http://www.collembola.org/taxa/proiinae.htm - Last updated on 2019.01.17 by Frans Janssens
Checklist of the Collembola: Proisotominae

Subfamilia Proisotominae Stach, 1947

Genus Number
of species
Ballistura25
Bellisotoma2
Burmisotoma1
Clavisotoma13
Cliforga1
Dimorphotoma2
Folsomides68
Folsomina5
Granisotoma5
Guthriella1
Isotopenola9
Mucrotoma1
Narynia4
Proisotoma78
Scutisotoma34
Strenzketoma1
Subisotoma16
Villusisotoma2
Proisotominae268
Genera with number of species



Proisotominae from Tasmania
2014.04.08 © Murray, A.
Proisotominae from Tasmania
2014.04.16 © Murray, A.


Genus Villusisotoma Christiansen, KA et Nascimbene, P, 2006:334, -

Species brevis (Christiansen, KA & Nascimbene, P, 2006:335, -) Christiansen, KA & Nascimbene, P, 2006:335, -, t.t.

Species longa Christiansen, KA et Nascimbene, P, 2006:336, -


Genus Burmisotoma Christiansen, KA et Nascimbene, P, 2006:340, -

Species lamellifera (Christiansen, KA & Nascimbene, P, 2006:342, -) Christiansen, KA & Nascimbene, P, 2006:342, -, t.t.


Genus Guthriella Börner, C, 1906:172

Species muskegis (Guthrie, JE, 1903:72) Börner, C, 1906:172, t.t.

Isotoma muskegis ♂ from the USA
After Guthrie JE, 1903 Plate XIII Fig.1
Isotoma muskegis ♀ from the USA
After Guthrie JE, 1903 Plate XIII Fig.2
Isotoma muskegis ♀ from the USA
After Guthrie JE, 1903 Plate XIII Fig.3
Guthriella muskegis ♀ from Panama
After Palacios-Vargas JG & Castaño-Meneses G, 2009 Fig.1A
Guthriella muskegis ♀ from Panama
Abdominal segments 5 and 6
After Palacios-Vargas JG & Castaño-Meneses G, 2009 Fig.1C
Guthriella muskegis ♂ from Panama
After Palacios-Vargas JG & Castaño-Meneses G, 2009 Fig.1B
 
Guthriella muskegis ♂ from Panama
Abdominal segments 5 and 6
After Palacios-Vargas JG & Castaño-Meneses G, 2009 Fig.1D
Guthriella muskegis ♂ from Canada
2009.07.17 Palacios-Vargas, J.G.
Guthriella muskegis ♂ from Canada
2013.12.05 © Proctor, H.
Guthriella muskegis ♂ from Canada
Lateral aspect
2013.12.05 © Proctor, H.
Guthriella muskegis ♂ from Canada
Ventral aspect
2013.12.05 © Proctor, H.



Genus Cliforga Wray, 1952

  Syn.: Isotomidae Ciforga(sic) Poole, RW in Poole, RW et Gentili, P, 1997:50 ISS

Species alleghaniensis Wray, 1952

             Syn.: Cliforga aleghaniensis(sic) Potapov, MB, Fjellberg, A et Bokova, A, 2017:96 ISS



Genus Proisotoma Börner, 1901 key

  Syn.: Degeeriadæ(sic) Isotoma Rainbow, WJ, 1907:313
  Syn.: ... Folsomia(lapsus) Dekinesh, 1991
  Syn.: Isotomidae Parisotoma(lapsus) Frenot & al, 2005


  Syn.: Proisotominae Proisotama(sic) Bano, R, 2016:55 ISS
  Syn.: Proisotominae Proisotama(sic) Bano, R, 2016:60 ISS

Species sp.1 Potapov, M, Babenko, A et Fjellberg, A, 2006:74

Species sp.2 Potapov, 2001

Species alticola (Loksa & Rubio, 1966) Mari Mutt, JA & Bellinger, PF, 1990:64

Species andina Rapoport & Rubio, 1968

Species anopolitana Schulz, H-J et Lymberakis, P, 2006:230

Proisotoma anopolitana from Crete
"Crown" on 5th abdominal segment
After Schulz H-J, 2010 Fig.2

Species atrata (Salmon, JT, 1941:314)

Species bayouensis Mills, HB, 1931:1

Species beta Christiansen & Bellinger, 1980

Species biseta Rapoport, 1963

Species brevidens Stach, J, 1947:204

Proisotoma brevidens from Poland
After Stach, J., 1947 Plate XX Fig.1.

             Syn.: Proisotoma brevicauda Potapov, M et Bogomolov, M, 2016:257 lapsus ISS


Species brevipenna (Mac Gillivray, 1896)

Species bulba Christiansen & Bellinger, 1980

Species burti Fernando, 1960

Species caerulea Salmon, 1955

Species carolii (Tarsia, 1940)

Species clavipila (Axelson, WM, 1903:7) Linnaniemi, WM, 1912:131

            Syn.: Isotoma minuta var. palermitana Linnaniemi, WM, 1912:131 =?


Species coeca da Gama, 1961

Species copiosa de Mendonça, MC, Queiroz, GC et da Silveira, TC, 2015:55

Proisotoma copiosa from Brazil
After de Mendonça MC & al., 2015 Fig.17

Species davidi Barra, J-A, 2001:23

Species decemoculata (Scherbakov, 1899) Denis, JR, 1931:118

             Syn.: Isotoma 10-oculata(sic) Absolon, K, 1901:33 ISS

Species dottrensi Gisin, 1949

Species douglasi de Mendonça, MC, Queiroz, GC et da Silveira, TC, 2015:52

Proisotoma douglasi from Brazil
After de Mendonça MC & al., 2015 Fig.1

Species dualis Potapov, M et Bogomolov, M, 2016:257

Proisotoma dualis from Buryat Republic
Upper: normal form. Lower: mobile form
After Potapov M & Bogomolov M, 2016 Fig.12-13

Species erasoi Izarra, 1968

Species fernandeziana Rubio, 1974

Species fraterna Rusek, J, 1967:193

Species frisoni Folsom, JW, 1937:55

Species fungi Selga, 1962

Species granulata (Salmon, JT, 1943:379) Lawrence

Species granulata (Stach J, 1947:227) Stach, 1955

Proisotoma granulata from Poland
After Stach, J., 1947 Plate XXV Fig.1.

Species haweaensis (Salmon, JT, 1941:314)

Species himalayana Baijal, HN, 1958:353

Proisotoma himalayana from the Himalaya
After Baijal HN, 1958 Fig.5A.

Species immersa (Folsom, JW, 1924:1) Denis, JR, 1931:119

Isotoma immersa from the USA
After Folsom JW, 1924 Fig.1

Species intermedia Izarra, 1969

Species intermixta Handschin, 1928

             Syn.: Proisotoma intermixa(sic) Denis, JR, 1931:118 ISS

Species juaniae Lucianez & Simón, 1992

Species judeana Palissa, 2006

Species koepckei Winter, 1967

            Syn.: Proisotoma koepckei Winter, 1963 nn


Species lutii Izarra, 1973

Species macedoi Winter, 1967

Species mauretanica Handschin, 1925

Species micrura Börner, 1907

Species minima (Absolon, K, 1901:33) Linnaniemi, 1907, B

            Syn.: Isotoma mirabilis Börner, C, 1901:44
            Syn.: Proisotoma americana Mills, HB, 1934:51
            Syn.: Folsomidiella inaequalis Bagnall, 1949


Proisotoma minima from Poland
After Stach, J., 1947 Plate XXI Fig.1.
Proisotoma minima from Russia
Upper: normal form. Lower: mobile form
After Potapov M & Bogomolov M, 2016 Fig.10-11

Species minuta (Tullberg, 1871) Linnaniemi, 1907, t.t., B

            Syn.: Isotoma pallida Moniez, 1894
            Syn.: Isotoma notabilis Scherbakov, AM, 1898:11 nec Schaeffer
Isotoma notabilis from Russia
After Scherbakov AM, 1898 Table II Fig.17


            Syn.: Isotoma stagnalis Willem, V, 1900:28,B
             Syn.: Isotoma tagnales(sic) Dekinesh, 1991 ISS
            Syn.: Isotoma stagnalis var? tenebricola Willem, V, 1900:29,B
            Syn.: Isotoma (Proisotoma) stagnalis f. principalis Börner, C, 1901:133,B
            Syn.: Isotoma stscherbakowi Börner, 1901
             Syn.: Isotoma tscherbakow(sic) Dekinesh, 1991 ISS
            Syn.: Isotoma clavipila Handschin, E in Dahl, F, 1929:61 =?
            Syn.: Isotoma clavipila Gisin, H, 1960:197 =?
            Syn.: Isotoma minima Guthrie, JE, 1903:63 nec Absolon, 1901
            Syn.: Isotoma minuta var. palermitana Börner, 1903
            Syn.: Isotoma lineata Wahlgren, E, 1906:6
            Syn.: Isotoma troglodytica Rainbow, WJ, 1907:313
Isotoma troglodytica from Australia
After Rainbow WJ, 1907 Fig.49

             Syn.: Isotoma troglodytes(sic) Wolf, 1934-38 ISS
            Syn.: Proisotoma guthriei Linnaniemi, WM, 1912:132
             Syn.: Folsomia guthrei(sic) Scott, 1942 ISS
             Syn.: Folsomia guthrieri(sic) Yosii, 1959 ISS
            Syn.: Proisotoma oeensis Caroli, 1914 =?
             Syn.: Isotomina oeenensis(sic) Stach, J, 1947:269 ISS
             Syn.: Isotomina ooensis(sic) Salmon, JT, 1964:356 ISS
            Syn.: Proisotoma oeensis var. mauretanica Potapov, M, 2001:409 =?
            Syn.: Proisotoma minuta f. inermis Denis, JR, 1931:119
            Syn.: Proisotoma minuta f. meridionalis Denis, JR, 1931:119
            Syn.: Proisotoma minuta f. principalis Denis, 1931
            Syn.: Isotoma wahlgreni Salmon, JT, 1964:408
            Syn.: Proisotoma minuta ssp. rhopalotricha Massoud & Rapoport, 1968
            Syn.: Proisotoma? sp. Greenslade, 1987
            Syn.: Proisotoma sp. Greenslade, 1990



Proisotoma minuta from Poland
After Stach, J., 1947 Plate XXII Fig.1.
Proisotoma minuta from China
After Stach J, 1964 Plate VII Fig.5

             Syn.: Proisotoma munuta(sic) Yoshii, R et Suhardjno, YR, 1992:12 ISS
             Syn.: Parisotoma(lapsus) minutea(sic) Frenot & al, 2005 ISS
Proisotoma minuta from the UK
2005 © Hopkin, S.P.
Proisotoma minuta from the UK
2005 © CEH.
Proisotoma minuta from the UK
1+1 anterior manubrial setae
6 anterior dental setae
2005 © Hopkin, S.P.
Proisotoma minuta from the USA
2007 © Murray, T.
Proisotoma minuta from the UK, juvenile
2007.10.03 © Valentine, B
Proisotoma minuta from the UK
2007.12.08 © Valentine., B
Proisotoma minuta from the UK
2007.12.28 © Valentine., B
Proisotoma minuta from the UK
2007.12.30 © Valentine., B
Proisotoma minuta from the UK
2008.07.03 © Valentine., B
Proisotoma minuta from the UK
2008.11.12 © Valentine., B
Proisotoma minuta from France
2008.12.07 © Jacquemin, G.
Proisotoma minuta juv. from the USA
2009.03.03 © Justis, S.
Proisotoma minuta juv. from the USA
2009.03.03 © Justis, S.
Proisotoma minuta juv. from Italy
After Beccati M et al, 2011 Fig.2
Proisotoma minuta from the USA
2012.02.15 © Kouri, J.
Proisotoma minuta from the USA
2012.04.30 © Murray, T.
Proisotoma minuta from the USA
2012.04.30 © Murray, T.
Proisotoma minuta juv. from Canada
2014.01.26 © Hochhalter, R.
Proisotoma minuta from France
2015.01.31 © Legrand, T.
Proisotoma minuta from the UK
2015.11.08 © Valentine., B
Proisotoma minuta juv. from the UK
Elongate eyepatch with ocelli in 2 groups
2015.11.10 © Valentine., B
Proisotoma minuta juv. from Canada
First instar (0.3mm)
2016.02.12 © Hochhalter, R.
Proisotoma minuta juv. from Canada
First instar (0.3mm)
Pale eyepatch: 8 ocelli, 2 anterior touching
2016.02.12 © Hochhalter, R.
Proisotoma minuta juv. from Canada
Second instar (0.4mm)
Pale eyepatch: 8 ocelli, 2 anterior touching
2016.02.12 © Hochhalter, R.
Proisotoma minuta from Germany
2017.01.28 © Melzer, L.
Proisotoma minuta from Belgium
2017.08.09 © Huskens, M.-L.
Proisotoma minuta from Belgium
2018.01.04 © Huskens, M.-L.
Proisotoma minuta from Belgium
Anterior 2 ocelli osculating (*)
2018.01.04 © Huskens, M.-L.
Proisotoma minuta from France
2018.10.25 © Garcelon, P.
Proisotoma minuta from France
Furca not reaching up to collophore
2018.10.25 © Garcelon, P.



Species multituberculata Martynova, 1971

Species musci Lawrence, PN, 1969:547

Proisotoma musci from the Solomon Islands
After Lawrence, P.N., 1969 Fig.1.

Species muscicola Stach, J, 1965:353

Species najtae Selga, 1971

Species niger (Carpenter, GH, 1925:8)

Species nigromaculosa Folsom, JW, 1932:62

Species nova Palissa, 1969

Species octojuga Salmon, JT, 1949:33


Cryptopygus okukensis from New Zealand
After Salmon, J.T. 1941 Pl.48 Fig.141.
Species okukensis (Salmon, JT, 1941:313)

Species oliveirae Deharveng, L, 1984:123

Species pajonalica Loksa & Rubio, 1966

Species pakurensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2017:98

Proisotoma pakurensis from India
After Mandal GP, Suman KK & Bhattacharya KK, 2017 Plate.1 Fig.1

Species papillosa Stach, 1937

Species paronai Börner, 1907

            Syn.: Isotoma palustris Parona, 1895


             Syn.: Proisotoma Paronai(sic) Denis, JR, 1931:119 ISS

Species patagonica (Wahlgren, 1900) Stach, J, 1947:217

Species perparva Jackson, 1927

Species pettersonae Christiansen, KA et Nascimbene, P, 2006:340, -

Species quinocellata Salmon, 1954

Species ramosi Arlé, R, 1960:1960:166

Species ridleyi Denis, 1933

Species ripicola Linnaniemi, WM, 1912:128

            Syn.: Isotoma agilis Scherbakov, 1899


Proisotoma ripicola from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573
Proisotoma ripicola from Corsica
After Schulz H-J, 2015 Abb.6

Species sagardoyi Izarra, 1981, n.n.

Species santosorum Palacios-Vargas, JG et Arbea, JI, 2009:449

Proisotoma santosorum from Mexico
After Palacios-Vargas JG & Arbea JI Fig.9

            Syn.: Proisotoma minima Palacios-Varga, 1991 ca



Species senetijohani Baijal & Chandra, 1970

Species sepulcralis (Folsom, JW, 1902:364) Folsom, JW, 1937:6

             Syn.: Isotoma sepulchuraris(sic) Bueker, 1939 ISS
             Syn.: Proisotoma (Proisotoma) sepalcralis(sic) Salmon, JT, 1964:357 ISS
             Syn.: Proisotoma sepulchralis(sic) Christiansen, K et Bellinger, P, 1980:562 ISS
             Syn.: Proisotoma (Proisotoma) sepulchralis(sic) Christiansen, K et Bellinger, P, 1980:575 ISS
             Syn.: Proisotoma sepulchralis(sic) Poole, RW in Poole, RW et Gentili, P, 1997:52 ISS
             Syn.: Proisotoma sepulchralis(sic) Christiansen, K et Bellinger, P, 1998:623 ISS
             Syn.: Proisotoma (Proisotoma) sepulchralis(sic) Christiansen, K et Bellinger, P, 1998:635 ISS
             Syn.: Proisotoma sepulchralis(sic) Robles & al, 2012 ISS
             Syn.: Proisotoma sepulchralis(sic) Palacios-Vargas, JG, Ortiz-Ceballos, AI et Salvador-Escobedo, I, 2016:457 ISS

Species subdeflexa Delamare Deboutteville, 1951

Species subminuta Denis, JR, 1931:112

            Syn.: Isotoma stagnalis Yoshii, 1992 =?
             Syn.: Isotoma tenebricosa(sic) Yoshii, 1992 =? ISS


Proisotoma subminuta from Spain
After Arbea, J.I., 2006 Fig.1

Species submuscicola Nguyen, 1995

Species tenella (Reuter, 1895) Linnaniemi, 1912

            Syn.: Isotoma clavata Schött, H, 1893:73
            Syn.: Proisotoma (Proisotoma) simplex Folsom, JW, 1937:61
            Syn.: Proisotoma alba Yosii, 1939


Proisotoma tenella from Poland
After Stach, J., 1947 Plate XXVI Fig.1.

Species terrigena (Salmon, JT, 1943:379)

Species turikana Palacios-Vargas, JG et Arbea, JI, 2009:446

Proisotoma turikana from Venezuela
After Palacios-Vargas JG & Arbea JI Fig.1

            Syn.: Proisotoma sp. Viloria & al, 1992



Species vestita (Brown, 1923)

Species violenta Banasco, 1985,cf

Species xanthella Salmon, JT, 1949:34


Genus Bellisotoma Soto-Adames, FN, Giordano, R et Christiansen, K, 2013:8

Species ewingi (Folsom in James, 1933) Soto-Adames, FN, Giordano, R & Christiansen, K, 2013:12

Species joycei (Soto-Adames & Giordano, 2011) Soto-Adames, FN, Giordano, R & Christiansen, K, 2013:10, t.t.


Genus Folsomides Stach, 1922 key

  Syn.: Proisotominae Varisotoma Salmon, JT, 1964:124
  Syn.: Proisotominae Varisotoma Salmon, JT, 1964:351


Folsomides sp. from Australia
Scale bar = 0.1 mm
1999 © Walter, D.E.
Folsomides sp. from Australia
Head; scale bar = 0.01 mm
1999 © Walter, D.E.
Folsomides sp. from Indonesia
2008.06.29 © Nurtjahya, E.

Species sp.1 Greenslade, P in Houston, WWK, 1994:72

            Syn.: Folsomides marchicus Greenslade P in Low, 1978


Species sp.2 Arbea, JI et Blasco-Zumeta, J, 2001:41

Species sp.3 Schneider, C et Deharveng, L, 2017:13

            Syn.: Folsomides americanus Schneider, C et Deharveng, L, 2017:2 nr

Species almanzorensis Lucianez & Simón, 1991

Species alvarezi Selga, 1960

Species analuisae Fjellberg, A, 1993:470

Species andinensis Díaz, A et Najt, J, 1984:139

Species angularis (Axelson, WM, 1905:791) Arbea, JI, 2015:413, B

            Syn.: Folsomides decemoculatus Mills, HB, 1935:134
             Syn.: Folsomides (angularis-group) decemoculata(sic) Arbea, JI, 2015:413 ISS
            Syn.: Proisotoma marchica Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:9
            Syn.: Folsomides millsi Paclt, 1954


Folsomides angularis from Russia
After Potapov M & Stebaeva S, 1997 Fig.18
Folsomides angularis from the UK
2018.10.04 © Shaw, P.

Species arenosus Martynova, 1979

Species arenus Potapov & Thibaud, 2003

Species aridoviator Potapov, M et Stebaeva, S, 1997:126

Folsomides aridoviator from Russia
After Potapov M & Stebaeva S, 1997 Fig.19

Species arnoldi Suhardjono & Greenslade P, 1994:22

Species asiaticus Martynova, 1970

Species ayllonensis Simón & Luciáñez, 1990

Species californicus Palacios-Vargas, JG et Villarreal-Rosas, J, 2013:481

            Syn.: Isotomidae Folsomides Villarreal-Rosas, J, Palacios-Vargas, JG et Maya, Y, 2014:516


Species cenkeh Yoshii, R et Suhardjno, YR, 1992:12

Species centralis (Denis, JR, 1931:111) Greenslade P & Thompson, 1981

            Syn.: Proisotoma sundana Handschin, 1932
            Syn.: Parisotoma canituda Salmon, 1951
            Syn.: Proisotoma centralis ssp. epiphyta Winter, 1967
            Syn.: Proisotoma (Proisotoma) woodgeri Rapoport & Maño, 1969
             Syn.: Proisotoma wooddgeri(sic) Gruia, M, 1983:195 ISS
            Syn.: Proisotoma sp. Heatwole, Done & Cameron, 1981


Folsomides centralis from Vanuatu
2006 © Boldsystems.org

Species chichinautzini Kovác, L et Palacios-Vargas, JG, 1996:601

            Syn.: Folsomides decemoculatus Palacios-Vargas, 1978
            Syn.: Folsomides variabilis Palacios-Vargas, 1979
            Syn.: Folsomides angularis Palacios-Vargas, JG, 1981:87


Species croci Potapov & Thibaud, 2003

Species cumbrosus Fjellberg, A, 1993:463

Species decemoculatus (Scherbakov, 1899) Ellis, WN, 1974:118

Species deflexus Schött, 1927

Species delamarei Thibaud, J-M, Najt, J et Jaquemart (sic), S (+), 1994:200

             Syn.: Folsomides dealmarei(sic) Pomorski, RJ, 2001:265 ISS

Species denisi (Womersley, H, 1935:215) Greenslade, P in Houston, WWK, 1994:72

Astephanus denisi from Australia
After Womersley H, 1935 Fig.4e

Species deserticolus Wood, TG, 1970:79

            Syn.: Folsomides deserticola(sic) Wood, TG, 1970:79 IOS
Folsomides deserticola(sic) from Australia
After Wood, T.G, 1970 Fig.1




             Syn.: Folsomides deserticola(sic) Wood, 1971 ISS
             Syn.: Folsomides deserticola(sic) Greenslade PJM & Greenslade P, 1973 ISS
             Syn.: Folsomides deserticola(sic) Ellis, WN, 1974:118 ISS
             Syn.: Folsomides deserticola(sic) Jacquemart, S, 1974:41 ISS
             Syn.: Folsomides deserticola(sic) Greenslade P, 1981 ISS
             Syn.: Folsomides deserticola(sic) Greenslade, P in Houston, WWK, 1994:72 ISS
             Syn.: Folsomides deserticola(sic) Thibaud, J-M, Najt, J et Jaquemart(sic), S (+), 1994:201 ISS
             Syn.: Folsomides deserticola(sic) Pomorski, RJ, 2001:265 ISS



Species ellisi Arbea, JI, 2015:416

Folsomides ellisi from Ibiza
After Arbea JI, 2015 Fig.4
            Syn.: Folsomides angularis Poinsot & Barra, 1982 partim
            Syn.: Folsomides marchicus Nosek, 1961
            Syn.: Folsomides marchicus Dallai, 1967
            Syn.: Folsomides marchicus Ellis, WN, 1974:129
            Syn.: Folsomides xerophilus Fjellberg, A, 1993:462 partim
            Syn.: Folsomides xerophilus da Gama & al., 1997
            Syn.: Folsomides xerophilus Potapov, 2001 partim


Species famarensis Fjellberg, A, 1993:467

Species fjellbergi Arbea, JI, 2015:414

Folsomides fjellbergi from Tenerife
After Arbea JI, 2015 Fig.2
            Syn.: Folsomides angularis Nosek, 1961
            Syn.: Folsomides angularis Poinsot-Ballaguer & Barra, 1978 partim
            Syn.: Folsomides angularis Poinsot & Barra, 1982
            Syn.: Folsomides angularis Arbea, JI et Jordana, R, 1990:195
            Syn.: Subisotoma variabilis ssp. psammophila Loksa & Bogojevic, 1970
            Syn.: Folsomides portucalensis Fjellberg, A, 1993:461
            Syn.: Folsomides portucalensis Potapov, M et Stebaeva, S, 1997:126
            Syn.: Folsomides portucalensis Arbea, JI et Blasco-Zumeta, J, 2001:35
            Syn.: Folsomides portucalensis Potapov, 2001
            Syn.: Folsomides portucalensis Dányi & Traser, 2008
            Syn.: Folsomides portucalensis Potapov, 2013


Species graminis Fjellberg, A, 1993:466

Species halshinicus Arbea, JI et Kahrarian, M, 2015:283

Folsomides halshinicus from Iran
After Arbea JI & Kahrarian M, 2015 Fig.1

Species insularis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:85

Species intermedius Fjellberg, A, 1993:468

Species lawrencei Gers & Deharveng, 1985

Species marchicus (Frenzel, 1941) Stach, 1947

            Syn.: Subisotoma petiti Delamare Deboutteville, 1948


             Syn.: Folsomides marchica(sic) Poole, RW in Poole, RW et Gentili, P, 1997:51 ISS
Folsomides aff. marchicus from Iran
After Kahrarian M & Arbea J, 2013 Fig.9



Species mediterraneus Arbea, JI et Jordana, R, 2002:2

Species monosetis Massoud & Rapoport, 1968

Species mosambicensis Cardoso, MA, 1966:87

Species nanus Ellis, WN, 1974:115

Species neozealandia Salmon, 1948

Species nepalicus Yosii, 1971

Species nigrocellatus Fjellberg, A, 1993:466

Species ononicolus Fjellberg, A, 1993:467

Species oromii Fjellberg, A, 1993:466

Species parvulus Stach, 1922, t.t.

            Syn.: Folsomides americanus Denis, 1931
             Syn.: Folsomides americana(sic) Womersley, H, 1935:214 ISS
             Syn.: Folsomides americana(sic) Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1999:162 ISS
            Syn.: Folsomides exiguus Folsom, JW, 1932:58
Folsomides exiguus from Australia
After Womersley H, 1935 Fig.4b
Folsomides exiguus
2006 © Boldsystems.org

            Syn.: Folsomides parvus Folsom JW in Mills, 1934:43-45
Folsomides parvus from Poland
After Stach, J., 1947 Plate X Fig.1.

             Syn.: Folsomides purvus(sic) Bano, R, 2016:59 ISS
            Syn.: Folsomides stachi Folsom JW in Mills, 1934:45-46
             Syn.: Folsomides Stachi(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:379 ISS
            Syn.: Folsomides exiguus ssp. eburneus Delamare Deboutteville, 1948
             Syn.: Folsomides anopthalamis(sic) Yayuk, RS(sic), 1989:126 ISS
            Syn.: Folsomides pseudoparvulus Martynova, 1978



Folsomides parvulus
2006 © Boldsystems.org
Folsomides parvulus from the USA
2012.05.16 © Kouri, J.D.
Folsomides parvulus from the USA
2012.09.30 © Kouri, J.D.
Folsomides parvulus from Iran
After Kahrarian M & Arbea J, 2013 Fig.8

             Syn.: Folsomides purvulus(sic) Bano, R, 2016:55 ISS
             Syn.: Folsomides purvulus(sic) Bano, R, 2016:59 ISS
Folsomides parvulus from India
After Bano R, 2016 p.59



Species petiti (Delamare Deboutteville, 1948) Wood, TG, 1970:81

Species pinicolus Fjellberg, A, 1993:468

Species pocosensillatus Fjellberg, A, 1993:465

Species portucalensis da Gama, 1961

            Syn.: Subisotoma psammophila Loksa & Bogojevic, 1970 =?


             Syn.: Folsomides portulacensis(sic) Petersen, H, 2011:475,cf ISS
Folsomides portucalensis from Portugal
After Arbea JI, 2015 Fig.1



Species pseudangularis Chen, 1985

Species quinocellatus Salmon, 1954

Species semiparvulus Fjellberg, A, 1993:460

Species sexophthalmus (Womersley, 1934) Wood, TG, 1970:82

             Syn.: Folsomides sexophthalma(sic) Wood, TG, 1970:82 ISS
             Syn.: Folsomides sexophthalma(sic) Greenslade, P, 1974:259 ISS
             Syn.: Folsomides sexophtalmus(sic) Pomorski, RJ, 2001:265 ISS

Species socorrensis Kovác, L et Palacios-Vargas, JG, 1996:598

            Syn.: Folsomides angularis Poinsot-Balaguer & Barra, 1982
            Syn.: Folsomides angularis Palacios-Vargas et al., 1982


Species subvinosus Arbea, JI et Kahrarian, M, 2015:285

Folsomides subvinosus from Iran
After Arbea JI & Kahrarian M, 2015 Fig.3

Species supranubius Fjellberg, A, 1993:469

Species teno Fjellberg, A, 1993:466

Species teres Christiansen & Bellinger, 1980

Species terrus Fjellberg, A, 1993:464

Species tonellus Fjellberg, A, 1993:468

Species troglobius (Rapoport & Maño, 1969:123) Palacios-Vargas, JG, 1999:36

Species unicus Fjellberg, A, 1993:464

Species urumqiensis Hao & Huang, 1995

Species vinosus Fjellberg, A, 1993:462

Folsomides vinosus from Iran
After Arbea JI & Kahrarian M, 2015 Fig.5

Species viridescens Pomorski, RJ, 2001:265

Species virungiensis Martynova, 1978

Species xerophilus Fjellberg, A, 1993:462 partim

Folsomides xerophilus from Tenerife
After Arbea JI, 2015 Fig.3

Species yucatanicus Kovác, L et Palacios-Vargas, JG, 1996:595

Species zairensis Martynova, 1978


Genus Folsomina Denis, 1931 key

Species infelicia Greenslade, P, 1999:4

Folsomina infelicia from Malaysia
After Greenslade, P., 1999 Fig.1.

Species lawrencei Greenslade, P, 1999:6

            Syn.: Folsomina onychiurina Lawrence, PN, 1969:550
Folsomina onychiurina from the Solomon Islands
After Lawrence, P.N., 1969 Fig.11.



Species onychiurina Denis, 1931, t.t., B

            Syn.: Folsomina yosiii Lawrence, PN, 1969:552
Folsomina yosiii from the Solomon Islands
After Lawrence, P.N., 1969 Fig.19.

             Syn.: Folsomina yosii(sic) Greenslade, P, 1974:260 ISS
             Syn.: Folsomina yosii(sic) Lasebikan, BA, 1974:266 ISS
             Syn.: Folsomina yosii(sic) Yayuk, RS(sic), 1989:119 ISS
             Syn.: Folsomina yosii(sic) Greenslade, P, 1999:10 ISS



             Syn.: Folsomina onychurina(sic) Potapov, MB et Greenslade, P, 2010:19 ISS

Species wuyanensis Zhao & Tamura, 1992

Remark by Greenslade, P. 1999:12: The status of this species is left unchanged, although it is possible that F. wuyanensis is a synonym of F. onychiurina.

Species yongxingensis Chen, 1987


Genus Scutisotoma Bagnall, 1949 key

  Syn.: Proisotoma Dimorphiella Grinbergs, A, 1968:139 nec Valkanov, 1928
  Syn.: Proisotoma Dimorphotoma Grinbergs, A, 1975:101

Species sp.1 Potapov, M, Babenko, A et Fjellberg, A, 2006:74

Species acorrelata Potapov, M, Babenko, A et Fjellberg, A, 2006:41

Species ananevae (Babenko & Bulavintsev, 1992) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species armeriae (Fjellberg, 1976) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species baica Potapov, M, Babenko, A et Fjellberg, A, 2006:49

Species bengei Huang & Potapov, 2012

Species champi Soto-Adames & Giordano, 2011

Species christianseni (Stach, 1959) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species dodecocellata Potapov, M, Babenko, A et Fjellberg, A, 2006:42

Species fjellbergi (Dunger, WG, 1982:54) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species indigirka Potapov, M, Babenko, A et Fjellberg, A, 2006:51

Species karadagi Potapov, M, Babenko, A et Fjellberg, A, 2006:24

Species kolymica Potapov, M, Babenko, A et Fjellberg, A, 2006:28

Species ladaki (Denis, JR, 1936:262) Potapov, M, Babenko, A & Fjellberg, A, 2006:21

Proisotoma ladaki from India
After Denis JR, 1936 Fig.5

Species longisensilla Potapov, M, Babenko, A et Fjellberg, A, 2006:30

Species matalini Potapov, M, Babenko, A et Fjellberg, A, 2006:36


Scutisotoma millimetrica from Russia
After Potapov, M. et al. 2006 Fig.80.
Species millimetrica Potapov, M, Babenko, A et Fjellberg, A, 2006:31

Species montana Potapov, M, Babenko, A et Fjellberg, A, 2006:45

Species muriphila (Grinbergs, A, 1968:139) Potapov, M, Babenko, A & Fjellberg, A, 2006:19

Species nicksmeti Potapov, M, Babenko, A et Fjellberg, A, 2006:52

Species oirota (Vilkamaa, 1988) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species potapovi Xie, G-L et Chen, J-X, 2008:?

Species postertriplex Babenko & Potapov, 2012

Species robustodens Huang & Potapov, 2012

Species schisti Potapov, M, Babenko, A et Fjellberg, A, 2006:34

Species stachanoremi Potapov, M, Babenko, A et Fjellberg, A, 2006:47

Species stepposa (Martynova, 1975) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Scutisotoma stepposa ♂ from Russia
'Super' male
After Chimitova A & Potapov M, 2011 Fig.2
Scutisotoma stepposa ♂ from Russia
'Modest' male
After Chimitova A & Potapov M, 2011 Fig.3
Scutisotoma stepposa ♂ from Russia
'Neutral' male
After Chimitova A & Potapov M, 2011 Fig.4
Scutisotoma stepposa ♀ from Russia
After Chimitova A & Potapov M, 2011 Fig.8

Species subarctica (Gisin, 1950) Potapov, M, Babenko, A & Fjellberg, A, 2006:16

Species tenuidentifera Potapov, M, Babenko, A et Fjellberg, A, 2006:39

Species tianshanica (Martynova, 1971) Potapov, M, Babenko, A & Fjellberg, A, 2006:17

Species titusi (Folsom, JW, 1937:49) Bagnall, 1949, t.t.

Scutisotoma titusi from the USA
2014.10.04 © Eiseman, C.
Scutisotoma titusi from the USA
2014.10.04 © Eiseman, C.
Scutisotoma titusi from the USA
Prothorax not greatly reduced
2014.10.04 © Eiseman, C.

Species tolerans (Babenko, 1994) Potapov, M, Babenko, A & Fjellberg, A, 2006:17

Species trichaetosa Huang & Potapov, 2012

Species variabilis (Gisin, 1949) Potapov, M, Babenko, A & Fjellberg, A, 2006:17

  Subspecies psammophila Loksa, I et Bogojevic,J, 1970:128


Genus Mucrotoma Rapoport, EH et Rubio, IR, 1963:122

Species leuca Rapoport, EH et Rubio, IR, 1963:122, t.t.

Mucrotoma leuca from Chile
After Rapoport EH & Rubio IR, 1963 Fig.81


Genus Subisotoma Stach, 1947 key

Species sp.1 Potapov, M, Babenko, A et Fjellberg, A, 2006:74

Species sp.2 Potapov, M, Babenko, A et Fjellberg, A, 2006:74

Species sp.3 Potapov, M, Babenko, A et Fjellberg, A, 2006:74

Species sp.4 Potapov, M, Babenko, A et Fjellberg, A, 2006:74


Species-group asiatica
Species asiatica (Martynova, 1970) Fjellberg, 1993

Species cruda Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:23

            Syn.: Subisotoma sp. Fjellberg, 1993


Species erratica Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:25


Species-group pusilla
Species bisensillata Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:15

Subisotoma bisensillata from ???
After Potapov M et al., 2009 Fig.13

Species guzeriplica Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:17

Species homonomica Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:13

Species multisensillata Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:20

            Syn.: Folsomides pusillus Stebeava, 1973


Species pomorskii Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:11

Species posteriomollis Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:19

Species pusilla (Schäffer, 1900) Stach, J, 1947:111, t.t.

Subisotoma pusilla from Poland
After Stach, J., 1947 Plate XII Fig.1.

Species quadrisensillata Gao, Y, Xiong, Y et Potapov, MB, 2009:59

Subisotoma quadrisensillata from China
After Gao Y & al., 2009 Fig.14

Species tenuis (Dunger, WG, 1982:51) Fjellberg, 1993


Genus Isotopenola Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:27 key

Species sp.1 Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:10

Species sp.2 Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:10

Species sp.3 Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:10

Species sp.4 Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:10

Isotopenola sp.4 from New Zealand
After Potapov M et al., 2009 Fig.14
            Syn.: Cryptopygus terrigenus Salmon, JT, 1943:379 =?


Species australis (Womersley, 1934) Potapov, M, Babenko, A, Fjellberg, A & Greenslade, P, 2009:28, t.t.

Species delicata Potapov, M, Babenko, A, Fjellberg, A et Greenslade, P, 2009:34

            Syn.: Cryptopygus sp. Farrow & Greenslade, 1992


Species loftyensis (Womersley, 1934) Potapov, M, Babenko, A, Fjellberg, A & Greenslade, P, 2009:33

Species nilgiris (Denis, 1947) Potapov, M, Babenko, A, Fjellberg, A & Greenslade, P, 2009:36

Species perterrens Greenslade & Potapov, 2012


Genus Ballistura Börner, C, 1906:172

  Syn.: Isotominae Ballistrura(sic) Salmon, JT, 1945:68 ISS
  Syn.: Isotomidae Ballistrura(sic) Poinsot, N, 1972:678 ISS
  Syn.: Isotomidae Ballistrura(sic) Hazra & Bhattacharya, 2003 ISS
Ballistura sp. from Estonia
2008.05.14 © Tartes, U.

  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:19 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:26 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:73 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:77 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:84 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:85 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:88 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:90 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:91 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:93 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:94 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:95 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:96 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:97 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:104 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:107 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:115 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:116 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:117 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:118 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:119 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:133 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:134 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:135 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:136 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:137 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP et Hazra, AK, 2009:180 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:44 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:46 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:47 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:48 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:50 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:53 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:54 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:55 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:56 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:57 ISS
  Syn.: Isotomidae Ballistrura(sic) Mandal, GP, Suman, KK et Hazra, AK, 2011:58 ISS
Ballistura sp. from Mexico
After Palacios-Vargas JG & Castaño-Meneses G, 2014 Fig.1
Ballistura sp. from France
2017.10.28 © Picard, J.
Ballistura sp. from France
2017.10.28 © Picard, J.



Species sp. González, V, Díaz, M et Palacios-Vargas, JG, 1997:47

Species alpa (Christiansen & Bellinger, 1980) Soto-Adames & Giordano, 2011

             Syn.: Folsomia alpha(sic) Christiansen, KA in Dindal, DL, 1990:985 ISS

Species albertinae (Ellis, 1976:291) Potapov, 2001

Species antiqua Folsom, JW, 1937:34


Proisotoma aqualata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.220.
Species aqualata (Salmon, JT, 1941:333) Salmon, JT, 1945:69


Species bengalensis Yosii, 1966

Ballistrura(sic) bengalensis from India
After Mandal GP & Hazra AK, 2009 Fig.25a

Species borealis (Axelson, WM, 1905:791) Potapov, 2001

Species corsica (Poinsot & Barra, 1982) Potapov, M, Babenko, A & Fjellberg, A, 2006:13

Species excavata (Folsom, JW, 1937:41)

Species extra (Christiansen & Bellinger, 1980) Fjellberg, A, 1984:86

Species fitchi (Denis, 1933) Potapov, 2001

            Syn.: Proisotoma fitchioides Denis, 1947

Species gisini (da Gama, 1964) Potapov, M, Babenko, A & Fjellberg, A, 2006:13

Species hankoi (Stach, J, 1930:288) Dányi, L & Traser, G, 2008:45

            Syn.: Proisotoma Hankói(sic) Stach, J, 1930:288 IOS


             Syn.: Proisotoma Hankoi(sic) Denis, JR, 1931:115 ISS



Species libra (Christiansen & Bellinger, 1980) Skidmore, 1995:54

Species meridionalis (Dallai, R, 1973:510) Potapov, M, Babenko, A & Fjellberg, A, 2006:13

Folsomides meridionalis from Italy
After Dallai R, 1973 Fig.7a

Species navacerradensis (Selga, 1962) Potapov, M, Babenko, A & Fjellberg, A, 2006:13

  Subspecies pratensis Zivadinovic, 1972

Species obtusicauda (Schäffer, 1897)

Species pratensis (Betsch, J-M et Massoud, Z, 1970:210)

Species rossi Soto-Adames & Giordano, 2011

Species schoetti (von Dalla Torre, 1895) Stach, J, 1947:237, t.t.

            Syn.: Isotoma Schöttii(sic) von Dalla Torre, 1895,tt IOS
            Syn.: Isotoma litoralis(sic) Schött, H, 1893:75
Isotoma litoralis(sic) from Sweden
After Schött H, 1893 Taf.VII Fig.1

            Syn.: Isotoma lacustris Schött, 1896



             Syn.: Isotoma Schötti(sic) Willem, 1902 ISS
             Syn.: Proisotoma schötti(sic) Linanniemi, 1907 ISS
             Syn.: Isotoma Schötti(sic) Imms, AD in Herdman, WA, 1906:77 ISS
             Syn.: Proisotoma Schötti(sic) Denis, JR, 1924:197 ISS
             Syn.: Proisotoma Schotti(sic) Denis, J-R, 1924:228 ISS
             Syn.: Proisotoma schötti(sic) Handschin, 1929 ISS
             Syn.: Proisotoma schötti(sic) Buitendijk, AM, 1930:58 ISS
             Syn.: Proisotoma Schötti(sic) Denis, JR, 1931:117 ISS
             Syn.: Proisotoma schötti(sic) Kseneman, M, 1932:15 ISS
             Syn.: Proisotoma (Ballistrura) schötti(sic) Folsom, 1937 ISS
             Syn.: Proisotoma schötti(sic) Kloet, GS et Hincks, WD, 1945:6 ISS
             Syn.: Ballistrura(sic) schötti(sic) Salmon, JT, 1945:69 ISS
             Syn.: Ballisrura(sic) schötti(sic) Christiansen, K, 1958:70 ISS
             Syn.: Ballistura schötti(sic) Yosii, R, 1959:5 ISS
             Syn.: Proisotoma schötti(sic) Paclt, J, 1961:230 ISS
             Syn.: Ballistura schoettii(sic) ssp. schoettii(sic) Greenslade P in Houston, 1994 ISS
             Syn.: Proisotoma schötti(sic) Greenslade, P, 1997-:www0312 ISS
Ballistura schoetti from Sweden
2006 © Hall, K.
Ballistura schoetti from Sweden
Abd.5 and 6 separate
2006 © Hall, K.
Ballistura schoetti from France
2008.06.25 © Lebeaux, P.
Ballistura schoetti from Holland
2011.07.28 © van Duinen, J.
Ballistura schoetti from South Africa
2017 © Janion-Scheepers, C.
?Ballistura schoetti? from Holland
2017.02.28 © van Leeuwen, J.
Ballistura schoetti from France
2017.09.29 © Tiky, S.
Ballistura schoetti from Holland
Dense cover of short setae
2017.09.30 © van Duinen, J.
Ballistura schoetti from Holland
Abd.5 and 6 separate
2017.09.30 © van Duinen, J.
Ballistura schoetti from Holland
Eyepatch with 8 ocelli; ocellus E distinctly largest
2017.09.30 © van Duinen, J.
Ballistura schoetti from France
2017.11.25 © Picard, J.
Ballistura schoetti from France
2018.04.01 © Picard, J.
Ballistura schoetti from France
2018.04.01 © Picard, J.
Ballistura schoetti juv. from France
2018.04.01 © Picard, J.
Ballistura schoetti juv. from France
2018.04.01 © Picard, J.
Ballistura schoetti? from the USA
2018.11.19 © Bairstow, A.

  Subspecies lutea Womersley, 1934





Species tuberculata (Stach, J, 1947:218) Skidmore, 1995:55

Proisotoma tuberculata from Poland
After Stach, J., 1947 Plate XXIII Fig.1.

Species veletensis (Steiner, 1959) Potapov, M, Babenko, A & Fjellberg, A, 2006:13

Species vernoga Wray, 1958

Species vetusta Folsom, JW, 1937:35

Species wrangeliensis Martynova, 1973


Genus Strenzketoma Potapov, M, Babenko, A et Fjellberg, A, 2006:63

Species buddenbrocki (Strenzke, 1954) Potapov, M, Babenko, A & Fjellberg, A, 2006:63, t.t.


Genus Dimorphotoma Grinbergs, A, 1975:101

Species huadongensis (Chen, 1985) Potapov, 2001

Species porcella Ellis, WN, 1976:295

Dimorphotoma porcella from Crete
After Schulz H-J, 2010 Fig.1


Genus Narynia Martynova, 1968

Species andreevae Martynova, 1975

Species kolymiensis Martynova, 1981

Species luanae Huang, Potapov & Gao, 2010

Species setosa Martynova, 1968


Genus Clavisotoma Ellis, 1970

Species africana (Womersley, H, 1934:457)

Species brisbanensis (Womersley, H, 1935:215)

Species fatonei (Rapoport, 1959)


Clavisotoma filifera from the UK
2005 © CEH
Species filifera (Denis, JR, 1931:112)


Species inaequalis (Schäffer, C, 1898:403)

Species lamelligera (Börner, C, 1909:108)

Species laticauda (Folsom, JW, 1937:42)

             Syn.: Ballistura laticaudata(sic) Vázquez & Palacios-Vargas, 1990 ISS
             Syn.: Ballistura laticuada(sic) Palacios-Vargas, JG et Arbea, JI, 2009:452 ISS

Species palustris (Cassagnau, P, 1959:82)

Species plicicauda (Handschin, 1927)

Species richardi (Denis, J-R, 1924:228)

            Syn.: Proisotoma Richardi(sic) Denis, J-R, 1924:228 IOS
            Syn.: Proisotoma Richardi(sic) Denis, J-R, 1924:229 IOS
            Syn.: Proisotoma Richardi(sic) Denis, J-R, 1924:230 IOS
Proisotoma Richardi(sic) from Monaco
After Denis J-R, 1924 Fig.39 left
Proisotoma Richardi(sic) from Monaco
After Denis J-R, 1924 Fig.39 right
             Syn.: Proisotoma Richardi(sic) Denis, JR, 1931:115 ISS




Species sjoestedti (Wahlgren, 1910)

             Syn.: Proisotoma Sjostedti(sic) Denis, JR, 1931:116 ISS

Species yamorensis (Uchida, 1949) Yoshii, R & Suhardjono, YR, 1992:76

Species yosiii (Stach, J, 1947:193)

            Syn.: Proisotoma yosii(sic) Stach, J, 1947:193 IOS
             Syn.: Proisotoma schötti(sic) Yosii, 1939 ISS
             Syn.: Proisotoma schötti(sic) Stach, J, 1947:193 ISS
             Syn.: Proisotoma schötti(sic) Denis, 1948 ISS
             Syn.: Ballistura schötti(sic) Yosii, R, 1959:6 ISS
            Syn.: Ballistura schoetti Yosii, R, 1977:154


             Syn.: Proisotoma yosii(sic) Yosii, R, 1959:6 ISS
             Syn.: Proisotoma (Clavisotoma) yosii(sic) Ellis, WN, 1970:22 ISS





Genus Granisotoma Stach, 1947

  Syn.: ... Panchaetoma Bagnall, 1949
  Syn.: ... Iostoma(sic) Fjellberg, A, 1988:965 ISS


Species communa (Mac Gillivray, 1896)

             Syn.: Pancheotoma(sic) comuna(sic) Cutz-Pool, LQ, Palacios-Vargas, JG et Vázquez, M, 2003:93 ISS
             Syn.: Isotoma comuna(sic) Vázquez, M et Palacios-Vargas, JG, 2004:86 ISS
             Syn.: Granisotoma comuna(sic) García-Gómez, A, Castaño-Meneses, G et Palacios-Vargas, JG, 2009:913 ISS
             Syn.: Isotoma comuna(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:110,ca ISS
             Syn.: Isotoma comuna(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:110 ISS

Species danilevskyi (Martynova, 1968) Greenslade, P & Potapov, M, 2015:340

Species rainieri (Folsom, JW, 1937:45) Stach, 1947

            Syn.: Isotoma kisoana Yosii, 1939


Species sadoana Yosii, 1965

Species sinensis Huang & Liang, 1991


References