensis Cassagnau, 1996
Species africana Cassagnau, 1996
Species angustior (Rusek, 1971) Cassagnau, P, 1982:6
Species anophthalma Cassagnau, 1991
Species anosyennica Cassagnau, 1996
Species badaga Cassagnau, P, 1988:28
Species barbata Cassagnau, P, 1988:30
Species bilinskii Pasnik & Weiner, 2013
Species carayoni Massoud & Thibaud, 1987
Species cassagnaui Weiner, WM et Najt, J, 1998:229
Species coalescens Cassagnau, 1996
Species colimana Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:96
            Syn.: Deutonura sp. Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:101
Species daniae Palacios-Vargas & Díaz, 1992:158
Species decorata Cassagnau, P, 1988:31
Species deharvengi Cassagnau, 1991
Species dejeani Cassagnau, 1996
Species dilatata Deharveng, L et Bedos, A, 1993:187
Species epiphytica Smolis, A et Deharveng, L, 2003:320
Species formosana (Yosii, 1965) Deharveng, L, 1989:104
Species ganesh Cassagnau, 1991
Species guadalcanarae (Yosii, 1960) Deharveng & Greenslade P, 1992:738
Species guae (Yoshii, 1976) Cassagnau, P, 1982:6
Species gurung Cassagnau, 1991
Species ili Christiansen, K et Bellinger, P, 1992:88
Species indrabahadouri Cassagnau, 1991
Species insularum (Carpenter, 1935) Cassagnau, P, 1982:6
Species irregularis Cassagnau, 1991
Species khumbica (Cassagnau, 1971) Cassagnau, P, 1982:6
Species lamjungensis Cassagnau, 1991
Species lanna Deharveng, L et Bedos, A, 1993:184
Species loebli Cassagnau, P, 1988:27
Species lonavlana (Yosii, 1966) Cassagnau, P, 1982:6
Species louisi Thibaud & Najt, 1993
Species macronychia Cassagnau, P, 1988:26
Species miniseta Massoud & Thibaud, 1987
             Syn.: Paleonura minisetae(sic) Palacios-Vargas & Díaz, 1992 ISS
             Syn.: Paleonura minisetae(sic) Palacios-Vargas & Gómez-Anaya, 1995 ISS
Species monochaeta Deharveng, L et Bedos, A, 1993:185
Species monophthalma (Yosii, 1966) Cassagnau, P, 1982:6
Species nuda Cassagnau & Oliveira, 1990
Species paurochaetosa Cassagnau, 1991
Species pescadorius Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:95
Species petebellingeri Palacios-Vargas, JG et Simón-Benito, JC, 2007:320
Species plumosa Cassagnau, 1996
Species reducta (Yosii, 1966) Cassagnau, P, 1982:6
Species rosacea (Schött, H, 1917:7) Deharveng, L & Greenslade, P, 1992:738
             Syn.: Achorutes roseus(sic) Sarawak YYYY lapsus ISS
Paleonura rosacea from Australia
2016.08.11 © Ball, D.
Paleonura rosacea from Australia
2016.08.11 © Ball, D.
Paleonura rosacea from Australia
2016.10.31 © Steve & Alison.
Paleonura rosacea from Australia
2016.10.31 © Steve & Alison.
Paleonura rosacea from Australia
Pointed mouthcone
2016.10.31 © Steve & Alison.

Species setikholensis Cassagnau, 1991
Species siva (Yosii, 1966) Cassagnau, P, 1982:6
Species sp. Najt & Thibaud, 1987
Species sp. Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:93
Species spectabilis Cassagnau, P, 1982:7, t.t.
Species trisetosa Cassagnau, 1991

Genus Speleonura Palacios-Vargas, JG et Simón-Benito, JC, 2007:318
Species kenchristianseni Palacios-Vargas, JG et Simón-Benito, JC, 2007:318, t.t.

Genus Ectonura Cassagnau, 1980
Ectonura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.15

Species araucariae (Yosii, 1960) Deharveng & Bedos, 2002
Species barrai Janion, C, Bedos, A et Deharveng, L, 2011:39
Species coatesi Barra, J-A, 1994:186
Remark by Janion, C., Bedos, A. & Deharveng, L. 2011:34: The only differential character separating unambiguously E. coatesi from E. oribiensis is the morphology of modified chaetae Ag in the male.

Species judithae Deharveng & Bedos, 2002
Species koumac Deharveng & Bedos, 2002
Species lata Deharveng, Weiner & Najt, 1997
Species longirostris Deharveng, Weiner & Najt, 1997
Species minima Deharveng, Weiner & Najt, 1997
Species monochaeta Janion, C, Bedos, A et Deharveng, L, 2011:34
Species natalensis (Womersley, 1934) Cassagnau, 1980, t.t.
            Syn.: Neanura montana Paclt, 1959
Species oribiensis (Coates, 1968) Cassagnau, 1980
Species paralata Deharveng, Weiner & Najt, 1997
Species proxima Deharveng, Weiner & Najt, 1997
Species pulchra Deharveng, Weiner & Najt, 1997
Species sensillata Deharveng & Bedos, 2002
Species snowdeni Queiroz, GC et Deharveng, L, 2015:136
Ectonura snowdeni from Brazil
After Queiroz GC et Deharveng L, 2015 Fig.2

Species sylvatica Deharveng, Weiner & Najt, 1997

Genus Vitronura Yosii, 1969 key
  Syn.: Neanurinae Matanura(sic) Yosii, R, 1977:149 ISS
Vitronura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.12

Species sp. Weiner & Najt, 1991
Species acuta Deharveng, L et Weiner, WM, 1984:52
Species dentata Deharveng, L et Weiner, WM, 1984:54
Species giselae (Gisin, 1950) Cassagnau, 1980
            Syn.: Neanura mandarina Yosii, 1954 tt
Neanura (Propeanura) mandarina from Japan
After Yosii R, 1956 Pl.XXIV Fig.139

            Syn.: Metanura pygmaea Yosii, 1954 =?
            Syn.: Anoura rosea Gervais, 1842
Anoura rosea from France
After Nicolet H, 1847 Pl.6 fig.16
Vitronura rosea from Switzerland
2008.03.17 © Haentjens, R.
Remark by Janssens, F. 2015.10.14: Given Yoshii (1995) synonymised rosea Gervais with giselae Gisin and mandarina Yosii, according to ICZN rules of priority, rosea Gervais, 1842 takes priority on giselae Gisin, 1950 and mandarina Yosii, 1954. However, this is not yet reflected in the current taxonomic hierarchy due to taxonomic complications with the type species mandarina.


Remark by Deharveng, L. & Bedos, A. 2000:210: Identification of the Vietnamese material has to be confirmed.

Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.27 © Justis, S.
Vitronura giselae from the USA
2009.02.20 © Houston, S.
Vitronura giselae from the USA
2009.10.25 © Bergen, L.
Vitronura giselae from the USA
2009.10.25 © Bergen, L.
Vitronura giselae from the USA
2010.06.18 © Montgomery, G.
Vitronura giselae from the USA
2011.12.24 © Fleming, K.
Vitronura giselae from the USA
2012.01.19 © Kouri, J.
Vitronura giselae from the USA
Dorsal head tubercles: 1+2+3+6
2015.07.29 © Wechsler, D.
Vitronura giselae from Malaysia
2015.12.07 © Krásenský, P.
Vitronura giselae from France
Dorsal head tubercles: 1+2+3+6
2016.02.08 © Garcelon, P.
Vitronura giselae from France
Dorsal head tubercles: 1+2+3+6
2016.02.11 © Garcelon, P.

  Subspecies citrina Latzel, 1922
             Syn.: Achorutes roseus-citrinus(sic) Handschin, E in Dahl, F, 1929:42,si ISS
Species gressitti Cassagnau, P et Deharveng, L, 1981:161
Species joanna (Coates, 1968) Greenslade, P, 1997-:www0312
Species kunigamiensis Tanaka & Hasegawa, 2010
Species latior (Rusek, J, 1967:189)
Species luzonica Yosii, 1976
Species macgillivrayi (Denis, 1933) Cassagnau & Palacios-Vargas, 1983
            Syn.: Biloba costaricae Stach, J, 1951:29
Remark by Palacios-Vargas, J.G. 2010.08.09: I was in Costa Rica collecting Neanurids and found some Vitronura specimens. I realized that what was described by Denis (1933) as Achorutes macgillivrayi and that described by Stach (1951) as Biloba costaricae, are the same species.
Species mascula Smolis & Deharveng, 2006
Species namhaeiensis Lee, 1974
Species pygmaea (Yosii, 1954) Yosii, 1969
Species qingchengensis Jiang & Yin, 2011
Species quartadecima Gao, Y, Bu, Y et Palacios-Vargas, JG, 2012:1147
Vitronura quartadecima from China
After Gao Y, Bu Y & Palacios-Vargas JG, 2012 Fig.15

Species setaebarbulata Gao, Y, Bu, Y et Palacios-Vargas, JG, 2012:1143
Vitronura setaebarbulata from China
After Gao Y, Bu Y & Palacios-Vargas JG, 2012 Fig.2

Species shaanxiensis Jiang & Yin, 2011
Species singaporiensis (Yosii, R, 1959:14) Yosii, 1976
Bilobella singaporiensis from Singapore
After Yosii R, 1959 Fig.8A

Species sinica Yosii, 1976
Species tubercula Lee & Kim, 1990

Genus Blasconura Cassagnau, P, 1983:19
Blasconura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.11

Species anamalensis Cassagnau, P, 1988:37
Species batai Bedos & Deharveng, 2000
Species ceylonica Cassagnau, P, 1988:35
Species hirtella (Börner, C, 1906:170) Cassagnau, P, 1983:16
            Syn.: Neanura hirtella var. schötti(sic) Womersley, 1935
Lobella (Propeanura) hirtella
from Singapore
After Yosii R, 1959 Fig.7A

Remark by Deharveng, L. & Bedos, A. 2000:210: Blasconura hirtella may correspond to a complex of closely related species. The identification of the Vietnamese specimens has to be checked.

Blasconura hirtella from Singapore
2008.03.29 © Ng, M.
Blasconura hirtella from Singapore
2008.07.21 © Ng, M.
Blasconura hirtella from Singapore
2008.07.21 © Ng, M.
Blasconura hirtella from Singapore
2011.06.11 © Koh, J.
Blasconura hirtella from Singapore
2011.07.16 © Koh, J.
Blasconura hirtella from Singapore
2012.10.27 © Bay, N.
Blasconura hirtella from Singapore
2015.11.27 © Bay, N.
Blasconura hirtella from Singapore
2015.11.27 © Bay, N.

  Subspecies novaezealandiae Salmon, JT, 1941:307
            Syn.: Neanura hirtella ssp. novae-zealandiae(sic) Salmon, JT, 1941:307 IOS
             Syn.: Neanura hirtella var. nova-zelandiae(sic) Massoud, Z, 1967:318,si ISS
Blasconura hirtella ssp. novaezealandiae juv. from New Zealand
2014.02.09 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.

Species palniensis Cassagnau, P, 1988:36
Species prabhooi Cassagnau, P, 1988:34
Species separata (Denis, 1934) Cassagnau, 1988
Remark by Cassagnau, 1988 in Deharveng, L. & Bedos, A. 2000:210: The attribution to the genus Blasconura is uncertain.

Species sholica Cassagnau, P, 1988:33
Species toda Cassagnau, P, 1988:32
Species vivianae Palacios-Vargas & Díaz, 1992:162

Genus Australonura Cassagnau in Dallai, 1980
Australonura from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Det. Deharveng, L., 2008.12.22
Australonura sp. from New Zealand
2014.03.02 © Murray, A.
Australonura sp. from New Zealand
2014.03.02 © Murray, A.
Australonura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.13
Australonura sp. from Tasmania
2016.02.02 © Tony.
Australonura sp. from Tasmania
Head tubercles
2016.02.02 © Tony.

Species sp.1 Palacios-Vargas, JG et Deharveng, L, 2014:42
Species ahimsa Greenslade P & Deharveng, 1990:572
Species beata Greenslade P & Deharveng, 1990:574
Species cirrata (Schött, H, 1917:8) Greenslade P & Deharveng, 1990:575
            Syn.: Achorutes hirtellus Handschin, 1926
            Syn.: Achorutes hirtellus ssp. noname Denis, 1933
            Syn.: Achorutes hirtellus Womersley, 1933
             Syn.: Neanura hirtellus(sic) var. cirratus(sic) Womersley, H, 1942:24 ISS
Species friasica (Cassagnau & Oliveira, 1990) Palacios-Vargas, JG & Deharveng, L, 2014:40
            Syn.: Neanura limnophila Cassagnau & Rapoport, 1962 partim
Species gili Queiroz & Deharveng, 2014
Species grandes Greenslade P & Deharveng, 1990:576
Species grossi (Yosii, 1966) Cassagnau, P, 1983:16, t.t.
Species lawrencei Deharveng & Greenslade P, 1990:598
Species limnophila (Cassagnau & Rapoport, 1962) Palacios-Vargas, JG & Deharveng, L, 2014:35
Australonura limnophila from Argentina
After Palacios-Vargas JG & Deharveng L, 2014 Fig.2

Species meridionalis (Stach, 1951, s.i.) King, Greenslade P & Hutchinson, 1985
             Syn.: Australonura meridonalis(sic) Greenslade P & Deharveng, 1990 ISS
Species neotropica Queiroz & Deharveng, 2014
Species norfolkensis Greenslade P & Deharveng, 1990:580
Species novaecaledoniae (Yosii, R, 1960:10) Deharveng, L in Tillier, S, 1988:46
            Syn.: Neanura novae-caledoniae(sic) Yosii, R, 1960:10 IOS
             Syn.: Neanura novacaledoniae(sic) Massoud, Z, 1967:307 ISS
Species paraguayensis Palacios-Vargas, JG et Deharveng, L, 2014:42
            Syn.: Paleonura sp. Palacios-Vargas & Mejía Recamier, 2009
Species popamanasiu Deharveng & Greenslade P, 1990:602
Species quarta Greenslade P & Deharveng, 1990:582
Species redita Greenslade P & Deharveng, 1990:584
Species rennellensis Deharveng & Greenslade P, 1990:602
Species sanguisugarum Greenslade P & Deharveng, 1990:584
Species scoparia Greenslade P & Deharveng, 1990:586
Species wellingtonia (Womersley, H, 1936:477) Greenslade, P & Deharveng, L, 1990:588
Achorutes hirtellus var. wellingtonia from Tasmania
After Womersley H, 1936 Fig.2a


Genus Zelandanura Deharveng, L et Wise, KAJ, 1987:143
  Syn.: Paleonurini Zealandanura(sic) Greenslade, P et Deharveng, L, 1991:853 ISS
Species bituberculata Deharveng, L et Wise, KAJ, 1987:144, t.t.

Genus Phradmon Greenslade P & Deharveng, 1991
Phradmon sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.14

Species australasiae (Womersley, H, 1935:208) Greenslade, P & Deharveng, L, 1991:841, t.t.
Paranura australasiae from Australia
After Womersley H, 1939 Fig.44AD
             Syn.: Paranura australasia(sic) Salmon, JT, 1964:283 ISS
Species floccosus Greenslade P & Deharveng, 1991:844
Species maralali Greenslade P & Deharveng, 1991:845
Species schotti (Womersley, H, 1935:210) Greenslade, P & Deharveng, L, 1991:847
             Syn.: Neanura hirtella ssp. schoetti(sic) Salmon, 1941 ISS
             Syn.: Paranura australasia(sic) ssp. schoetti(sic) Salmon, JT, 1964:283 nn ISS
             Syn.: Paranura hirtella ssp. schoetti(sic) Yosii, 1966 ISS
             Syn.: Australonura schoetti(sic) Greenslade, P et Deharveng, L, 1990:591 ISS
             Syn.: Phradmon schoetti(sic) Greenslade, P et Deharveng, L, 1991:847 ISS
             Syn.: Phradmon schoetii(sic) Greenslade, P et Deharveng, L, 1991:837 ISS
             Syn.: Phradmon schoetti(sic) Greenslade, P in Houston, WWK, 1994:36 ISS
Species tasmaniae Greenslade P & Deharveng, 1991:849
Species trisetosus Greenslade P & Deharveng, 1991:851

Genus Blasconurella Deharveng, L et Bedos, A, 1992:300 key
Species arcuata Deharveng, L et Bedos, A, 1992:307, t.t.
Species intermedia Deharveng, L et Bedos, A, 1992:305
Species lobata Deharveng, L et Bedos, A, 1992:306
Species palmata Deharveng, L et Bedos, A, 1992:309
Species quinquesetosa Deharveng, L et Bedos, A, 1992:302

Genus Paramanura Cassagnau, 1986
Species najtae Cassagnau, 1986
Species problematica Cassagnau, 1991

Genus Hazaranura Cassagnau, 1991
  Syn.: Paleonurini Hazanura(sic) Smolis, A, 2000:412 ISS
Species elongata Cassagnau, 1991

Genus Nepalanura Yosii, 1966
Species paranuroides Yosii, 1966
             Syn.: Nepalanura (Nepalanura) paranurodes(sic) Mandal, GP et Hazra, AK, 2009:2 ISS

Genus Thaianura Yosii, 1961
Species umesaoi Yosii, 1961, t.t.

Genus Womersleya Denis, 1948
Species formosana Lee & Kim, 1990
Species marhia Baijal, HN, 1958:351
Womerselya(lapsus) marhia from the Himalaya
After Baijal HN, 1958 Fig.2A.

Species vicina (Denis, 1934) Denis, 1948, t.t.

Genus Afrobella Cassagnau, P, 1983:19
Species delamarei Cassagnau, 1996
Species guineensis (Murphy, 1965) Cassagnau, P, 1983:16, t.t.
             Syn.: Afrobella guineense(sic) Cassagnau, P, 1983:16 ISS
Species kirivatensis Cassagnau, 1996
Species mamillata Cassagnau, 1996
Species pauliani (Massoud, 1963) Cassagnau, P, 1983:21
Species scoparia Cassagnau, 1996
Species simplex (Massoud, 1963) Cassagnau, P, 1983:21
Species thibaudi Cassagnau, 1996
Species wittei Cassagnau, 1996

Genus Bilobella Caroli, E, 1912:367 key
  Syn.: ... Lathriopyga Caroli, 1910 partim
Bilobella carpatica from Ukraine
Prothoracic dorsal internal tubercles absent (*)
After Smolis A & Kaprus IJ, 2008 Fig.1

Species albanica Cassagnau, P et Peja, N, 1979:208
Species aurantiaca (Caroli, E, 1912:367) Stach, J, 1951:88, t.t.
Achorutes aurantiacus
After Caroli E, 1912 Fig.5
            Syn.: Neanura aurantiaca Caroli, 1910 nn
            Syn.: Bilobella burgalensis Simon & Pozo, 1987
            Syn.: Neanura (Bilobella) pyrenaea Selga, 1959
Remark by Maynard, E.A. 1951:70: Although the arrangement of the tubercles does not seem to agree with the meager description of this European species at my disposal, I am tentatively identifying my specimens as N. aurantiaca.

             Syn.: Bilobella auratiaca(sic) Stach, J, 1951:88 ISS
             Syn.: Neanura aurantica(sic) Rusek, J in Dlabola, J, 1977:13 ISS
Remark by Christiansen, K.A. &Bellinger, P.F. 1998:411: The occurence of this species in the Nearctic region is dubious. According to Maynard's description of his "aurantiaca", the head has 10 tubercles instead of the, 18(sic) [must be 8] described by Caroli; if this is correct, his species, at least, is probably distinct.

Bilobella aurantiaca
2005 © Domene, X.
Bilobella aurantiaca from Czechia
2007.07.05 © Krásenský, P.
Bilobella aurantiaca from Czechia
2007.07.05 © Krásenský, P.
Bilobella aurantiaca from Italy
2008.12.22 © Gregg, S.
Bilobella aurantiaca from Italy
2008.12.22 © Gregg, S.
Bilobella aurantiaca from France
2012.09.30 © Lebeaux, P.
Bilobella aurantiaca from Spain
After Arbea JI, 2013 Fig.7
Bilobella aurantiaca from Spain
2013.02.01 © Montoro, A.
Bilobella aurantiaca from Portugal
2014.04.13 © Wergen., B
Bilobella aurantiaca from Slovenia
2017.03.04 © Pisec, N.
Bilobella aurantiaca from Slovenia
2017.03.10 © Pisec, N.

Species braunerae Deharveng, L, 1981:96
Bilobella braunerae from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.7
Bilobella aurantiaca(sic) from Holland
Legit Berg, M.
2007.03.13 © Heijerman, T.
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
 
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
Bilobella braunerae from Holland
After Berg M, 2009 Fig.1 © Heijerman T
Bilobella braunerae from Germany
2010.04.05 © Marquard, F.
Bilobella braunerae from Germany
2010.04.05 © Marquard, F.
Bilobella braunerae from Germany
2010.04.23 © Marquard, F.
Bilobella braunerae from Germany
2010.05.25 © Marquard, F.
Bilobella braunerae from Germany
2014.08.16 © Beck, K.
Bilobella braunerae from Germany
2014.08.16 © Beck, K.
Bilobella braunerae from Germany
2014.08.16 © Kluck, A.
Bilobella braunerae from Germany
1+3+4 cephalic tubercles
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
Ocular tubercle with 2 ocelli
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
Flower shaped texture of dorsal tubercles
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
2014.10.10 © Kluck, A.
Bilobella braunerae from Germany
Pointed mouthcone
2014.10.10 © Kluck, A.
Bilobella braunerae from Germany
Ocular tubercle with 2 ocelli
2014.10.21 © Kluck, A.
Bilobella braunerae from Germany
2014.10.21 © Kluck, A.
Bilobella braunerae from the UK
Leg.: Boardman, P. Det.: Shaw, P.
2015.01.28 © Shaw, P.
Bilobella braunerae from the UK
2015.01.dd © Boardman, P.
Bilobella braunerae from Germany
Sharply pointed mouth-cone
2015.08.08 © Kluck, A.
Bilobella braunerae juv. from Germany
2015.08.08 © Kluck, A.
Bilobella braunerae from Germany
2015.08.31 © Beck, K.
Bilobella braunerae from Belgium
Abd.6 partly hidden under abd.5
2017.03.08 © Huskens, M.-L.
Bilobella braunerae from Belgium
2 ocelli without pigment
2017.03.08 © Huskens, M.-L.
Bilobella braunerae from Belgium
Sharply pointed mouthcone
2017.03.10 © Huskens, M.-L.
Bilobella braunerae from Slovenia
2017.03.10 © Pisec, N.

Species carpatica Smolis, A et Kaprus, IJ, 2008:510
Species coiffaiti Cassagnau, 1968
Species digitata Cassagnau, 1967
Bilobella digitata
After Potapov M in Babenko A, 1988

Species excolorata (Loksa, I et Rubio, I, 1966:142)
Neanura excolorata from Hungary
After Loksa I & Rubio I, 1966 Abb.17

Species ligurica Deharveng, L, 1981:97
Species mahunkai Dányi, L, 2010:29
Bilobella mahunkai from Montenegro
After Dányi, L., 2010 Fig.2
Bilobella mahunkai from Montenegro
After Dányi, L., 2010 Fig.2

Species massoudi Cassagnau, 1968
             Syn.: Bilobella massoudis(sic) Frati, F et Dell'Ampio, E, 2000:345 ISS
Species matsakisi Cassagnau, 1967
Bilobella matsakisi from Albania
After Traser G & Kontschán J, 2004 Fig.7

Species proxima Cassagnau, P et Peja, N, 1979:209
Species subaurantiaca Cassagnau, P et Peja, N, 1979:210
Species zekoi Cassagnau, P et Peja, N, 1979:210

Genus Chaetobella Cassagnau, P, 1983:19
Species buddha Cassagnau, 1993
Species eurygaster Cassagnau, 1993
Species multiseta Cassagnau, 1993
Species numatai (Yosii, 1966) Cassagnau, P, 1983:16, t.t.
Species rhodophila Cassagnau, 1993
Species sigillata Cassagnau, 1993
Species sylvatica Cassagnau, 1993
Species tonglica Cassagnau, 1993
Species verticicola Cassagnau, 1993

Genus Travura Cassagnau, P et Deharveng, L, 1980:2
Species betschi Cassagnau, P et Deharveng, L, 1980:6
Species divergens Cassagnau, P et Deharveng, L, 1980:3, t.t.
Species mikenensis Cassagnau, 2000

Genus Adbiloba Stach, 1951
Species intermedia Cassagnau, 2000
Species leleupi Cassagnau, 2000
Species plurichaetosa Cassagnau, P et Peja, N, 1979:206
Species prodoni Cassagnau, 1993
Species sikkimensis Yosii, 1966
Neanura sikkimensis from India
After Mandal GP & Hazra AK, 2009 Fig.8a

Species sokolowi (Philiptschenko, 1926) Stach, 1951, t.t.

Genus Parvatinura Cassagnau, P, 1982:7
Species colcheni Cassagnau, P, 1984:9
Species dobremezi Cassagnau, P, 1982:8, t.t.
Species loebli Cassagnau, P, 1984:8
Species numica Cassagnau, P, 1984:8
Species taounjensis Cassagnau, P, 1984:8

Genus Inameria Cassagnau, P, 1983:20
Species corallina (Imms, 1912) Cassagnau, P, 1983:16, t.t.
            Syn.: Lobella (Propeanura) subcorallina Prabhoo, 1971
Species kenyensis Cassagnau, P, 1984:9

Genus Nilgirella Cassagnau, P, 1983:20
Species amboinensis Yoshii, R et Suhardjono, YR, 1992:3, i.s.
Species indica (Handschin, E, 1929:235) Cassagnau, P, 1983:16, t.t.
Achorutes indicus from South-India
After Handschin E, 1929 Fig.7

Species longiseta Cassagnau, P, 1988:44
Species palniensis Cassagnau, P, 1988:43
Species piljainae Cassagnau, P, 1988:42

Genus Phylliomeria Delamare Deboutteville, 1948
Species africana Delamare Deboutteville, 1948, t.t.
Species bebourensis Cassagnau, P, 1984:27
Species ruwenzorensis Cassagnau, P, 1984:26
Species simplex Cassagnau, P, 1984:26

Genus Gnatholonche Börner, C, 1906:168
  Syn.: Achorutes Gnatolonche(sic)) Schött, H, 1917:8 ISS
  Syn.: Paleonurini Gnatolonche(sic) Cassagnau, P, 1991:223 ISS
Species angularis (Salmon, JT, 1944:140) Stach, J, 1951:93
Gnatholonche angularis from New Zealand
2009 © Te Papa Museum
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.

Species annapurnensis Cassagnau, P, 1984:10
Species anomala (Yosii, 1966) Cassagnau, P, 1984:4, s.i.
Species coineaui Cassagnau, P, 1984:11
Species dentata Cassagnau, 1993
Species dobatensis Cassagnau, P, 1984:10
Species durrieui Cassagnau, P, 1984:9
Species elongata Cassagnau, P, 1984:13
Species ensifer Cassagnau, 1993
Species impariseta Cassagnau, 1993
Species intermedia (Imms, 1912) Stach, J, 1951:94
Species jesti Cassagnau, P, 1984:14
Species lamjurica Cassagnau, 1993
Species limbu Cassagnau, P, 1984:13
Species lipaspis (Börner, C, 1906:170) Handschin, 1926, t.t.
Species mahabaratensis Cassagnau, P, 1984:10
Species marginata Cassagnau, P, 1984:12
Species montana (Handschin, 1929) Stach, 1951:94
Species polychaetosa Cassagnau, P, 1984:14
Species prodoni Cassagnau, P, 1984:12
Species reducta Cassagnau, P, 1984:11
Species sarkimani Cassagnau, 1993
Species sensilla Salmon, 1948
Species setensis Cassagnau, 1993
Species sherpa Cassagnau, 1993

Genus Synameria Cassagnau, P, 1983:20
Species intermedia Cassagnau, P, 1990:22
Species miranda (Yosii, 1966) Cassagnau, P, 1983:16, t.t.
Species orientalis Cassagnau, P, 1990:20
Synameria orientalis from Nepal
After Cassagnau P in Thibaud J-M & D'Haese C, 2010 Fig.32


Genus Nepalimeria Cassagnau, P, 1984:5
Species coccinea Cassagnau, P, 1984:26
Species dal Cassagnau, P, 1984:24, t.t.
Species ganash Cassagnau, 1993
Species heterochaeta Cassagnau, P, 1984:25
Species khorensis Cassagnau, P, 1984:25
Species lepchana (Yosii, 1966) Cassagnau, P, 1984:4, s.i.

Genus Elgonura Cassagnau, P, 1984:5
Species erinacea Cassagnau, P, 1984:28, t.t.

Genus Singalimeria Cassagnau, P, 1984:5
Species pachyderma Cassagnau, P, 1984:24, t.t.

Genus Himalmeria Cassagnau, P, 1984:4
Subgenus Himalmeria Cassagnau, P, 1984:4 ss
Species gurung Cassagnau, P, 1984:15, t.t.
Species karmapa Cassagnau, P, 1984:16
Species ornata Cassagnau, P, 1984:16
Subgenus Yetimeria Cassagnau, P, 1984:4
Species armata Cassagnau, P, 1984:18
Species arunensis Cassagnau, P, 1984:21
Species cervicornis Cassagnau, P, 1984:21
Species cuneata Cassagnau, 1993
Species digitata Cassagnau, P, 1984:17
Species elegans Cassagnau, P, 1984:23
Species flabellifera Cassagnau, P, 1984:20
Species foliata Cassagnau, 1993
Species himalayana (Yosii, 1966) Cassagnau, P, 1984:4, s.i.
Species jaljalensis Cassagnau, P, 1984:19
Species lama Cassagnau, P, 1984:18, t.t.
Species lanata Cassagnau, P, 1984:19
Species musifera Cassagnau, P, 1984:20
Species rostrata Cassagnau, P, 1984:19
Species sikkimensis Cassagnau, P, 1984:18
Species smetanai Cassagnau, P, 1984:22
Species spatulata Cassagnau, P, 1984:20

Genus Calvinura Cassagnau, P, 1988:39
Species besucheti Cassagnau, P, 1988:39, t.t.
Species reducta Cassagnau, P, 1988:40

Genus Camerounura Cassagnau, P, 1991:221
Species delamarei Cassagnau, P, 1991:222, t.t.

Genus Digitanura Deharveng, L, 1987:145
Species quadrilobata Deharveng, L, 1987:146, t.t.
Digitanura quadrilobata from Thailand
2010.08.15 © Damrongsiri, S.


Genus Rambutanura Deharveng, 1988
Rambutanura sp. from Malaysia
2015.12.07 © Krásenský, P.

Species carcharia Smolis, A, 2007:?
Species dawydoffi (Denis, 1934) Deharveng, 1988
Species malayana (Yosii, 1976)
Species yoshiiana Deharveng, 1988

Genus Stenomeria Cassagnau, P, 1990:29
Species deharvengi Cassagnau, P, 1990:29, t.t.

Genus Tamulmeria Cassagnau, P, 1988:46
Species callipygos Cassagnau, P, 1988:46, t.t.
Species keralensis Cassagnau, P, 1988:47

Tribus Paranurini

Genus Paranura Axelson, 1902 key
Species anops Christiansen, K et Bellinger, P, 1980:346
Species bisetosa Deharveng, L, 1989:113
             Syn.: Paranura Bisetosa(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species chiangdaoensis Deharveng, L, 1989:105
             Syn.: Paranura Chiangdaoensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species coenobita Cassagnau, 1991
             Syn.: Paranura cenobita(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:685 ISS
             Syn.: Paranura coenotiba(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:685 ISS
Species colombiana Palacios-Vargas, JG et Peñaranda-Parada, MR, 2005:308
Species colorata Mills, HB, 1934:28
             Syn.: Paranura 6-punctata Mills, HB, 1934:128 ISS
            Syn.: Paranura sexpunctata Neartic records
             Syn.: Paranura Colorata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Paranura colorata from the USA
2016.12.12 © Dobbs, Br.
Paranura colorata from the USA
Pointed mouthcone
2016.12.29 © Dobbs, Br.

Species dalgeri Deharveng, L, 1989:111
Species garoensis Cassagnau, 1991
Species globulifer Deharveng, L, 1989:118
Species godavarica Cassagnau, 1991
Species ieti (Yosii, 1966) Deharveng, L, 1989:104
            Syn.: Paranura najtae Deharveng, L et Weiner, WM, 1984:11
             Syn.: Paranura Ieti(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species impedita Palacios-Vargas, JG et Deharveng, L, 1987:111
Species johorensis (Yosii, 1976) Deharveng, L, 1989:104
Species jorgei Palacios-Vargas, JG et Deharveng, L, 1987:106
             Syn.: Paranura Jorgei(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species koryoi Deharveng, L et Weiner, WM, 1984:7
             Syn.: Paranura Koryoi(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species leclerci Deharveng, L, 1989:115
             Syn.: Paranura Leclerci(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species longisensillata Palacios-Vargas, JG et Deharveng, L, 1987:111
             Syn.: Paranura Longisensillata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species magdalenae Simón-Benito, JC et Palacios-Vargas, JG, 2008:685
             Syn.: Paranura Magdalenae(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species meo Deharveng, L, 1989:118
Species mjohjangensis Deharveng, L et Weiner, WM, 1984:9
             Syn.: Paranura Mjohjangensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species modesta Deharveng, L, 1989:105
Species nalo Christiansen, K et Bellinger, P, 1992:68
Species nudifera Yoshii, R et Suhardjono, YR, 1992:8
Species quadrilobata Hammer, 1953
Species quadripunctata Yosii, 1966
Species rooensis Simón-Benito, JC et Palacios-Vargas, JG, 2008:683
             Syn.: Paranura Rooensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species rosea Lee, B-H et Kim, J-T, 1984:20
Species sarukhani Palacios-Vargas, JG et Deharveng, L, 1987:106
             Syn.: Paranura Sarukhani(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species sexpunctata Axelson, 1902, t.t.
            Syn.: Börneria(sic) clavisetis Axelson, 1902
             Syn.: Paranura sexpunctata var. claviseta(sic) Womersley, H, 1935:208 ISS
            Syn.: Paranura clavisetis var. quadrimaculata Linnaniemi, 1907
            Syn.: Paranura sexpunctata ssp. colorata Mills, 1934
             Syn.: Paranura Sexpunctata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Species sitchensis Fjellberg, 1985
Species squamosa Cassagnau, 1991
Species tamul Cassagnau, P, 1988:25
Species tapatia Palacios-Vargas, JG et Peñaranda-Parada, MR, 2005:311
Species tibiotarsalis Deharveng, L, 1989:110
Species timorensis Yoshii, R et Suhardjono, YR, 1992:76
Paranura timorensis from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.1F

Species tridentata Lee, B-H et Kim, J-T, 1984:19

Genus Oregonanura Smolis, 2008
Species cascadensis Smolis, 2008 tt

Tribus Sensillanurini Cassagnau P, 1983:10 key
            Syn.: ... Neanurini Massoud, Z, 1967:298 partim
             Syn.: Neanurinae Sensillanurine(sic) Palacios-Vargas & Najt, 1986 ISS
             Syn.: Neanurinae Sensillanurine(sic) Palacios-Vargas, JG, 1996:822 ISS
             Syn.: Neanurinae Sensillanurina(sic) Palacios-Vargas, JG et Ríos, G, 1996:442 ISS
Genus Sensillanura Deharveng, 1981 key
Species sp.1 Palacios-Vargas, JG, 1990:19
Species austriaca (da Gama, 1963) Cassagnau, P, 1983:11
Species barberi (Handschin, 1928) Cassagnau & Palacios-Vargas, 1983, t.t.
            Syn.: Achorutes Barberi(sic) Handschin, 1928,tt IOS
            Syn.: Neanura quadrioculata Guthrie, JE, 1903:101
            Syn.: Biloba pseudoquadrioculata Stach, J, 1951:31
Sensillanura barberi from the USA
2008.11.10 © Justis, S.
Sensillanura barberi from the USA
2008.11.10 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi juv. from the USA
2012.02.25 © Murray, T.
Sensillanura barberi from the USA
2012.12.18 © Montgomery, G.

Species bullsa (Wray, 1953) Cassagnau & Palacios-Vargas, 1983
             Syn.: Sensillanura bulsa(sic) Palacios-Vargas, JG et Catalán, E, 2010:162 ISS
Species caeca (Folsom, 1916) Cassagnau & Palacios-Vargas, 1983
             Syn.: Sensillanura coeca(sic) Cassagnau & Palacios-Vargas, 1983 ISS
Species illina (Christiansen & Bellinger, 1980:378) Cassagnau & Palacios-Vargas, 1983
Species millsi (Christiansen & Bellinger, 1980:349) Cassagnau & Palacios-Vargas, 1983
Species santizaragozai Palacios-Vargas, JG et Catalán, E, 2010:161

Genus Americanura Cassagnau, P, 1983:11 key
  Syn.: ... Mexicanura Palacios-Vargas, 1983 nn
  Syn.: Sensillanurini Palmanura Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:1 =?
Remark by Ellis, W.N. & Bellinger, P.F. 1984:2: Both the generic name and that of the type-species are attributed to a paper in print by Cassagnau & Palacios-Vargas (1983). As the published information contains a nomenclaturally valid diagnosis of the new genus (and by implication the type-species as well), the publication of Cassagnau (1983) takes precedence.

  Syn.: Neanurinae Americaneura(sic) Peck, SB, 2001:106 ISS
  Syn.: Collembola Americaneura(sic) Peck, SB et Jacquemart, J, 2011:4 ISS
Species banksi (Denis, 1933) Cassagnau & Palacios-Vargas, 1983
Species bara (Christiansen, K & Bellinger, P, 1980:373) Palacios-Vargas, JG & Najt, J, 1986:24
Americanura bara? from the USA
2017.03.26 © Lee, C.

Species basseti Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:439
Americanura basseti from Panama
After Palacios-Vargas & al, 2009 Fig.14-15

Species castagnorum Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:439
Americanura castagnorum from Panama
After Palacios-Vargas & al, 2009 Fig.12-13

Species castillorum Palacios-Vargas & Najt, 1986:15
Species guatemalteca Cassagnau & Palacios-Vargas, 1983:2
Species imitator Cassagnau & Palacios-Vargas, 1983:2
Species interrogator Cassagnau & Palacios-Vargas, 1983:2
Species izabalana Palacios-Vargas, JG et Ríos, G, 1996:440
Species iztac Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:437
Americanura iztac from Mexico
After Palacios-Vargas & al, 2009 Fig.10-11

Species janssensi Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:435
Americanura janssensi from Mexico
After Palacios-Vargas & al, 2009 Fig.8-9

Species macgregori Cassagnau & Palacios-Vargas, 1983:2
Species medellini Palacios-Vargas, JG et Ríos, G, 1996:437
Species mexicana Cassagnau, P, 1983:11, t.t.
Species najtae Arango, A, Villagomez, F et Palacios-Vargas, JG, 2017:52
Americanura najtae from Mexico
After Arango A, Villagomez F & Palacios-Vargas JG, 2017 Fig.1AC

Species nova (Christiansen, K & Reddell, JR, 1986:158) Palacios-Vargas, JG & Najt, J, 1986:23
Species palaciosi Paniagua Nucamendi, J, 2012:88
Species prima Cassagnau, P et Palacios-Vargas, JG, 1983:6
Species sardinasensis Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:433
Americanura sardinasensis from Mexico
After Palacios-Vargas & al, 2009 Fig.1-2

Species setafoliacea Cassagnau & Palacios-Vargas, 1983:2
             Syn.: Americanura setaefoliacea(sic) Palacios-Vargas, 1996 ISS
             Syn.: Americanura setaefoliacea(sic) Palacios-Vargas & Ríos, 1996 ISS
Species sotanophila Cassagnau & Palacios-Vargas, 1983:2
Species unguimitl Palacios-Vargas & Najt, 1986:11
             Syn.: Americanura unguimitli(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:360 ISS

Genus Palmanura Cassagnau, P, 1983:11 key
Species colotlipa Palacios-Vargas, JG, 1996:820
Species curta Palacios-Vargas, JG, 1996:812
Species goyai Palacios-Vargas, JG et Simón-Benito, JC, 2009:268
Palmanura goyai from Mexico
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.6,9
            Syn.: Palmanura sp.2 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2

Species guilleni Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:73
Species hieronimi Palacios-Vargas, JG et Simón-Benito, JC, 2009:268
            Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:268 IOS
Palmanura hieronimus(sic) from Mexico
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.12
            Syn.: Palmanura sp.3 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2
             Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:10 ISS
             Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:74 ISS
Species lacandona Palacios-Vargas, JG, 1996:805
            Syn.: Palmanura lacandonicus(sic) Palacios-Vargas, JG, 1996:805 IOS
            Syn.: Palmanura sp. Palacios-Vargas, JG et Ríos-Saís, GA, 1995:266
             Syn.: Palmanura lancadona(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:269 ISS
Species lindquisti Palacios-Vargas, JG, 1996:807
Species matildae Palacios-Vargas, JG, 1996:806
             Syn.: Palmanura matildeae(sic) Palacios-Vargas, JG, 1996:806 ISS
Species mirabilis Cassagnau, P, 1983:11, t.t.
            Syn.: ... sp. Lavelle, Maury & Serrano, 1981
  Subspecies longa Palacios-Vargas, JG, 1996:821
Species normae Palacios-Vargas, JG, 1996:818
Species pascuali Palacios-Vargas, JG, 1996:814
Species primigenia Palacios-Vargas, JG, 1986:107
Species quetzalae Palacios-Vargas, JG, 1984:4
Species saham Vázquez, MM et Palacios-Vargas, JG, 2005:75
Species schusteri Palacios-Vargas, JG, 1996:809
Species sernai Palacios-Vargas, JG et Simón-Benito, JC, 2009:266
            Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:266 IOS
Palmanura sernaius(sic) from Colombia
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.4-5
            Syn.: Palmanura sp.1 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2
            Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:10
             Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:74 ISS
Species wilsoni Palacios-Vargas, JG, 1996:816
             Syn.: Palmanura wilson(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:270 ISS

Genus Tabasconura Palacios-Vargas, JG et Catalán, E, 2015:131
Species tapijulapana Palacios-Vargas, JG et Catalán, E, 2015:137, t.t.
            Syn.: Tabasconura tapajulapana(sic) Palacios-Vargas, JG et Catalán, E, 2015:137 IOS
             Syn.: Tabasconura tapiculapana(sic) Palacios-Vargas, JG et Catalán, E, 2015:137 ISS


References