http://www.collembola.org/taxa/neelidae.htm - Last updated on 2019.01.28 by Frans Janssens
Checklist of the Collembola: Neelidae

Familia Neelidae Folsom JW, 1896:391 key

Genus Number
of species
Acanthoneelidus1
Megalothorax34
Neelides7
Neelus13
Spinaethorax3
Zelandothorax1
Neelidae59
Genera with number of species
  Syn.: Sminthuridae Megalothoracini Börner, 1900


Neelus? sp.
2005 © Cheung, D. & Schmidt, J.
Megalothorax? sp. nov. from the UK
2006 © Brocklehurst, K.
Neelidae from the UK
2008.04.13 © Kilford., B
Neelidae from Hungary
2010.03.09 © Pfliegler, W.


Genus Acanthoneelidus Bretfeld, G et Griegel, A, 2006:46

  Syn.: Neelidae Acanthothorax Bretfeld, G et Griegel, A, 1999:212
  Syn.: Neelidae Acanthothorax Bretfeld & Griegel, 1999, nec Gaede, 1832
  Syn.: Neelidae Acanthothorax Bretfeld & Griegel, 1999, nec Costa, 1839


Species pratensis (Bretfeld, G & Griegel, A, 1999:212) Bretfeld, G & Griegel, A, 2006:46, t.t.

Acanthoneelidus pratensis
Outline of 3rd foot
After Schneider C, 2017 in Schneider C & Deharveng L, 2017 Fig.9L


Genus Neelus Folsom, JW, 1896:391 key  key

Remark by ICZN. 2007:544: Placed on the Official List of Names in Zoology. Opinion 291.

Species sp.1 Papác, V, Lukic, M et Kovác, L, 2016:71

Species sp.2 Papác, V, Lukic, M et Kovác, L, 2016:71

Species sp.3 Papác, V, Lukic, M et Kovác, L, 2016:71

Species sp.4 Schneider, C, Zon, SD et D'Haese, CA, 2018:6

Species sp.5 Schneider, C, Zon, SD et D'Haese, CA, 2018:6

Species cvitanovici Papác, V, Lukic, M et Kovác, L, 2016:52

Neelus cvitanovici from Croatia
After Papác V, Lukic M & Kovác L, 2016 Fig.2

Species desantisi Najt, 1971

Species fimbriatus Bretfeld & Trinklein, 2000

             Syn.: Neelus fimriatus(sic) Papác, V et Palacios-Vargas, JG, 2016:50 ISS

Species klisurensis Kovác, L et Papác, V, 2010:44

Species koseli Kovác, L et Papác, V, 2010:37

            Syn.: Neelus sp. Kovác & Krchová, 2007


Neelus koseli from Slovakia
2012 © Kovác, L.

Species labralisetosus Massoud & Vannier, 1967

Species lackovici Papác, V, Lukic, M et Kovác, L, 2016:61

Neelus lackovici from Croatia
After Papác V, Lukic M & Kovác L, 2016 Fig.3

Species murinus Folsom, JW, 1896:391, n.c., t.t., B

            Syn.: Neelus minimus Womersley, 1932
            Syn.: Megalothorax minimus Womersley, 1939
            Syn.: Megalothorax murinus f. bolivari Bonet, 1944


Neelus murinus from the USA
After Folsom JW, 1896 Plate 8 Fig.5
Neelus murinus from the USA
Sagittal section
After Folsom JW, 1901 Plate 2 Fig.2
Neelus murinus from the UK
Antenna
2005 © Hopkin, S.P.
Neelus murinus from Belgium
2004 © Janssens, F.
Neelus murinus from the UK
2012.03.20 © Nurcombe, E.
Neelus murinus from the UK
2012.03.20 © Nurcombe, E.
Neelus murinus from the UK
2012.08.21 © Murray, A.
Neelus murinus from the UK
2012.08.21 © Murray, A.
Neelus murinus from the UK
Note tubular wax secretions
2012.08.21 © Murray, A.
Neelus murinus from the UK
2012.09.05 © Murray, A.
Neelus murinus from the UK
2012.10.03 © Murray, A.
Neelus murinus from the UK
2013.01.06 © Murray, A.
Neelus murinus from the UK
2013.01.06 © Murray, A.
Neelus murinus from the UK
2013.09.08 © Murray, A.
Neelus murinus from the UK
Furca with long slender mucrones
2013.12.10 © Nurcombe, E.
Neelus murinus from the UK
2013.12.12 © Nurcombe, E.
Neelus murinus from the UK
Wax tubes
2013.12.12 © Nurcombe, E.
Neelus murinus from the UK
Cephalic sensillar pits
2013.12.12 © Nurcombe, E.
Neelus murinus from Tasmania
2014.03.05 © Murray, A.
Neelus murinus? from Japan
2016.10.22 © Hara, Y.
Neelus murinus? from Japan
Long sharply pointed mucrones
2016.10.22 © Hara, Y.
Neelus murinus? from Japan
Long and narrow ant.3+4
2016.10.22 © Hara, Y.
Neelus murinus from France
2016.12.17 © Garcelon, P.
Neelus murinus from France
2017.01.28 © Picard, J.
Neelus murinus
Outline of 3rd foot
After Schneider C, 2017 in Schneider C & Deharveng L, 2017 Fig.9H
Neelus murinus from France
2018.01.06 © Picard, J.
Neelus murinus from France
2018.01.06 © Picard, J.
Neelus murinus from France
Distinctly slender antenna
2018.01.06 © Picard, J.
Neelus murinus from France
2018.01.10 © Garcelon, P.
Neelus murinus from France
Distinctly slender antenna
2018.02.08 © Garcelon, P.
Neelus murinus from France
2019.01.26 © Garcelon, P.


Genus Neelides Caroli, 1912

Species sp.1 Greenslade P, 1994

Species sp.2 Janssens F & Murray A, 2014

Neelides "gigeri" from Tasmania
2014.03.09 © Murray, A.
Neelides "gigeri" from Tasmania
2014.03.09 © Murray, A.

Species bisetosus Bretfeld & Trinklein, 2000

Species dianae (Christiansen, K & Bellinger, P, 1981:1050) Cutz-Pool, LQ, Palacios-Vargas, JG & Vázquez, M, 2003:98

Species folsomi Caroli, 1912:2, t.t.

Neelides folsomi
Outline of 3rd foot
After Schneider C, 2017 in Schneider C & Deharveng L, 2017 Fig.9K

Species minutus (Folsom, JW, 1901:221) Bonet, 1947

Neelus minutus from the USA
After Folsom JW, 1901 Plate 2 Fig.3
            Syn.: Neelus sp. Becker, 1905
            Syn.: Neelides folsomi Caroli, 1912
            Syn.: Neelides folsomi Yosii, 1965
            Syn.: Neelides folsomi Yosii, R, 1977:163



Neelides minutus from the USA
2006 © Bernard, E.C.
Neelides minutus from the UK
Antenna
2005 © Hopkin, S.P.
Neelides minutus from Germany
2014.10.19 © Landgraf, A.
Neelides minutus from Germany
2014.10.19 © Landgraf, A.
Neelides minutus from the UK
2015.02.12 © Murray, A.
Neelides minutus from the UK
Characteristic shape of antennae
2015.02.12 © Murray, A.
Neelides minutus from the UK
Middorsal longitudinal stripe
2015.02.12 © Murray, A.
Neelides minutus from the UK
Ant.4 thinner then ant.3
2015.03.19 © Murray, A.
Neelides minutus from the UK
2015.03.19 © Murray, A.
Neelides minutus from the UK
Subadult
2015.03.19 © Murray, A.
Neelides minutus from Canada
2015.07.16 © Murray, A.

Species snideri Bernard, EC, 1975:183

Neelides snideri from the USA
After Bernard EC, 1975 Fig.1


Genus Megalothorax Willem, V, 1900:7 key  key

  Syn.: ... Neelus Folsom, 1901, partim
  Syn.: ... Neelus Womersley, 1932
  Syn.: Neelidae Amerus Collinge, WE et Shoebotham, JW, 1909:46 nec Berlese, 1883


Megalothorax sp. from Australia
1999 © Walter, D.E.
Megalothorax? sp. nov. from the UK
2006.11.01 © Brocklehurst, K.




  Syn.: Neelidae Megolothorax(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS


  Syn.: Neelidae megalothorax(sic) Schneider, C, Zon, SD et D'Haese, CA, 2018:21 ISS

Species sp.1 Schneider, C, Porco, D et Deharveng, L, 2016:62

            Syn.: Megalothorax sp.ARG.2 Schneider, C, Zon, SD et D'Haese, CA, 2018:19
            Syn.: Megalothorax sp.CHL.1 Schneider, C, Zon, SD et D'Haese, CA, 2018:19

Species sp.2 Schneider, C, Porco, D et Deharveng, L, 2016:64


Species-group minimus Schneider C & D'Haese C, 2013:361
Species aquaticus Stach, 1951

Species carpaticus Papác, V et Kovác, L, 2013:551

            Syn.: Megalothorax sp.nov. Kovác & al., 2006
            Syn.: Megalothorax sp.2 Papác & Kovác, 2012

Species minimus Willem, V, 1900:9, t.t., B

            Syn.: Amerus normani Collinge, WE et Shoebotham, JW, 1909:47
Amerus normani from the UK
After Collinge WE & Shoebotham JW, 1909 Pl.VI Fig.1

             Syn.: Amerus normanni(sic) Gisin, H, 1960:264 ISS
            Syn.: Megalothorax tristani Bonet, 1944
            Syn.: Neelus (Megalothorax) maculosus Maynard, EA, 1951:202
Neelus (Megalothorax) maculosus from the USA
After Maynard EA, 1951 Pl.XXII Fig.356




Megalothorax minimus
After Börner, C., 1906
Megalothorax minimus
After Börner, C., 1906
Neelus minimus from Australia
After Womersley H, 1932 Fig.1a
Megalothorax minimus from the UK
2005 © Hopkin, S.P.
Megalothorax minimus from the UK
In front of Pogonognathellus longicornis
2005 © Hopkin, S.P.
Megalothorax minimus from Sweden
2006 © Hall, K.
Megalothorax minimus from Sweden
2006 © Hall, K.
Megalothorax minimus from Sweden
2006 © Hall, K.

             Syn.: Megalothorax miniumus(sic) Fjellberg, A, Nygaard, PH et Stabbetorp, OE in Halldorsson, G, Oddsdottir, ES et Eggertsson, O, 2007:119 ISS
Megalothorax minimus from the USA
With diverticulate gut
2011.01.07 © Gerber, G.K.

             Syn.: Megolothorax(sic) mimimus(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS
Megalothorax minimus juv. from the UK
2012.06.24 © Murray, A.
Megalothorax minimus from the UK
2012.06.24 © Murray, A.
Megalothorax minimus from the UK
2012.06.24 © Murray, A.
Megalothorax minimus from the UK
2012.06.24 © Murray, A.
Megalothorax minimus from the UK
2012.06.26 © Murray, A.
Megalothorax minimus from the UK
2012.07.08 © Nurcombe, E.
Megalothorax minimus from the UK
2012.07.10 © Murray, A.
Megalothorax minimus from the UK
2012.07.10 © Murray, A.
Megalothorax minimus from the UK
2012.11.18 © Lewington, A.
Megalothorax minimus from Belgium (bar=0.1mm)
After Schneider C & D'Haese CA, 2013 Fig.1D
Megalothorax minimus from Belgium
Abdominal sensory field with T-shaped sensillum (bar=0.01mm)
After Schneider C & D'Haese CA, 2013 Fig.3F
Megalothorax minimus from France (bar=0.1mm)
After Schneider C & D'Haese CA, 2013 Fig.1C
Megalothorax minimus from France (bar=0.1mm)
After Schneider C & D'Haese CA, 2013 Fig.1F
Megalothorax minimus juv. from the UK
Pre-jump posture: uplifted prolegs
2013.01.12 © Nurcombe, E.
Megalothorax minimus from the UK
2013.01.19 © Murray, A.
Megalothorax minimus from the UK
Mesothoracic, metacoxal and 4th abdominal sensory pits
2013.04.02 © Murray, A.
Megalothorax minimus from the UK
2013.09.07 © Nurcombe, E.
Megalothorax minimus from Sweden
2014.01.21 © Hall, K.
Megalothorax minimus juv. from Canada
2014.02.01 © Hochhalter, R.
Megalothorax minimus juv. from Canada
Feeding instar size 0.21 mm
2014.02.01 © Hochhalter, R.
Megalothorax minimus juv. from Canada
Feeding instar size 0.23 mm
2014.02.01 © Hochhalter, R.
Megalothorax minimus from Holland
Feeding instar
2014.05.04 © van Duinen, J.
Megalothorax minimus juv. from Germany
2014.09.27 © Beck, K.
Megalothorax minimus from the UK
2015.03.14 © Murray, A.
Megalothorax minimus juv. from the UK
With diverticulate gut
2015.03.24 © Garlick, T.
Megalothorax minimus from the USA
2015.05.06 © Dankowicz, E.
Megalothorax minimus from Belgium
2015.10.29 © Janssens, F.
Megalothorax minimus from Belgium
2016.09.21 © Huskens, M.-L.
Megalothorax minimus
Outline of 3rd foot
After Schneider C, 2017 in Schneider C & Deharveng L, 2017 Fig.9I
Megalothorax minimus from France
2018.01.15 © Picard, J.



Species potapovi Schneider, C, Porco, D et Deharveng, L, 2016:41

Megalothorax potapovi from FE Russia
After Schneider C, Porco D & Deharveng L, 2016 Fig.1A
Megalothorax potapovi from FE Russia
Head dorsal (left), ventral (right)
After Schneider C, Porco D & Deharveng L, 2016 Fig.14AB

Species sanctistephani Christian, E, 1998:15

Species sanguineus Schneider, C, Porco, D et Deharveng, L, 2016:50

Megalothorax sanguineus from France
Head dorsal (left), ventral (right)
After Schneider C, Porco D & Deharveng L, 2016 Fig.14CD

Species svalbardensis Schneider, C et D'Haese, CA, 2013:345

Megalothorax svalbardensis from Svalbard (bar=0.1mm)
After Schneider C & D'Haese CA, 2013 Fig.1B

Species tatrensis Papác, V et Kovác, L, 2013:546

            Syn.: Megalothorax sp.n. Mock & al., 2003
            Syn.: Megalothorax sp.1 Papác & Kovác, 2012

Species tuberculatus Deharveng & Beruete, 1993

Species willemi Schneider, C et D'Haese, CA, 2013:347, B


Species-group incertus Schneider C & D'Haese C, 2013:361
Species sp.3 Schneider, C, Porco, D et Deharveng, L, 2016:62

            Syn.: Megalothorax (incertus-group) sp.USA.1 Schneider, C, Zon, SD et D'Haese, CA, 2018:19

Species sp.4 Schneider, C, Porco, D et Deharveng, L, 2016:62

            Syn.: Megalothorax (incertus-group) sp.ARG.1 Schneider, C, Zon, SD et D'Haese, CA, 2018:19

Species sp.5 Schneider, C, Porco, D et Deharveng, L, 2016:62

            Syn.: Megalothorax (incertus-group) sp.FRA.8 Schneider, C, Zon, SD et D'Haese, CA, 2018:19

Species draco Papác, V et Kovác, L, 2013:562

            Syn.: Megalothorax sp.4 Papác & Kovác, 2012


Species hipmani Papác, V et Kovác, L, 2013:557

            Syn.: Megalothorax incertus Mock & al., 2002 cf
            Syn.: Megalothorax sp.1 Papác, 2009
            Syn.: Megalothorax sp.1 Papác, 2011
            Syn.: Megalothorax sp.3 Papác & Kovác, 2012


Species interruptus Hüther, 1967

Species laevis Denis, 1948

            Syn.: Megalothorax minimus Deharveng & al, 2009
            Syn.: Megalothorax interruptus Schneider, C et D'Haese, CA, 2013:352,cf
Megalothorax cf interruptus from French Guiana (bar=0.1mm)
After Schneider C & D'Haese CA, 2013 Fig.1A




Megalothorax laevis from Ivory Coast
After Schneider C, Zon SD & D'Haese CA, 2018 Fig.2A
Megalothorax laevis from Ivory Coast
Head dorsal (left), ventral (right)
After Schneider C, Zon SD & Deharveng L, 2018 Fig.11AB

Species massoudi Deharveng, 1978

Species nigropunctatus Schneider, C et D'Haese, CA, 2013:334

Species perspicillum Schneider, C et D'Haese, CA, 2013:340

Megalothorax perspicillum from France (bar=0.1mm)
Head extended
After Schneider C & D'Haese CA, 2013 Fig.1E

Species-group other Schneider C & D'Haese CA, 2013:359
Species granulosus Schneider, C et D'Haese, CA, 2013:333


Species-group unresolved Schneider C & D'Haese CA, 2013:359
Species albus (Maynard, EA, 1951:201) Stach, J, 1957:13, s.d.

Species australis Delamare Deboutteville, C et Massoud, Z, 1963:174

Species boneti Stach, J, 1960:548

Species gabonensis Massoud & Vannier, 1965

Species incertus Börner, C, 1903:160

            Syn.: Neelus swani Womersley, H, 1932:12
Neelus swani from Australia
After Womersley H, 1932 Fig.1f
Megalothorax swani from New Zealand
After Salmon JT, 1941 Pl.70 Fig.480

            Syn.: Megalothorax remyi Denis, 1933
             Syn.: Megalothorax Remyi(sic) Bonet, 1947 ISS
             Syn.: Megalothorax Remyi(sic) Delamare Deboutteville, 1963 ISS
            Syn.: Megalothorax incertoides Mills, HB, 1934:85
Megalothorax incertoides from the USA
After Mills, H.B., 1934 Plate 2 Fig.12

             Syn.: Megalothorax incertoïdes(sic) Schneider, C et D'Haese, CA, 2013:357 ISS
            Syn.: Megalothorax minimus Womersley, 1939



Megalothorax incertus juv. from New Zealand
2014.02.13 © Murray, A.
Megalothorax incertus juv. from New Zealand
2014.02.13 © Murray, A.
Megalothorax incertus juv. from New Zealand
2014.02.13 © Murray, A.

Species piloli (Christiansen, K & Bellinger, P, 1992:328) Schneider, C & D'Haese, CA, 2013:331

Species poki (Christiansen, K & Bellinger, P, 1992:330) Schneider, C & D'Haese, CA, 2013:331

Species rapoporti Salmon, JT, 1964:557

            Syn.: Megalothorax Boneti(sic) Rapoport, 1962, nec Stach, 1960
            Syn.: Megalothorax bonetella Najt & Rapoport, 1965


Species rubidus Salmon, JT, 1946:29

Megalothorax rubidus from New Zealand
2009 © Te Papa Museum

Species subtristani Massoud & Vannier, 1965

            Syn.: Megalothorax subtristani Delamare Deboutteville, 1948 nn
            Syn.: Megalothorax subtristani Delamare Deboutteville, 1951 nn


Species tristani Denis, 1933

             Syn.: Megalothorax Tristani(sic) Delamare Deboutteville & Massoud, 1963 ISS



Genus Spinaethorax Papác, V et Palacios-Vargas, JG, 2016:39

Species adamantis Schneider, C et Deharveng, L, 2017:4

Spinaethorax adamantis from Vietnam
After Schneider C & Deharveng L, 2017 Fig.1
Spinaethorax adamantis from Vietnam
Outline of 3rd foot
After Schneider C & Deharveng L, 2017 Fig.9G

Species spinotricosus (Palacios-Vargas, JG & Sánchez, A, 1999:56) Papác, V & Palacios-Vargas, JG, 2016:39, t.t.

            Syn.: Megalothorax sp.nov. Palacios-Vargas, JG et Gamboa-Vargas, JA, 1997:90
            Syn.: Megalothorax spinsus(sic) Palacios-Vargas, JG, 2006:9 lapsus


             Syn.: Spinaethorax spinotrichosus(sic) Papác, V et Palacios-Vargas, JG, 2016:47 ISS

Species tonoius (Palacios-Vargas, JG & Sánchez, A, 1999:60) Papác, V & Palacios-Vargas, JG, 2016:47


Genus Zelandothorax Delamare Deboutteville, C et Massoud, Z, 1963:172

Species novaezealandiae (Salmon, JT, 1944:172) Delamare Deboutteville, C & Massoud, Z, 1963:172, t.t.

            Syn.: Megalothorax novae-zealandiae(sic) Salmon, JT, 1944:172 IOS
Megalothorax novae-zealandiae(sic) from New Zealand
After Salmon JT, 1944 Pl.64 Fig.184




             Syn.: Megalothorax novae-zealandiae(sic) Salmon, JT, 1946:28 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Bonet, 1947 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Salmon, 1948 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Salmon, 1949 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Delamare Deboutteville, 1950 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Barlet, J, 1951:143 ISS
             Syn.: Megalothorax novae-zealandiae(sic) Stach, J, 1957:16 ISS
             Syn.: Zelandothorax novae-zealandiae(sic) Delamare Deboutteville, C et Massoud, Z, 1963:172,tt ISS
             Syn.: Megalothorax novae-zealandiae(sic) Salmon, JT, 1964:557 ISS
             Syn.: Megalothorax novozealandiae(sic) Papác, V et Palacios-Vargas, JG, 2016:49 ISS
Zelandothorax novaezealandiae from New Zealand
Outline of 3rd foot
After Bonet F, 1947 in Schneider C & Deharveng L, 2017 Fig.9J

References