http://www.collembola.org/taxa/paronell.htm - Last updated on 2021.04.07 by Frans Janssens
Checklist of the Collembola: Paronellinae

Subfamilia Paronellinae Börner C, 1906:162 sensu Zang F et al, 2019:252 key  key  key  key

Genus Number
of species
Bromacanthus12
Calobatinus4
Campylothorax14
Cephalophilus3
Cyphoderinus1
Cyphoderodes7
Cyphoderus82
Delamarerus2
Lepidonella16
Megacyphoderus5
Mimoderus4
Paracyphoderus1
Paronella3
Paronellidaegennov1
Paronellidaegennov21
Pseudocyphoderus4
Serroderus26
Trichorypha1
Troglobius4
Paronellinae191
Genera with number of species
  Syn.: Entomobryidae Paronelliae(sic) Mitra, SK, 1984:101 ISS
Paronellinae from China
2008.07.18 © NCode, A.
Paronellinae from Taiwan
2009.02.04 © Wu, S.
Paronellinae from Taiwan
2010.06.23 © Chien, H.-C.
Paronellinae from Australia
2012.06.23 © Kathy & Marissa.




Genus Paronellidaegennov Deharveng, L, Brehier, F, Bedos, A, Tian, M-Y, Li, Y-B, Zhang, F Qin, W-G et Tan, X-F, 2008:78

Species sp.1 Deharveng, L, Brehier, F, Bedos, A, Tian, M-Y, Li, Y-B, Zhang, F Qin, W-G et Tan, X-F, 2008:78

Paronellidae gen. nov. sp. nov. from China
After Deharveng L et al, 2008 Fig.5


Genus Paronellidaegennov2 Culver & Pipan, 2009

Species sp.1 Culver & Pipan, 2009


Tribus Paronellini Börner, 1906:162 sensu Zhang F et al, 2019:252 key

            Syn.: Paronellinae Troglopedetini Soto-Adames, FN, Jordana, R et Baquero, E, 2014:1


Paronellini from Indonesia
Composed eye with Extra Ocular Structure (EOS)
2013.09.21 © Bertner, P.
Paronellini from Indonesia
Hyaline oval apically rounded scales
2013.09.21 © Bertner, P.
Paronellini from Malaysia
Composed eye with Extra Ocular Structure (EOS)
2014.02.12 © Bertner, P.
Paronellini from Malaysia
Apically knobbed dorsal thoracic macrosetae
2014.02.12 © Bertner, P.
Paronellini from Malaysia
Collophore with extended vesicles pressed in the surface waterfilm (drinking)
2014.02.12 © Bertner, P.
Remark by Zhang, F & al. 2015:7: Molecular phylogeny supports the closer relationship of Cyphoderini and Troglopedetini [= Paronellini], both of which have been clustered with Lepidocyrtinae.
Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The presence of pseudopores behind the posterior row of setae on the fourth abdominal segment, as seen in Troglopedetes, Trogolaphysa, Cyphoderus and Troglobius, corroborates the close relationship of Cyphoderus ([Cyphoderini]) to Trogolaphysa (Paronellini).


Genus Paronella Schött, H, 1893:14 F

  Syn.: Paronellidae Trogonella Massoud in Wilson JM, 1982:218
  Syn.: Paronellidae Trogonella Wilson, JM, 1982:218
  Syn.: ... Trichorypha Thibaud, J-M et Najt, J, 1988:722
  Syn.: ... Trichorypha Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15
  Syn.: ... Microparonella Salmon, 1964


Remark by ICZN. 2007:608: Placed on the Official List of Names in Zoology. Opinion 291.

Species fusca Schött, H, 1893:16, t.t.

Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1A
Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1B
Paronella fusca
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.29

Species montana Delamare Deboutteville, 1950

Species purpurea Barra, 1969



Genus Campylothorax Schött, H, 1893:18 key

  Syn.: Dicranocentroides Dicranocentropha Wray, DL, 1953:144
  Syn.: Entomobryidae Dicranocentrapha(sic) Mari Mutt, JA, 1976:121 ISS


Campylothorax
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.33

  Syn.: Paronellini Campylothorx(sic) Mitra, SK, 1993:65 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:19 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:20 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:21 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:24 ISS
Campylothorax sp. from Mexico
2016.10.06 © Murray, A.
Campylothorax sabanus from Brazil
Modified after Mari Mutt, 1987 in Bellini BC & Cipola NG, 2017 Fig.2


  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:165 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:166 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:167 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:168 ISS


Species camelinus Womersley, 1930

            Syn.: Campylothorax camelina(sic) Womersley, 1930 IOS


             Syn.: Campylothorax camelina(sic) Mitra, SK et Dallai, R, 1980:275 ISS



Species cassagnaui Mitra, SK et Dallai, R, 1980:313

            Syn.: Campylothorax cassagnaus(sic) Mitra, SK et Dallai, R, 1980:313 IOS
            Syn.: Campylothorax cinctus Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:131
             Syn.: Campylothorax cincta(sic) Soto-Adames, FN, 2016:2 ISS


             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & Najt, 1995 ISS
             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1996:171 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & al., 1997 ISS
             Syn.: Campylothorax cassagnaus(sic) Culik, MP et Zeppelini, DF, 2003:1129 ISS
Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Leg. McAlister, E.; Det. Culik, M.
Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Mucro
Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.
Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.

             Syn.: Campylothorax cassagnaus(sic) da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:1 ISS
             Syn.: Campylothorax cassagnaus(sic) Brito, RA, de Lima, ECA, Ferreira, AS, de Oliveira, JVLC, Amorim, TPL, Machado, TG, Lopes, BCH, Costa, IA, Alves, JLS, Brito, NP, Zeppelini, D, 2017:11 ISS




Species cinctus Schött, 1927

Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.45A
Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.46A

Species cubanus Gruia, M, 1983:203

            Syn.: Campylothorax cubana(sic) Gruia, M, 1983:203 IOS
             Syn.: Campylothorax cubana(sic) Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:94 ISS
             Syn.: Camphylotorax(sic) cubana(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS
Campylothtorax(sic) cubanus from Haiti
After Soto-Adames FN, 2016 Fig.12A
Campylothtorax(sic) cubanus from Hispaniola
After Soto-Adames FN, 2016 Fig.12B





Species dominicanus Soto-Adames, FN, 2016:20

Campylothtorax(sic) dominicanus from Hispaniola
After Soto-Adames FN, 2016 Fig.10
             Syn.: Campylothorax domincanus(sic) Soto-Adames, FN, 2016:28 ISS


Species hexosetosus Soto-Adames, FN, 2016:16

Campylothorax hexosetosus from Hispaniola
After Soto-Adames FN, 2016 Fig.7
             Syn.: Campylothorax hexosetosa(sic) Soto-Adames, FN, 2016:16 ISS


Species longicornis Schött, H, 1893:20, t.t.

Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.1A
From syntype
Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.20
Mucro

Species melanocephalus Mitra, SK et Dallai, R, 1980:298

Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.29
Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.30

Species mitrai Bellini & Meneses, 2012

Species notidanus Soto-Adames, FN, 2016:11

Campylothorax notidanus from Hispaniola
After Soto-Adames FN, 2016 Fig.4A

Species plagatus Cipola, NG et de Lima Oliveira, FG, 2016:488

Campylothorax plagatus from Brazil
After Cipola NG & de Lima Oliveira FG, 2016 Fig.1

Species sabanus (Wray, DL, 1953:144) Mitra, 1975

            Syn.: Dicranocentroides (Dicranocentropha) sabana(sic) Wray, DL, 1953:144 IOS
            Syn.: Dicranocentroides fasciculatus Rapoport, 1971 lapsus
            Syn.: Dicranocentroides fasciculatus Mari Mutt, JA et Bellinger, PF, 1990:125


             Syn.: Campylothorax sabana(sic) Mitra, 1975 ISS
             Syn.: Campylothorax sabana(sic) Mitra, SK et Dallai, R, 1980:300 ISS
Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.32A
Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.33A

             Syn.: Campylothorax sabana(sic) Cipola, NG et de Lima Oliveira, FG, 2016:494 ISS




Species schaefferi Börner, C, 1906:179

            Syn.: Campylothorax schäfferi(sic) Börner, C, 1906:179 IOS

             Syn.: S(sic) Schäfferi(sic) Schött, H, 1903:XIII nn ISS


             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:2 ISS
             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:12 ISS
Campylothorax aff. schaefferi from Brazil
After Cipola NG & al, 2019 Fig.9





Species viruaensis Santos IP, Cipola NG et Bellini BC in da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:2

Campylothorax viruaensis from Brazil
After da Silva Santos IP, Cipola NG & Bellini BC in da Silva Santos IP & al, 2016 Fig.1



Tribus Cyphoderini Börner C, 1906:162 sensu Zhang F & al, 2019:252 key




             Syn.: Entomobryidae Cyphoderitae(sic) Delamare Deboutteville, C, 1945:48 ISS
             Syn.: Mydoniidae Cyphodurinae(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
Cyphoderus albinus from the UK
With Myrmica rubra
2007 © Cornwall, N.J.
Cyphoderus albinus from France
With Pheidole pallidula
2008.03.15 © Lebeaux, P.

             Syn.: Lepidocyrtidae Cyphoderidae(sic) Castaño-Meneses, G, Palacios-Vargas, JG, Delabie, JHC, Zeppelini, D et Mariano, CSF, 2017:741 ISS



Genus Cyphoderus Nicolet H, [1842:63], n.c. Opinion 291 key  key  key M

  Syn.: Podurelles (Corps linéaires) Cyphodeirus(sic) Nicolet H, 1842:63, nr Opinion291 IOS
  Syn.: ... Podura Fabricius, 1783 partim
  Syn.: ... Lepidocyrthus(sic) Bourlet ISS
  Syn.: Podurelles(sic) Cyphodurus Nicolet, H in Desor, E, 1841:384 nn
  Syn.: Thysanura Beckia Lubbock, J, 1870:279
  Syn.: ... Cyphoderella Börner, 1913 nn
  Syn.: ... Borecus Folsom, 1923
  Syn.: Cyphoderidae Cyphoda Delamare Deboutteville, C, 1948:360
  Syn.: Entomobryidae Cyphoda Gisin, H, 1960:258
  Syn.: Collembola Cyphodems(sic) da Gama, MM, 1988:74 ISS
  Syn.: Collembola Cyphodem(sic) da Gama, MM, 1988:84 ISS
  Syn.: Cyphoderinae Cyphoda Soto-Adames, FN, Barra, J-A, Christiansen, K et Jordana R, 2008:512
  Syn.: Cyphoderinae Cyphoda de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47
  Syn.: Cyphoderinae Cyphoda Cipola, NG, 2021:il
  Syn.: Troglopedetini Camponotus(sic) Salmon, JT, 1964:533 lapsus ISS


  Syn.: Podurellen(sic) Cyphodeirus(sic) Elditt, HL, 1854:40 ISS
  Syn.: Podura Cyphodeirus(sic) Elditt, HL, 1854:43 ISS
  Syn.: Collembola Cyphodeirus(sic) Ellis, WN et Bellinger, PF, 1973:14 Opinion IOS
Remark by ICZN. 2007:226: Emendation of Cyphodeirus. Placed on the Official List of Names in Zoology. Opinion 291.


  Syn.: Cyphoderidae Cyphyoderus(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS
Cyphoderus sp. from South Africa
Ant.3 as 'pearly bead' (*)
2020.04.25 © Robertson, H.





Species sp.1 Janssens F sensu Zeppelini & Castaño, 1995

            Syn.: Cyphoderus n.sp. Zeppelini Filho, D et Castaño Meneses, G, 1995:10


Species sp.2 (Janssens F sensu Arbea & Blasco-Zumeta, 2001) Soto-Adames & al, 2008

            Syn.: Cyphoda n.sp. Arbea, JI et Blasco-Zumeta, J, 2001:47


            Syn.: Cyphoderus sp. Soto-Adames & al, 2008

Species sp.3 Janssens F sensu Bellini & Zeppelini, 2004

            Syn.: Cyphoderus undescribed species Bellini, BC et Zeppelini, D, 2004:588


Species adelaideae Womersley, 1934

Species affinis Giard, 1895

Species africanus Womersley, 1929

Species agnotus Börner, C, 1906:180

             Syn.: Beckia albinos(sic) Parona, 1895 ISS
             Syn.: Beckia albinos(sic) Bonet, F, 1933:123 ISS
             Syn.: Beckia albinos(sic) Bonet, F, 1933:124 ISS


             Syn.: Cyphoderus aenotus(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:48 ISS
Cyphoderus cf. agnotus from the Galapagos
After Katz AD & al, 2016 Fig.21B



Species albinus Nicolet, H, 1842:67, t.t., B

            Syn.: Cyphodeirus(sic) albinos(sic) Nicolet, H, 1842:67,tt IOS
Cyphodeirus(sic) albinos(sic) from Switzerland
After Nicolet H., 1842 Pl.7 Fig.7

            Syn.: crystalpoduren Fabricius O, 1783
            Syn.: Lepidocyrtus argentatus Bourlet, C, 1845:67
            Syn.: Beckia argentea Lubbock, J, 1870:279,tt
Beckia argentea from the UK
After Lubbock J., 1873 Pl.23

             Syn.: Beckia argenta(sic) Tömösváry, 1883 ISS
             Syn.: Beckia argenta(sic) Dalla Torre, 1888 ISS
             Syn.: Cyphodeirus(sic) argentus(sic) Vellay, 1899 ISS
             Syn.: Cyphodeirus(sic) (Beckia) argenteus Vellay, 1900 ISS
            Syn.: Cyphoderus affinis Giard, 1895
            Syn.: Tullbergia immaculata Lie-Pettersen, 1896
            Syn.: Cyphodeirus(sic) albus Carl, 1899



             Syn.: Lepidocyrtus albinos(sic) Gervais in Walckenaër, 1844 ISS
             Syn.: Cyphodeirus(sic) Albinos(sic) Bourlet, C, 1845:67 ISS
             Syn.: Cyphoderus albinos(sic) Nicolet, H, 1847:383 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Elditt, HL, 1854:40 ISS
             Syn.: Lepidocyrtus albinos(sic) Lubbock, J, 1870:301 ISS
             Syn.: Beckia albinos(sic) Lubbock, J, 1870:279 ISS
             Syn.: Beckia albinos(sic) Lubbock, J, 1873:149 ISS
Beckia albinos(sic) from the UK
After Lubbock J, 1873 Pl.24
Beckia albinos(sic) from the UK
After Beck J in Lubbock J, 1873 Pl.72

             Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 ISS
             Syn.: Beckia albinos(sic) Parona, 1878 ISS
             Syn.: Beckia albinos(sic) Parona, 1882 ISS
             Syn.: Beckia albinos(sic) Tömösváry, 1883 ISS
             Syn.: Beckia albinos(sic) Brook, 1884 ISS
             Syn.: Beckia (Cyphoderus) albinos(sic) Reuter, 1886 ISS
             Syn.: Cyphoderus albinos(sic) Oudemans, 1887 ISS
             Syn.: Beckia albinos(sic) Parona, 1887 ISS
             Syn.: Beckia albinos(sic) Dalla Torre, 1888 ISS
             Syn.: Beckia albinos(sic) Parona, 1888 ISS
             Syn.: Cyphoderus albinos(sic) Uzel, 1890 ISS
             Syn.: Cyphoderus albinos(sic) Moniez, 1891 ISS
             Syn.: Cyphoderus albinos(sic) Moniez, 1891 ISS
             Syn.: Cyphoderus albinos(sic) Uzel, 1891 ISS
             Syn.: Beckia albinos(sic) Moniez, 1894 ISS
             Syn.: Cyphoderus albinos(sic) Lameere, A, 1895:591,B ISS
             Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1896 ISS
             Syn.: Cyphoderus albinos(sic) Poppe & Schäffer, 1897 ISS
             Syn.: Beckia albinos(sic) Parona, 1897 ISS
             Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1898 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 ISS
             Syn.: Cyphoderus albinos(sic) Carl, J, 1899:336 ISS
             Syn.: Cyphoderus albinos(sic) Carl, 1900 ISS
             Syn.: Cyphoderus albinos(sic) Börner, C, 1901:71 ISS
             Syn.: Cyphoderus albinos(sic) Börner, C, 1901:136,B ISS
             Syn.: Cyphoderus albinos(sic) Carl, 1901 ISS
             Syn.: Cyphoderus albinos(sic) Ågren, 1903 ISS
             Syn.: Cyphoderus albinos(sic) Axelson, 1907 ISS
             Syn.: Cyphoderus albinos(sic) Evans, 1908 ISS
             Syn.: Cyphoderus albinos(sic) ssp. borneri(sic) Schille, 1908 ISS
             Syn.: Cyphoderus albinos(sic) Collinge, WE et Shoebotham, JW, 1909:89 ISS
             Syn.: Cyphoderus albinos(sic) Bagnall, 1910 ISS
             Syn.: Cyphoderus albinos(sic) Bagnall, 1910 ISS
             Syn.: Cyphoderus albinos(sic) Axelson, 1911 ISS
             Syn.: Cyphoderus albinos(sic) Carpenter, GH, 1916:47 ISS
             Syn.: Cyphoderus albinos(sic) Womersley, 1923 ISS
             Syn.: Cyphoderus albinos(sic) Denis, 1924 ISS
             Syn.: Cyphoderus albinos(sic) Brown, 1926 ISS
             Syn.: Cyphodeirus(sic) albinos(sic) Coutelen, F, 1928:854 ISS
             Syn.: Cyphoderus albinos(sic) Turk, 1938 ISS
             Syn.: Cyphoderus albinos(sic) Geyskens, J, 1945:16,B ISS
             Syn.: Cyphodurus albinos(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
             Syn.: Cyphoderus albinos(sic) Chopard, 1955 ISS
             Syn.: Camponotus(sic) albinos(sic) Salmon, JT, 1964:533 ISS
             Syn.: Cyphoderus albidus(sic) Berg, MP, 2001:xiv ISS
             Syn.: Cyphoderus albinos(sic) Hopkin, SP, 2002:1 ISS
             Syn.: Cyphoderus albinos(sic) Henderickx, H et Vets, V, 2003:9 ISS
Cyphoderus albinus from the UK
2007 © Cornwall, N.J.
Cyphoderus albinus from the UK
2007 © Cornwall, N.J.
Cyphoderus albinus from France
2008.03.15 © Lebeaux, P.
Cyphoderus albinus from Czechia
2008.03.28 © Krásenský, P.
Cyphoderus albinus from Germany
With Platyarthrus hoffmannseggi
2009.04.22 © Schlüter, H.
Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.
 
Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.
Cyphoderus albinus from Sweden
2011.04.11 © Engström, A.
Cyphoderus albinus from France
2012.08.07 © Quintin, C.
Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.
Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.
Cyphoderus albinus from the UK
2012.11.27 © Murray, A.
Cyphoderus albinus from the UK
2012.11.27 © Murray, A.
Cyphoderus albinus juv. from the UK
2012.11.28 © Murray, A.
Cyphoderus albinus from the UK
With Lasius flavus
2012.11.28 © Murray, A.
Cyphoderus albinus from France
Ant.3 as 'pearly bead'
2013.05.12 © Quintin, C.
Cyphoderus albinus from France
Head in dorsal aspect semi-circular
2014.07.28 © Quintin, C.

             Syn.: Cyphoderus albinos(sic) Manu, M, Szekely, L, Oromulu, LV, Barbuceanu, D, Honciuc, V, Maican, S, Fiera, C, Purice, D et Ion, M, 2015:299 ISS
Cyphoderus albinus from Holland
2015.11.14 © van Leeuwen, J.

             Syn.: Cyphoderus albinos(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45,tt ISS
Cyphoderus albinus from Spain
2016.04.22 © Tizon, J.
Cyphoderus albinus from Belgium
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from Belgium
Ant.3 shaped as a 'pearly bead' (*)
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from Belgium
Frontal aspect
2018.05.27 © Huskens, M.-L.
Cyphoderus albinus from France
With Lasius flavus
2018.12.19 © Lebeaux, P.
Cyphoderus albinus from France
2019.01.14 © Tiky, S.
Cyphoderus albinus from the UK
2020.05.14 © Uncle, F.
Cyphoderus albinus from the UK
Ant.3 with narrow intersegmental rings (*)
2020.05.14 © Uncle, F.
Cyphoderus albinus from the UK
Furca
2020.05.14 © Uncle, F.
Cyphoderus albinus from the UK
Mucro distinctly narrow and long
Dens with distinctly large fringed scales
2020.05.14 © Uncle, F.
Cyphoderus albinus from the UK
Unguis & unguiculus with wing-like teeth
2020.05.14 © Uncle, F.
Cyphoderus albinus from the UK
Mucro apically bidentate
2020.05.14 © Uncle, F.
Cyphoderus albinus from France
2020.04.02 © Reinach, Br.
Cyphoderus albinus from France
Ant.3 with narrow intersegmental rings
2020.04.02 © Reinach, Br.
Cyphoderus albinus from France
2021.02.01 © Vol, F.

  Subspecies apatelus Börner, 1913


             Syn.: Cyphoderus albinus ssp. apaletus(sic) Jacquemart, S(+), 1980:2 ISS

  Subspecies boerneri Schille, 1906


            Syn.: Cyphoderus albinus ssp. Börneri(sic) Schille, 1906 IOS
            Syn.: Cyphoderus albinus Stach, 1919
            Syn.: Cyphoderus albinus Stach, 1922


             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1908 ISS
             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1912 ISS
             Syn.: Cyphoderus Börneri(sic) Börner, 1913 ISS
             Syn.: Cyphoderus albinus ssp. Börneri(sic) Delamare Deboutteville, C, 1948:342 ISS
             Syn.: Cyphoderus albinus f. Boerneri(sic) Cassagnau, 1963 ISS
             Syn.: Cyphoderus albinus ssp. börneri(sic) Jacquemart, S(+), 1980:2 ISS



Species amaranus Brown, JM, 1926:215

Species antennatus Delamare Deboutteville, C, 1945:39

Species antigone Fernando, 1959

Species arcuatus Wahlgren, E, 1906:20

Species arlei Cassagnau, 1963:138

            Syn.: Cyphoderus Arlei(sic) Cassagnau, 1963 IOS

Species asiaticus Yosii, R, 1959:56

Species assimilis Börner, C, 1906:181

            Syn.: Cyphoderus simulans Imms, 1912
            Syn.: Cyphoderus similis Denis, 1931
            Syn.: Cyphoderus subsimilis Paclt, 1959
            Syn.: Cyphoderus assimilis ssp. empodialis Nuñez, 1962


Species bidenticulatus (Parona, 1888) Börner, 1903

            Syn.: Cyphoderus bidenticulatus var. veneris Denis, 1924
            Syn.: Cyphoderus jugoslavicus Denis, 1936


             Syn.: Cyphoderus bidenticulata(sic) Handschin, 1942 ISS
             Syn.: Cyphoderus bidenticulati(sic) Delamare Deboutteville, 1948 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Jantarit & al, 2014 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:263,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:640,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:641 ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:3,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:5,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:6,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:8,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:9,gr ISS

Species bidenticulatus2 Womersley, 1934 nec Parona, 1888 sd

Species buxtoni Brown, 1920

            Syn.: Cyphoderus Buxtoni(sic) Brown, 1920 IOS


             Syn.: Cyphoderus Buxtoni(sic) Brown, 1926 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, C, 1945:36 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, 1948 ISS



Species caetetus Zeppelini, D et Oliveira, JVLC, 2016:561

Cyphoderus caetetus from Brazil
After Zeppelini D & Oliveira JVLC & al2016 Fig.2

Species canariensis da Gama, MM, 1988:82

Species colurus Börner, C in Schultze, 1908:57

             Syn.: Cyphoda colurus(sic) Delamare Deboutteville, 1948 ISS

Species cuthbertsoni Womersley, 1929

             Syn.: Cyphoderus Cuthbertsoni(sic) Delamare Deboutteville, C, 1945:36 ISS

Species dorsti Jacquemart, 1974 in Peck SB & Jacquemart J, 2011:1 u

Species empodialis Rapoport, 1962

Species equidenticulati Nunes, RC et Bellini, BC, 2018:72

Species folsomi Handschin, 1927

            Syn.: Cyphoderus Folsomi(sic) Handschin, 1927 IOS
            Syn.: Cyphoderus Folsomi(sic) Delamare Deboutteville, 1948 IOS

Species galapagoensis Jacquemart, S, 1976:153

Species ganeensis Tyagi & Baijal, 1979

Species genneserae Carpenter, 1913

Species gisini Gruia, 1967

Species grassei (Cassagnau, P & Delamare Deboutteville, C, 1955:384) Christiansen, K, 1957:79

            Syn.: Cyphoda Grassei(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:384 IOS

Species handschini (Delamare Deboutteville, C, 1948:346) Dallai, R, 1973:536

            Syn.: Cyphoderus Handschini Delamare Deboutteville, C, 1948:346 IOS
            Syn.: Cyphoderus assimilis Handschin, 1926




Species heymonsi Börner, 1906

            Syn.: Cyphoderus Heymonsi(sic) Börner, 1906 IOS
            Syn.: Cyphoderus Heymonsi(sic) Börner, 1913 IOS
            Syn.: Cyphoda Heymonsi(sic) Delamare Deboutteville, 1948 IOS

Species holonycha Silvestri, 1918

Species hrdyi Rusek, 1971

Species inaequalis Folsom, JW, 1927:12

Species indicus Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45

Cyphoderus indicus from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.45 Img.1

Species innominatus Mills, HB in Pearse, AS, 1938:186

             Syn.: Cyphoderus inomminatus(sic) Zeppelini, D et Oliveira, JVLC, 2016:568 ISS

Species insularum Carpenter, GH, 1916:47

             Syn.: Cyphoderus bidenticula Handschin, 1942, partim ISS

Species javanus Börner, C, 1906:180

            Syn.: Cyphoderus agnotus Handschin, 1942
            Syn.: Cyphoderus assimilis Yosii, 1953
            Syn.: Cyphoderus assimilis Yosii, R, 1977:162
            Syn.: Cyphoderus javanus f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143
            Syn.: Cyphoderus borneensis Yoshii, 1980


             Syn.: Cyphoderus javanicus(sic) Schött, H, 1917:51 ISS
Cyphoderus javanus from India
After Mandal GP & Hazra AK, 2009 Fig.41a



Species jharkhandensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:47

Cyphoderus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.47 Img.1

Species kawari Jacquemart, S, 1974:33

Species khaochakanus Jantarit, Satasook & Deharveng, 2014

             Syn.: Cyphoderus (bidenticulati(sic)-group) khaochanus(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:58 ISS

Species komareki Rusek, 1961

Species lantohi Yoshii, 1987

Species laticlavatus (Stach, J, 1960:545) Wilson, JM, 1982:218

             Syn.: Cyphoderopsis lavaticus(sic) Massoud in Wilson JM, 1982:218 lapsus ISS
             Syn.: Cyphoderopsis lavaticus(sic) Wilson, JM, 1982:218 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:218 ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:219 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Thibaud, J-M et Najt, J, 1988:724 lapsus ISS

Species limboxiphius Börner, 1913

            Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Rapoport & Izarra, 1962 IOS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Bernava-Laborde & Palacios-Vargas, 2008 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Salazar Martínez, AE, Narváez Beinhorn, MJ et Bernava Laborde, V, 2017:110 ISS




Species manuneru Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:259

Species maroccanus (Delamare Deboutteville, 1948:361) Jacquemart, 1974:43, t.t.

            Syn.: Cyphoda marocanus(sic) Delamare Deboutteville, 1948 IOS
             Syn.: Cyphoda marocanus(sic) Jacquemart, 1974 ISS
             Syn.: Cyphoderus marocanus(sic) Jacquemart, 1974 ISS



Species mitumbis Tshelnokov, 1977

Species monopterus Rusek, 1981

Species mucrominimus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:48

Species mucrostrimenus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:54

Species napoensis Thibaud, J-M et Najt, J, 1987:939

Species natalensis Börner, 1913

Species nichollsi Womersley, 1934

            Syn.: Cyphoderus Nichollsi(sic) Womersley, 1934 IOS

Species nigeri Jacquemart, 1980:1

Species nitricolus (Rapoport, 1962) Soto-Adames & al, 2008

Species omoensis Delamare Deboutteville, C, 1945:35

Species paralbinus Jacquemart, S(+), 1980:3

Species pinnatus (Folsom, 1923) Folsom, JW, 1927:13

Species propinquus Stach, 1963

Species pseudalbinus Schött, H, 1917:50

Species reuteri Loksa, I, 1978:61

Species rubiae Baijal, HN, 1955:536

Species saheli Jacquemart, 1980:3

Species sarojinii Bhattacharjee, 1985

Species similis Folsom, JW, 1927:12

            Syn.: Cyphoderus assimilis Folsom, JW, 1932:71
            Syn.: Cyphoderus similis f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143
            Syn.: Cyphoderus javanus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 =?


Cyphoderus similis from Canada
2007.04.29 © Luk, S.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.05.11 © Reed, D.
Cyphoderus similis from the USA
2007.09.03 © McClarin, J.
Cyphoderus similis from the USA
2007.09.03 © McClarin, J.
Cyphoderus similis from the USA
2008.04.05 © Murray, T.
Cyphoderus similis from the USA
2008.05.17 © Montgomery, G.
Cyphoderus similis from the USA
2009.12.06 © Justis, S.
Cyphoderus similis from the USA
2009.12.06 © Justis, S.
Cyphoderus similis from the USA
On Camponotus casteneus
2010.10.31 © Hart, J.
Cyphoderus similis from the USA
On Camponotus casteneus
2010.10.31 © Hart, J.
Cyphoderus similis from the USA
2011.12.11 © Kouri, J.
Cyphoderus similis from the USA
2013.05.12 © Liberta, A.
Cyphoderus similis from the USA
In nest of Tapinoma sessile
2015.12.07 © Keim., B
Cyphoderus similis from Canada
With Lasius sp.
2017.06.04 © Guay, P.
Cyphoderus similis from the USA
2019.12.29 © Joll, C.
Cyphoderus similis from the USA
With ants
2019.12.29 © Joll, C.
Cyphoderus similis from Canada
2020.02.21 © Strickland, O.
Cyphoderus similis from the USA
In ant nest
2020.09.28 © Shahan, T.

Species songkhlaensis Jantarit, Satasook & Deharveng, 2014

Species squamidives Silvestri, 1918

Species subserratus Delamare Deboutteville, 1948

Species subsimilis Delamare Deboutteville, C, 1948:346

Cyphoderus subsimilis from the Ivory Coast
2016.05.10 © Parmentier, T.
Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.
Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.
Distal dental scale as long as bidentate mucro
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Two unpair ungual teeth (*)
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Straight bidentate mucro
Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Dens with 4 internal and 6 external scales

Species sudanensis Wahlgren, E, 1906:18

Species sumatranus Yoshii, 1987

Species termitellus Delamare Deboutteville, C, 1948:339

Species trinervoides Paclt, 1965

Species variabilis Jacquemart, 1980:2

Species veneris Delamare Deboutteville, 1923

Species virungiensis Tshelnokov, 1977

Species yoshiiorum Thibaud, J-M et Najt, J, 1987:937

Species yugoslavicus Denis, 1933

Species zoroastris Yosii, 1963


Genus Pseudocyphoderus Imms, 1912 M

  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:132 ISS

Species annandalei Imms, 1912, t.t.

            Syn.: Pseudocyphoderus Annandalei(sic) Imms, 1912,tt IOS
             Syn.: Pseudocyphoderus Annandalei(sic) Börner, 1913 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Schött, H, 1917:2 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Delamare Deboutteville, C, 1948:391 ISS
Pseudocyphoderus annandalei from India
After Mandal GP & Hazra AK, 2009 Fig.45a

             Syn.: Pseudocyphoderus annendalei(sic) Mandal, GP et Hazra, AK, 2009:42 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:132 ISS




Species melittophilus Mari Mutt, JA et Bellinger, PF, 1990:130

             Syn.: Pseudocyphoderus melittophilus(sic) Mari Mutt, JA et Bellinger, PF, 1990:130 ISS
             Syn.: Pseudocyphoderus mellitophilous(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:52 ISS

Species squamicaudus Silvestri, 1917

Species wasmanni Börner, 1913

             Syn.: Pseudocyphoderus Wassmanni(sic) Delamare Deboutteville, 1948 ISS



Genus Cyphoderodes Silvestri, 1910 M

Species ceylonicus Silvestri, 1910

Species dubius Börner, 1913

Species mitrai Yoshii, 1987

Species punjabicus Yosii & Ashraf, 1964

Species singaporicus Murphy & Yoshii, 1992

Species xenopus (Börner, 1913) Culik, MP & Zeppelini, DF, 2003:1130

Species yayukae (Yoshii, 1987)


Genus Paracyphoderus Delamare Deboutteville, C, 1948:365 M

Species dimorphus (Silvestri, 1910) Delamare Deboutteville, 1948, t.t.


Genus Megacyphoderus Delamare Deboutteville, C, 1948:365 M

Species gallicus Delamare Deboutteville, 1948:368

Species oraniensis Delamare Deboutteville, 1948:366

Species orientalis (Folsom, 1924) Castaño-Meneses, G, Palacios-Vargas, JG & do Carmo, AFR, 2015:394

Species silvestrii (Denis, J-R, 1924:199) Delamare Deboutteville, C, 1948:367

            Syn.: Cyphoderus Silvestrii(sic) Denis, J-R, 1924:199 IOS


Species termitum (Wahlgren, E, 1906:19) Delamare Deboutteville, 1948


Genus Serroderus Delamare Deboutteville, C, 1948:369 M

Species sp.1 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Serrodus sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:20

Species acanthotermiensis Delamare Deboutteville, 1948:382

Species alfredi (Yosii, R, 1959:56) Yoshii, 1987

Species alvaradoi Selga, 1962

Species aurantius Delamare Deboutteville, 1948

Species bilobatus Delamare Deboutteville, 1948:375

Species brevimucronatus (Silvestri, 1917)

Species delamarei Cassagnau, 1963:147

Species dicuspiditermitis (Yoshii, 1980) Yayuk, 1989:18

Species distinctus (Denis, 1942) Delamare Deboutteville, 1948, t.t.

Species durio Yoshii, 1987

Species guanobia Paulian & Delamare Deboutteville, 1948

Species hozawai (Kinoshita, 1917) Yosii, R, 1977:162

Serroderus hozawai from Japan
2014.10.02 © Hara, Y.

Species humilis Delamare Deboutteville, 1948:371

  Varietas guanobia Delamare Deboutteville & Paulian, 1948

  Varietas aurantia Delamare Deboutteville, 1948

Species interpositus (Denis, 1942) Delamare Deboutteville, 1948

  Forma termitophila Delamare Deboutteville, 1948

Species mango Yoshii, 1992

Species multidentatus Delamare Deboutteville, 1948:356

Species protermiensis Delamare Deboutteville, 1948:381

Species quadridentatus (Delamare Deboutteville, 1948:355)

             Syn.: Cyphoderus quadridentalus Jacquemart, 1980 ISS

Species sabahnus (Yoshii, 1980) Yayuk, 1989:18

Species socialis Delamare Deboutteville, 1948:379

Species spinatus Christiansen, 1957:79

Species spinicaudus Delamare Deboutteville, 1948

Species sublimis Delamare Deboutteville, 1948:374

Species termitophilus Delamare Deboutteville, 1948

Species tridenticulatus Denis, 1948


Genus Cephalophilus Delamare Deboutteville, C, 1948:382 M

Species anoplotermiensis Barra, 1969

Species protermiensis Delamare Deboutteville, 1948:386

Species rectangularis Delamare Deboutteville, 1948:383, t.t.


Genus Calobatinus Silvestri, 1918 M

  Syn.: Calobatina Denis, 1942 Calobatina Denis, 1942 Calobatina Denis, 1942 ISS

Species erythraeus Silvestri, 1918

Species grassei (Denis, 1942) Delamare Deboutteville, 1948

            Syn.: Calobatina Grassei Denis, 1942 IOS
            Syn.: Calobatinus Grassei Delamare Deboutteville, 1948 IOS

Species occidentalis Silvestri, 1918

Species rhadinopus (Börner, 1913) Silvestri, 1918, t.t.


Genus Cyphoderinus Denis, 1942 M

Species fraterculus Denis, 1942, t.t.


Genus Mimoderus Yoshii, 1980 M

Species diusi Yoshii, 1980

Species novaeguinae (Uchida, 1949) Yayuk, RS (sic), 1989:18)

            Syn.: Cyphoderus novaguinae(sic) Uchida, 1949 IOS
            Syn.: Mimoderus novaguinae(sic) Yayuk, RS(sic), 1989:18) IOS

Species saikehi Yoshii, 1980

Species serratus (Schött, H, 1917:52) Greenslade, P in Houston, WWK, 1994:109


Genus Delamarerus Mitra, SK, 1976:645 M

  Syn.: Cyphoderidae Delamareus(sic) Mari Mutt, JA, 1977:230 ISS

Species immsi Mitra, SK, 1976:647, t.t.

Delamarerus immsi from India
After Mitra SK, 1976 Fig.1A
Delamarerus immsi from India
After Mandal GP & Hazra AK, 2009 Fig.46a

Species machadoi (Delamare Deboutteville, 1958) Mari Mutt, JA, 1977:234


Genus Troglobius Palacios-Vargas, JG et Wilson, J, 1990:68 key M

Species albertinoi Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2016:659

Troglobius albertinoi from Brazil
After Cipola NG & Bellini BC in Cipola NG & al, 2016 Fig.1
             Syn.: Troglobius albertonoi(sic) Cipola, NG, de Morais, JW et Bellini, BC, 2016:658 ISS


Species brasiliensis Palacios-Vargas, JG et Zeppelini, D, 1995:173

Species coprophagus Palacios-Vargas, JG et Wilson, J, 1990:69, t.t.

Species ferroicus Zeppelini, D, da Silva, DD et Palacios-Vargas, JG, 2014:3

Troglobius ferroicus from Brazil
Candle-shaped antennal sensilla
After Zeppelini D & al, 2014 Fig.1-2


Tribus Bromacanthini Mitra, 1993 sensu Zhang F & al, 2019:252 key

Remark by Mitra, S.K. 1993:62: General chaetal pattern of Lepidonella and Bromacanthus comes close to some genera under Lepidocyrtini.
             Syn.: Paronellinae Bromocanthini(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15 ISS
             Syn.: Paronellidae Bromocanthini(sic) Soto-Adames, FN et Bellini, BC, 2015:331 ISS
Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The tribe Bromacanthini is more similar to some Entomobryidae than to Paronellini+Cyphoderini.




Genus Lepidonella Yosii, 1960

  Syn.: Paronellini Microparonella Carpenter, GH, 1916:43


Lepidonella
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.37
Lepidonella sp. from Vietnam
2009 © Boldsystems.org
Lepidonella sp. from Vanuatu
2010 © Deharveng, L. & Bedos, A.
Lepidonella? sp. from New Zealand
2017.12.08 © Navest, M.P.
Lepidonella? sp. from New Zealand
2018.10.22 © Navest, M.P.

Species sp.1 Bellini, BC et Zeppelini, D, 2004:588

Species annulicornis (Oudemans, JT in Weber, M, 1890:87) Mitra, SK, 1992:212

Paronella annulicornis from Java
After Schött H, 1903 Taf.II Fig.5
Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.36
Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.37

Species ceylonica (Yosii, 1966) Deharveng, L & Bedos, A, 1995:22

Cyphoderopsis ceylonica from India
After Mandal GP & Hazra AK, 2009 Fig.43a
Lepidonella ceylonica from Ceylon
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.35

Species doveri (Carpenter, 1932) Deharveng & al, 2018

Species duodecimoculata (Prabhoo, 1971) Deharveng, L & Bedos, A, 1995:22

Lepidonella duodecimoculata from India
After Prabhoo in Soto-Adames FN & Bellini BC, 2015 Fig.31

Species flava (Carpenter, GH, 1916:44) Deharveng, L & Bedos, A, 1995:22

            Syn.: Paronella seychellesiensis Salmon, JT, 1964:545
             Syn.: Microparonella seychellesensis(sic) Yoshii, 1981 ISS
             Syn.: Microparonella seychellensis(sic) Yoshii, R et Suhardjno, YR, 1992:48 ISS
             Syn.: Paronella seychelliensis(sic) Nguyen, M et Soto-Adames, FN, 2018:193 ISS


Species incerta (Handschin, 1925) Mitra, SK, 1992:265

            Syn.: Lepidonella oudemansi Soto-Adames, FN et Bellini, BC, 2015:337 =?


Species kei (Yoshii, R & Suhardjno, YR, 1992:47) Deharveng, L & Bedos, A, 1995:22

Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A right
Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A left

             Syn.: Lepidonella key(sic) Soto-Adames, FN et Bellini, BC, 2015:339 ISS


Species lecongkieti Deharveng, L et Bedos, A, 1995:23

Species marimutti Soto-Adames, FN et Bellini, BC, 2015:334

            Syn.: Lepidonella marimuti(sic) Soto-Adames, FN et Bellini, BC, 2015:334 IOS
            Syn.: Microparonella incerta Mari Mutt, JA, 1987:404


             Syn.: Lepidonella marimuti(sic) Bellini, BC et Cipola, NG, 2017:176 ISS
             Syn.: Lepidonella marimuti(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS



Species nigrofasciata (Handschin, 1928) Deharveng, L & Bedos, A, 1995:22

Lepidonella nigrofasciata from Indonesia
After Handschin in Soto-Adames FN & Bellini BC, 2015 Fig.33

Species oudemansi (Yoshii, 1983) Deharveng, L & Bedos, A, 1995:22

Lepidonella oudemansi from Malaysia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.32

Species picta (Schäffer, C, 1898:410) Mitra, SK, 1992:212

Species subcarpenteri (Denis, 1948) Deharveng, L & Bedos, A, 1995:22

Lepidonella subcarpenteri from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.38

Species tokiokai Yosii, 1960 tt

Lepidonella tokiokai from the Solomon Islands
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.34

Species zeppelinii Soto-Adames, FN et Bellini, BC, 2015:332

Lepidonella zeppelinii from Brazil
After Soto-Adames FN & Bellini BC, 2015 Fig.1
             Syn.: Lepidonella zeppelini(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS
             Syn.: Lepidonella zepellinii(sic) Nguyen, M et Soto-Adames, FN, 2018:192 ISS
             Syn.: Lepidonella zeppelini(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS



Genus Bromacanthus Schött, 1925

  Syn.: Paronellinae Glacialoca Salmon, JT, 1941:405
  Syn.: ... Handschinella Yosii, R, 1959:53


Bromacanthus
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.35

Species sp.1 Yoshii, 1983

Species sp.2 Yoshii, 1983

Species elongatus Yoshii, 1981

Species flavidulus Yoshii, 1981

Species halmaherae Yoshii, 1983

Pseudoparonella (Irianella) halmaherae from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.24A

Species handschini Schött, 1925, t.t.

             Syn.: Pseudoparonella (Bromacanthus) Handschini(sic) Absolon, K et Ksenemann, M, 1942:14 ISS

Species orientalis (Handschin, 1930)

Species palawanicus Yoshii, 1983

Species sago Yoshii, R et Suhardjno, YR, 1992:49

Bromacanthus sago from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.28A

Species seramensis Yoshii, 1989

Species setigerus (Börner, C, 1906:178) Yoshii, 1981

             Syn.: Bromacanthus setigera(sic) Mitra, SK, 2003:84 ISS

Species wallacei Yoshii, R et Greenslade, P, 1993:82


Genus Trichorypha Schött, H, 1893:16

Species atrofasciata Schött, H, 1893:18, t.t.

            Syn.: Trichorypha atro-fasciata(sic) Schött, H, 1893:18 IOS
            Syn.: Trogolaphysa lamottei Nguyen, M et Soto-Adames, FN, 2018:184 =?



References