Checklist of the Collembola: Paronellinae

http://www.collembola.org/taxa/paronell.htm - Last updated on 2023.04.28 by Frans Janssens

Checklist of the Collembola: Paronellinae

Subfamilia Paronellinae Börner C, 1906:162 sensu Zang F et al, 2019:252

Genera with number of species
Genus	Number of species
Bromacanthus	12
Calobatinus	4
Campylothorax	14
Cephalophilus	3
Cyphoderinus	1
Cyphoderodes	7
Cyphoderus	85
Delamarerus	2
Lepidonella	16
Megacyphoderus	5
Mimoderus	4
Paracyphoderus	1
Paronella	3
Paronellidaegennov	1
Paronellidaegennov2	1
Pseudocyphoderus	4
Serroderus	26
Trichorypha	1
Troglobius	4
Trogolaphysa	83
Paronellinae	277

Syn.: Entomobryidae Paronelliae(sic) Mitra, SK, 1984:101 ISS

Paronellinae from China
2008.07.18 © NCode, A.

Paronellinae from Taiwan
2009.02.04 © Wu, S.

Paronellinae from Taiwan
2010.06.23 © Chien, H.-C.

Paronellinae from Australia
2012.06.23 © Kathy & Marissa.

Genus Paronellidaegennov Deharveng, L, Brehier, F, Bedos, A, Tian, M-Y, Li, Y-B, Zhang, F Qin, W-G et Tan, X-F, 2008:78

Species sp.1 Deharveng, L, Brehier, F, Bedos, A, Tian, M-Y, Li, Y-B, Zhang, F Qin, W-G et Tan, X-F, 2008:78

Paronellidae gen. nov. sp. nov. from China
After Deharveng L et al, 2008 Fig.5

Genus Paronellidaegennov2 Culver & Pipan, 2009

Species sp.1 Culver & Pipan, 2009

Tribus Paronellini Börner, 1906:162 sensu Zhang F et al, 2019:252

Syn.: Paronellinae Troglopedetini Soto-Adames, FN, Jordana, R et Baquero, E, 2014:1

Paronellini from Indonesia
Composed eye with Extra Ocular Structure (EOS)
2013.09.21 © Bertner, P.

Paronellini from Indonesia
Hyaline oval apically rounded scales
2013.09.21 © Bertner, P.

Paronellini from Malaysia
Composed eye with Extra Ocular Structure (EOS)
2014.02.12 © Bertner, P.

Paronellini from Malaysia
Apically knobbed dorsal thoracic macrosetae
2014.02.12 © Bertner, P.

Paronellini from Malaysia
Collophore with extended vesicles pressed in the surface waterfilm (drinking)
2014.02.12 © Bertner, P.

Remark by Zhang, F & al. 2015:7: Molecular phylogeny supports the closer relationship of Cyphoderini and Troglopedetini [= Paronellini], both of which have been clustered with Lepidocyrtinae.

Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The presence of pseudopores behind the posterior row of setae on the fourth abdominal segment, as seen in Troglopedetes, Trogolaphysa, Cyphoderus and Troglobius, corroborates the close relationship of Cyphoderus ([Cyphoderini]) to Trogolaphysa (Paronellini).

Genus Paronella Schött, H, 1893:14 F

  Syn.: Paronellidae Trogonella Massoud in Wilson JM, 1982:218
  Syn.: Paronellidae Trogonella Wilson, JM, 1982:218
  Syn.: ... Trichorypha Thibaud, J-M et Najt, J, 1988:722
  Syn.: ... Trichorypha Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15
  Syn.: ... Microparonella Salmon, 1964

Remark by ICZN. 2007:608: Placed on the Official List of Names in Zoology. Opinion 291.

Species fusca Schött, H, 1893:16, t.t.

Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1A

Paronella fusca from Cameroons
From "Lectotype"
After Mitra SK, 1992 Fig.1B

Paronella fusca
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.29

Species montana Delamare Deboutteville, 1950

Species purpurea Barra, 1969

Genus Campylothorax Schött, H, 1893:18

Syn.: Dicranocentroides Dicranocentropha Wray, DL, 1953:144
Syn.: Entomobryidae Dicranocentrapha(sic) Mari Mutt, JA, 1976:121 ISS

Campylothorax
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.33

  Syn.: Paronellini Campylothorx(sic) Mitra, SK, 1993:65 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:19 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:20 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:21 ISS
  Syn.: Paronellini Campylothtorax(sic) Soto-Adames, FN, 2016:24 ISS

Campylothorax sp. from Mexico
2016.10.06 © Murray, A.

Campylothorax sabanus from Brazil
Modified after Mari Mutt, 1987 in Bellini BC & Cipola NG, 2017 Fig.2

  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:165 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:166 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:167 ISS
  Syn.: Paronellini Campylothoraxi(sic) Bellini, BC et Cipola, NG, 2017:168 ISS

Species camelinus Womersley, 1930

Syn.: Campylothorax camelina(sic) Womersley, 1930 IOS

Syn.: Campylothorax camelina(sic) Mitra, SK et Dallai, R, 1980:275 ISS

Species cassagnaui Mitra, SK et Dallai, R, 1980:313

            Syn.: Campylothorax cassagnaus(sic) Mitra, SK et Dallai, R, 1980:313 IOS
            Syn.: Campylothorax cinctus Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:131
             Syn.: Campylothorax cincta(sic) Soto-Adames, FN, 2016:2 ISS

             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & Najt, 1995 ISS
             Syn.: Campylothorax cassagnaus(sic) Mari Mutt, JA et Bellinger, PF, 1996:171 ISS
             Syn.: Campylothorax cassagnaus(sic) Díaz & al., 1997 ISS
             Syn.: Campylothorax cassagnaus(sic) Culik, MP et Zeppelini, DF, 2003:1129 ISS

Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Leg. McAlister, E.; Det. Culik, M.

Campylothorax cf. cassagnaui from Costa Rica
2009.mm.dd © Shaw, P.
Mucro

Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.

Campylothorax cassagnaui from Panama
2014.03.dd © Bowley, S.

Syn.: Campylothorax cassagnaus(sic) da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:1 ISS
Syn.: Campylothorax cassagnaus(sic) Brito, RA, de Lima, ECA, Ferreira, AS, de Oliveira, JVLC, Amorim, TPL, Machado, TG, Lopes, BCH, Costa, IA, Alves, JLS, Brito, NP, Zeppelini, D, 2017:11 ISS

Species cinctus Schött, 1927

Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.45A

Campylothorax cinctus from Cameroon
After Mitra SK & Dallai R, 1980 Fig.46A

Species cubanus Gruia, M, 1983:203

            Syn.: Campylothorax cubana(sic) Gruia, M, 1983:203 IOS
             Syn.: Campylothorax cubana(sic) Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:94 ISS
             Syn.: Camphylotorax(sic) cubana(sic) Velázquez, DR et Novoa, NM, 2002:7 ISS

Campylothtorax(sic) cubanus from Haiti
After Soto-Adames FN, 2016 Fig.12A

Campylothtorax(sic) cubanus from Hispaniola
After Soto-Adames FN, 2016 Fig.12B

Species dominicanus Soto-Adames, FN, 2016:20

Campylothtorax(sic) dominicanus from Hispaniola
After Soto-Adames FN, 2016 Fig.10

Syn.: Campylothorax domincanus(sic) Soto-Adames, FN, 2016:28 ISS

Species hexosetosus Soto-Adames, FN, 2016:16

Campylothorax hexosetosus from Hispaniola
After Soto-Adames FN, 2016 Fig.7

Syn.: Campylothorax hexosetosa(sic) Soto-Adames, FN, 2016:16 ISS

Species longicornis Schött, H, 1893:20, t.t.

Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.1A
From syntype

Campylothorax longicornis from Cameroon
After Mitra SK & Dallai R, 1980 Fig.20
Mucro

Species melanocephalus Mitra, SK et Dallai, R, 1980:298

Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.29

Campylothorax melanocephalus from Zaire
After Mitra SK & Dallai R, 1980 Fig.30

Species mitrai Bellini & Meneses, 2012

Species notidanus Soto-Adames, FN, 2016:11

Campylothorax notidanus from Hispaniola
After Soto-Adames FN, 2016 Fig.4A

Species plagatus Cipola, NG et de Lima Oliveira, FG, 2016:488

Campylothorax plagatus from Brazil
After Cipola NG & de Lima Oliveira FG, 2016 Fig.1

Species sabanus (Wray, DL, 1953:144) Mitra, 1975

            Syn.: Dicranocentroides (Dicranocentropha) sabana(sic) Wray, DL, 1953:144 IOS
            Syn.: Dicranocentroides fasciculatus Rapoport, 1971 lapsus
            Syn.: Dicranocentroides fasciculatus Mari Mutt, JA et Bellinger, PF, 1990:125

             Syn.: Campylothorax sabana(sic) Mitra, 1975 ISS
             Syn.: Campylothorax sabana(sic) Mitra, SK et Dallai, R, 1980:300 ISS

Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.32A

Campylothorax sabana(sic) from Puerto Rico
After Mitra SK & Dallai R, 1980 Fig.33A

Syn.: Campylothorax sabana(sic) Cipola, NG et de Lima Oliveira, FG, 2016:494 ISS

Species schaefferi Börner, C, 1906:179

            Syn.: Campylothorax schäfferi(sic) Börner, C, 1906:179 IOS

             Syn.: S(sic) Schäfferi(sic) Schött, H, 1903:XIII nn ISS

             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:2 ISS
             Syn.: Campylothorax shaefferi(sic) Soto-Adames, FN, 2016:12 ISS

Campylothorax aff. schaefferi from Brazil
After Cipola NG & al, 2019 Fig.9

Species viruaensis Santos IP, Cipola NG et Bellini BC in da Silva Santos, IP, Cipola, NG, de Morais, JW et Bellini, BC, 2016:2

Campylothorax viruaensis from Brazil
After da Silva Santos IP, Cipola NG & Bellini BC in da Silva Santos IP & al, 2016 Fig.1

Genus Trogolaphysa Mills, HB in Pearse, AS, 1938:184

  Syn.: Paronellinae Dicranocentruga Wray, DL, 1953:141
  Syn.: ... Dicranocentrus(sic) Mari Mutt, JA, 1979:60 ISS
  Syn.: Paronellini Diranocentruga(sic) Mitra, SK, 1993:42 ISS

Dicranocentruga
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.31

  Syn.: Paronellini Dicraanocentruga(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellinae Dicranocentuga(sic) Mitra, SK, 2002:109 ISS
  Syn.: Paronellinae Metaparonella Mitra, 1972 u
  Syn.: Collembola Troglopodetes(sic) Elliott, WR, Palacios-Vargas, JG, Medellín, RA et Calva, O, 2023:406 ISS

Trogolaphysa
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.40

  Syn.: Troglopedetini Trogolaphyusa(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellidae Troglolaphysa(sic) Palacios-Vargas, JG et Zeppelini, D, 1995:173 ISS
  Syn.: Paronellini Trogolpahysa Soto-Adames, FN, 2015:23 ISS
  Syn.: Collembola Troglolaphysa(sic) Duarte Núñez, DA, Pinzón Florián, OP et Palacios-Vargas, JG, 2020:1205 ISS

Species sp.1 (Mari Mutt, JA, 1987:403)

Species sp.2 Palacios-Vargas, JG et Thibaud, J-M, 1997:328

Species sp.3 Bellini, BC et Zeppelini, D, 2004:588

Species sp.4 (Bellini, BC et Zeppelini, D, 2004:588)

Species sp.5 Palacios-Vargas, JG, 2008:46

Species sp.6 Bellini, 2014

Species sp.7 de Oliveira, MPA, de Oliveira Bernardi, LF, Zeppelini, D et Ferreira, RL, 2016:133

Trogolaphysa new sp. from Brazil
Parasitised by mite (Parasitengona, Microtrombidiidae)
After de Oliveira MPA & al, 2016 Fig.2A

Species sp.8 Mari Mutt & al, 2016

Species sp.9 Soto-Adames, FN et Anderson, EW, 2017:40

Species aelleni Yoshii, R, 1988:453

Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:36 ISS
Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:39 ISS

Species amazoccidentalensis Cipola NG in Cipola, NG, de Morais, JW et Bellini, BC, 2022:253

Trogolaphysa amazoccidentalensis from Brazil
After Cipola NG & al, 2022 Fig.2A

Trogolaphysa amazoccidentalensis from Brazil
After Cipola NG & al, 2022 Fig.2B

Species balteata Soto-Adames, FN, 2015:27

Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4C

Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4D

Species beguei (Delamare Deboutteville, 1951) Nguyen, M & Soto-Adames, FN, 2018:182

Syn.: Paronella beguei Delamare Deboutteville, 1948 nn

Trogolaphysa beguei from the Ivory Coast
After Nguyen M & Soto-Adames FN, 2018 Fig.5J

Species belizeana Palacios-Vargas, JG et Thibaud, J-M, 1997:326

Species berlandi (Denis, 1925) Thibaud, J-M & Najt, J, 1988:726

Syn.: Paronella Berlandi(sic) Denis, J-R, 1933:268 ISS
Syn.: Paronella Berlandi(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:134 ISS

Species bessoni Thibaud, J-M et Najt, J, 1988:724

Species caripensis (Gruia, M, 1987:155) Thibaud, J-M & Najt, J, 1988:723

Species carpenteri (Denis, 1925) Yoshii, 1988

            Syn.: Dicranocentruga geminata Mari Mutt, JA, 1987:402 =?
            Syn.: Dicranocentruga jataca Wray, DL, 1953:143 =?

             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:262 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:263 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:264 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:265 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:266 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:267 ISS
             Syn.: Paronella Carpenteri(sic) Denis, J-R, 1933:319 ISS
             Syn.: Paronella Carpenteri(sic) Cassagnau, 1963 ISS

Species clarencei Soto-Adames, FN, 2015:21

Trogolaphysa clarencei from Mexico
After Soto-Adames FN, 2015 Fig.3E

Species cotopaxiana Thibaud, J-M et Najt, J, 1988:726

Syn.: Lepidonella sp. Lima & Zeppelini, 2015
Syn.: Lepidonella sp. de Lima, ECA, Lopes, BCH, Oliveira-Neto, MA, de Mendonça, MC et Zeppelini, D, 2022:25

Species dimorphica Soto-Adames, FN, 2015:14

Trogolaphysa dimorphica from Mexico
After Soto-Adames FN, 2015 Fig.3B

Species distinguenda (Denis, 1931) Thibaud, J-M & Najt, J, 1988:726

Species ecuatorica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:22) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa ecuatoricus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa ecuatoricus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS
             Syn.: Trogolaphysa ecuatoriana(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 ISS

Species entreriosensis Soto-Adames, FN, 2015:23

Trogolaphysa entreriosensis from Argentina
After Soto-Adames FN, 2015 Fig.3F

Species ernesti Cipola NG et Bellini BC in Bellini, BC et Cipola, NG, 2017:170

Trogolaphysa ernesti from Brazil
After Bellini BC & Cipola NG, 2017 Fig.62

Species formosensis da Silva & Bellini, 2015

Species fuelleborni (Börner, 1903) Nguyen, M & Soto-Adames, FN, 2018:182

            Syn.: Paronella fülleborni(sic) Börner, 1903 IOS
             Syn.: Paronella fülleborni(sic) Wahlgren, 1908 ISS
             Syn.: Paronella fülleborni(sic) Handschin, 1929 ISS
             Syn.: Paronella Fuelleborni(sic) Denis, J-R, 1933:263 ISS
             Syn.: Paronella Fuelleborni(sic) Denis, J-R, 1933:268 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:441 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:471 ISS
             Syn.: Paronella fülleborni(sic) Absolon, K et Ksenemann, M, 1942:13 ISS

Species geminata (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species ghesquierei (Marlier, 1945) Nguyen, M & Soto-Adames, FN, 2018:182

             Syn.: Dicranocentruga ghesourieri(sic) Mitra, SK, 2002:114 ISS
             Syn.: Paronella ghesquièrei(sic) Thibaud, J-M, 2013:238 ISS
             Syn.: Trogolaphysa ghesquieri(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS
             Syn.: Dicranocentruga ghesouerei(sic) Nguyen, M et Soto-Adames, FN, 2018:183 ISS

Species giordanoae Soto-Adames, FN et Taylor, SJ, 2013:40

Trogolaphysa giordanoae from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.1

Species grassei (Delamare Deboutteville, 1952) Nguyen, M & Soto-Adames, FN, 2018:183

Syn.: Paronella Grassei(sic) Delamare Deboutteville, 1951 nn IOS

Species guacharo Yoshii, R, 1988:455

Species guayabambensis (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species haitica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:16) Thibaud, J-M & Najt, J, 1988:723

Syn.: Trogolaphysa haiticus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
Syn.: Trogolaphysa haiticus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS

Species hauseri Yoshii, R, 1988:457

Species hirtipes (Handschin, 1924) Thibaud, J-M & Najt, J, 1988:725

Syn.: Paronella carpenteri Denis, 1925 =?

Trogolaphysa cf. hirtipes from Brazil
After Cipola NG & al, 2019 Fig.10

Species hondurasensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

Syn.: Trogolaphysa honduraensis(sic) Soto-Adames, FN, 2015:36 ISS

Species innominata (Denis, J-R, 1933:263) Thibaud, J-M & Najt, J, 1988:726

            Syn.: Paronella innominata f. I Denis, J-R, 1933:263
            Syn.: Paronella innominata f. II Denis, J-R, 1933:263
            Syn.: Paronella innominata f. III Denis, J-R, 1933:263

Remark by Mitra, S.K. 2009.07.31:in litt.: Originally, Denis (1933) described 3 forms of "innominata": C.R.I, II, and III. They were considered as different species by Salmon (1964:544), but are infraspecific morphs or varieties.

Species jacobyae Soto-Adames, FN et Taylor, SJ, 2013:47

Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 IOS

Trogolaphysa jacobyi(sic) from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.24

Trogolaphysa jacobyi(sic) from Belize
Ant.4 with constriction
After Soto-Adames FN & Taylor SJ, 2013 Fig.25

Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:2 ISS
Syn.: Trogolaphysa jacobyi(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS

Species jamaicana (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:19) Thibaud, J-M & Najt, J, 1988:723

Syn.: Trogolaphysa jamaicanus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species jataca (Wray, DL, 1953:143) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Dicranocentruga jacata(sic) Mari Mutt, JA, 1979:60 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:28 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas et al., 1986 ISS

Species judithnajtae Nguyen, M et Soto-Adames, FN, 2018:184

Trogolaphysa judithnajtae from Madagascar
After Nguyen M & Soto-Adames FN, 2018 Fig.1

Species laterolineata Soto-Adames, FN, 2015:16

Trogolaphysa laterolineata from Mexico
After Soto-Adames FN, 2015 Fig.3C

Species luquillensis (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species marielouiseae Soto-Adames, FN, 2015:19

             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:1 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:2 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:20 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:10 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:17 ISS

Trogolaphysa marielouisea(sic) from Mexico
After Soto-Adames FN, 2015 Fig.3D

Species marimutti (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

Species maya Mills, HB in Pearse, AS, 1938:184, t.t.

Syn.: Troglopedetes delamarei Massoud, Z et Gruia, M, 1973:339

Troglopedetes delamarei from Cuba
After Massoud Z & Gruia M, 1973 Fig.7E

Species millsi Arlé, 1939

Species monomaculata (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

Species nacionalica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

Syn.: Trogolaphysa nacionalicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species neoannulicornis (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species nigrocephala Cipola NG in Cipola, NG, de Morais, JW et Bellini, BC, 2022:260

Trogolaphysa nigrocephala from Brazil
After Cipola NG & al, 2022 Fig.8

Species nigrolineata Cipola NG in Cipola, NG, de Morais, JW et Bellini, BC, 2022:266

Trogolaphysa nigrolineata from Brazil
After Cipola NG & al, 2022 Fig.13

Species nigromaculata (Schött, H, 1903:VII) Nguyen, M & Soto-Adames, FN, 2018:189

Paronella nigromaculata from Kamerun
Schematic outline of habitus
After Schött H, 1903 Taf.I Fig.10

             Syn.: Paronella fülleborni(sic) Philiptschenko, 1926 ISS
             Syn.: Paronella fülleborni(sic) Martynova, 1961 ISS
            Syn.: Neophorella dubia Paclt, 1959

Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Pauliani, 1952

Syn.: Paronella nigromaculata ssp. pallida Delamare Deboutteville, 1951 nn

Species ocellata Soto-Adames, FN, 2015:25

Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4A

Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4B

Species octosetosa Soto-Adames, FN, 2015:32

Trogolaphysa octosetosa from Mexico
After Soto-Adames FN, 2015 Fig.4F

Species oztotlica (Ojeda, M & Palacios-Vargas, JG, 1984:16) Thibaud, J-M & Najt, J, 1988:723

Syn.: Troglopedetes sp. Palacios-Vargas, 1982

Troglopedetes oztotlicus from Mexico
After Hoffmann A & al, 1986 Fig.18

Syn.: Trogolaphysa oztotlicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species palaciosi Soto-Adames, FN, 2015:32

Species paracarpenteri Soto-Adames, FN, 2015:29

Trogolaphysa paracarpenteri from Mexico
After Soto-Adames FN, 2015 Fig.4E

Syn.: Trogolaphysa paracapenteri(sic) Soto-Adames, FN, 2015:31 ISS

Species penicillata (Schött, 1926) Nguyen, M & Soto-Adames, FN, 2018:190

Subspecies obscura Delamare Deboutteville, 1950

Syn.: Paronella penicillata ssp. obscura Delamare Deboutteville, 1951 nn

Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Paulian, 1952

Species piracurucaensis Nunes, RC et Bellini, BC, 2018:87

Trogolaphysa-piracurucaensis from Brazil
After Nunes RC & Bellini BC, 2018 Fig.44

Species quinquesetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species pseudomaculata Cipola NG in Cipola, NG, de Morais, JW et Bellini, BC, 2022:278

Trogolaphysa pseudomaculata from Brazil
Typical pattern
After Cipola NG & al, 2022 Fig.21A

Trogolaphysa pseudomaculata from Brazil
Vestigial pattern
After Cipola NG & al, 2022 Fig.21B

Trogolaphysa pseudomaculata juv. from Brazil
After Cipola NG & al, 2022 Fig.21C

Species quinquedentomucronata Cipola NG in Cipola, NG, de Morais, JW et Bellini, BC, 2022:284

Trogolaphysa quinquedentomucronata from Brazil
After Cipola NG & al, 2022 Fig.27A

Trogolaphysa quinquedentomucronata from Brazil
After Cipola NG & al, 2022 Fig.27B

Species quisqueyana Soto-Adames, FN, Jordana, R et Baquero, E, 2014:10

Trogolaphysa quisqueyana from the Dominican Republic
After Soto-Adames & al., 2014 Fig.43

Species relicta (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:26) Thibaud, J-M & Najt, J, 1988:723

Syn.: Trogolaphysa relictus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species riopedrensis (Mari Mutt, JA, 1987:403)

Species sauroni Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12

Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12 IOS

Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 left

Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 right

Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 left

Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 right

             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:17 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:29 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN et Anderson, EW, 2017:40 ISS

Species separata (Denis, J-R, 1933:262) Thibaud, J-M & Najt, J, 1988:726

Syn.: Dicranocentruga riopedrensis Mari Mutt, JA, 1987:403 =?

Species sexsetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species stannardi Soto-Adames, FN, 2015:8

Trogolaphysa stannardi from Mexico
After Soto-Adames FN, 2015 Fig.3A

Species strinatii Yoshii, R, 1988:452

Species subterranea (Mari Mutt, JA, 1987:403) Mari Mutt, JA & Bellinger, PF, 1996:171

Syn.: Dicranocentruga subterranean(sic) Mari Mutt, JA, 1987:403 IOS

Species tijucana (Arlé & Guimaraes, 1979) Mari Mutt, JA & Bellinger, PF, 1990:124

Syn.: Troglopedetes tijucana(sic) Palacios-Vargas et al., 1989 ISS
Syn.: Trogolaphysa tijuacana Soto-Adames, FN, 2015:39 ISS

Species toroi (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:12) Thibaud, J-M & Najt, J, 1988:723

Species trioculata Soto-Adames, FN, 2015:35

Trogolaphysa trioculata from Mexico
After Soto-Adames FN, 2015 Fig.3G

Species trisetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species variabilis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

Species villiersi (Delamare Deboutteville, 1950) Soto-Adames, FN, 2014:il

Syn.: Paronella villiersi Delamare Deboutteville & Paulian, 1947 nn
Syn.: Paronella villersi(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

Subspecies anomala Delamare Deboutteville, 1951

Subspecies pallida Delamare Deboutteville, 1950

Species vivieni (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:181 ISS
Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

Species wahlgreni (Mitra, SK, 2002:109) Nguyen, M & Soto-Adames, FN, 2018:192

Syn.: Paronella nigromaculata Wahlgren, 1908

Dicranocentruga wahlgreni from Kenia
After Mitra SK, 2002 Fig.1A

Syn.: Dicranocentruga whalgreni(sic) Nguyen, M et Soto-Adames, FN, 2018:189 ISS
Syn.: Trogolaphysa wahlbergi(sic) Nguyen, M et Soto-Adames, FN, 2018:194 lapsus ISS

Species xtolokensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:10) Thibaud, J-M & Najt, J, 1988:723

Species yoshiii (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:14) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetes yoshiius(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:14 IOS

             Syn.: Trogolaphysa yoshius(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa yoshiia(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Trogolaphysa yoshiia(sic) Palacios-Vargas, JG, 2006:10 ISS
             Syn.: Trogolaphysa yoshiia(sic) Soto-Adames, FN et Taylor, SJ, 2013:37 ISS

Tribus Cyphoderini Börner C, 1906:162 sensu Zhang F & al, 2019:252

Syn.: Entomobryidae Cyphoderitae(sic) Delamare Deboutteville, C, 1945:48 ISS
Syn.: Mydoniidae Cyphodurinae(sic) Kloet, GS et Hincks, WD, 1945:8 ISS

Cyphoderus albinus from the UK
With Myrmica rubra
2007 © Cornwall, N.J.

Cyphoderus albinus from France
With Pheidole pallidula
2008.03.15 © Lebeaux, P.

Syn.: Lepidocyrtidae Cyphoderidae(sic) Castaño-Meneses, G, Palacios-Vargas, JG, Delabie, JHC, Zeppelini, D et Mariano, CSF, 2017:741 ISS

Genus Cyphoderus Nicolet H, [1842:63] nc Opinion 291

M

  Syn.: Podurelles (Corps linéaires) Cyphodeirus(sic) Nicolet H, 1842:63, nr Opinion291 IOS
  Syn.: ... Podura Fabricius, 1783 partim
  Syn.: ... Lepidocyrthus(sic) Bourlet ISS
  Syn.: Podurelles(sic) Cyphodurus Nicolet, H in Desor, E, 1841:384 nn
  Syn.: Thysanura Beckia Lubbock, J, 1870:279
  Syn.: ... Cyphoderella Börner, 1913 nn
  Syn.: ... Borecus Folsom, 1923
  Syn.: Cyphoderidae Cyphoda Delamare Deboutteville, C, 1948:360
  Syn.: Entomobryidae Cyphoda Gisin, H, 1960:258
  Syn.: Collembola Cyphodems(sic) da Gama, MM, 1988:74 ISS
  Syn.: Collembola Cyphodem(sic) da Gama, MM, 1988:84 ISS
  Syn.: Cyphoderinae Cyphoda Soto-Adames, FN, Barra, J-A, Christiansen, K et Jordana R, 2008:512
  Syn.: Cyphoderinae Cyphoda de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47
  Syn.: Cyphoderinae Cyphoda Cipola, NG, 2021:il
  Syn.: Troglopedetini Camponotus(sic) Salmon, JT, 1964:533 lapsus ISS

  Syn.: Podurellen(sic) Cyphodeirus(sic) Elditt, HL, 1854:40 ISS
  Syn.: Podura Cyphodeirus(sic) Elditt, HL, 1854:43 ISS
  Syn.: Collembola Cyphodeirus(sic) Ellis, WN et Bellinger, PF, 1973:14 Opinion IOS

Remark by ICZN. 2007:226: Emendation of Cyphodeirus. Placed on the Official List of Names in Zoology. Opinion 291.

Syn.: Cyphoderidae Cyphyoderus(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS

Cyphoderus sp. from South Africa
Ant.3 as 'pearly bead' (*)
2020.04.25 © Robertson, H.

Cyphoderus sp. from France
2021.08.26 © Picard, J.

Cyphoderus sp. from France
Antennae
Ant.3 distinctly shorter then ant.2
2021.08.26 © Picard, J.

Cyphoderus sp. from France
Distinctly long mucro
2021.08.26 © Picard, J.

Cyphoderus sp. from Taiwan
2021.09.11 © Cheng, H.-J.

Cyphoderus sp. from Taiwan
Long mucro with 2 apical teeth
2021.09.11 © Cheng, H.-J.

Species sp.1 Janssens F sensu Zeppelini & Castaño, 1995

Syn.: Cyphoderus n.sp. Zeppelini Filho, D et Castaño Meneses, G, 1995:10

Species sp.2 (Janssens F sensu Arbea & Blasco-Zumeta, 2001) Soto-Adames & al, 2008

Syn.: Cyphoda n.sp. Arbea, JI et Blasco-Zumeta, J, 2001:47

Syn.: Cyphoderus sp. Soto-Adames & al, 2008

Species sp.3 Janssens F sensu Bellini & Zeppelini, 2004

Syn.: Cyphoderus undescribed species Bellini, BC et Zeppelini, D, 2004:588

Species adelaideae Womersley, 1934

Species africanus Womersley, 1929

Species amaranus Brown, JM, 1926:215

Species antennatus Delamare Deboutteville, C, 1945:39

Species antigone Fernando, 1959

Species arcuatus Wahlgren, E, 1906:20

Species asiaticus Yosii, R, 1959:56

Species assimilis Börner, C, 1906:181

            Syn.: Cyphoderus simulans Imms, 1912
            Syn.: Cyphoderus similis Denis, 1931
            Syn.: Cyphoderus subsimilis Paclt, 1959
            Syn.: Cyphoderus assimilis ssp. empodialis Mari Mutt, JA et Bellinger, PF, 1990:128

Species buxtoni Brown, 1920

            Syn.: Cyphoderus Buxtoni(sic) Brown, 1920 IOS

             Syn.: Cyphoderus Buxtoni(sic) Brown, 1926 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, C, 1945:36 ISS
             Syn.: Cyphoderus Buxtoni(sic) Delamare Deboutteville, 1948 ISS

Species canariensis da Gama, MM, 1988:82

Species colurus Börner, C in Schultze, 1908:57

Syn.: Cyphoda colurus(sic) Delamare Deboutteville, 1948 ISS
Syn.: Cyphoda colorus(sic) Thibaud, J-M, 2013:240 ISS

Species cuthbertsoni Womersley, 1929

Syn.: Cyphoderus Cuthbertsoni(sic) Delamare Deboutteville, C, 1945:36 ISS

Species dorsti Jacquemart, 1974 in Peck SB & Jacquemart J, 2011:1 u

Species ganeensis Tyagi & Baijal, 1979

Species genneserae Carpenter, 1913

Species gisini Gruia, 1967

Species grassei (Cassagnau, P & Delamare Deboutteville, C, 1955:384) Christiansen, K, 1957:79

Syn.: Cyphoda Grassei(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:384 IOS

Species handschini (Delamare Deboutteville, C, 1948:346) Dallai, R, 1973:536

Syn.: Cyphoderus Handschini Delamare Deboutteville, C, 1948:346 IOS
Syn.: Cyphoderus assimilis Handschin, 1926

Species heymonsi Börner, 1906

            Syn.: Cyphoderus Heymonsi(sic) Börner, 1906 IOS
            Syn.: Cyphoderus Heymonsi(sic) Börner, 1913 IOS
            Syn.: Cyphoda Heymonsi(sic) Delamare Deboutteville, 1948 IOS

Species holonycha Silvestri, 1918

Species hrdyi Rusek, 1971

Species indicus Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45

Cyphoderus indicus from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.45 Img.1

Species insularum Carpenter, GH, 1916:47

Syn.: Cyphoderus bidenticula Handschin, 1942, partim ISS

Species jharkhandensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:47

Cyphoderus jharkhandensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2016 p.47 Img.1

Species kawari Jacquemart, S, 1974:33

Species komareki Rusek, 1961

Species lantohi Yoshii, 1987

Species laticlavatus (Stach, J, 1960:545) Wilson, JM, 1982:218

             Syn.: Cyphoderopsis lavaticus(sic) Massoud in Wilson JM, 1982:218 lapsus ISS
             Syn.: Cyphoderopsis lavaticus(sic) Wilson, JM, 1982:218 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:218 ISS
             Syn.: Cyphoderus lavaticus(sic) Wilson, JM, 1982:219 lapsus ISS
             Syn.: Cyphoderus lavaticus(sic) Thibaud, J-M et Najt, J, 1988:724 lapsus ISS

Species limboxiphius Börner, 1913

            Syn.: Cyphoderus limboxiphius ssp. nitricolus Mari Mutt, JA et Bellinger, PF, 1990:129
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Bernava-Laborde & Palacios-Vargas, 2008 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Salazar Martínez, AE, Narváez Beinhorn, MJ et Bernava Laborde, V, 2017:110 ISS

Species maroccanus (Delamare Deboutteville, C, 1948:361) Jacquemart, S, 1974:43, t.t.

            Syn.: Cyphoda marocanus(sic) Delamare Deboutteville, C, 1948:361 IOS
             Syn.: Cyphoda maroccanus(sic) Delamare Deboutteville, C, 1948:360,tt ISS
            Syn.: Cyphoda marocanus(sic) Jacquemart, S, 1974:19 IOS
            Syn.: Cyphoda marocanus(sic) Jacquemart, S, 1974:35 IOS
            Syn.: Cyphoderus marocanus(sic) Jacquemart, S, 1974:43 IOS
            Syn.: Cyphoda marocanus(sic) Thibaud, J-M, 2013:234 IOS
            Syn.: Cyphoda marocanus(sic) Thibaud, J-M, 2013:237 IOS

Species mitumbis Tshelnokov, 1977

Species monopterus Rusek, 1981

Species natalensis Börner, 1913

Species nichollsi Womersley, 1934

Syn.: Cyphoderus Nichollsi(sic) Womersley, 1934 IOS

Species nigeri Jacquemart, 1980:1

Species omoensis Delamare Deboutteville, C, 1945:35

            Syn.: Cyphoderus arcuatus var. aethiopicus Handschin, 1929
             Syn.: Cyphoderus genneserae var. aethiopica(sic) Delamare Deboutteville, C, 1948:337 ISS
             Syn.: Cyphoderus genneserae ssp. aethiopica(sic) Thibaud, J-M, 2013:239 ISS
             Syn.: Cyphoderus genneserae ssp. aethiopica(sic) Thibaud, J-M, 2013:241 ISS
            Syn.: Cyphoderus bidenticulatus var. veneris Gisin, 1944
            Syn.: Cyphoderus bidenticulatus var. veneris Gisin, 1944

Species paralbinus Jacquemart, S(+), 1980:3

Species propinquus Stach, 1963

Species pseudalbinus Schött, H, 1917:50

Species reuteri Loksa, I, 1978:61

Species rubiae Baijal, HN, 1955:536

Species saheli Jacquemart, 1980:3

Species sarojinii Bhattacharjee, 1985

Species squamidives Silvestri, 1918

Syn.: Cyphoderus Cuthbertsoni(sic) Womersley, 1929 =?

Syn.: Cyphoderus arcuatus ssp. sqamides(sic) Thibaud, J-M, 2013:240 ISS

Species subsimilis Delamare Deboutteville, C, 1948:346

Cyphoderus subsimilis from the Ivory Coast
2016.05.10 © Parmentier, T.

Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.

Cyphoderus subsimilis from the Ivory Coast
2016.05.19 © Janssens, F.
Leg.: Parmentier, T.
Distal dental scale as long as bidentate mucro

Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Two unpair ungual teeth (*)

Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Straight bidentate mucro

Cyphoderus subsimilis from the Ivory Coast
2016.06.15 © Janssens, F.
Leg.: Parmentier, T.
Dens with 4 internal and 6 external scales

Species sudanensis Wahlgren, E, 1906:18

Species sumatranus Yoshii, 1987

Species termitellus Delamare Deboutteville, C, 1948:339

            Syn.: Cyphoderus termitum Silvestri, 1918
            Syn.: Cyphoderus termitum Denis, 1942
            Syn.: Cyphoderus termitum Handschin, 1942

Species trinervoides Paclt, 1965

Species variabilis Jacquemart, 1980:2

Species virungiensis Tshelnokov, 1977

Species yugoslavicus Denis, 1933

Species zoroastris Yosii, 1963

Species-group inermes(sic) Delamare-Deboutteville, 1948
            Syn.: Cyphoderus inermes(sic)-group Delamare-Deboutteville, 1948 IOS
             Syn.: Cyphoderus inerme(sic)-group Zeppelini, D et Oliveira, JVLC, 2016:560 ISS
            Syn.: Cyphoderus inermes(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 IOS
             Syn.: Cyphoderus inerme(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:58 ISS
             Syn.: Cyphoderus Inermes(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 ISS
            Syn.: Cyphoderus inermes(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:38 IOS
            Syn.: Cyphoderus inermes(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 IOS
            Syn.: Cyphoderus inermes(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:41 IOS

Species adversus Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:30

Species innominatus Mills, HB in Pearse, AS, 1938:186

             Syn.: Cyphoderus similis f. inermes(sic) Cassagnau, 1963 ISS
             Syn.: Cyphoderus similis f. inermes(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 ISS
             Syn.: Cyphoderus similis f. inerme(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:41 ISS

             Syn.: Cyphoderus inomminatus(sic) Zeppelini, D et Oliveira, JVLC, 2016:568 ISS

Cyphoderus innominatus from Brazil
In colonies of Atta sexdens
After Mota Filho TMM & al, 2021 Fig.1b

Cyphoderus innominatus from Brazil
In colonies of Atta sexdens
After Mota Filho TMM & al, 2021 Fig.1c

Species nitricolus Rapoport & de Izarra, 1962

            Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Rapoport & de Izarra, 1962 IOS

            Syn.: Cyphoderus javanus f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143 IOS
            Syn.: Cyphoderus javanus f. inerme(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:41 IOS

             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:1 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:2 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:41 ISS
             Syn.: Cyphoderus limboxiphius ssp. nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:42 ISS
             Syn.: Cyphoderus (inermes(sic)-group) nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:27 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:1 ISS
             Syn.: Cyphoderus (Inermes(sic)-group) nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 ISS
             Syn.: Cyphoderus (inermes(sic)-group) nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:13 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:14 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:26 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:28 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:29 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:30 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:34 ISS
             Syn.: Cyphoderus (inermes(sic)-group) nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:41 ISS

Species-group bidenticulati(sic) Delamare-Deboutteville, 1948
            Syn.: Cyphoderus bidenticulati(sic)-group Delamare-Deboutteville, 1948 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Zeppelini, D et Oliveira, JVLC, 2016:560 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:640 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Mateos, E, Winkler, D, Riutort, M et Alvarez-Presas, M, 2021:474 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Mateos, E, Zhang, B et Alvarez-Presas, M, 2023:308 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Mateos, E, Zhang, B et Alvarez-Presas, M, 2023:310 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Mateos, E, Zhang, B et Alvarez-Presas, M, 2023:311 IOS
             Syn.: Cyphoderus Bidenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 ISS
            Syn.: Cyphoderus bidenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:38 IOS
            Syn.: Cyphoderus bidenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 IOS

Species agnotus Börner, C, 1906:180

             Syn.: Beckia albinos(sic) Parona, 1895 ISS
             Syn.: Beckia albinos(sic) Bonet, F, 1933:123 ISS
             Syn.: Beckia albinos(sic) Bonet, F, 1933:124 ISS

             Syn.: Cyphoderus aenotus(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:48 ISS

Cyphoderus cf. agnotus from the Galapagos
After Katz AD & al, 2016 Fig.21B

Species albinus Nicolet, H, 1842:67, t.t., B

Syn.: Cyphodeirus(sic) albinos(sic) Nicolet, H, 1842:67,tt IOS

Cyphodeirus(sic) albinos(sic) from Switzerland
After Nicolet H., 1842 Pl.7 Fig.7

            Syn.: crystalpoduren Fabricius O, 1783
            Syn.: Lepidocyrtus argentatus Bourlet, C, 1845:67
            Syn.: Beckia argentea Lubbock, J, 1870:279,tt

Beckia argentea from the UK
After Lubbock J., 1873 Pl.23

             Syn.: Beckia argenta(sic) Tömösváry, 1883 ISS
             Syn.: Beckia argenta(sic) Dalla Torre, 1888 ISS
             Syn.: Cyphodeirus(sic) argentus(sic) Vellay, 1899 ISS
             Syn.: Cyphodeirus(sic) (Beckia) argenteus Vellay, 1900 ISS
            Syn.: Cyphoderus affinis Giard, 1895
            Syn.: Cyphoderus affinis Schött, 1921 =?
            Syn.: Cyphoderus affinis Heckman, CW, 2001:294
            Syn.: Cyphoderus affinis Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5
            Syn.: Tullbergia immaculata Lie-Pettersen, 1896
            Syn.: Cyphodeirus(sic) albus Carl, 1899

            Syn.: Lepidocyrtus albinos(sic) Gervais in Walckenaër, 1844 IOS
            Syn.: Cyphodeirus(sic) Albinos(sic) Bourlet, C, 1845:67 IOS
            Syn.: Cyphoderus albinos(sic) Nicolet, H, 1847:383 IOS
            Syn.: Cyphodeirus(sic) albinos(sic) Elditt, HL, 1854:40 IOS
            Syn.: Lepidocyrtus albinos(sic) Lubbock, J, 1870:301 IOS
            Syn.: Beckia albinos(sic) Lubbock, J, 1870:279 IOS
            Syn.: Beckia albinos(sic) Lubbock, J, 1873:149 IOS

Beckia albinos(sic) from the UK
After Lubbock J, 1873 Pl.24

Syn.: Beckia albinos(sic) Beck, J in Lubbock, J, 1873:253 IOS

Beckia albinos(sic) from the UK
After Beck J in Lubbock J, 1873 Pl.72

            Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 IOS
            Syn.: Cyphodeirus(sic) albinos(sic) Parona, 1875 IOS
            Syn.: Beckia albinos(sic) Parona, 1878 IOS
            Syn.: Beckia albinos(sic) Parona, 1882 IOS
            Syn.: Beckia albinos(sic) Tömösváry, 1883 IOS
            Syn.: Beckia albinos(sic) Brook, 1884 IOS
            Syn.: Beckia (Cyphoderus) albinos(sic) Reuter, 1886 IOS
            Syn.: Cyphoderus albinos(sic) Oudemans, 1887 IOS
            Syn.: Beckia albinos(sic) Parona, 1887 IOS
            Syn.: Beckia albinos(sic) Dalla Torre, 1888 IOS
            Syn.: Beckia albinos(sic) Parona, 1888 IOS
            Syn.: Cyphoderus albinos(sic) Uzel, 1890 IOS
            Syn.: Cyphoderus albinos(sic) Moniez, 1891 IOS
            Syn.: Cyphoderus albinos(sic) Moniez, 1891 IOS
            Syn.: Cyphoderus albinos(sic) Uzel, 1891 IOS
            Syn.: Beckia albinos(sic) Moniez, 1894 IOS
            Syn.: Cyphoderus albinos(sic) Lameere, A, 1895:591,B IOS
            Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1896 IOS
            Syn.: Cyphoderus albinos(sic) Poppe & Schäffer, 1897 IOS
            Syn.: Beckia albinos(sic) Parona, 1897 IOS
            Syn.: Cyphoderus albinos(sic) Lie-Pettersen, 1898 IOS
            Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 IOS
            Syn.: Cyphodeirus(sic) albinos(sic) Scherbakov, 1898 IOS
            Syn.: Cyphoderus albinos(sic) Carl, J, 1899:336 IOS
            Syn.: Cyphoderus albinos(sic) Carl, 1900 IOS
            Syn.: Cyphoderus albinos(sic) Börner, C, 1901:71 IOS
            Syn.: Cyphoderus albinos(sic) Börner, C, 1901:136,B IOS
            Syn.: Cyphoderus albinos(sic) Carl, 1901 IOS
            Syn.: Cyphoderus albinos(sic) Ågren, 1903 IOS
            Syn.: Cyphoderus albinos(sic) Axelson, 1905 IOS
            Syn.: Cyphoderus albinos(sic) Axelson, 1907 IOS
            Syn.: Cyphoderus albinos(sic) Evans, 1908 IOS
            Syn.: Cyphoderus albinos(sic) ssp. borneri(sic) Schille, 1908 IOS
            Syn.: Cyphoderus albinos(sic) Collinge, WE et Shoebotham, JW, 1909:89 IOS
            Syn.: Cyphoderus albinos(sic) Bagnall, 1910 IOS
            Syn.: Cyphoderus albinos(sic) Bagnall, 1910 IOS
            Syn.: Cyphoderus albinos(sic) Axelson, 1911 IOS
            Syn.: Cyphoderus albinos(sic) Carpenter, GH, 1916:47 IOS
            Syn.: Cyphoderus albinos(sic) Womersley, 1923 IOS
            Syn.: Cyphoderus albinos(sic) Denis, 1924 IOS
            Syn.: Cyphoderus albinos(sic) Brown, 1926 IOS
            Syn.: Cyphodeirus(sic) albinos(sic) Coutelen, F, 1928:854 IOS
            Syn.: Cyphoderus albinos(sic) Denis, J-R, 1933:270 IOS
            Syn.: Lepidocyrtus albinos(sic) Perrier, 1934 IOS
            Syn.: Cyphoderus albinos(sic) Turk, 1938 IOS
            Syn.: Cyphoderus albinos(sic) Geyskens, J, 1945:16,B IOS
            Syn.: Cyphodurus albinos(sic) Kloet, GS et Hincks, WD, 1945:8 IOS
            Syn.: Cyphoderus albinos(sic) Steel, 1946 IOS
            Syn.: Cyphoderus albinos(sic) Chopard, 1955 IOS
            Syn.: Camponotus(sic) albinos(sic) Salmon, JT, 1964:533 IOS
             Syn.: Cyphoderus albidus(sic) Berg, MP, 2001:xiv ISS
            Syn.: Cyphoderus albinos(sic) Hopkin, SP, 2002:1 IOS
            Syn.: Cyphoderus albinos(sic) Henderickx, H et Vets, V, 2003:9 IOS

Cyphoderus albinus from the UK
2007 © Cornwall, N.J.

Cyphoderus albinus from the UK
2007 © Cornwall, N.J.

Cyphoderus albinus from France
2008.03.15 © Lebeaux, P.

Cyphoderus albinus from Czechia
2008.03.28 © Krásenský, P.

Cyphoderus albinus from Germany
With Platyarthrus hoffmannseggi
2009.04.22 © Schlüter, H.

Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.

Cyphoderus albinus from Holland
2011.07.31 © van Duinen, J.

Cyphoderus albinus from Sweden
2011.04.11 © Engström, A.

Cyphoderus albinus from France
2012.08.07 © Quintin, C.

Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.

Cyphoderus albinus from Portugal
With partly recovered antennae
2012.10.11 © Murray, A.

Cyphoderus albinus from the UK
2012.11.27 © Murray, A.

Cyphoderus albinus from the UK
2012.11.27 © Murray, A.

Cyphoderus albinus juv. from the UK
2012.11.28 © Murray, A.

Cyphoderus albinus from the UK
With Lasius flavus
2012.11.28 © Murray, A.

Cyphoderus albinus from France
Ant.3 as 'pearly bead'
2013.05.12 © Quintin, C.

Cyphoderus albinus from France
Head in dorsal aspect semi-circular
2014.07.28 © Quintin, C.

Syn.: Cyphoderus albinos(sic) Manu, M, Szekely, L, Oromulu, LV, Barbuceanu, D, Honciuc, V, Maican, S, Fiera, C, Purice, D et Ion, M, 2015:299 ISS

Cyphoderus albinus from Holland
2015.11.14 © van Leeuwen, J.

Syn.: Cyphoderus albinos(sic) Mandal, GP, Suman, KK et Bhattacharya, KK, 2016:45,tt IOS

Cyphoderus albinus from Spain
2016.04.22 © Tizon, J.

Cyphoderus albinus from Belgium
2018.05.27 © Huskens, M.-L.

Cyphoderus albinus from Belgium
Ant.3 shaped as a 'pearly bead' (*)
2018.05.27 © Huskens, M.-L.

Cyphoderus albinus from Belgium
Frontal aspect
2018.05.27 © Huskens, M.-L.

Cyphoderus albinus from France
With Lasius flavus
2018.12.19 © Lebeaux, P.

Cyphoderus albinus from France
2019.01.14 © Tiky, S.

Cyphoderus albinus from the UK
2020.05.14 © Uncle, F.

Cyphoderus albinus from the UK
Ant.3 with narrow intersegmental rings (*)
2020.05.14 © Uncle, F.

Cyphoderus albinus from the UK
Furca
2020.05.14 © Uncle, F.

Cyphoderus albinus from the UK
Mucro distinctly narrow and long
Dens with distinctly large fringed scales
2020.05.14 © Uncle, F.

Cyphoderus albinus from the UK
Unguis & unguiculus with wing-like teeth
2020.05.14 © Uncle, F.

Cyphoderus albinus from the UK
Mucro apically bidentate
2020.05.14 © Uncle, F.

Cyphoderus albinus from France
2020.04.02 © Reinach, Br.

Cyphoderus albinus from France
Ant.3 with narrow intersegmental rings
2020.04.02 © Reinach, Br.

Cyphoderus albinus from France
2021.02.01 © Vol, F.

Cyphoderus albinus from France
With Myrmica sp.
2021.08.26 © Vol, F.

Cyphoderus albinus from Belgium
Ant.3 distinctly shorter then ant.2 and ant.4
2021.11.14 © Huskens, M.-L.

Cyphoderus albinus from Belgium
Ant.3 with apical and basal constriction
Resembles pearly bead
2021.11.14 © Huskens, M.-L.

Cyphoderus albinus from Belgium
Distinctly long mucro
Apically bidentate
2021.11.14 © Huskens, M.-L.

Subspecies apatelus Börner, 1913

Syn.: Cyphoderus albinus ssp. apaletus(sic) Jacquemart, S(+), 1980:2 ISS

Subspecies boerneri Schille, 1906

            Syn.: Cyphoderus albinus ssp. Börneri(sic) Schille, 1906 IOS
            Syn.: Cyphoderus albinus Stach, 1919
            Syn.: Cyphoderus albinus Stach, 1922

             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1908 ISS
             Syn.: Cyphoderus albinos(sic) ssp. börneri(sic) Schille, 1912 ISS
             Syn.: Cyphoderus Börneri(sic) Börner, 1913 ISS
             Syn.: Cyphoderus albinus ssp. Börneri(sic) Delamare Deboutteville, C, 1948:342 ISS
             Syn.: Cyphoderus albinus f. Boerneri(sic) Cassagnau, 1963 ISS
             Syn.: Cyphoderus albinus ssp. börneri(sic) Jacquemart, S(+), 1980:2 ISS

Species amazonicus Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:34

Syn.: Cyphoderus amanozicus(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:42 ISS

Species empodialis Rapoport, 1962

Syn.: Cyphoderus assimilis ssp. empodialis Nuñez, 1962

Species equidenticulati Nunes, RC et Bellini, BC, 2018:72

            Syn.: Cyphoderus equidenticulati(sic) Nunes, RC et Bellini, BC, 2018:72 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:3 IOS
            Syn.: Cyphoderus (Bidenticulati(sic)-group) equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 IOS
            Syn.: Cyphoderus (bidenticulati(sic)-group) equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:12 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:16 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:38 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 IOS
            Syn.: Cyphoderus (bidenticulati(sic)-group) equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:40 IOS
            Syn.: Cyphoderus equidenticulati(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:42 IOS

Remark by Janssens, F. 2023.02.04: The new name means "having small equal teeth", but the etymology refers to the large inner paired basal teeth of the unguis. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). The closest match with the etymology is the name "equidentatus".

Species folsomi Handschin, 1927

Syn.: Cyphoderus Folsomi(sic) Handschin, 1927 IOS
Syn.: Cyphoderus Folsomi(sic) Delamare Deboutteville, 1948 IOS

Species galapagoensis Jacquemart, S, 1976:153

Species inaequalis Folsom, JW, 1927:12

Species javanus Börner, C, 1906:180

            Syn.: Cyphoderus agnotus Handschin, 1942
            Syn.: Cyphoderus assimilis Yosii, 1953
            Syn.: Cyphoderus assimilis Yosii, R, 1977:162
            Syn.: Cyphoderus borneensis Yoshii, 1980

             Syn.: Cyphoderus javanicus(sic) Schött, H, 1917:51 ISS

Cyphoderus javanus from India
After Mandal GP & Hazra AK, 2009 Fig.41a

Species khaochakanus Jantarit, Satasook & Deharveng, 2014

Syn.: Cyphoderus (bidenticulati(sic)-group) khaochanus(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:58 ISS

Species mucrostrimenus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:54

Species palaciosi Oliveira, Brito & Zeppelini, 2021

Species parafolsomi Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:13

Species pataxo Oliveira, Brito & Zeppelini, 2021

Species similis Folsom, JW, 1927:12

            Syn.: Cyphoderus assimilis Folsom, JW, 1932:71
             Syn.: Cyphoderus nitricola(sic) Rapoport & Izarra, 1962 cf ISS
             Syn.: Cyphoderus nitricola(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 ISS
            Syn.: Cyphoderus similis f. inerme(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:143
             Syn.: Cyphoderus javanus f. inermes(sic) Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:5 ISS
            Syn.: Cyphoderus javanus Lima & Zeppelini, 2015
            Syn.: Cyphoderus javanus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 =?
            Syn.: Cyphoderus javanus de Lima, ECA, Lopes, BCH, Oliveira-Neto, MA, de Mendonça, MC et Zeppelini, D, 2022:25

Cyphoderus similis from the USA
2007.05.11 © Reed, D.

Cyphoderus similis from the USA
2007.05.11 © Reed, D.

Cyphoderus similis from the USA
2007.05.11 © Reed, D.

Cyphoderus similis from the USA
2007.09.03 © McClarin, J.

Cyphoderus similis from the USA
2008.04.05 © Murray, T.

Cyphoderus similis from the USA
2008.05.17 © Montgomery, G.

Cyphoderus similis from the USA
2009.12.06 © Justis, S.

Cyphoderus similis from the USA
2009.12.06 © Justis, S.

Cyphoderus similis from the USA
On Camponotus casteneus
2010.10.31 © Hart, J.

Cyphoderus similis from the USA
2011.12.11 © Kouri, J.

Cyphoderus similis from the USA
2013.05.12 © Liberta, A.

Cyphoderus similis from the USA
In nest of Tapinoma sessile
2015.12.07 © Keim., B

Cyphoderus similis from Canada
With Lasius sp.
2017.06.04 © Guay, P.

Cyphoderus similis from the USA
2019.12.29 © Joll, C.

Cyphoderus similis from the USA
With ants
2019.12.29 © Joll, C.

Cyphoderus similis from Canada
2020.02.21 © Strickland, O.

Cyphoderus similis from the USA
In ant nest
2020.09.28 © Shahan, T.

Cyphoderus similis from the USA
2023.01.21 © Gruver, Be.

Cyphoderus similis from the USA
Mesothorax distinctly wide
With collar of clavate setae
2023.01.21 © Gruver, Be.

Cyphoderus similis from the USA
Ant.3 shaped as pearly bead (*)
2023.01.21 © Gruver, Be.

Species songkhlaensis Jantarit, Satasook & Deharveng, 2014

Species-group tridenticulati(sic) Delamare-Deboutteville, 1948
            Syn.: Cyphoderus tridenticulati(sic)-group Delamare-Deboutteville, 1948 IOS
            Syn.: Cyphoderus tridenticulati(sic)-group Thibaud, J-M et Najt, J, 1987:937 IOS
            Syn.: Cyphoderus tridenticulati(sic)-group Zeppelini & Oliveira, 2016 IOS
            Syn.: Cyphoderus tridenticulati(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 IOS
             Syn.: Cyphoderus Tridenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 ISS
            Syn.: Cyphoderus tridenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:38 IOS
            Syn.: Cyphoderus tridenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 IOS
            Syn.: Cyphoderus tridenticulati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:40 IOS

Species arlei Cassagnau, 1963:138

Syn.: Cyphoderus Arlei(sic) Cassagnau, 1963 IOS

Species bidenticulatus (Parona, 1888) Börner, 1903

            Syn.: Cyphoderus veneris Denis, J-R, 1923:57
            Syn.: Cyphoderus veneris Gisin, H, 1960:259
            Syn.: Cyphoderus bidenticulatus var. veneris Dallai, R, 1973:532
            Syn.: Cyphoderus veneris Moursi & al, 1983
            Syn.: Cyphoderus jugoslavicus Denis, 1936

             Syn.: Cyphoderus bidenticulata(sic) Handschin, 1942 ISS
             Syn.: Cyphoderus bidenticulati(sic) Delamare Deboutteville, 1948 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Jantarit & al, 2014 gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:263,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:640,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Mateos, E, Escuer, P, Busmachiu, G, Riutort, M et Alvarez-Presas, M, 2018:641 ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:3,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:5,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:6,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:8,gr ISS
             Syn.: Cyphoderus bidenticulati(sic) Winkler, D, Mateos, E, Traser, G, Lakatos, F et Tóth, V, 2020:9,gr ISS

Species bidenticulatus2 Womersley, 1934 nec Parona, 1888 sd

Species manuneru Bernard, EC, Soto-Adames, FN et Wynne, JJ, 2015:259

Species mucrominimus de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:48

Species napoensis Thibaud, J-M et Najt, J, 1987:939

Species yoshiiorum Thibaud, J-M et Najt, J, 1987:937

Species-group multidentati(sic) Delamare-Deboutteville, 1948
            Syn.: Cyphoderus multidentati(sic)-group Delamare-Deboutteville, 1948 IOS
            Syn.: Cyphoderus multidentati(sic)-group Cassagnau, 1963 IOS
            Syn.: Cyphoderus multidentati(sic)-group Zeppelini, D et Oliveira, JVLC, 2016:560 IOS
            Syn.: Cyphoderus multidentati(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:47 IOS
             Syn.: Cyphoderus multidentate(sic)-group de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:59 ISS
             Syn.: Cyphoderus Multidentati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:4 ISS
            Syn.: Cyphoderus multidentati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:38 IOS
            Syn.: Cyphoderus multidentati(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:39 IOS
             Syn.: Cyphoderus multidentate(sic)-group Oliveira, JVLC, Zeppelini, D, Castaño-Meneses, G et Palacios-Vargas, JG, 2023:40 ISS

Species caetetus Zeppelini, D et Oliveira, JVLC, 2016:561

Cyphoderus caetetus from Brazil
After Zeppelini D & Oliveira JVLC & al2016 Fig.2

Species pinnatus (Folsom, 1923) Folsom, JW, 1927:13

Species subserratus Delamare Deboutteville, 1948

Genus Pseudocyphoderus Imms, 1912 M

  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
  Syn.: Cyphoderinae Pesudocyphoderus(sic) Mandal, GP et Hazra, AK, 2009:132 ISS
  Syn.: Collembola Pseudocyrphoderus(sic) Thibaud, J-M, 2013:240 ISS

Species annandalei Imms, 1912, t.t.

            Syn.: Pseudocyphoderus Annandalei(sic) Imms, 1912,tt IOS
             Syn.: Pseudocyphoderus Annandalei(sic) Börner, 1913 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Schött, H, 1917:2 ISS
             Syn.: Pseudocyphoderus Annandalei(sic) Delamare Deboutteville, C, 1948:391 ISS

Pseudocyphoderus annandalei from India
After Mandal GP & Hazra AK, 2009 Fig.45a

             Syn.: Pseudocyphoderus annendalei(sic) Mandal, GP et Hazra, AK, 2009:42 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:75 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:79 ISS
             Syn.: Pesudocyphoderus(sic) annendalei(sic) Mandal, GP et Hazra, AK, 2009:132 ISS

Species melittophilus Mari Mutt, JA et Bellinger, PF, 1990:130

Syn.: Pseudocyphoderus melittophilus(sic) Mari Mutt, JA et Bellinger, PF, 1990:130 ISS
Syn.: Pseudocyphoderus mellitophilous(sic) de Oliveira, JVLC, Alves, JLS et Zeppelini, D, 2017:52 ISS

Species squamicaudus (Silvestri, 1917) Thibaud, J-M, 2013:238

Syn.: Pseudocyphoderus sqamicauda(sic) Thibaud, J-M, 2013:238 ISS
Syn.: Pseudocyphoderus squamicauda(sic) Thibaud, J-M, 2013:239 ISS

Species wasmanni Börner, 1913

Syn.: Pseudocyphoderus Wassmanni(sic) Delamare Deboutteville, 1948 ISS
Syn.: Pseudocyphoderus wassmanni(sic) Thibaud, J-M, 2013:240 ISS

Genus Cyphoderodes Silvestri, 1910 M

Species ceylonicus Silvestri, 1910

Species dubius Börner, 1913

Species mitrai Yoshii, 1987

Species punjabicus Yosii & Ashraf, 1964

Species singaporicus Murphy & Yoshii, 1992

Species xenopus (Börner, 1913) Culik, MP & Zeppelini, DF, 2003:1130

Species yayukae (Yoshii, 1987)

Genus Paracyphoderus Delamare Deboutteville, C, 1948:365 M

Species dimorphus (Silvestri, 1910) Delamare Deboutteville, 1948, t.t.

Genus Megacyphoderus Delamare Deboutteville, C, 1948:365 M

Species gallicus Delamare Deboutteville, 1948:368

Species oraniensis Delamare Deboutteville, 1948:366

Species orientalis (Folsom, 1924) Castaño-Meneses, G, Palacios-Vargas, JG & do Carmo, AFR, 2015:394

Species silvestrii (Denis, J-R, 1924:199) Delamare Deboutteville, C, 1948:367

Syn.: Cyphoderus Silvestrii(sic) Denis, J-R, 1924:199 IOS

Species termitum (Wahlgren, E, 1906:19) Delamare Deboutteville, 1948

Genus Serroderus Delamare Deboutteville, C, 1948:369 M

Species sp.1 Janssens F sensu Ospina-Sánchez, 2019

Syn.: Serrodus sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:20

Species acanthotermiensis Delamare Deboutteville, 1948:382

Species alfredi (Yosii, R, 1959:56) Yoshii, 1987

Species alvaradoi Selga, 1962

Species aurantius Delamare Deboutteville, 1948

Syn.: Serroderus humilis var. aurantia(sic) Delamare Deboutteville, 1948 IOS
Syn.: Serroderus humilis var. aurantia(sic) Thibaud, J-M, 2013:236 IOS

Species bilobatus Delamare Deboutteville, 1948:375

Species brevimucronatus (Silvestri, 1917)

Syn.: Cyphoderus arcuatus var. brevimucronata(sic) Silvestri, 1917 IOS
Syn.: Cyphoderus arcuatus var. brevimucronata(sic) Thibaud, J-M, 2013:236 IOS

Species delamarei Cassagnau, 1963:147

Species dicuspiditermitis (Yoshii, 1980) Yayuk, 1989:18

Species distinctus (Denis, 1942) Delamare Deboutteville, 1948, t.t.

Species durio Yoshii, 1987

Species guanobius (Paulian & Delamare Deboutteville, 1948)

Syn.: Cyphoderus humilis var. guanobia(sic) Paulian & Delamare Deboutteville, 1948 IOS
Syn.: Cyphoderus humilis var. guanobia(sic) Thibaud, J-M, 2013:236 IOS

Species hozawai (Kinoshita, 1917) Yosii, R, 1977:162

Serroderus hozawai from Japan
2014.10.02 © Hara, Y.

Species humilis Delamare Deboutteville, C, 1948:371

Species interpositus (Denis, 1942) Delamare Deboutteville, 1948

Species mango Yoshii, 1992

Species multidentatus Delamare Deboutteville, 1948:356

Species protermiensis Delamare Deboutteville, 1948:381

Species quadridentatus (Delamare Deboutteville, 1948:355)

Syn.: Cyphoderus quadridentalus Jacquemart, 1980 ISS

Species sabahnus (Yoshii, 1980) Yayuk, 1989:18

Species socialis Delamare Deboutteville, 1948:379

Species spinatus Christiansen, 1957:79

Species spinicaudus Delamare Deboutteville, 1948

Syn.: Serroderus spinicauda(sic) Thibaud, J-M, 2013:238 ISS

Species sublimis Delamare Deboutteville, 1948:374

Species termitophilus Delamare Deboutteville, C, 1948:378

Syn.: Serroderus interpositus f. termitophila(sic) Delamare Deboutteville, C, 1948:378 IOS
Syn.: Serroderus interpositus f. termitophila(sic) Thibaud, J-M, 2013:236 IOS

Species tridenticulatus Denis, 1948

Genus Cephalophilus Delamare Deboutteville, C, 1948:382 M

Species anoplotermiensis Barra, 1969

Species protermiensis Delamare Deboutteville, 1948:386

Species rectangularis Delamare Deboutteville, 1948:383, t.t.

Genus Calobatinus Silvestri, 1918 M

  Syn.: ... Calobatella Börner, 1913 nec Mik, 1898
  Syn.: Collembola Calobathella(sic) Thibaud, J-M, 2013:240 ISS
  Syn.: ... Calobatana Strand, 1928

  Syn.: ... Calobatina Denis, 1942 ISS

Species erythraeus Silvestri, 1918

            Syn.: Calobatinus rhadinopus var. erythraea(sic) Silvestri, 1918 IOS
            Syn.: Calobatinus erythraea(sic) Delamare Deboutteville, 1948 IOS
            Syn.: Cyphoderus termitum ssp. hadinopus(sic) var. erythraea(sic) Thibaud, J-M, 2013:235 IOS
            Syn.: Calobatinus rhadinopus ssp. erythraea(sic) Thibaud, J-M, 2013:238 IOS
            Syn.: Calobatinus erythraea(sic) Thibaud, J-M, 2013:239 IOS

Species grassei (Denis, 1942) Delamare Deboutteville, 1948

Syn.: Calobatina Grassei Denis, 1942 IOS
Syn.: Calobatinus Grassei Delamare Deboutteville, 1948 IOS

Species occidentalis Silvestri, 1918

Species rhadinopus (Börner, 1913) Silvestri, 1918, t.t.

Syn.: Calobathella(sic) rhadinopus(sic) Thibaud, J-M, 2013:240 ISS

Genus Cyphoderinus Denis, 1942 M

Species fraterculus Denis, 1942, t.t.

Genus Mimoderus Yoshii, 1980 M

Species diusi Yoshii, 1980

Species novaeguinae (Uchida, 1949) Yayuk, RS (sic), 1989:18)

Syn.: Cyphoderus novaguinae(sic) Uchida, 1949 IOS
Syn.: Mimoderus novaguinae(sic) Yayuk, RS(sic), 1989:18) IOS

Species saikehi Yoshii, 1980

Species serratus (Schött, H, 1917:52) Greenslade, P in Houston, WWK, 1994:109

Genus Delamarerus Mitra, SK, 1976:645 M

Syn.: Cyphoderidae Delamareus(sic) Mari Mutt, JA, 1977:230 ISS

Species immsi Mitra, SK, 1976:647, t.t.

Delamarerus immsi from India
After Mitra SK, 1976 Fig.1A

Delamarerus immsi from India
After Mandal GP & Hazra AK, 2009 Fig.46a

Species machadoi (Delamare Deboutteville, 1958) Mari Mutt, JA, 1977:234

Genus Troglobius Palacios-Vargas, JG et Wilson, J, 1990:68

Species albertinoi Cipola NG et Bellini BC in Cipola, NG, de Morais, JW et Bellini, BC, 2016:659

Troglobius albertinoi from Brazil
After Cipola NG & Bellini BC in Cipola NG & al, 2016 Fig.1

Syn.: Troglobius albertonoi(sic) Cipola, NG, de Morais, JW et Bellini, BC, 2016:658 ISS

Species brasiliensis Palacios-Vargas, JG et Zeppelini, D, 1995:173

Species coprophagus Palacios-Vargas, JG et Wilson, J, 1990:69, t.t.

Species ferroicus Zeppelini, D, da Silva, DD et Palacios-Vargas, JG, 2014:3

Troglobius ferroicus from Brazil
Candle-shaped antennal sensilla
After Zeppelini D & al, 2014 Fig.1-2

Tribus Bromacanthini Mitra, 1993 sensu Zhang F & al, 2019:252

Remark by Mitra, S.K. 1993:62: General chaetal pattern of Lepidonella and Bromacanthus comes close to some genera under Lepidocyrtini.

Syn.: Paronellinae Bromocanthini(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:15 ISS
Syn.: Paronellidae Bromocanthini(sic) Soto-Adames, FN et Bellini, BC, 2015:331 ISS

Remark by Zeppelini D & Oliveira J.L.V.C. 2016:569: The tribe Bromacanthini is more similar to some Entomobryidae than to Paronellini+Cyphoderini.

Genus Lepidonella Yosii, 1960

Syn.: Paronellini Microparonella Carpenter, GH, 1916:43

Lepidonella
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.37

Lepidonella sp. from Vietnam
2009 © Boldsystems.org

Lepidonella sp. from Vanuatu
2010 © Deharveng, L. & Bedos, A.

Lepidonella? sp. from New Zealand
2017.12.08 © Navest, M.P.

Lepidonella? sp. from New Zealand
2018.10.22 © Navest, M.P.

Species sp.1 Bellini, BC et Zeppelini, D, 2004:588

Species annulicornis (Oudemans, JT in Weber, M, 1890:87) Mitra, SK, 1992:212

Paronella annulicornis from Java
After Schött H, 1903 Taf.II Fig.5

Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.36

Lepidonella annulicornis from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.37

Species ceylonica (Yosii, 1966) Deharveng, L & Bedos, A, 1995:22

Cyphoderopsis ceylonica from India
After Mandal GP & Hazra AK, 2009 Fig.43a

Lepidonella ceylonica from Ceylon
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.35

Species doveri (Carpenter, 1932) Deharveng & al, 2018

Species duodecimoculata (Prabhoo, 1971) Deharveng, L & Bedos, A, 1995:22

Lepidonella duodecimoculata from India
After Prabhoo in Soto-Adames FN & Bellini BC, 2015 Fig.31

Species flava (Carpenter, GH, 1916:44) Deharveng, L & Bedos, A, 1995:22

            Syn.: Paronella seychellesiensis Salmon, JT, 1964:545
             Syn.: Microparonella seychellesensis(sic) Yoshii, 1981 ISS
             Syn.: Microparonella seychellensis(sic) Yoshii, R et Suhardjno, YR, 1992:48 ISS
             Syn.: Paronella seychelliensis(sic) Nguyen, M et Soto-Adames, FN, 2018:193 ISS

Species incerta (Handschin, 1925) Mitra, SK, 1992:265

Syn.: Lepidonella oudemansi Soto-Adames, FN et Bellini, BC, 2015:337 =?

Species kei (Yoshii, R & Suhardjno, YR, 1992:47) Deharveng, L & Bedos, A, 1995:22

Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A right

Microparonella kei from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.27A left

Syn.: Lepidonella key(sic) Soto-Adames, FN et Bellini, BC, 2015:339 ISS

Species lecongkieti Deharveng, L et Bedos, A, 1995:23

Species marimutti Soto-Adames, FN et Bellini, BC, 2015:334

            Syn.: Lepidonella marimuti(sic) Soto-Adames, FN et Bellini, BC, 2015:334 IOS
            Syn.: Microparonella incerta Mari Mutt, JA, 1987:404

             Syn.: Lepidonella marimuti(sic) Bellini, BC et Cipola, NG, 2017:176 ISS
             Syn.: Lepidonella marimuti(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS

Species nigrofasciata (Handschin, 1928) Deharveng, L & Bedos, A, 1995:22

Lepidonella nigrofasciata from Indonesia
After Handschin in Soto-Adames FN & Bellini BC, 2015 Fig.33

Species oudemansi (Yoshii, 1983) Deharveng, L & Bedos, A, 1995:22

Lepidonella oudemansi from Malaysia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.32

Species picta (Schäffer, C, 1898:410) Mitra, SK, 1992:212

Species subcarpenteri (Denis, 1948) Deharveng, L & Bedos, A, 1995:22

Lepidonella subcarpenteri from Indonesia
After Yoshii in Soto-Adames FN & Bellini BC, 2015 Fig.38

Species tokiokai Yosii, 1960 tt

Lepidonella tokiokai from the Solomon Islands
After Yosii in Soto-Adames FN & Bellini BC, 2015 Fig.34

Species zeppelinii Soto-Adames, FN et Bellini, BC, 2015:332

Lepidonella zeppelinii from Brazil
After Soto-Adames FN & Bellini BC, 2015 Fig.1

             Syn.: Lepidonella zeppelini(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS
             Syn.: Lepidonella zepellinii(sic) Nguyen, M et Soto-Adames, FN, 2018:192 ISS
             Syn.: Lepidonella zeppelini(sic) Nguyen, M et Soto-Adames, FN, 2018:194 ISS

Genus Bromacanthus Schött, 1925

  Syn.: Paronellinae Glacialoca Salmon, JT, 1941:405
  Syn.: ... Handschinella Yosii, R, 1959:53

Bromacanthus
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.35

Species sp.1 Yoshii, 1983

Species sp.2 Yoshii, 1983

Species elongatus Yoshii, 1981

Species flavidulus Yoshii, 1981

Species halmaherae Yoshii, 1983

Pseudoparonella (Irianella) halmaherae from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.24A

Species handschini Schött, 1925, t.t.

Syn.: Pseudoparonella (Bromacanthus) Handschini(sic) Absolon, K et Ksenemann, M, 1942:14 ISS

Species orientalis (Handschin, 1930)

Species palawanicus Yoshii, 1983

Species sago Yoshii, R et Suhardjno, YR, 1992:49

Bromacanthus sago from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.28A

Species seramensis Yoshii, 1989

Species setigerus (Börner, C, 1906:178) Yoshii, 1981

Syn.: Bromacanthus setigera(sic) Mitra, SK, 2003:84 ISS

Species wallacei Yoshii, R et Greenslade, P, 1993:82

Genus Trichorypha Schött, H, 1893:16

Species atrofasciata Schött, H, 1893:18, t.t.

            Syn.: Trichorypha atro-fasciata(sic) Schött, H, 1893:18 IOS
            Syn.: Trogolaphysa lamottei Nguyen, M et Soto-Adames, FN, 2018:184 =?

            Syn.: Trichorypha atro-fasciata(sic) Thibaud, J-M, 2013:237 IOS

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