http://www.collembola.org/taxa/pseuinae.htm - Last updated on 2019.10.04 by Frans Janssens
Checklist of the Collembola: Pseudachorutinae

Subfamilia Pseudachorutinae Börner, 1906

Genus Number
of species
Acanthanura5
Aethiopella23
Aethiopellina4
Anurachorutes1
Anurida79
Anuridella5
Anuritelsa1
Arlesia6
Arlesiella6
Brasilimeria4
Cassagnaudina4
Cassagnaurida1
Cephalachorutes16
Ceratrimeria11
Delamarellina3
Forsteramea1
Furculanurida18
Gamachorutes1
Gastranurida1
Grananurida5
Halachorutes2
Handschinurida3
Holacanthella5
Hylaeanura4
Intermediurida1
Israelimeria1
Kenyura7
Koreanurina4
Lanzhotia1
Linnaniemia2
Megalanura1
Meganurida1
Micranurida28
Minotaurella3
Najtafrica1
Neotropiella19
Notachorudina2
Oudemansia7
Paranurida1
Philotella6
Platanurida3
Pongeia1
Pratanurida8
Protachorutes2
Pseudachorudina14
Pseudachorutella11
Pseudachorutes119
PseudachorutinaeGennov1
Pseudanurida6
Quatacanthella1
Rusekella7
Sernatropiella2
Simonachorutes3
Stachorutes18
Tasmanura1
Tijucameria3
Venezuelida1
Womersleymeria1
Pseudachorutinae496
Genera with number of species
  Syn.: Collembola Neanurini(sic) Becker, E, 1905:72
  Syn.: Neanuridae Pseudachorutine(sic) Skarzynski, D et Pomorski, RJ, 1994:364 ISS
Pseudachorutinae from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutinae from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutinae from Australia
2017.03.04 © Steve & Alison.
Pseudachorutinae from the USA
2017.10.16 © Johnson, S.T.




Genus Cephalachorutes Bedos, A et Deharveng, L, 1991:145

Species sp.1 Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178

            Syn.: Cephalachorutes minimus Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178 nr

Species anneae Queiroz, GC et Mendonça, MC, 2016:47

Cephalachorutes anneae from Brazil
After Queiroz GC & de Mendonça MC, 2016 Fig.8A

Species asiaticus Bedos, A et Deharveng, L, 1991:152, t.t.

Species barthae Bedos, A et Deharveng, L, 1991:149

Species caecus Bedos, A et Deharveng, L, 1991:149

Species centurionis Bedos, A et Deharveng, L, 1991:149

Species deharvengi Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41

Species delamarei (Murphy, 1965) Bedos, A & Deharveng, L, 1991:147

Species judithae Shveenkova, YB, 2017:82

            Syn.: Cephalachorutes sp.n. Shveenkova, 2011
Cephalachorutes judithae from Vietnam
After Shveenkova YB, 2017 Fig.4A



Species masi Thibaud, J-M, 2008:507

Remark by Janssens, F. 2009.02.06: Given the name is derived from the surname "Masy", strictly spoken the correct latinisation "of Masy" is "masyi".

Species microphthalmus (Barra, 1969) Bedos, A & Deharveng, L, 1991:147

Species minimus (Massoud, 1963) Bedos, A & Deharveng, L, 1991:147

Species murphyi Bedos, A et Deharveng, L, 1991:151

Species nakaoi (Yosii, 1966) Bedos, A & Deharveng, L, 1991:152

Species pestilentiae Bedos, A et Deharveng, L, 1991:151

Species pillaii (Prabhoo, 1971) Bedos, A & Deharveng, L, 1991:147


Genus Pseudachorutes Tullberg, 1871 key

  Syn.: Poduridæ(sic) Brachysius MacGillivray, AD, 1893:317
  Syn.: Poduridæ(sic) Gnathocephalus MacGillivray, AD, 1893:318
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Sphragiphora Houlbert, 1924
  Syn.: ... Sphagiphorura(sic) Pomorski, RJ et Skarzynski, D, 1992:57 ISS
  Syn.: ... Logacanura Wray, 1958
  Syn.: ... Metachorudina Massoud, 1963
  Syn.: ... Neoafricella Salmon, 1964


Pseudachorutes sp. from Germany
Mandibula
After Anonymous Fig.B

  Syn.: Collembola Pseudachotures(sic) Verlinden, 1977 ISS
Pseudachorutes sp. from the USA
2010.01.17 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Ocular area
2010.01.18 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Furcula
2010.02.12 © Ten Eyck, C.
Pseudachorutes sp. from the USA
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Pointed buccal cone
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Furcula
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from Portugal
2012.10.10 © Murray, A.
Pseudachorutes sp. from the USA
2014.05.22 © Murray, T.
Pseudachorutes sp. from Holland
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Holland
Pointed mouthcone
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Germany
Large round post-antennal organ (PAO)
2015.02.20 © Mattew, M.
Pseudachorutes sp. from the UK
2015.02.26 © Murray, A.
Pseudachorutes sp. from the UK
Pointed mouth-cone
2015.02.26 © Murray, A.
Pseudachorutes sp. from France
2016.03.03 © Garcelon, P.
Pseudachorutes sp. from Australia
2016.04.25 © Murray, A.

  Syn.: Neanuridae Pseudahorutes(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:241 ISS
Pseudachorutes sp. from France
2018.01.06 © Picard, J.
Pseudachorutes sp. from France
Unguiculus reduced (*)
2018.01.06 © Picard, J.
Pseudachorutes sp. from France
Anal spines absent (*)
2018.01.06 © Picard, J.
Pseudachorutes sp. from the USA
2018.03.20 © Montgomery, G.
Pseudachorutes sp. from Tenerife
2018.03.30 © Huskens, M.-L.
Pseudachorutes sp. from Tenerife
Large ring-shaped PAO with many vesicles
2018.03.30 © Huskens, M.-L.
Pseudachorutes sp. from the USA
2018.12.09 © Owl, Bo.
Pseudachorutes sp. from the USA
Pointed mouthcone (*)
2018.12.09 © Owl, Bo.
Pseudachorutes sp. from the USA
Post-antennal organ roundish (PAO)
2018.12.09 © Owl, Bo.
Pseudachorutes sp. from the USA
2018.12.14 © Owl, Bo.
Pseudachorutes sp. from the UK
Large cauliform post-antennal organ (PAO)
2019.02.25 © Smart, J.





Species sp.1 Denis, J-R, 1924:223

Species sp.2 Denis, J-R, 1924:223

Species sp.3 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8

Species sp.4 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8

Species sp.5 Christiansen, K et Bellinger, P, 1980:297bis

Species sp.6 Christiansen, K et Bellinger, P, 1980:297bis

Species sp.7 Christiansen, K et Bellinger, P, 1980:297bis

Species sp.8 Soto-Adames, FN et Taylor, SJ, 2013:153

Species sp.9 Queiroz, GC et Zeppelini, D, 2017:132

Species algidensis Carpenter, GH, 1925:4

             Syn.: Pseudachorutes aldgidenssi(sic) Womersley, H, 1936:327 ISS

Species alluaudi Delamare Deboutteville, 1945

             Syn.: Pseudachorutes allauaudi(sic) Massoud, 1967 ISS

Species americanus Stach, 1949

            Syn.: Pseudachorutes boerneri Nearctic records
            Syn.: Pseudachorutes corticolus Mills, 1934
            Syn.: Pseudachorutes corticicolus f. globulisetosa Gisin, 1944, Nearctic records

Species amucronatus Díaz & Najt, 1995

Species andrei Weiner, WM et Najt, J, 1985:260

Species anomalus Imms, 1912

Species aphysus Christiansen & Bellinger, 1980:304

Species aureofasciatus (Mac Gillivray, 1893) Mills, HB, 1934:23

            Syn.: Pseudachorutes aureofasciatus f. millsi Maynard, EA, 1951:58
Pseudachorutes aureofasciatus from the USA
Non-banded juvenile
2007.02.22 © Bernard, E.C.

            Syn.: Pseudachorutes (Pseudachorutes) bifidus Christiansen & Bellinger, 1980



Pseudachorutes aureofasciatus
1980 © Norton, R.A. in Dindal.
Pseudachorutes aureofasciatus from the USA
Banded juvenile
2007.02.22 © Bernard, E.C.
Pseudachorutes aureofasciatus from the USA
"Butts up colony"
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
Badly smelling defensive secretion
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2012.05.01 © Murray, T.

Species auricularius Cassagnau & Rapoport, 1962

Species beta Najt, J et Weiner, WM, 1991:106

Species bifasciatus Oliveira & Deharveng, 1994

Species bifidus Christiansen & Bellinger, 1980:307

Species bobeitio Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32

Species cacaoiensis Najt, Thibaud & Weiner, 1990

Species caerulescens Schött, H, 1893:24

            Syn.: Pseudachorutes cærulescens(sic) Schött, H, 1893:24 IOS
             Syn.: Pseudachorutes coerulescens(sic) Wahlgren, E, 1906:4 ISS



Species caramel Kaprus, IJ, Weiner, WM et Nevo, E, [2005]:263

Pseudachorutes caramel from Israel
After Kaprus, I.J., Weiner, W.M. & Nevo, E., [2005] Fig.1

Species chazeaui Najt, J et Weiner, WM, 1991:100

Species cheni Shi, S, Jiang, J et Pan, Z-X, 2008:57

Pseudachorutes cheni from China
After Shi S et al., 2008 Fig.1-2

Species columbicus Rusek, J, 1991:120

Species complexus (MacGillivray, AD, 1893:318) Folsom, 1916

Species conicus Lee & Kim, 1984

Species conspicuatus Salmon, JT, 1944:137

            Syn.: Pseudachorutes conspicuatus ssp. principalis Massoud, Z, 1967:156


Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.
Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.

  Subspecies flavus Salmon, JT, 1944:139

  Subspecies lineatus Salmon, JT, 1944:138

Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2009 © Te Papa Museum
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.

  Subspecies maximus Salmon, JT, 1944:138

Species corticicolus (Schäffer, 1897) Krausbauer, 1898

             Syn.: Pseudachorutes börneri(sic) Schött, 1902 ISS
            Syn.: Pseudachorutes corticicolus f. globulisetosa Gisin, 1944 partim


             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1898 ISS
             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1901 ISS
             Syn.: Pseudachorutes corticicola(sic) Handschin, E in Dahl, F, 1929:34 ISS
             Syn.: Pseudachorutes corticola(sic) Kloet, GS et Hincks, WD, 1945:4 ISS
             Syn.: Pseudachorutes (Pseudachorutes) corticicola(sic) Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:83 ISS
             Syn.: Pseudachorutes corticolus(sic) Snider, RJ, 1967:192 ISS
             Syn.: Pseudachorutes corticicola(sic) Loksa, I, 1978:55 ISS
             Syn.: Pseudachorutes corticolus(sic) Palacios-Vargas, JG, 1990:19 ISS
             Syn.: Pseudachorutes corticolus(sic) Palacios-Vargas, JG, 1990:19 nr ISS
             Syn.: Pseudachorutes corticolus(sic) Szeptycki, A et Weiner, WM in Razowski J, 1990:20 ISS
             Syn.: Pseudachorutes corticolus(sic) Barra, J-A, 1993:71 ISS
             Syn.: Pseudachorutes corticicola(sic) Farkas, J, 1995:46 ISS
             Syn.: Pseudachorutes corticolus(sic) Kopeszki, H et Meyer, E, 1996:224 ISS
             Syn.: Pseudachorutes corticicola(sic) Arbea, JI et Jordana, R in Ramos, MA et al, 1997:323 ISS
Pseudachorutes corticicolus from France
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Pointed mouthcone
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Ant.3+4 fused
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Abd.4 chaetotaxy
2018.06.16 © Tiky, S.



Species crassus da Gama, 1964

Species curtus Christiansen & Bellinger, 1980:232

Species dalensi Weiner, WM et Najt, J, 1985:258

Species decussus (Salmon, JT, 1941:298)

Micranurida decussa from New Zealand
After Salmon JT, 1941 Pl.43 Fig.72

Species deficiens Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:558

Pseudachorutes deficiens from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.1A,E

Species deserticus Palacios-Vargas, 1990:11

Species difficilis Denis, 1931

Species dilatatus (MacGillivray, AD, 1893:317) Christiansen & Bellinger, 1980

Species dubius Krausbauer, 1898

            Syn.: Pseudachorutes lapponicus Ågren, H, 1904:8
            Syn.: Schöttella(sic) poppei Schäffer, 1897


             Syn.: Pseudahorutes(sic) dibius(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:241 ISS
             Syn.: Pseudachorutes dibius(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:242 ISS

Species galapagoensis Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:157

Species gilvus Oliveira & Deharveng, 1995

Species geronensis (Massoud, 1963) Massoud, 1967:158

Species gilvus Oliveira & Deharveng, 1994

Species guyanensis Najt, Thibaud & Weiner, 1990

Species herberti Arlé & Rufino, 1976

Species hitakamiensis Tamura, 2001

Species hodeberti Thibaud, J-M, 2008:508

Species indiana Christiansen, K et Bellinger, P, 1980:312

Species innominatus Denis, 1931

Species insularis Yosii, 1965

Species isawaensis Tamura, 2001

Species janstachi Kaprus, IJ et Weiner, WM, 2009:9

Pseudachorutes janstachi from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.32

Species japonicus Kinoshita, 1916

Species javanicus (Handschin, 1920, i.s.) Handschin, 1920

Species jeonjuensis Lee & Kim, 1984

Species jianxiucheni Gao, Yin & Palacios-Vargas, 2008

Species kangchenjungae Yosii, 1966

Species labiatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:565

Pseudachorutes labiatus from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.5A

Species laricis Arbea, JI et Jordana, R, 1989:161

Species legrisi Thibaud & Massoud, 1983

Species libanensis (Cassagnau, P et Delamare Deboutteville, C, 1955:372)

Species libanensis Ellis, WN, 1976:269

Remark by Kaprus, I.J., Weiner, W.M. & Nevo, E. [2005]:263: P. libanensis was described from Lebanon as Aetiopella(sic) libanensis by CASSAGNAU and DELAMARE (1955) and subsequently redescribed by ELLIS (1976) on the basis of the material from Central Crete. The comparison of both descriptions shows, that they concern two separate species.

Species lishanensis Gao, Yin & Palacios-Vargas, 2008

Species longisetis Yosii, 1961

             Syn.: Pseudachorutes longisetus(sic) Cassagnau, P, 1974:309 ISS

Species lunatus Folsom, 1916

            Syn.: Pseudachorutes (Pseudachorudina) ignotus Christiansen, K et Bellinger, P, 1980:301 =?


Species macrochaetus Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:158

Species massoudi Arlé, R, 1966:7

Species medjugorjensis Barra, 1993:69

Species minor Peja, 1985

Species multidentatus Najt, Thibaud & Weiner, 1990

Species murphyi Massoud, 1965

             Syn.: Pseudachorutes murphy(sic) Peck, SB et Jacquemart, J, 2011:5,cf ISS

Species namphoensis Weiner, WM et Najt, J, 1985:262

Species nica Palacios-Vargas, JG et Mejía-Madrid, HH, 2012:64

Species niloticus Wahlgren, E, 1906:3

Species octosensillatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:561

Species orghidani Massoud, Z et Gruia, M, 1973:329

Species orientalis Loring, 1984

Species ouatilouensis Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32

Species palmiensis Börner, 1903

            Syn.: Pseudachorutes palmiensis f. secundo Denis, 1948
            Syn.: Pseudachorudina conjungens Stach, 1949
            Syn.: Pseudachorutes cantabricus Simón & Pozo, 1979


             Syn.: Pseudachorutes palmatus(sic) Wahlgren, E, 1906:4 lapsus ISS
Pseudachorutes palmiensis from France
2006 © Boldsystems.org
Pseudachorutes palmiensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.42
Pseudachorutes palmiensis from Spain
2016.04.24 © Tizon, J.



Species parvulus Börner, 1901, B

            Syn.: Pseudachorutes parvulus f. infuscata Yosii, 1954
            Syn.: Pseudachorutes corticicolus Selga, 1971
            Syn.: Pseudachorutes subcrassus Selga, 1971, partim Arbea & Jordana in Ramos, 1997


Pseudachorutes parvulus from Spain
Modified after Arbea JI & Jordana R, 1997 Fig.144A
Pseudachorutes parvulus from Spain
Axis (G-A) of eyepatch directed to the side
Modified after Arbea JI & Jordana R, 1997 Fig.144A
Pseudachorutes parvulus from Spain
In state of anhydrobiosis
Arbea JI & Blasco-Zumeta J, 2001 Fig.2
Pseudachorutes parvulus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.52
Pseudachorutes parvulus from Belgium
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Post-antennal organ (PAO) with 6 vesicles
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus ♂ from Belgium
Genital plate with star-shaped orifice
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Axis (G-A) of eyepatch directed to the side
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Post-antennal organ (PAO) with 6 vesicles
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Pointed mouthcone (*)
2019.09.14 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Furcula with short dentes and mucrones
2019.09.14 © Huskens, M.-L.

Species pasurus (Wray, 1958) Christiansen & Bellinger, 1980

Species pauliani Delamare Deboutteville, 1952

Species periyarensis Prabhoo, 1971

Species plurichaetosus Arbea, JI et Jordana, R, 1991:78

Species poahi Christiansen, K et Bellinger, P, 1992:65

Species polychaetosus Gao, Y et Palacios-Vargas, J, 2008:54

Species ponmudiensis (Prabhoo, 1971, i.s.) Prabhoo, 1971

Species pratensis Rusek, 1973

Pseudachorutes pratensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.65

Species prosimplex Weiner, WM et Najt, J, 1985:256

Species pseudoniloticus Massoud, 1963

Species puniceus Salmon, JT, 1944:139

Species reductus Thibaud & Massoud, 1983

Species rhaeticus (Carl, J, 1899:298) Handschin, E, 1924:96

            Syn.: Schöttella(sic) rhätica(sic) Carl, J, 1899:298 IOS
            Syn.: Schöttella(sic) rhætica(sic) Carl, J, 1899:298 IOS

Species rugatus Wray, 1952

Species saxatilis Macnamara, 1920

Species scythicus Kaprus, IJ et Weiner, WM, 2009:3

Pseudachorutes scythicus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.1

Species sexsetosus Denis, 1934

Species shiragamiensis Hisamatsu & Tamura, 1998

Species sibiricus Reuter, 1891

Species sibiricus (Rusek J, 1991:123) Rusek, J, 1991:123

Species simplex Maynard, EA, 1951:59

Pseudachorutes simplex from the USA
2006 © Bernard, E.C.
Pseudachorutes simplex from the USA
2007.02.22 © Bernard, E.C.

Species solaris da Silveira, TC et de Mendonça, MC, 2018:2

Pseudachorutes solaris from Brazil
After da Silveira TC & de Mendonça MC, 2018 Fig.1i

Species subabdominalis Steiner, 1958

Species subcrassoides Mills, HB, 1934:24

             Syn.: Pseudachorutes subcrassoïdes Massoud, 1967 ISS

Species subcrassus Tullberg, 1871, t.t., B

            Syn.: Pseudachorutes tuberculatus Schött, 1902


Pseudachorutes subcrassus from Russia
After Scherbakov AM, 1898 Table I Fig.4
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from Holland
2015.02.21 © Baas, A.H.

Species subcrassus Schäffer, 1897, nec Tullberg, 1871

Species tamajonensis Simón, 1986

Species tasmaniensis Womersley, H, 1936:475

Species tavignani Poinsot-Balaguer, 1978

Species tillieri Najt, J et Weiner, WM, 1991:104

Species unidentatus Salmon, 1956, i.s.

Species uniformis Thibaud & Massoud, 1983

Species univesicatus Weiner & Najt, 1991

             Syn.: Pseudachorutes universicatus(sic) Greenslade, P, 1997-:www0312 ISS

Species vasylii Kaprus, IJ et Weiner, WM, 2009:7

Pseudachorutes vasylii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.22

Species vitalii Kaprus, IJ et Weiner, WM, 2009:5

Pseudachorutes vitalii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.11

Species wandae Gao, Yin & Palacios-Vargas, 2008

Species yoshiii Najt, J et Weiner, WM, 1991:102

            Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM, 1991:102 IOS
             Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36 ISS
             Syn.: Pseudachorutes yoshii(sic) Gao, Y et Palacios-Vargas, J, 2008:55 ISS
             Syn.: Pseudachorutes yoshii(sic) da Silveira, TC et de Mendonça, MC, 2018:4 ISS




Genus Simonachorutes Skarzynski, D, Arbea, JI et Piwnik, A, 2016:172

Species agrensis (Arbea, JI & Jordana, R, 1989:164) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:175

Simonachorutes agrensis from Spain
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.18

Species romeroi (Simón, 1986) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:172, t.t.

Simonachorutes romeroi from Spain
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.1-2

Species weinerae Skarzynski, D, Arbea, JI et Piwnik, A, 2016:178


Genus Pratanurida Rusek, 1973

  Syn.: ... Pratanuria(sic) Rusek, 1973 IOS
Pratanurida sp. from Spain
After Arbea JI, 2013 Fig.8



Species boerneri (Schött, 1902) Jordana, R & Arbea, JI in Ramos, MA & al, 1997:345, B

            Syn.: Pseudachorutes corticicolus Mills, 1934
            Syn.: Pseudachorutes forsslundi Agrell, 1943
            Syn.: Pseudachorutes corticicola f. globulisetosa Gisin, 1944
            Syn.: Pseudachorutes boerneri ssp. americanus Stach, 1949


             Syn.: Pseudachorutes börneri(sic) Wahlgren, E, 1906:4 ISS
Pratanurida boerneri from Mexico
1997 © Palacios-Vargas, J.G.



Species cassagnaui (Rusek, 1973) Kaprus, IJ & Weiner, WM, 2002:283, t.t.

Species foxi Fjellberg, 1985

Species harti Christiansen & Bellinger, 1980:340

Species minuta (Selga, 1966) Arbea, JI & Jordana, R, 1990:189

  Subspecies menorquina Arbea, JI et Jordana, R, 1990:190

Species mucronata (Ellis, WN, 1976:272) Kaprus, IJ & Weiner, WM, 2002:283

Species podolica Kaprus, IJ et Weiner, WM, 2002:283

Pratanurida podolica from Ukraine
After Kaprus, I.J. & Weiner, W.M., 2002 Fig.1

Species tananensis Fjellberg, 1985


Genus Pseudachorutella Stach, J, 1949:79

  Syn.: ... Pseudachorutes Tullberg, 1871 partim
  Syn.: Pseudachorutidae Montachorutes Stach, J, 1949:55
  Syn.: ... Notachoridina Cassagnau & Rapoport, 1962 partim


Pseudachorutella gigas from Chile
Ant.3+4 dorsally fused, ventrally separate (*)
2016.09.02 © Invertebrados, C.

Species africana Weiner & Najt, 1991

Species asigillata (Börner, C, 1901:36) Stach, J, 1949:55, t.t.

            Syn.: Pseudachorutes clavatus Börner, C, 1901:36
            Syn.: Pseudachorutes silvaticus Ågren, 1903


Species balcanica Cassagnau, P et Peja, N, 1979:205

Species bescidica Smolis & Skarzynski, 2007

Species boudinoti Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:39

Species castrii (Rapoport, EH & Rubio, IR, 1963:111) Massoud, Z, 1967:170

Notachorudina castrii from Chile
After Rapoport EH & Rubio IR, 1963 Fig.49

Species clavata (Börner, C, 1901:36)

Remark by Dányi, L. & Traser, G. 2008:18: According to da Gama (1964) the species is a junior synonym of Pseudachorutella asigillata (Börner), but it is a valid, separate taxon in our opinion.

Species gigas (Rapoport, EH & Rubio, IR, 1963:112) Massoud, Z, 1967:169

Pseudachorutella gigas from Chile
2016.09.02 © Invertebrados, C.
Pseudachorutella gigas from Chile
Unguis with lateral apical teeth (*)
2016.09.02 © Invertebrados, C.

Species maxima Najt, Thibaud & Weiner, 1990

Species remyi (Denis, 1933) Massoud, Z, 1967:170, t.t.

            Syn.: Pseudachorutes Remyi(sic) Denis, 1933 IOS
             Syn.: Pseudachorutes Remyi(sic) Stach, J, 1949:56,tt ISS
  Subspecies alpina Stach, J, 1949:65





Species stachi Massoud, 1965


Genus Pseudanurida Schött, 1901 key

  Syn.: ... Neachorutes Womersley, 1933
  Syn.: Neanurini(sic) Pseudachorutides Becker, E, 1905:72
  Syn.: Neanurini(sic) Pseudochorutides(sic) Becker, E, 1905:72 ISS
  Syn.: Neanurini(sic) Pseudochorutides(sic) Becker, E, 1906:72 ISS
  Syn.: ... Pseudachorutoides(sic) Denis, 1938 ISS
  Syn.: Pseudachorutini Pseudachorutoides(sic) Womersley, H, 1939:315 ISS
  Syn.: Collembola Pseudachorutoides(sic) Ellis, WN et Bellinger, PF, 1973:45 ISS


Pseudanurida sp. from Singapore
2011.01.04 © Koh, J.

Species billitonensis Schött, 1901, t.t.

Pseudanurida billitonensis from Japan
After Uchida H, 1962 Fig.A
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.

Species bogoyawlenskyi (Becker, E, 1905:72) Salmon, 1951

            Syn.: Pseudachorutides Bogoyawlensky(sic) Becker, E, 1905:72 IOS
            Syn.: Pseudachorutides clysmae Jackson, 1927
            Syn.: Pseudachorutoides(sic) dollfusi Denis, 1938


             Syn.: Pseudochorutides(sic) Bogoyawlensky(sic) Becker, E, 1905:72 ISS
             Syn.: Pseudachorutides Bogoyawlensky(sic) Becker, E, 1906:72 ISS
             Syn.: Pseudochorutides(sic) Bogoyawlensky(sic) Becker, E, 1906:72 ISS
             Syn.: Pseudachorutoides(sic) bogoyawlensky(sic) Womersley, H, 1939:315 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Stach, J, 1949:54 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Gisin, H, 1960:69 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Salmon, JT, 1964:277 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Massoud, Z, 1967:172 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Massoud, Z, 1967:174 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Massoud, Z, 1967:175 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Massoud, Z, 1967:175 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:49 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:50 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:53 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:55 ISS
             Syn.: Pseudachorutes bogoyawlensky(sic) Murphy, DH, 1971:55 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:56 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:57 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Queiroz, GC et Zeppelini, D, 2017:150 ISS
Pseudanurida bogoyawlenskyi from Pakistan
After Aslam S, Arbea JI & Siddiqui G, 2018 Fig.3
Pseudanurida bogoyawlenskyi from Pakistan
With orange eggs
After Aslam S, Arbea JI & Siddiqui G, 2018 Fig.10





Species glauerti (Womersley, 1933) Murphy, DH, 1971:56

            Syn.: Pseudanurida australica Salmon, 1955
            Syn.: Pseudanurida billitonensis Womersley, 1939


Species longisetosa Lee, B-H et Kim, J-T, 1994:208

Species sawayana Schuster, 1965

             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, 1965 cf ISS


Species yini Murphy, DH, 1971:62


Genus Halachorutes Arlé, R, 1967:368

  Syn.: ... Holachorutes(sic) Jacquemart, S et Jacques, J-M, 1980:14 ISS

Species quadrisetosus (Snider, RJ, 1981:244) Queiroz, GC & Zeppelini, D, 2017:151

Species schusteri Arlé, R, 1967:370, t.t.

Halachorutes schusteri from Brazil
After Queiroz GC & Zeppelini D, 2017 Fig.17C
Halachorutes schusteri from Brazil
After Queiroz GC & Zeppelini D, 2017 Fig.17B


Genus Oudemansia Schött, 1893

Species barnardi (Womersley, H, 1934:447) Uchida, H, 1965:87

Species coerulea Schött, 1893, t.t.

            Syn.: Oudemansia subcoerulea Denis, 1948
             Syn.: Oudemansia subcoerula Prabhoo, 1970 ISS
            Syn.: Polyacanthella oceanica Uchida, 1955 si
            Syn.: Oudemansia hirashimai Uchida, H, 1965:86
            Syn.: Oudemansia hirashimai Yosii, R, 1977:146


             Syn.: Oudemansia cærulea(sic) MacGillivray, AD, 1893:318 ISS
             Syn.: Friesea coerula Salmon, 1964 ISS

Species dubia Denis, 1947

Species esakii (Kinoshita, 1932) Yosii, 1958

             Syn.: Oudemansia esaki(sic) Park, K-H et Lee, B-H, 2000:228 ISS
             Syn.: Oudemansia esaki(sic) Park, K-H, Kim, K-G, Park, S-J, Kim, B-J et Lee, B-H, 2000:415 ISS

Species georgia Christiansen & Bellinger, 1980:295

Species petiti Delamare Deboutteville, C et Massoud, Z, 1964:387

Species shotti Denis, 1948

             Syn.: Friesea shötti(sic) Salmon, 1964 ISS
             Syn.: Oudemansia shötti(sic) Massoud, 1967 ISS



Genus Tasmanura Womersley, H, 1937:154

Species evansi Womersley, H, 1937:154, t.t.

Tasmanura evansi from Tasmania
After Womersley H, 1937 Fig.1a
Tasmanura evansi from Tasmania
After Womersley H, 1939 Fig.44M


Genus Ceratrimeria Börner, C, 1906:167

  Syn.: ... Platanurida Carpenter, 1925
  Syn.: ... Tasmanura Womersley, 1937
  Syn.: Pseudachorutidae Zealandmeria Stach, J, 1949:59


  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1924:220 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1924:222 ISS
  Syn.: ... Ceratimeria(sic) Denis, 1931 ISS
  Syn.: Pseudachorutinae Caratrimeria(sic) Yosii, R, 1977:147 ISS
Ceratrimeria sp. from New Zealand
2006 © Minor, M. & Robertson, A.
Ceratrimeria sp. from Tasmania
2012.10.28 © Zosterops, K.
Ceratrimeria sp. from Australia
2015.07.27 © Porch, N.



Species aurea Salmon, JT, 1944:133

Species crassa (Oudemans, JT in Weber, M, 1890:74) Yayuk, 1989:5

Species harrisi Salmon, JT, 1942:258

Ceratrimeria harrisi from New Zealand
After Salmon J.T., 1942 Pl.43 Fig.4
Ceratrimeria harrisi from New Zealand
2015 © TePapa Museum

Species indica (Handschin, E, 1929:231) Massoud, Z, 1967:180

Species longicornis Handschin, 1926

Species lydiae Najt, J et Weiner, WM, 1991:108

Species maxima (Schött, 1901) Börner, C, 1906:167, t.t.

Ceratrimeria maxima from Australia
After Schött H, 1917 Fig.4
             Syn.: Ceratimeria(sic) maxima Denis, J-R, 1924:222 ISS


Species novaezealandiae (Womersley, H, 1936:318) Womersley, 1937

            Syn.: Pseudachorutes novae-zealandiae(sic) Womersley, H, 1936:318 IOS
             Syn.: Ceratrimeria novae-zealandiae(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1941:302 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1942:254 ISS
             Syn.: Zealandmeria novae-zealandiae(sic) Stach, J, 1949:59,si ISS
             Syn.: Aoteareria novae-zealandiae(sic) Salmon, 1964 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Massoud, Z, 1967:182 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Zealandmeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Pseudachorutes novazealandiae(sic) Greenslade, P, 2015:6 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, P, 2015:6 ISS
Ceratrimeria novaezealandiae from New Zealand
2017.12.01 © Maurice
Ceratrimeria novaezealandiae from New Zealand
Eyepatch with 8 ocelli
2017.12.01 © Maurice





Species pulchella Handschin, 1926

Species takaoensis (Kinoshita, 1916) Yosii, R, 1977:147

Species yasumatsui (Uchida, 1950) Yosii, R, 1977:147


Genus PseudachorutinaeGennov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105

Species spnov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105

Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.
Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.


Genus Aethiopellina Delamare Deboutteville, C, 1951:284

Species bimaculata Delamare Deboutteville, C, 1951:284

Species hebrardi Poinsot-Balaguer, 1978

Species proboscina Delamare Deboutteville, C, 1951:284, t.t.

Species sphaeroculata Massoud, 1963


Genus Aethiopella Handschin, 1942 key

  Syn.: Arthropleona Certrimeria(sic) Womersley, H, 1934:443 ISS
  Syn.: Protachorutidae Ruwenzoriella Salmon, JT, 1964:269


  Syn.: Neanuridae Aethiopiella(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS

Species africana (Delamare Deboutteville, 1945) Massoud, Z, 1967:188

  Subspecies nigra Delamare Deboutteville, 1945

Species ariana Najt, Thibaud & Weiner, 1990

Species basilewskyi (Salmon, 1956) Massoud, Z, 1967:188

Species caraibensis Thibaud & Massoud, 1983

            Syn.: Aethiopella caraïbensis(sic) Thibaud & Massoud, 1983 IOS
             Syn.: Aethiopella caraïbensis(sic) Mari Mutt & Bellinger, 1990 ISS



Species condei Massoud, 1963

Species delamarei Arlé, R, 1960:164

Species flavoantennata (Philiptschenko, 1926 tt) Handschin, 1942

            Syn.: Certrimeria(sic) flavoantennatus(sic) var. capensis Womersley, H, 1934:448


             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, H, 1934:448 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1939 ISS

Species guineensis Handschin, 1942

Species handschini (Denis, J-R, 1924:221) Handschin, 1942

            Syn.: Pseudachorutes Handschini(sic) Denis, J-R, 1924:221 IOS
Pseudachorutes Handschini(sic) from Ethiopia
After Denis J-R, 1924 Fig.31



Species jeanneli Massoud, Z, 1963:60

Species kuolo Christiansen, K et Bellinger, P, 1992:63

Species littoralis Fernandes, LH et de Mendonça, MC, 2003:1

Species machadoi Massoud, 1963

Species maculata (Delamare Deboutteville, 1945) Massoud, Z, 1967:188

Species mandibulata (Delamare Deboutteville, 1945) Salmon, 1956

Species mangenoti Massoud, 1963

Species pedifalx (Salmon, 1956) Massoud, Z, 1967:190

Species pilarandresae Palacios-Vargas, JG et Montejo-Cruz, M, 2014:1039

Species ricardoi da Paz, RV, Queiroz, GC et Bellini, BC, 2019:41

Aethiopella ricardoi from Brazil
After da Paz et al, 2019 Fig.1

Species silvestris Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36

Species tournieri Delamare Deboutteville, 1950

            Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, 1950 IOS


             Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, C, 1951:282 ISS



Species ugandensis Salmon, 1954

Species villiersi Massoud, 1963


Genus Protachorutes Cassagnau, P, 1955:154

Species bicolor (Christiansen & Bellinger, 1980:343) Christiansen, KA in Dindal, DL, 1990:979

Species pyrenaeus Cassagnau, P, 1955:154, t.t.


Genus Arlesiella Delamare Deboutteville, C, 1951:285

Species amazonica Arlé, R, 1966:8

Species camerounensis Massoud, 1963

Species delamarei Massoud, Z, 1963:64

Species monodi (Delamare Deboutteville, 1952) Massoud, 1967:198

Species saprophila Delamare Deboutteville, C, 1951:285, t.t.

Species simulans Massoud, Z, 1963:67


Genus Linnaniemia Philiptschenko, 1926

Species bicolor (Delamare Deboutteville, 1945) Massoud, Z, 1967:202

Species gigas Philiptschenko, 1926:202, t.t.


Genus Neotropiella Handschin, 1942

  Syn.: ... Americotrimeria Stach, 1949


Species sp.1 Bellini, BC et Zeppelini, D, 2004:588

Species arlei Najt, Thibaud & Weiner, 1990

Species barbatae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:43

Species carli (Denis, J-R, 1924:222) Arlé, 1960

            Syn.: Pseudachorutes Carli(sic) Denis, J-R, 1924:222 IOS

            Syn.: Ceratrimeria pentoculata Womersley, 1930
            Syn.: Neotropiella araguaensis Rapoport & Maño, 1969


             Syn.: Pseudachorutes Carli(sic) Denis, J-R, 1924:223 ISS
Neotropiella carli from French Guyana
Pointed mouthcone
After Thibaud J-M & D'Haese C, 2010 Fig.2
2009 © D'Haese, C.
Neotropiella carli from Ecuador
Eyepatch with 5 ocelli; PAO present
2014.01.31 © McClarin, J.
Neotropiella carli from Ecuador
Body segments trilobed
2014.01.31 © McClarin, J.





Species denisi (Arlé, 1939) Stach, 1949

Species digitomucronata Massoud & Thibaud, 1982

Species duranti Díaz & Najt, 1995

Species gordae Díaz & Najt, 1995

Species insularis Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:47

Species macunaimae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:45

Species meridionalis (Arlé, 1939) Stach, 1949

Neotropiella meridionalis from Brazil
After Cipola NG & al, 2019 Fig.14

Species minima Thibaud & Oliveira, 2010

Species mirabilis (Handschin, 1929) Massoud, 1967:206

Species murphyi Massoud, 1965

Species pedisensilla Najt, Thibaud & Weiner, 1990

Species plurichaetosa Thibaud & Oliveira, 2010

Species quinqueoculata (Denis, 1931) Arlé, 1962

            Syn.: Pseudachorutes 5-oculatus(sic) Denis, 1931 IOS


             Syn.: Americotrimeria 5-oculata(sic) Stach, 1949 ISS
             Syn.: Neotropiella quinque-oculata(sic) Arlé, R, 1966:13 ISS
             Syn.: Neotropiella 5-oculata(sic) Arlé, R, 1966:15 ISS
             Syn.: Neotropiella pentoculata(sic) Massoud, 1967 ISS



Species silvestrii (Denis, 1929) Massoud, 1967:208, t.t.

Species vanderdrifti Massoud, 1963


Genus Sernatropiella Palacios-Vargas, JG, 2019:374

Species malkini (Arlé, 1981) Palacios-Vargas, JG, 2019:379

Species pinzonae Palacios-Vargas, JG, 2019:374, t.t.

Sernatropiella pinzonae from Colombia
After Palacios-Vargas JG, 2019 Fig.1
Sernatropiella pinzonae from Colombia
After Palacios-Vargas JG, 2019 Fig.2
Sernatropiella pinzonae from Colombia
With cephalic pseudocelli (*)
After Palacios-Vargas JG, 2019 Fig.5


Genus Grananurida Yosii, 1954

  Syn.: Neanuridae Agranurida(sic) Kim, JT et Lee, B-H, 2000:180 ISS
Remark by Janssens F. & Bernard, E.C. 2007:in litt.: Agranurida(sic) Kim, J.T. & Lee.-H., 2000:180 is an incorrect subsequent spelling of Grananurida.


Species alba (Yosii, 1966) Weiner, WM & Najt, J, 1985:281, i.s.

Species baicalica Rusek, J, 1991:126

Species koreana Lee, B-H et Kim, J-T, 1984:21

Species minuta Kim, JT et Lee, B-H, 2000:180

Species tuberculata Yosii, 1954, t.t.


Genus Najtafrica Barra, J-A, 2002:178

Species riebi (Barra, J-A, 1994:184) Barra, J-A, 2002:178


Genus Stachorutes Dallai, 1973 key

  Syn.: Neanuridae Chihuahuachorutes Palacios-Vargas, JG, 1990:7


Species cabagnerensis Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150

Species cuihuaensis Gao & Yin, 2007

Species dallaii Weiner, WM et Najt, J, 1998:226

             Syn.: Stachorutes dallai(sic) Kovác, L, 1999:135 ISS
             Syn.: Stachorutes dallai(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS
             Syn.: Stachorutes dallai(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS

Species dematteisi Dallai, 1973, t.t.

             Syn.: Stachorutes dematteisae(sic) Palacios-Vargas, 1990 ISS
             Syn.: Stachorutes demattiesi(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Stachorutes dematteisae(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:108 ISS

Species escobarae (Palacios-Vargas, JG, 1990:8) Thibaud, J-M & Palacios-Vargas, JG, 2000:107

Species gracilis Smolis, A et Shvejonkova, JB, 2006:44

Species jizuensis Tamura & Zhao, 1997

Species longirostris Deharveng & Lienhard, 1983

Species maya Thibaud, J-M et Palacios-Vargas, JG, 2000:108

Species najtae Fanciulli, PP, Dallai, R, Frati, F et Carapelli, A, 2017:32

Stachorutes najtae from Italy
After Fanciulli PP & al, 2017 Fig.3A

Species navajellus Fjellberg, 1984

             Syn.: Stachorutes navajelus(sic) Slawska, M, 1996:329 ISS
             Syn.: Stachorutes novajellus(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS

Species ruseki Kovác, L, 1999:135

Species scherae Deharveng & Lienhard, 1983

Species sphagnophilus Slawska, M, 1996:325

             Syn.: Stachorutes sphagnophillus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:156 ISS

Species tatricus Smolis, A et Skarzynski, D, [2002]:407

             Syn.: Stachorutes tatricusa(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS
             Syn.: Stachorutes tatricusb(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS

Species tieni Pomorski, RJ et Smolis, A, 1999:?

Species triocellatus Pomorski, RJ et Smolis, A, 1999:?

             Syn.: Stachorutes triocelatus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS

Species valdeaibarensis Arbea, JI et Jordana, R, 1991:80

             Syn.: Stachorutes valdeibarensis(sic) Slawska, M, 1996:329 ISS



Genus Platanurida Carpenter, GH, 1925:5

Platanurida sp. from New Zealand
2016.03.18 © Murray, A.
Platanurida sp. from New Zealand
2016.03.18 © Murray, A.

Species lata Carpenter, GH, 1925:5, t.t.

Ceratrimeria lata from New Zealand
After Salmon JT, 1941 Pl.44 Fig.73

Species marplesi (Salmon, JT, 1941:301) Massoud, Z, 1967:212

Ceratrimeria marplesi from New Zealand
After Salmon JT, 1941 Pl.44 Fig.79
Platanurida marplesi from New Zealand
2014 © TePapa Museum
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.

Species marplesoides Massoud, Z, 1967:210

            Syn.: Platanurida marplesoïdes(sic) Massoud, Z, 1967:210 IOS
            Syn.: Ceratrimeria marplesi Salmon, 1942, nec Salmon, 1941





Genus Israelimeria Weiner, WM et Kaprus, IJ, 2005:49

Species poliakovae Weiner, WM et Kaprus, IJ, 2005:50, t.t.

Israelimeria poliakovae from Israel
After Weiner W.M. & Kaprus, I.J, 2005 Fig.1


Genus Arlesia Handschin, 1942

  Syn.: ... Arlésia(sic) Handschin, 1942 IOS
  Syn.: Neanuridae Portachorutes Wray, DL, 1953:147
  Syn.: ... Pseudarlesia Salmon, 1964
  Syn.: Pseudachorutinae Handschinia Massoud, Z, 1967:214




Species albipes (Folsom, 1927) Stach, J, 1949:57, t.t.

            Syn.: Portachorutes mambatus Wray, DL, 1953:147
            Syn.: Arlesia albipes f. decorata Mari Mutt, JA et Bellinger, PF, 1990:41


             Syn.: Arlesia albibes(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:45 ISS

Species arleana de Mendonça, MC et Fernandes, LH, 1999:1196

            Syn.: Arlesia albipes f. decorata Arlé & Rufino, 1976


Species cochabambensis Cassagnau & Rapoport, 1962

Species innominata Salmon, 1964

Species intermedia Fernandes, LH et de Mendonça, MC, 2004:21

Species variabilis Thibaud & Massoud, 1983


Genus Intermediurida Najt, Thibaud & Weiner, 1990

Species anophthalma Najt, Thibaud & Weiner, 1990, t.t.

             Syn.: Intermediurida anophtalma(sic) Queiroz, GC et Zeppelini, D, 2017:152,tt ISS
             Syn.: Intermediurida anophtalma(sic) Queiroz, GC et Zeppelini, D, 2017:153 ISS



Genus Venezuelida Díaz & Najt, 1995

Species polylepiana Díaz & Najt, 1995, t.t.


Genus Handschinurida Queiroz, GC, 2015:499

  Syn.: Pseudachorutidae Handschinia Stach, J, 1949:56 nec Lallemand, 1935
  Syn.: Collembola Handschinia Ellis, WN et Bellinger, PF, 1973:22 nec Lallemand, 1935
  Syn.: Pseudachorutinae Handschinia Queiroz, GC et de Mendonça MC, 2014:171 nec Lallemand, 1935


Species fluminensis (Arlé, 1939 tt) Queiroz, GC, 2015:499, t.t.

             Syn.: Arlesia flaminensis(sic) Palacios-Vargas, JG in Alvarez-Sánchez, J et Naranjo-García, E, 2003:221,ca ISS
Handschinia fluminensis from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.15



Species proxima (Arlé, 1939) Queiroz, GC, 2015:499

Handschinia proxima from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.19

Species rauli (Queiroz, GC & de Mendonça MC, 2014:172) Queiroz, GC, 2015:499

Handschinia rauli from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.23


Genus Brasilimeria Stach, J, 1949:56 key

Species sp. Queiroz, GC et Zeppelini, D, 2017:133

Species anura (Arlé, 1939) Stach, J, 1949:56, t.t.

Brasilimeria anura from Brazil
Dorsal aspect
Hemolymphatic defense bleeding (arrow)
After Zeppelini D et al, 2019 Fig.1a
Brasilimeria anura from Brazil
Ventral aspect
Hemolymphatic defense bleeding (arrows)
After Zeppelini D et al, 2019 Fig.1b

Species assua Zeppelini, D, Queiroz, GC, Lopes, NP et Mendonça-Junior, FJB, 2019:5

            Syn.: Brasilimeria assu(sic) Zeppelini, D, Queiroz, GC, Lopes, NP et Mendonça-Junior, FJB, 2019:5 IOS

Species wygodzinskyi (Arlé, R, 1943:110) Arlé, 1959

            Syn.: Ceratrimeria Wygodzinskyi(sic) Arlé, R, 1943:110 IOS
Brasilimeria wygodzinskyi from Brazil
After Massoud Z, 1967 Fig.69G




Genus Tijucameria de Mendonça, MC et Fernandes, LH, 2005:699

  Syn.: Pseudachorutinae Tijucameira(sic) Queiroz, GC et Zeppelini, D, 2017:131 ISS

Species sp. Queiroz, GC et Zeppelini, D, 2017:133

            Syn.: Neotropiella denisi Arlé, 1981


Species gabrieli de Mendonça & Silveira, 2012

Species mame de Mendonça, MC et Fernandes, LH, 2005:699, t.t.


Genus Kenyura Salmon, 1954

  Syn.: Neanuridae Paranurella Arlé, R, 1960:162
  Syn.: Pseudachorutini Pseudoparanura Massoud, Z, 1963:70 =?


Species africana (Massoud, Z, 1963:70) Massoud, Z, 1967:230

Species delicata Arlé, R, 1966:4

Species monoculata (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.

Species multisensa Salmon, 1954, t.t.

Species nicaraguensis Palacios-Vargas & Deharveng, 2010

Species porcula (Arlé, R, 1960:163) Arlé, R, 1966:2

            Syn.: Paranurella porculus(sic) Arlé, R, 1960:163 IOS


             Syn.: Kenyura porculus(sic) Arlé, R, 1966:2 ISS
             Syn.: Kenyura porculus(sic) Massoud, Z, 1967:229 ISS
             Syn.: Kenyura porculus(sic) Arlé, 1981 ISS
             Syn.: Kenyura porculus(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
             Syn.: Kenyura porculus(sic) Culik, MP et Zeppelini, DF, 2003:1124 ISS



Species xinguensis Arlé, R, 1966:3


Genus Furculanurida Massoud, Z, 1967:231 key

Species africana (Massoud, Z, 1963:68) Massoud, 1967:231, t.t.

Species arawakensis Thibaud & Massoud, 1983

Species arlei Thibaud & Massoud, 1980

Species ashrafi (Yosii, 1966) Thibaud, J-M & Palacios-Vargas, JG, 2000:112

Species belemensis Arlé & Rufino, 1976

Species boiunae da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:4

            Syn.: Furculanurida boiuna(sic) da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:4 IOS
            Syn.: Furculanurida (Furculanirida) boiuna(sic) da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:6 IOS

Species duodecimoculata Thibaud & Massoud, 1980

Species emucronata Zon, Tano & Deharveng, 2014

Species furculata (Salmon, 1956) Massoud, 1967:233

Species goeldiana Arlé & Rufino, 1976

Species grandcolasorum Weiner, WM et Najt, J, 1998:223

Furculanurida grandcolasorum from Tanzania
Bicolour form
After Weiner W.M. & Najt J, 1998 Fig.17

Species guatemalensis Palacios-Vargas, JG et Gao, Y, 2009:55

Furculanurida guatemalensis from Guatemala
After Palacios JG & Gao Y, 2009 Fig.1
             Syn.: Furculanurida guatemaltesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS
             Syn.: Furculanurida guatemalesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS


Species langdoni Bernard, 2007

Species longisensillata Najt, Thibaud & Weiner, 1990

Species nessimiani (Fernandes, LH & de Mendonça, MC, 2003:4) Palacios-Vargas, JG & Gao, Y, 2009:55

             Syn.: Furculanurida nessiamiani(sic) Palacios-Vargas, JG et Gao, Y, 2009:55 ISS

Species perplexa (Salmon, 1956) Massoud, 1967:233

Species septemoculata Palacios-Vargas, JG et Gao, Y, 2009:56

Furculanurida septemoculata from Guatemala
After Palacios JG & Gao Y, 2009 Fig.12

Species tropicalia Queiroz & Fernandes, 2011


Genus Micranurida Börner, 1901

Species agenjoi Simón, 1981

Species anini Christiansen, K et Bellinger, P, 1992:68

Species anophthalmica Stach, J, 1946:26

Species balta Fjellberg, A, 1998:77

Species bescidica Smolis, A et Skarzynski, D, 2004:1

Species candida Cassagnau, P, 1952:310

Species fluminensis Fernandes, LH et de Mendonça, MC, 2004:22

Species forsslundi Gisin, 1949, B

Species furcifera Mills, HB, 1934:27

Species grana Lee & Kim, 1984

Species hygrophila Rusek, 1973

Species intermedia Dallai, 1973

Species japonica Tamura, 1998

Species lanceolata Najt, J et Weiner, WM, 1985:35

Species meridionalis Cassagnau, P, 1952:310

Species montana Lee & Kim, 1984

Species papillosa Axelson, 1902, B

Species porcella Fjellberg, 1985

Species pygmaea Börner, 1901, t.t., B

            Syn.: Micranurida pygmaea f. hasai Kseneman, 1936
            Syn.: Micranurida meridionalis Arbea & Jordana, 1985


Micranuridae pygmaea from Italy
After Dallai R, 1973 Fig.6n
             Syn.: Micranurida pygmea(sic) Slawska, M, 1996:328 ISS


Species retezatica Gruia & Harsia, 1990

Species rostrata Babenko, AB, 2007:364

Species sensillata (Gisin, 1953) Dunger, 1974

Species sp. nov. Palacios-Vargas, 1981:89

Species sp. nov. Palacios-Vargas & Gómez-Anaya, 1993:8

Species spirillifera Hammer, 1953

             Syn.: Micranurida spirilifera(sic) Massoud, Z, 1967:236 ISS

Species tuberculata Lee & Kim, 1984

Species valiana Fjellberg, 1985

Species wladimiri Najt & Rubio, 1978


Genus Meganurida Carpenter, 1935

Species mumfordi Carpenter, 1935, t.t.


Genus Pseudachorudina Stach, 1949

Species alpina Stach, 1949

Species angelieri Cassagnau, P, 1959:76

Species berninii (Dallai, R, 1970:438) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:5


Pseudachorutes brunneus from New Zealand
After Salmon, J.T. 1941 Pl.46 Fig.101.
Species brunnea (Carpenter, GH, 1925:2) Massoud, Z, 1967:243


Species dahli (Schäffer, C, 1898:400) Yoshii, R & Suhardjno, YR, 1992:2

            Syn.: Schöttella(sic) dahlii(sic) Schäffer, C, 1898:400 IOS
            Syn.: Pseudachorudina delamarei Massoud, 1965


             Syn.: Pseudachorutes dahlii(sic) Börner, 1901 ISS
             Syn.: Schöttella(sic) dahlii(sic) Schött, 1901 ISS
             Syn.: Schöttella(sic) dahlii(sic) Stach, 1929 ISS
             Syn.: Pseudachorutes dahlii(sic) Womersley, 1937 ISS
             Syn.: Schoettella dahlii(sic) Salmon, JT, 1964:200 ISS
             Syn.: Pseudachorutes dahlii(sic) Massoud, Z, 1967:165 ISS
             Syn.: Pseudachorutes? dahlii(sic) Yayuk, RS(sic), 1989:4 ISS
             Syn.: Pseudachorudina dahlii(sic) Yoshii, R et Suhardjno, YR, 1992:2 ISS
Pseudachoridina dahlii(sic) from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.1A





Species evansi (Womersley, H, 1936:475) Stach, 1949

Species falteronensis (Denis, 1926) Massoud, 1967:243, t.t.

Species ignota Christiansen & Bellinger, 1980:300

Species mabirensis (Philiptschenko, 1926) Salmon, 1954

Species meridionalis (Bonet, 1929) Jordana, R, Arbea, JI, Simón, C & Luciáñez, MJ in Ramos, MA & al, 1997:777

            Syn.: Pseudachorudina bougisi Delamare Deboutteville, 1951
            Syn.: Pseudachorudina bougisi var. catalonica Massoud, 1963
            Syn.: Pseudachorudina patriciae Gough, 1976
            Syn.: Pseudachorudina najtae Simon & Pozo, 1989


Species nepalica Yosii, 1966


Brachystomella osextara from New Zealand
After Salmon, J.T. 1941 Pl.45 Fig.88.
Species osextara (Salmon, JT, 1941:303) Massoud, Z, 1967:243

Species pacifica (Womersley, H, 1936:318) Massoud, 1967:241

Species texensis Christiansen & Bellinger, 1980:301


Genus Cassagnaudina Massoud, 1967

Species coiffaiti (Cassagnau, P, 1955:145) Massoud, Z, 1967:245

            Syn.: Pseudachorudina Coiffaiti(sic) Cassagnau, P, 1955:145 IOS

Species khumbuensis Yosii, 1971

Species perisi Simón, 1981

Species tourratensis (Cassagnau, P, 1964:470)


Genus Cassagnaurida Salmon, 1964

Species dentata (Cassagnau & Rapoport, 1962) Massoud, 1967:246, t.t.


Genus Notachorudina Cassagnau & Rapoport, 1962

  Syn.: Collembola Nothachorudina(sic) Rapoport, EH et Rubio, IR, 1963:111 ISS

Species patagonica Cassagnau & Rapoport, 1962, t.t.

Species rapoporti Queiroz, GC et Zeppelini, D, 2017:138

Notachorudina rapoporti from Chile
With everted anal sacs
After Queiroz GC & Zeppelini D, 2017 Fig.11B


Genus Forsteramea Salmon, JT, 1965:645

Species megacephala (Salmon, JT, 1954:213) Salmon, JT, 1965:645, t.t.

            Syn.: Montachorutes megacephala(sic) Salmon, JT, 1954:213 IOS
Montachorutes megacephala(sic) from New Zealand
After Salmon JT, 1954 Text-Fig.1

             Syn.: Montachorutes? megacephala(sic) Rapoport, EH et Rubio, IR, 1963:113 ISS
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.






Genus Delamarellina Rapoport, EH et Rubio, IR, 1963:113

  Syn.: Pseudachorutinae Dellamarelina Queiroz, GC et Zeppelini, D, 2017:129 ISS

Species globulosa (Cassagnau & Rapoport, 1962) Massoud, Z, 1967:253

Species guilleni Rapoport, EH et Rubio, IR, 1963:113, t.t.

Delamarellina guilleni from Chile
After Rapoport EH & Rubio IR, 1963 Fig.62

Species ubiquata (Salmon, JT, 1944:133) Massoud, Z, 1967:251


Genus Quatacanthella Salmon, JT, 1945:68


Polyacanthella proprieta from New Zealand
After Salmon, J.T. 1941 Pl.42 Fig.57.
Species proprieta (Salmon, JT, 1941:296) Salmon, JT, 1945:68, t.t.


Genus Anurida Laboulbène, 1865 key  key

  Syn.: ... Anoura(sic) Gervais, 1844
  Syn.: Lipuridæ(sic) Anoura(sic) Lubbock, J, 1862:601 partim ISS
  Syn.: Aphoruridæ(sic) Aphoromma MacGillivray, AD, 1893:314
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: Hypanurida Denis, 1931 Hypanurida Denis, 1931 Hypanurida Denis, 1931
  Syn.: Pseudachorutini Anuridella Mills, HB, 1934:26
  Syn.: Neanurinae Anuridella Gisin, H, 1960:87
  Syn.: Neanurinae Anuridella Poole, RW in Poole, RW et Gentili, P, 1997:46
  Syn.: Neanurinae Gastranurida Gisin, H, 1960:87


Anurida sp. from Holland
2018.09.30 © van Leeuwen, J.



Species abashiriensis Babenko, A et Nakamori, T, 2018:377

Anurida abashiriensis from Japan
After Babenko A & Nakamori T, 2018 Fig.12-13

Species alpina Agrell, 1939

Species amorita Folsom, JW, 1902:90

Anurida amorita from Alaska
After Folsom JW, 1902 Pl.IV Fig.3

Species annatshagi Tshelnokov, 1988

Species ariesi Harsia & Gruia, 1992


Anurida ashbyae from the Lesser Antilles
Det. Christiansen, K.
2006 © De Silva, M.
Species ashbyae Christiansen, K et Bellinger, P, 1988:228


Species asiatica Babenko, AB, 2002:374

Species assimilis Yosii, R, 1956:28

  Subspecies persimilis Yosii, R, 1956:28

Species azurea Babenko, AB, 1997:513

Species balatovae Rusek, 1970

Species beringi Fjellberg, 1985

            Syn.: Anurida tullbergi Hammer, 1953
            Syn.: Anurida (Anurida) tullbergi Christiansen, K et Bellinger, P, 1980:323


Species bisetosa Bagnall, 1949

Species bondarenkoae Tshelnokov, 1988

Species bonneti Cassagnau, P, 1955:147

            Syn.: Anurida Bonneti(sic) Cassagnau, P, 1955:147 IOS

Species borealis Martynova in Stebaeva, 1976, n.n.

Species callainii Dallai, R et Martinozzi, I, 1980:8

Species caprariensis Dallai, 1969

Species carpatica Babenko, AB, 1998:658

Species clavata Schäffer, 1897

Species confinis Babenko, AB, 1998:651

Species decemoculata Hammer, 1953

Species dentata Christiansen & Bellinger, 1980:326

            Syn.: Anurida (Anurida) decemoculata Christiansen & Bellinger, 1980,si


Species desnuda Yosii, R, 1956:25

Species diabolica Yosii, R, 1956:26

Species dynkengda Babenko, AB, 1998:649

Species frigida Fjellberg, 1973

Species granaria (Nicolet, H, 1847:387) Tullberg, 1869, t.t., B

            Syn.: Anurida bagnalli Stach, J, 1949:14
            Syn.: Anurida comellinii Gisin, 1949
            Syn.: Anurida pseudogranaria Stach, J, 1949:30


Anoura granaria from the UK
After Lubbock J, 1873 Pl.49
Anurida granaria from the UK
2005 © Hopkin, S.P.
Anurida granaria from Sweden
2006 © Hall, K.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria juv. from the UK
2013.01.29 © Murray, A.
Anurida granaria from the UK
Circular post-antennal organ
2013.01.29 © Murray, A.
Anurida granaria from the UK
Granulated cuticula
2013.01.29 © Murray, A.
Anurida granaria from the UK
Antennae and mouthcone
2013.03.23 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from Holland
2014.03.14 © van Duinen, J.
Anurida granaria from Canada
Anal spines absent
2015.08.12 © Murray, A.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
Cuticle coarsely granulate
Abd.6 not bilobed
2015.09.26 © Tony.
Anurida granaria from Tasmania
Postantennal organ circular
Tapering antenna
2015.09.26 © Tony.
Anurida granaria from New Zealand
Mouth: labrum, maxillae heads, hypopharynx, split labium
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Ant.4: apical eversible trilobed sensillar vesicle
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Rudiment of furcula
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Post-antennal organ
2016.02.10 © Koken, M.

Species granulata Agrell, 1943, B

            Syn.: Micranurida sexpunctata Handschin, 1924, nec Axelson, 1902
            Syn.: Hypanurida mirabilis Denis, 1931
            Syn.: Hypanurida flavescens Kseneman, 1934
            Syn.: Micranurida handschini Gisin, 1944
            Syn.: Anurida hexophthalmica Stach, J, 1949:26


Species hammerae Christiansen, KA, 1952:135

            Syn.: Anurida hammeri(sic) Christiansen, KA, 1952:135 IOS
Anurida hammeri(sic) from Alaska
After Christiansen K, 1952 Pl.12 Fig.1-2

            Syn.: Anurida polychaetosa Tshelnokov, 1988



             Syn.: Anurida hammeri(sic) Massoud, Z, 1967:265 ISS
             Syn.: Anurida (Anurida) hammeri(sic) Christiansen, K et Bellinger, P, 1980:330 ISS
             Syn.: Anurida hammeri(sic) Christiansen, KA in Dindal, DL, 1990:978 ISS
             Syn.: Anurida hammeri(sic) Poole, RW in Poole, RW et Gentili, P, 1997:47 ISS



Species interior Fjellberg, 1985

Species iriei Yosii, R, 1970:4

Species kaya Babenko, AB, 1998:656

Species kazukoae Tamura & Zhao, 1997

             Syn.: Anurida kazokoae(sic) Yue, Q-Y et Yin, W-Y, 1999:222 ISS

Species kolymensis Tshelnokov, 1988

Species komi Babenko, AB, 1998:653

Species kyshyensis Babenko, A et Nakamori, T, 2018:575

Anurida kyshyensis from Russia
After Babenko A & Nakamori T, 2018 Fig.2-5

Species latosensillata Dunger, 1975

Species luciae Fjellberg, A, 1985:359

Species lvivska Babenko, AB, 1998:659

Species mandibulata Tshelnokov, 1988

Species mara Christiansen, K et Bellinger, P, 1988:230

Species marchena Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:161

             Syn.: Anurida marchana(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS

Species maritima (Guérin-Méneville, 1836 tt) Laboulbène, 1865, t.t., B

            Syn.: Podura ambulatoria Ström in Plateau, 1890
            Syn.: Anurida bisetosa Bagnall, 1949
            Syn.: Anurida megalops Bagnall, 1949


Anurida maritima from Germany
Capitulum of maxilla
After Anonymous fig.C

             Syn.: Anurida maritime(sic) D'Haese, CA, 2003:581 ISS
Anurida maritima from France
2003 © De Bruyn, L.
Anurida maritima from the UK
2005 © Hopkin, S.P.
Anurida maritima from the UK
Post-antennal organ and eye with 5 ocelli
2005 © Hopkin, S.P.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Aggregation on tidal pool
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Circular post-antennal organ (PAO) with max 10 vesicles
2007.08.11 © Reed,D.E.
Anurida maritima from the UK
2008.01.06 © Roberts, S.
Anurida maritima from the UK
2008.03.30 © Roberts, S.
Anurida maritima from the UK
2008.06.29 © Ramone, J.
Anurida maritima from the UK
2008.07.27 © Roberts, S.
Anurida maritima from the USA
2009.02.21 © Edmonds, M.
Anurida maritima from France
2009.08.09 © Henderickx, H.
Anurida maritima from France
Egg cluster
2009.08.09 © Henderickx, H.
Anurida maritima from the USA
Aggregation on tidal pool
2013.07.21 © Kynd, F.
Anurida maritima from Portugal
Eyepatch with 5 ocelli
2016.08.06 © Almeida, J.
Anurida maritima from Holland
Mouthcone moderately pointed
2017.07.21 © van Duinen, J.
Anurida maritima from Holland
2017.08.04 © Wieringa, P.
Anurida maritima from Holland
With protruded mouthcone
2017.08.04 © Wieringa, P.
Anurida maritima from France
2019.08.31 © Tiky, S.
Anurida maritima from France
With protruded mouthcone
2019.08.31 © Tiky, S.
Anurida maritima from France
Anal spines absent
2019.08.31 © Tiky, S.
Anurida maritima from France
Unguiculus absent (*)
2019.08.31 © Tiky, S.



Species maritima2 Arbea, JI, 2001:39

Species martynovae Fjellberg, 1985

Species nadezhdae Babenko, AB, 1998:654

Species narli Fjellberg, 1985

Species neanuriformis Babenko, AB, 2002:369

Species nuda Yosii, 1956

            Syn.: Anurida granaria Uchida, 1952
            Syn.: Anurida granaria Yosii, R, 1977:147


Species okamotoi Yosii, R, 1970:2

Species orientalis Martynova in Stebaeva, 1976, n.n.

Species palustris Babenko, AB, 1997:522

Species papillosa (Axelson, 1902) Fjellberg, 1985

Species papillosoides (Hammer, 1953) Massoud, Z, 1967:258

             Syn.: Anurida papillosoïdes Massoud, Z, 1967:258 ISS

Species parapapillosa Tshelnokov, 1988

Species plurichaetotica Yosii, R, 1966:544

Species polaris (Hammer, 1954) Fjellberg, 1985

Species pusilla da Gama, 1964

Species reducta Fjellberg, 1985

Species riverina Sterzynska, M, Kaprus, I et Ehrnsberger, R, 2006:255

Anurida riverina from Poland
After Sterzynska M, Kaprus I & Ehrnsberger R, 2006 Fig.1

Species sakhensis Babenko, AB, 1997:529

Species setosa Yosii, R, 1956:27

Species similis Fjellberg, 1985

Species speobia Yosii, R, 1954:64

Species stereoodorata Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49

Anurida stereoodorata from Abkhazi
After Jordana R & al, 2012 Fig.11A




             Syn.: Anurida stereoodarata(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49 ISS

Species subarctica Fjellberg, 1985

            Syn.: Anurida tajmyrica Tshelnokov, 1988


Species tamurae Yue, Q-Y et Yin, W-Y, 1999:224

            Syn.: Anurida tamurea(sic) Yue, Q-Y et Yin, W-Y, 1999:224 IOS

Species tatianae Babenko, AB, 1997:530

Species thalassophila (Bagnall, 1939) Stach, J, 1949:14

            Syn.: Anurida granaria Remy, 1928
            Syn.: Anurida remyi Denis, 1947


Species trioculata Kinoshita, 1916

Species troglodyta Lee & Park, 2016

Species tullbergi Schött, 1891:192

            Syn.: Anurida crassicornis Reuter L & Reuter OM, 1879-1880 =?
            Syn.: Anurida ellipsoides Stach, J, 1949:34
             Syn.: Anurida ellipsoïdes(sic) Massoud, Z, 1967:260 ISS
             Syn.: Anurida elipsoides(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS
             Syn.: Anurida ellipsoids(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:260 ISS
            Syn.: Anurida brunsvigiensis Hüther, 1964
             Syn.: Anurida brusvigiensis(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS


             Syn.: Anurida Tullbergi(sic) Schött, H, 1893:91 ISS
Anurida Tullbergii(sic) from Sweden
After Schött H, 1893 Taf.VII Fig.16


             Syn.: Anurida Tullbergi(sic) Schött, H, 1917:6 ISS
             Syn.: Anurida tulbergii(sic) Shaw, P et Benefer, CM, 2015:200 ISS
Anurida tullbergi from Belgium
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Post-antennal organ (PAO) oval
Eyepatch with 5 ocelli
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Ant.3+4 fused
2017.05.20 © Huskens, M.-L.



Species uniformis Gisin, 1953

             Syn.: Anurida uniformes(sic) Babenko, AB, 1998:654 ISS

Species vicaria Yosii, R, 1956:30

Species vontoernei Nosek, 1962

Species vulcanica Fjellberg, A, 1985:357

Species weberi Christiansen & Bellinger, 1980:335

Species wrangelensis Tshelnokov, 1988

Species zanokhae Babenko, AB, 1998:649


Genus Minotaurella Weiner, WM, 1999:373

Species banyulensis (Denis, 1947) Weiner, WM, 1999:376

Species edaphica Kaprus, IJ et Nevo, E, 2003:67

Minotaurella edaphica from Israel
After Kaprus I.J. & Nevo E, 2003 Fig.2a

Species isabellae Weiner, WM, 1999:374, t.t.

             Syn.: Minotaurella isabelle(sic) Kaprus, IJ et Nevo, E, 2003:69 ISS



Genus Anuridella Willem, 1906

  Syn.: ... Anurodes Bagnall, 1949
Anuridella sp. from the UK
2013.08.23 © Fenwick, D.



Species sp. Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:892

Species calcarata Denis, 1925:267

            Syn.: Anuridella immsiana Bagnall, 1939


Species germanica (Gisin, 1951)

Species marina Willem, V, 1906:247, t.t.

Anuridella marina
After Willem V, 1906
            Syn.: Anuridella hintoni Bagnall, 1949



Species submarina Bagnall, 1934


Genus Gastranurida Bagnall, 1949

Species denisi (Bagnall, 1939) Massoud, Z, 1967:268, t.t.

            Syn.: Anurida trioculata Denis, 1922, nec Kinoshita, 1916
            Syn.: Paranura hibernica Bagnall, 1941
             Syn.: Gastranurida gisini(sic) Babenko, A et Nakamori, T, 2018:578 lapsus ISS



Genus Hylaeanura Arlé, R, 1966:5 key

  Syn.: Pseudachorutinae Hylaenura(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
  Syn.: Neanuridae Hyleanura(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS

Species infima (Arlé, R, 1960:163) Arlé, R, 1966:5, t.t.

Species mendoncae Zeppelini, D et Palacios-Vargas, JG, 2013:1402

Species nepalensis (Yosii, 1966) Vázquez, M, Cutz-Pool, LQ & Palacios-Vargas, JG, 1998:370

Species nohbecana Vázquez, M, Cutz-Pool, LQ et Palacios-Vargas, JG, 1998:367


Genus Anurachorutes Kuznetsova & Potapov, 1988

Species martynovae Kuznetsova & Potapov, 1988


Genus Anuritelsa Womersley, H, 1939:315

Species maritima Womersley, H, 1939:316, t.t.


Genus Gamachorutes Cassagnau, 1978

Species verrucosus Cassagnau, 1978


Genus Koreanurina Najt & Weiner, 1992

Species alba Wu, Bedos & Deharveng, 2014

Species inexpectata Najt & Weiner, 1992

Species szeptyckii Najt & Weiner, 1992

Species weinerae Wu, Bedos & Deharveng, 2014


Genus Lanzhotia Rusek, 1985

Species brachycera Rusek, 1985


Genus Paranurida Skarzynski, D et Pomorski, RJ, 1994:363

Species kopetdagi Skarzynski, D et Pomorski, RJ, 1994:364


Genus Philotella Najt, J et Weiner, WM, 1985:29

Species deharvengi Najt, J et Weiner, WM, 1985:30, t.t.

Species miracli Najt, J et Weiner, WM, 1985:32

             Syn.: Philotella miracle(sic) Babenko, AB, 2007:364 ISS

Species obesa Babenko, AB, 2007:362

Species olgae Kniss & Thibaud, 1995

Species sp. Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41

Species tertia Najt, J et Weiner, WM, 1985:34


Genus Rusekella Deharveng, 1982

Species cantabrica Deharveng, 1982

Species cassagnaui Deharveng, 1982

Species flagellata (Arbea, JI et Jordana, R, 1985:64)

Species gamae Arbea, JI et Jordana, R, 1991:98

Species nivalis Deharveng, 1982

Species peyrei (Cassagnau, P, 1955:157)

            Syn.: Micranurida Peyrei(sic) Cassagnau, P, 1955:157 IOS

Species similis Deharveng, 1982

Rusekella similis from France
2006 © Boldsystems.org


Genus Pongeia Najt & Weiner, 2002

Species asturianensis Najt & Weiner, 2002, t.t.


Genus Holacanthella Börner, C, 1906:169 key

  Syn.: Pseudachorutinae Holocanthella(sic) Greenslade, P, 2015:5 ISS

Species brevispinosa (Salmon, JT, 1942:256) Massoud, Z, 1967:222

Ceratrimeria brevispinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.1
Holacanthella brevispinosa from New Zealand
2008 Minor, M. & Robertson, A.© SoilBugs

Species duospinosa (Salmon, JT, 1942:257) Massoud, Z, 1967:222

Ceratrimeria duospinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.9
Holacanthella duospinosa from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Holacanthella duospinosa from New Zealand
2007 © Stevens, M.
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
Ant.1 & 2 with apically pointed digitations
2014.05.26 © Murray, A.
Holacanthella duospinosa from New Zealand
2016.04.03 © Sullivan, J.
Holacanthella duospinosa from New Zealand
2017.06.13 © Maurice.
Holacanthella duospinosa from New Zealand
2017.12.13 © Maurice.

Species laterospinosa (Salmon, JT, 1944:132) Massoud, Z, 1967:222

Species paucispinosa (Salmon, JT, 1941:300) Massoud, Z, 1967:222

Ceratrimeria paucispinosa from New Zealand
After Salmon JT, 1941 Pl.44 Fig.81
Holacanthella paucispinosa from New Zealand
2006 Minor, M. & Robertson, A.© SoilBugs
Holacanthella paucispinosa from New Zealand
2006 © Stevens, M.
Holacanthella paucispinosa from New Zealand
After Thibaud J-M & D'Haese C, 2010 Fig.33
2009 © D'Haese, C.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.01.28 © Thorpe, S.E.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2013.08.31 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2013.09.22 © Jewell, T.
Holacanthella paucispinosa from New Zealand
2014.mm.dd © Stevens, M.
Holacanthella paucispinosa from New Zealand
2014.02.09 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2016.05.15 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
2016.06.22 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
Head frontoventral aspect
2016.06.22 © Navest, M.P.

Species spinosa (Lubbock, 1899) Börner, C, 1906:169, t.t.

Ceratrimeria spinosa from New Zealand
After Salmon, J.T., 1941 Pl.44 Fig.77
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
Pointed mouthcone
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
Tubercles
2014.05.15 © Murray, A.


Genus Acanthanura Börner, 1906

  Syn.: ... Anoura(sic) Lubbock, 1899 partim IOS
  Syn.: ... Ceratrimeria Börner, 1906 partim


Acanthanura sp. from Australia
2007.11.19 © Midtgaard, R.



Species sp.1 Garrick, RC, Sands, CJ, Rowell, DM, Hillis, DM et Sunnucks, P, 2006:1

Acanthanura sp. from Australia
2006 © Sunnucks, P.
Acanthanura sp. from Australia
2011.05.30 © Selemba, A.

Species sp.2 Janssens F & Murray A, 2014

Acanthanura "of Henderickx" from Tasmania
2007 © Henderickx, H.
Acanthanura "of Henderickx" from Tasmania
2012.12.29 © Zosterops, K.
Acanthanura "of Henderickx" from Tasmania
4th antennal segment
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
Pointed mouth-cone
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
Post-antennal organ
2014.09.25 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2014.11.09 © Grove, S.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.

Species sp.3 Greenslade P, 2015:il

Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. juv. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
Tuberculate cuticula
2014.09.13 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.10.28 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.11.14 © Murray, A.
Det.: Greenslade, P., 2015.06.01

Species sp.4 Janssens F & Huang D, 2019

Acanthanura sp. from Tasmania
2017.11.30 © Huang, D.
Acanthanura sp. from Tasmania
2017.11.30 © Huang, D.
Acanthanura sp. from Tasmania
Head and thorax
2017.11.30 © Huang, D.

Species dendyi (Lubbock, 1899) Börner, 1906, t.t.

            Syn.: Anoura(sic) Dendyi(sic) Lubbock, 1899 IOS
             Syn.: Neanura Dendyi(sic) Absolon, K, 1901:576 ISS
             Syn.: Acanthanura Dendyi(sic) Börner, 1906 ISS
             Syn.: Anoura(sic) Dendyi(sic) Schött, H, 1917:1 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1939 ISS
             Syn.: Acanthanura Dendyi(sic) Handschin, 1942 tt ISS
Acanthanura dendyi from Tasmania
2010.07.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010.12.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010 © Morris, R.






Genus Megalanura Ellis, WN et Bellinger, PF, 1973:30

  Syn.: ... Meganura Handschin, 1942, nec Szepligéti, 1906

Species tasmaniae (Lubbock, 1899 tt) Ellis, WN & Bellinger, PF, 1973:30

            Syn.: Anoura(sic) tasmania(sic) Lubbock, 1899 tt IOS
             Syn.: Anoura(sic) tasmaniæ(sic) Dendy, A, 1900:98 ISS
             Syn.: Neanura Tasmaniae(sic) Absolon, K, 1901:576 ISS
             Syn.: Anoura(sic) Tasmaniae(sic) Schött, H, 1917:1 ISS
Megalanura tasmaniae from Tasmania
2010 Morris, R.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Ventral aspect
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Composed eye with 8 ocelli; PAO elliptic
2014.09.26 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.






Genus Womersleymeria Stach, J, 1949:64

Womersleymeria sp. juv. from Australia
2015.07.27 © Porch, N.

Species bicornis (Womersley, H, 1940:137) Stach, J, 1949:64, t.t.

Ceratrimeria bicornis from Tasmania
After Womersley H, 1940 Fig.A
Womersleymeria bicornis from Tasmania
2010 Morris, R.
Womersleymeria bicornis from Tasmania
Aberration with fused digitations of abd.5
2014.09.22 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Pointed mouth-cone
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Post-antennal organ (PAO)
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.11.14 © Murray, A.

References