http://www.collembola.org/taxa/pseuinae.htm - Last updated on 2017.05.28 by Frans Janssens
Checklist of the Collembola: Pseudachorutinae

Subfamilia Pseudachorutinae Börner, 1906
Genus Number
of species
Acanthanura4
Aethiopella22
Aethiopellina4
Anurachorutes1
Anurida76
Anuridella5
Anuritelsa1
Arlesia6
Arlesiella6
Brasilimeria2
Cassagnaudina4
Cassagnaurida1
Cephalachorutes16
Ceratrimeria11
Delamarellina3
Forsteramea1
Furculanurida16
Gamachorutes1
Gastranurida1
Grananurida5
Halachorutes1
Handschinurida3
Holacanthella5
Hylaeanura4
Intermediurida1
Israelimeria1
Kenyura7
Koreanurina2
Lanzhotia1
Linnaniemia2
Megalanura1
Meganurida1
Micranurida28
Minotaurella3
Najtafrica1
Neotropiella20
Notachorudina1
Oudemansia7
Paranurida1
Philotella6
Platanurida3
Pongeia1
Pratanurida8
Protachorutes2
Pseudachorudina14
Pseudachorutella11
Pseudachorutes116
PseudachorutinaeGennov1
Pseudanurida6
Quatacanthella1
Rusekella7
Simonachorutes3
Stachorutes18
Tasmanura1
Tijucameria2
Venezuelida1
Womersleymeria1
Pseudachorutinae478
Genera with number of species
  Syn.: Neanuridae Pseudachorutine(sic) Skarzynski, D et Pomorski, RJ, 1994:364 ISS
Pseudachorutinae from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutinae from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutinae from Australia
2017.03.04 © Steve & Alison.


Genus Cephalachorutes Bedos, A et Deharveng, L, 1991:145
Species sp.1 Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178
            Syn.: Cephalachorutes minimus Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178 nr
Species anneae Queiroz, GC et Mendonça, MC, 2016:47
Cephalachorutes anneae from Brazil
After Queiroz GC & de Mendonça MC, 2016 Fig.8A

Species asiaticus Bedos, A et Deharveng, L, 1991:152, t.t.
Species barthae Bedos, A et Deharveng, L, 1991:149
Species caecus Bedos, A et Deharveng, L, 1991:149
Species centurionis Bedos, A et Deharveng, L, 1991:149
Species deharvengi Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41
Species delamarei (Murphy, 1965) Bedos, A & Deharveng, L, 1991:147
Species judithae Shveenkova, YB, 2017:82
            Syn.: Cephalachorutes sp.n. Shveenkova, 2011
Cephalachorutes judithae from Vietnam
After Shveenkova YB, 2017 Fig.4A

Species masi Thibaud, J-M, 2008:507
Remark by Janssens, F. 2009.02.06: Given the name is derived from the surname "Masy", strictly spoken the correct latinisation "of Masy" is "masyi".

Species microphthalmus (Barra, 1969) Bedos, A & Deharveng, L, 1991:147
Species minimus (Massoud, 1963) Bedos, A & Deharveng, L, 1991:147
Species murphyi Bedos, A et Deharveng, L, 1991:151
Species nakaoi (Yosii, 1966) Bedos, A & Deharveng, L, 1991:152
Species pestilentiae Bedos, A et Deharveng, L, 1991:151
Species pillaii (Prabhoo, 1971) Bedos, A & Deharveng, L, 1991:147

Genus Pseudachorutes Tullberg, 1871 key
  Syn.: Poduridæ(sic) Brachysius MacGillivray, AD, 1893:317
  Syn.: Poduridæ(sic) Gnathocephalus MacGillivray, AD, 1893:318
  Syn.: ... Sphragiphora Houlbert, 1924
  Syn.: ... Sphagiphorura(sic) Pomorski, RJ et Skarzynski, D, 1992:57 ISS
  Syn.: ... Logacanura Wray, 1958
  Syn.: ... Metachorudina Massoud, 1963
  Syn.: ... Neoafricella Salmon, 1964
Pseudachorutes sp. from Germany
Mandibula
After Anonymous Fig.B

  Syn.: Collembola Pseudachotures(sic) Verlinden, 1977 ISS
Pseudachorutes sp. from the USA
2010.01.17 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Ocular area
2010.01.18 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Furcula
2010.02.12 © Ten Eyck, C.
Pseudachorutes sp. from the USA
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Pointed buccal cone
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Furcula
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from Portugal
2012.10.10 © Murray, A.
Pseudachorutes sp. from the USA
2014.05.22 © Murray, T.
Pseudachorutes sp. from Holland
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Holland
Pointed mouthcone
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Germany
Large round post-antennal organ (PAO)
2015.02.20 © Mattew, M.
Pseudachorutes sp. from the UK
2015.02.26 © Murray, A.
Pseudachorutes sp. from the UK
Pointed mouth-cone
2015.02.26 © Murray, A.
Pseudachorutes sp. from France
2016.03.03 © Garcelon, P.
Pseudachorutes sp. from Australia
2016.04.25 © Murray, A.

Species sp.1 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8
Species sp.2 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8
Species sp.3 Christiansen, K et Bellinger, P, 1980:297bis
Species sp.4 Christiansen, K et Bellinger, P, 1980:297bis
Species sp.5 Christiansen, K et Bellinger, P, 1980:297bis
Species sp.6 Soto-Adames, FN et Taylor, SJ, 2013:153
Species algidensis Carpenter, GH, 1925:4
             Syn.: Pseudachorutes aldgidenssi(sic) Womersley, H, 1936:327 ISS
Species alluaudi Delamare Deboutteville, 1945
             Syn.: Pseudachorutes allauaudi(sic) Massoud, 1967 ISS
Species americanus Stach, 1949
            Syn.: Pseudachorutes boerneri Nearctic records
            Syn.: Pseudachorutes corticolus Mills, 1934
            Syn.: Pseudachorutes corticicolus f. globulisetosa Gisin, 1944, Nearctic records
Species amucronatus Díaz & Najt, 1995
Species andrei Weiner, WM et Najt, J, 1985:260
Species anomalus Imms, 1912
Species aphysus Christiansen & Bellinger, 1980:304
Species aureofasciatus (Mac Gillivray, 1893) Mills, HB, 1934:23
            Syn.: Pseudachorutes aureofasciatus f. millsi Maynard, EA, 1951:58
Pseudachorutes aureofasciatus from the USA
Non-banded juvenile
2007.02.22 © Bernard, E.C.

            Syn.: Pseudachorutes (Pseudachorutes) bifidus Christiansen & Bellinger, 1980
Pseudachorutes aureofasciatus
1980 © Norton, R.A. in Dindal.
Pseudachorutes aureofasciatus from the USA
Banded juvenile
2007.02.22 © Bernard, E.C.
Pseudachorutes aureofasciatus from the USA
"Butts up colony"
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
Badly smelling defensive secretion
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2012.05.01 © Murray, T.

Species auricularius Cassagnau & Rapoport, 1962
Species beta Najt, J et Weiner, WM, 1991:106
Species bifasciatus Oliveira & Deharveng, 1994
Species bifidus Christiansen & Bellinger, 1980:307
Species bobeitio Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32
Species cacaoiensis Najt, Thibaud & Weiner, 1990
Species caerulescens Schött, H, 1893:24
            Syn.: Pseudachorutes cærulescens(sic) Schött, H, 1893:24 IOS
             Syn.: Pseudachorutes coerulescens(sic) Wahlgren, E, 1906:4 ISS
Species caramel Kaprus, IJ, Weiner, WM et Nevo, E, [2005]:263
Pseudachorutes caramel from Israel
After Kaprus, I.J., Weiner, W.M. & Nevo, E., [2005] Fig.1

Species chazeaui Najt, J et Weiner, WM, 1991:100
Species cheni Shi, S, Jiang, J et Pan, Z-X, 2008:57
Pseudachorutes cheni from China
After Shi S et al., 2008 Fig.1-2

Species columbicus Rusek, J, 1991:120
Species complexus (MacGillivray, AD, 1893:318) Folsom, 1916
Species conicus Lee & Kim, 1984
Species conspicuatus Salmon, JT, 1944:137
            Syn.: Pseudachorutes conspicuatus ssp. principalis Massoud, Z, 1967:156
Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.
Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.

  Subspecies flavus Salmon, JT, 1944:139
  Subspecies lineatus Salmon, JT, 1944:138
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2009 © Te Papa Museum
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.

  Subspecies maximus Salmon, JT, 1944:138
Species corticicolus (Schäffer, 1897) Krausbauer, 1898
             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1898 ISS
             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1901 ISS
             Syn.: Pseudachorutes börneri(sic) Schött, 1902 ISS
             Syn.: Pseudachorutes corticola(sic) Kloet, GS et Hincks, WD, 1945:4 ISS
             Syn.: Pseudachorutes corticolus(sic) Snider, RJ, 1967:192 ISS
             Syn.: Pseudachorutes corticolus(sic) Palacios-Vargas, JG, 1990:19 ISS
             Syn.: Pseudachorutes corticolus(sic) Szeptycki, A et Weiner, WM in Razowski J, 1990:20 ISS
             Syn.: Pseudachorutes corticolus(sic) Barra, J-A, 1993:71 ISS
             Syn.: Pseudachorutes corticicola(sic) Farkas, J, 1995:46 ISS
             Syn.: Pseudachorutes corticolus(sic) Kopeszki, H et Meyer, E, 1996:224 ISS
             Syn.: Pseudachorutes corticicola(sic) Arbea, JI et Jordana, R in Ramos, MA et al, 1997:323 ISS
  Forma globulisetosa Gisin, 1944
Species crassus da Gama, 1964
Species curtus Christiansen & Bellinger, 1980:232
Species dalensi Weiner, WM et Najt, J, 1985:258
Species decussus (Salmon, JT, 1941:298)
Micranurida decussa from New Zealand
After Salmon JT, 1941 Pl.43 Fig.72

Species deficiens Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:558
Pseudachorutes deficiens from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.1A,E

Species deserticus Palacios-Vargas, 1990:11
Species difficilis Denis, 1931
Species dilatatus (MacGillivray, AD, 1893:317) Christiansen & Bellinger, 1980
Species dubius Krausbauer, 1898
            Syn.: Pseudachorutes lapponicus Ågren, H, 1904:8
            Syn.: Schöttella(sic) poppei Schäffer, 1897
Species galapagoensis Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:157
Species gilvus Oliveira & Deharveng, 1995
Species geronensis (Massoud, 1963) Massoud, 1967:158
Species gilvus Oliveira & Deharveng, 1994
Species guyanensis Najt, Thibaud & Weiner, 1990
Species herberti Arlé & Rufino, 1976
Species hitakamiensis Tamura, 2001
Species hodeberti Thibaud, J-M, 2008:508
Species indiana Christiansen, K et Bellinger, P, 1980:312
Species innominatus Denis, 1931
Species insularis Yosii, 1965
Species isawaensis Tamura, 2001
Species janstachi Kaprus, IJ et Weiner, WM, 2009:9
Pseudachorutes janstachi from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.32

Species japonicus Kinoshita, 1916
Species javanicus (Handschin, 1920, i.s.) Handschin, 1920
Species jeonjuensis Lee & Kim, 1984
Species jianxiucheni Gao, Yin & Palacios-Vargas, 2008
Species kangchenjungae Yosii, 1966
Species labiatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:565
Pseudachorutes labiatus from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.5A

Species laricis Arbea, JI et Jordana, R, 1989:161
Species legrisi Thibaud & Massoud, 1983
Species libanensis (Cassagnau, P et Delamare Deboutteville, C, 1955:372)
Species libanensis Ellis, WN, 1976:269
Remark by Kaprus, I.J., Weiner, W.M. & Nevo, E. [2005]:263: P. libanensis was described from Lebanon as Aetiopella(sic) libanensis by CASSAGNAU and DELAMARE (1955) and subsequently redescribed by ELLIS (1976) on the basis of the material from Central Crete. The comparison of both descriptions shows, that they concern two separate species.

Species lishanensis Gao, Yin & Palacios-Vargas, 2008
Species longisetis Yosii, 1961
             Syn.: Pseudachorutes longisetus(sic) Cassagnau, P, 1974:309 ISS
Species lunatus Folsom, 1916
            Syn.: Pseudachorutes (Pseudachorudina) ignotus Christiansen, K et Bellinger, P, 1980:301 =?
Species macrochaetus Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:158
Species massoudi Arlé, R, 1966:7
Species medjugorjensis Barra, 1993:69
Species minor Peja, 1985
Species multidentatus Najt, Thibaud & Weiner, 1990
Species murphyi Massoud, 1965
             Syn.: Pseudachorutes murphy(sic) Peck, SB et Jacquemart, J, 2011:5,cf ISS
Species namphoensis Weiner, WM et Najt, J, 1985:262
Species nica Palacios-Vargas, JG et Mejía-Madrid, HH, 2012:64
Species niloticus Wahlgren, E, 1906:3
Species octosensillatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:561
Species orghidani Massoud & Gruia, 1974
Species orientalis Loring, 1984
Species ouatilouensis Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32
Species palmiensis Börner, 1903
            Syn.: Pseudachorutes palmiensis f. secundo Denis, 1948
            Syn.: Pseudachorudina conjungens Stach, 1949
            Syn.: Pseudachorutes cantabricus Simón & Pozo, 1979
             Syn.: Pseudachorutes palmatus(sic) Wahlgren, E, 1906:4 lapsus ISS
Pseudachorutes palmiensis from France
2006 © Boldsystems.org
Pseudachorutes palmiensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.42
Pseudachorutes palmiensis from Spain
2016.04.24 © Tizon, J.

Species parvulus Börner, 1901, B
            Syn.: Pseudachorutes parvulus f. infuscata Yosii, 1954
            Syn.: Pseudachorutes corticicolus Selga, 1971
            Syn.: Pseudachorutes subcrassus Selga, 1971, partim Arbea & Jordana in Ramos, 1997
Pseudachorutes parvulus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.52

Species pasurus (Wray, 1958) Christiansen & Bellinger, 1980
Species pauliani Delamare Deboutteville, 1952
Species periyarensis Prabhoo, 1971
Species plurichaetosus Arbea, JI et Jordana, R, 1991:78
Species poahi Christiansen, K et Bellinger, P, 1992:65
Species polychaetosus Gao, Y et Palacios-Vargas, J, 2008:54
Species ponmudiensis (Prabhoo, 1971, i.s.) Prabhoo, 1971
Species pratensis Rusek, 1973
Pseudachorutes pratensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.65

Species prosimplex Weiner, WM et Najt, J, 1985:256
Species pseudoniloticus Massoud, 1963
Species puniceus Salmon, JT, 1944:139
Species quadrisetosus Snider, RJ, 1981:244
Species reductus Thibaud & Massoud, 1983
Species rhaeticus (Carl, J, 1899:298) Handschin, E, 1924:96
            Syn.: Schöttella(sic) rhätica(sic) Carl, J, 1899:298 IOS
            Syn.: Schöttella(sic) rhætica(sic) Carl, J, 1899:298 IOS
Species rugatus Wray, 1952
Species saxatilis Macnamara, 1920
Species scythicus Kaprus, IJ et Weiner, WM, 2009:3
Pseudachorutes scythicus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.1

Species sexsetosus Denis, 1934
Species shiragamiensis Hisamatsu & Tamura, 1998
Species sibiricus Reuter, 1891
Species sibiricus (Rusek J, 1991:123) Rusek, J, 1991:123
Species simplex Maynard, EA, 1951:59
Pseudachorutes simplex from the USA
2006 © Bernard, E.C.
Pseudachorutes simplex from the USA
2007.02.22 © Bernard, E.C.

Species subabdominalis Steiner, 1958
Species subcrassoides Mills, HB, 1934:24
             Syn.: Pseudachorutes subcrassoïdes Massoud, 1967 ISS
Species subcrassus Tullberg, 1871, t.t., B
            Syn.: Pseudachorutes tuberculatus Schött, 1902
Pseudachorutes subcrassus from Russia
After Scherbakov AM, 1898 Table I Fig.4
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from Holland
2015.02.21 © Baas, A.H.

Species subcrassus Schäffer, 1897, nec Tullberg, 1871
Species tamajonensis Simón, 1986
Species tasmaniensis Womersley, H, 1936:475
Species tavignani Poinsot-Balaguer, 1978
Species tillieri Najt, J et Weiner, WM, 1991:104
Species unidentatus Salmon, 1956, i.s.
Species uniformis Thibaud & Massoud, 1983
Species univesicatus Weiner & Najt, 1991
             Syn.: Pseudachorutes universicatus(sic) Greenslade, P, 1997-:www0312 ISS
Species vasylii Kaprus, IJ et Weiner, WM, 2009:7
Pseudachorutes vasylii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.22

Species vitalii Kaprus, IJ et Weiner, WM, 2009:5
Pseudachorutes vitalii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.11

Species wandae Gao, Yin & Palacios-Vargas, 2008
Species yoshiii Najt, J et Weiner, WM, 1991:102
            Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM, 1991:102 IOS
             Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36 ISS
             Syn.: Pseudachorutes yoshii(sic) Gao, Y et Palacios-Vargas, J, 2008:55 ISS

Genus Simonachorutes Skarzynski, D, Arbea, JI et Piwnik, A, 2016:172
Species agrensis (Arbea, JI & Jordana, R, 1989:164) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:175
Simonachorutes agrensis from Poland
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.18

Species romeroi (Simón, 1986) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:172, t.t.
Simonachorutes romeroi from Poland
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.1-2

Species weinerae Skarzynski, D, Arbea, JI et Piwnik, A, 2016:178

Genus Pratanurida Rusek, 1973
  Syn.: ... Pratanuria(sic) Rusek, 1973 IOS
Pratanurida sp. from Spain
After Arbea JI, 2013 Fig.8

Pratanurida boerneri from Mexico
1997 © Palacios-Vargas, J.G.

Species boerneri (Schött, 1902) Jordana, R & Arbea, JI in Ramos, MA & al, 1997:345, B

             Syn.: Pseudachorutes börneri(sic) Wahlgren, E, 1906:4 ISS
Species cassagnaui (Rusek, 1973) Kaprus, IJ & Weiner, WM, 2002:283, t.t.
Species foxi Fjellberg, 1985
Species harti Christiansen & Bellinger, 1980:340
Species minuta (Selga, 1966) Arbea, JI & Jordana, R, 1990:189
  Subspecies menorquina Arbea, JI et Jordana, R, 1990:190
Species mucronata (Ellis, WN, 1976:272) Kaprus, IJ & Weiner, WM, 2002:283
Species podolica Kaprus, IJ et Weiner, WM, 2002:283
Pratanurida podolica from Ukraine
After Kaprus, I.J. & Weiner, W.M., 2002 Fig.1

Species tananensis Fjellberg, 1985

Genus Halachorutes Arlé, R, 1967:368
  Syn.: ... Holachorutes(sic) Jacquemart, S et Jacques, J-M, 1980:14 ISS
Species schusteri Arlé, R, 1967:370, t.t.

Genus Pseudachorutella Stach, J, 1949:79
  Syn.: ... Pseudachorutes Tullberg, 1871 partim
  Syn.: Pseudachorutidae Montachorutes Stach, J, 1949:55
  Syn.: ... Notachoridina Cassagnau & Rapoport, 1962 partim
Pseudachorutella gigas from Chile
Ant.3+4 dorsally fused, ventrally separate (*)
2016.09.02 © Invertebrados, C.

Species africana Weiner & Najt, 1991
Species asigillata (Börner, C, 1901:36) Stach, J, 1949:55, t.t.
            Syn.: Pseudachorutes clavatus Börner, C, 1901:36
            Syn.: Pseudachorutes silvaticus Ågren, 1903
Species balcanica Cassagnau, P et Peja, N, 1979:205
Species bescidica Smolis & Skarzynski, 2007
Species boudinoti Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:39
Species castrii (Rapoport, EH & Rubio, IR, 1963:111) Massoud, Z, 1967:170
Notachorudina castrii from Chile
After Rapoport EH & Rubio IR, 1963 Fig.49

Species clavata (Börner, C, 1901:36)
Remark by Dányi, L. & Traser, G. 2008:18: According to da Gama (1964) the species is a junior synonym of Pseudachorutella asigillata (Börner), but it is a valid, separate taxon in our opinion.

Species gigas (Rapoport, EH & Rubio, IR, 1963:112) Massoud, Z, 1967:169
Pseudachorutella gigas from Chile
2016.09.02 © Invertebrados, C.
Pseudachorutella gigas from Chile
Unguis with lateral apical teeth (*)
2016.09.02 © Invertebrados, C.

Species maxima Najt, Thibaud & Weiner, 1990
Species remyi (Denis, 1933) Massoud, Z, 1967:170, t.t.
            Syn.: Pseudachorutes Remyi(sic) Denis, 1933 IOS
             Syn.: Pseudachorutes Remyi(sic) Stach, J, 1949:56,tt ISS
  Subspecies alpina Stach, J, 1949:65
Species stachi Massoud, 1965

Genus Pseudanurida Schött, 1901 key
  Syn.: ... Neachorutes Womersley, 1933
  Syn.: ... Pseudachorutides Becker, 1905
  Syn.: ... Pseudachorutoides(sic) Denis, 1938 ISS
Pseudanurida sp. from Singapore
2011.01.04 © Koh, J.

Species billitonensis Schött, 1901, t.t.
Pseudanurida billitonensis from Japan
After Uchida H, 1962 Fig.A.
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.

Species bogoyawlenskyi (Becker, 1905) Massoud, Z, 1967:173
            Syn.: Pseudachorutides Bogoyawlensky(sic) Becker, 1905 IOS
            Syn.: Pseudachorutides clysmae Jackson, 1927
            Syn.: Pseudachorutoides(lapsus) dollfusi Denis, 1938
             Syn.: Pseudanurida bogoyawlensky(sic) Massoud, Z, 1967:173 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:55 ISS
Species glauerti (Womersley, 1933) Murphy, DH, 1971:56
Species longisetosa Lee, B-H et Kim, J-T, 1994:208
Species sawayana Schuster, 1965
Species yini Murphy, DH, 1971:62

Genus Oudemansia Schött, 1893
Species barnardi (Womersley, 1934) Uchida, H, 1965:87
Species coerulea Schött, 1893, t.t.
            Syn.: Oudemansia subcoerulea Denis, 1948
             Syn.: Oudemansia subcoerula Prabhoo, 1970 ISS
            Syn.: Polyacanthella oceanica Uchida, 1955 si
            Syn.: Oudemansia hirashimai Uchida, H, 1965:86
            Syn.: Oudemansia hirashimai Yosii, R, 1977:146
             Syn.: Oudemansia cærulea(sic) MacGillivray, AD, 1893:318 ISS
             Syn.: Friesea coerula Salmon, 1964 ISS
Species dubia Denis, 1947
Species esakii (Kinoshita, 1932) Yosii, 1958
             Syn.: Oudemansia esaki(sic) Park, K-H et Lee, B-H, 2000:228 ISS
             Syn.: Oudemansia esaki(sic) Park, K-H, Kim, K-G, Park, S-J, Kim, B-J et Lee, B-H, 2000:415 ISS
Species georgia Christiansen & Bellinger, 1980:295
Species petiti Delamare Deboutteville, C et Massoud, Z, 1964:387
Species shotti Denis, 1948
             Syn.: Friesea shötti(sic) Salmon, 1964 ISS
             Syn.: Oudemansia shötti(sic) Massoud, 1967 ISS

Genus Tasmanura Womersley, H, 1937:154
Species evansi Womersley, H, 1937:154, t.t.
Tasmanura evansi from Tasmania
After Womersley H, 1937 Fig.1a
Tasmanura evansi from Tasmania
After Womersley H, 1939 Fig.44M


Genus Ceratrimeria Börner, C, 1906:167
  Syn.: ... Platanurida Carpenter, 1925
  Syn.: ... Tasmanura Womersley, 1937
  Syn.: Pseudachorutidae Zealandmeria Stach, J, 1949:59
  Syn.: Pseudachorutinae Caratrimeria(sic) Yosii, R, 1977:147 ISS
Ceratrimeria sp. from New Zealand
2006 © Minor, M. & Robertson, A.
Ceratrimeria sp. from Tasmania
2012.10.28 © Zosterops, K.
Ceratrimeria sp. from Australia
2015.07.27 © Porch, N.

Species aurea Salmon, JT, 1944:133
Species crassa (Oudemans, JT in Weber, M, 1890:74) Yayuk, 1989:5
Species harrisi Salmon, JT, 1942:258
Ceratrimeria harrisi from New Zealand
After Salmon J.T., 1942 Pl.43 Fig.4
Ceratrimeria harrisi from New Zealand
2015 © TePapa Museum

Species indica (Handschin, E, 1929:231) Massoud, Z, 1967:180
Species longicornis Handschin, 1926
Species lydiae Najt, J et Weiner, WM, 1991:108
Species maxima (Schött, 1901) Börner, C, 1906:167, t.t.
Ceratrimeria maxima from Australia
After Schött H, 1917 Fig.4

Species novaezealandiae (Womersley, H, 1936:318) Womersley, 1937
            Syn.: Pseudachorutes novae-zealandiae(sic) Womersley, H, 1936:318 IOS
             Syn.: Ceratrimeria novae-zealandiae(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1941:302 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1942:254 ISS
             Syn.: Zealandmeria novae-zealandiae(sic) Stach, J, 1949:59,si ISS
             Syn.: Aoteareria novae-zealandiae(sic) Salmon, 1964 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Massoud, Z, 1967:182 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Zealandmeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Pseudachorutes novazealandiae(sic) Greenslade, P, 2015:6 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, P, 2015:6 ISS
Species pulchella Handschin, 1926
Species takaoensis (Kinoshita, 1916) Yosii, R, 1977:147
Species yasumatsui (Uchida, 1950) Yosii, R, 1977:147

Genus PseudachorutinaeGennov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105
Species spnov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105
Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.
Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.


Genus Aethiopellina Delamare Deboutteville, C, 1951:284
Species bimaculata Delamare Deboutteville, C, 1951:284
Species hebrardi Poinsot-Balaguer, 1978
Species proboscina Delamare Deboutteville, C, 1951:284, t.t.
Species sphaeroculata Massoud, 1963

Genus Aethiopella Handschin, 1942
  Syn.: Protachorutidae Ruwenzoriella Salmon, JT, 1964:269
  Syn.: Neanuridae Aethiopiella(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS
Species africana (Delamare Deboutteville, 1945) Massoud, Z, 1967:188
  Subspecies nigra Delamare Deboutteville, 1945
Species ariana Najt, Thibaud & Weiner, 1990
Species basilewskyi (Salmon, 1956) Massoud, Z, 1967:188
Species caraibensis Thibaud & Massoud, 1983
            Syn.: Aethiopella caraïbensis(sic) Thibaud & Massoud, 1983 IOS
             Syn.: Aethiopella caraïbensis(sic) Mari Mutt & Bellinger, 1990 ISS
Species condei Massoud, 1963
Species delamarei Arlé, R, 1960:164
Species flavoantennata (Philiptschenko, 1926 tt) Handschin, 1942
            Syn.: Ceratrimeria flavoantennatus(sic) var. capensis Womersley, 1934
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1934 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1939 ISS
Species guineensis Handschin, 1942
Species handschini (Denis, 1924) Handschin, 1942
Species jeanneli Massoud, Z, 1963:60
Species kuolo Christiansen, K et Bellinger, P, 1992:63
Species littoralis Fernandes, LH et de Mendonça, MC, 2003:1
Species machadoi Massoud, 1963
Species maculata (Delamare Deboutteville, 1945) Massoud, Z, 1967:188
Species mandibulata (Delamare Deboutteville, 1945) Salmon, 1956
Species mangenoti Massoud, 1963
Species pedifalx (Salmon, 1956) Massoud, Z, 1967:190
Species pilarandresae Palacios-Vargas, JG et Montejo-Cruz, M, 2014:1039
Species silvestris Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36
Species tournieri Delamare Deboutteville, 1950
            Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, 1950 IOS
             Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, C, 1951:282 ISS
Species ugandensis Salmon, 1954
Species villiersi Massoud, 1963

Genus Protachorutes Cassagnau, P, 1955:154
Species bicolor (Christiansen & Bellinger, 1980:343) Christiansen, KA in Dindal, DL, 1990:979
Species pyrenaeus Cassagnau, P, 1955:154, t.t.

Genus Arlesiella Delamare Deboutteville, C, 1951:285
Species amazonica Arlé, R, 1966:8
Species camerounensis Massoud, 1963
Species delamarei Massoud, Z, 1963:64
Species monodi (Delamare Deboutteville, 1952) Massoud, 1967:198
Species saprophila Delamare Deboutteville, C, 1951:285, t.t.
Species simulans Massoud, Z, 1963:67

Genus Linnaniemia Philiptschenko, 1926
Species bicolor (Delamare Deboutteville, 1945) Massoud, Z, 1967:202
Species gigas Philiptschenko, 1926:202, t.t.

Genus Neotropiella Handschin, 1942
  Syn.: ... Americotrimeria Stach, 1949
Species sp.1 Bellini, BC et Zeppelini, D, 2004:588
Species arlei Najt, Thibaud & Weiner, 1990
Species barbatae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:43
Species carli (Denis, 1924) Arlé, 1960
            Syn.: Ceratrimeria pentoculata Womersley, 1930
            Syn.: Neotropiella araguaensis Rapoport & Maño, 1969
Neotropiella carli from French Guyana
Pointed mouthcone
After Thibaud J-M & D'Haese C, 2010 Fig.2
2009 © D'Haese, C.
Neotropiella carli from Ecuador
Eyepatch with 5 ocelli; PAO present
2014.01.31 © McClarin, J.
Neotropiella carli from Ecuador
Body segments trilobed
2014.01.31 © McClarin, J.

Species denisi (Arlé, 1939) Stach, 1949
Species digitomucronata Massoud & Thibaud, 1982
Species duranti Díaz & Najt, 1995
Species gordae Díaz & Najt, 1995
Species insularis Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:47
Species macunaimae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:45
Species malkini Arlé, 1981
Species meridionalis (Arlé, 1939) Stach, 1949
Species minima Thibaud & Oliveira, 2010
Species mirabilis (Handschin, 1929) Massoud, 1967:206
Species murphyi Massoud, 1965
Species pedisensilla Najt, Thibaud & Weiner, 1990
Species plurichaetosa Thibaud & Oliveira, 2010
Species quinqueoculata (Denis, 1931) Arlé, 1962
            Syn.: Pseudachorutes 5-oculatus(sic) Denis, 1931 IOS
             Syn.: Americotrimeria 5-oculata(sic) Stach, 1949 ISS
             Syn.: Neotropiella quinque-oculata(sic) Arlé, R, 1966:13 ISS
             Syn.: Neotropiella 5-oculata(sic) Arlé, R, 1966:15 ISS
             Syn.: Neotropiella pentoculata(sic) Massoud, 1967 ISS
Species silvestrii (Denis, 1929) Massoud, 1967:208, t.t.
Species vanderdrifti Massoud, 1963

Genus Grananurida Yosii, 1954
  Syn.: Neanuridae Agranurida(sic) Kim, JT et Lee, B-H, 2000:180 ISS
Remark by Janssens F. & Bernard, E.C. 2007:in litt.: Agranurida(sic) Kim, J.T. & Lee.-H., 2000:180 is an incorrect subsequent spelling of Grananurida.
Species alba (Yosii, 1966) Weiner, WM & Najt, J, 1985:281, i.s.
Species baicalica Rusek, J, 1991:126
Species koreana Lee, B-H et Kim, J-T, 1984:21
Species minuta Kim, JT et Lee, B-H, 2000:180
Species tuberculata Yosii, 1954, t.t.

Genus Najtafrica Barra, J-A, 2002:178
Species riebi (Barra, J-A, 1994:184) Barra, J-A, 2002:178

Genus Stachorutes Dallai, 1973 key
  Syn.: Neanuridae Chihuahuachorutes Palacios-Vargas, JG, 1990:7
Species cabagnerensis Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150
Species cuihuaensis Gao & Yin, 2007
Species dallaii Weiner, WM et Najt, J, 1998:226
             Syn.: Stachorutes dallai(sic) Kovác, L, 1999:135 ISS
             Syn.: Stachorutes dallai(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS
             Syn.: Stachorutes dallai(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS
Species dematteisi Dallai, 1973, t.t.
             Syn.: Stachorutes dematteisae(sic) Palacios-Vargas, 1990 ISS
             Syn.: Stachorutes demattiesi(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Stachorutes dematteisae(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:108 ISS
Species escobarae (Palacios-Vargas, JG, 1990:8) Thibaud, J-M & Palacios-Vargas, JG, 2000:107
Species gracilis Smolis, A et Shvejonkova, JB, 2006:44
Species jizuensis Tamura & Zhao, 1997
Species longirostris Deharveng & Lienhard, 1983
Species maya Thibaud, J-M et Palacios-Vargas, JG, 2000:108
Species najtae Fanciulli, PP, Dallai, R, Frati, F et Carapelli, A, 2017:32
Stachorutes najtae from Italy
After Fanciulli PP & al, 2017 Fig.3A

Species navajellus Fjellberg, 1984
             Syn.: Stachorutes navajelus(sic) Slawska, M, 1996:329 ISS
             Syn.: Stachorutes novajellus(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS
Species ruseki Kovác, L, 1999:135
Species scherae Deharveng & Lienhard, 1983
Species sphagnophilus Slawska, M, 1996:325
             Syn.: Stachorutes sphagnophillus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:156 ISS
Species tatricus Smolis, A et Skarzynski, D, [2002]:407
             Syn.: Stachorutes tatricusa(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS
             Syn.: Stachorutes tatricusb(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS
Species tieni Pomorski, RJ et Smolis, A, 1999:?
Species triocellatus Pomorski, RJ et Smolis, A, 1999:?
             Syn.: Stachorutes triocelatus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS
Species valdeaibarensis Arbea, JI et Jordana, R, 1991:80
             Syn.: Stachorutes valdeibarensis(sic) Slawska, M, 1996:329 ISS

Genus Intermediurida Najt, Thibaud & Weiner, 1990
Species anophthalma Najt, Thibaud & Weiner, 1990

Genus Venezuelida Díaz & Najt, 1995
Species polylepiana Díaz & Najt, 1995

Genus Tijucameria de Mendonça, MC et Fernandes, LH, 2005:699
Species gabrieli de Mendonça & Silveira, 2012
Species mame de Mendonça, MC et Fernandes, LH, 2005:699, t.t.

Genus Platanurida Carpenter, GH, 1925:5
Platanurida sp. from New Zealand
2016.03.18 © Murray, A.
Platanurida sp. from New Zealand
2016.03.18 © Murray, A.

Species lata Carpenter, GH, 1925:5, t.t.
Ceratrimeria lata from New Zealand
After Salmon JT, 1941 Pl.44 Fig.73

Species marplesi (Salmon, JT, 1941:301) Massoud, Z, 1967:212
Ceratrimeria marplesi from New Zealand
After Salmon JT, 1941 Pl.44 Fig.79
Platanurida marplesi from New Zealand
2014 © TePapa Museum
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.

Species marplesoides Massoud, Z, 1967:210
            Syn.: Platanurida marplesoïdes(sic) Massoud, Z, 1967:210 IOS
            Syn.: Ceratrimeria marplesi Salmon, 1942, nec Salmon, 1941

Genus Brasilimeria Stach, 1949
Species anura (Arlé, 1939) Arlé, 1959, t.t.
Species wygodzinskyi (Arlé, R, 1943:110) Arlé, 1962
            Syn.: Ceratrimeria Wygodzinskyi(sic) Arlé, R, 1943:110 IOS

Genus Israelimeria Weiner, WM et Kaprus, IJ, 2005:49
Species poliakovae Weiner, WM et Kaprus, IJ, 2005:50, t.t.
Israelimeria poliakovae from Israel
After Weiner W.M. & Kaprus, I.J, 2005 Fig.1


Genus Arlesia Handschin, 1942
  Syn.: ... Arlésia(sic) Handschin, 1942 IOS
  Syn.: Neanuridae Portachorutes Wray, DL, 1953:147
  Syn.: ... Pseudarlesia Salmon, 1964
  Syn.: Pseudachorutinae Handschinia Massoud, Z, 1967:214
Species albipes (Folsom, 1927) Stach, J, 1949:57, t.t.
            Syn.: Portachorutes mambatus Wray, DL, 1953:147
            Syn.: Arlesia albipes f. decorata Mari Mutt, JA et Bellinger, PF, 1990:41
             Syn.: Arlesia albibes(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:45 ISS
Species arleana de Mendonça, MC et Fernandes, LH, 1999:1196
            Syn.: Arlesia albipes f. decorata Arlé & Rufino, 1976
Species cochabambensis Cassagnau & Rapoport, 1962
Species innominata Salmon, 1964
Species intermedia Fernandes, LH et de Mendonça, MC, 2004:21
Species variabilis Thibaud & Massoud, 1983

Genus Handschinurida Queiroz, GC, 2015:499
  Syn.: Pseudachorutidae Handschinia Stach, J, 1949:56 nec Lallemand, 1935
  Syn.: Collembola Handschinia Ellis, WN et Bellinger, PF, 1973:22 nec Lallemand, 1935
  Syn.: Pseudachorutinae Handschinia Queiroz, GC et de Mendonça MC, 2014:171 nec Lallemand, 1935
Species fluminensis (Arlé, 1939 tt) Queiroz, GC, 2015:499, t.t.
             Syn.: Arlesia flaminensis(sic) Palacios-Vargas, JG in Alvarez-Sánchez, J et Naranjo-García, E, 2003:221,ca ISS
Handschinia fluminensis from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.15

Species proxima (Arlé, 1939) Queiroz, GC, 2015:499
Handschinia proxima from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.19

Species rauli (Queiroz, GC & de Mendonça MC, 2014:172) Queiroz, GC, 2015:499
Handschinia rauli from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.23


Genus Kenyura Salmon, 1954
Species africana (Massoud, Z, 1963:70) Massoud, Z, 1967:230
Species delicata Arlé, R, 1966:4
Species monoculata (Winter, 1963) Mari Mutt & Bellinger, 1990, n.n.
Species multisensa Salmon, 1954, t.t.
Species nicaraguensis Palacios-Vargas & Deharveng, 2010
Species porcula (Arlé, R, 1960:163) Arlé, R, 1966:2
            Syn.: Paranurella porculus(sic) Arlé, R, 1960:163 IOS
             Syn.: Kenyura porculus(sic) Arlé, R, 1966:2 ISS
             Syn.: Kenyura porculus(sic) Massoud, Z, 1967:229 ISS
             Syn.: Kenyura porculus(sic) Arlé, 1981 ISS
             Syn.: Kenyura porculus(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
             Syn.: Kenyura porculus(sic) Culik, MP et Zeppelini, DF, 2003:1124 ISS
Species xinguensis Arlé, R, 1966:3

Genus Furculanurida Massoud, Z, 1967:231 key
Species africana (Massoud, Z, 1963:68) Massoud, 1967:231, t.t.
Species arawakensis Thibaud & Massoud, 1983
Species arlei Thibaud & Massoud, 1980
Species ashrafi (Yosii, 1966) Thibaud, J-M & Palacios-Vargas, JG, 2000:112
Species belemensis Arlé & Rufino, 1976
Species duodecimoculata Thibaud & Massoud, 1980
Species furculata (Salmon, 1956) Massoud, 1967:233
Species goeldiana Arlé & Rufino, 1976
Species grandcolasorum Weiner, WM et Najt, J, 1998:223
Furculanurida grandcolasorum from Tanzania
Bicolour form
After Weiner W.M. & Najt J, 1998 Fig.17

Species guatemalensis Palacios-Vargas, JG et Gao, Y, 2009:55
Furculanurida guatemalensis from Guatemala
After Palacios JG & Gao Y, 2009 Fig.1
             Syn.: Furculanurida guatemaltesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS
             Syn.: Furculanurida guatemalesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS
Species langdoni Bernard, 2007
Species longisensillata Najt, Thibaud & Weiner, 1990
Species nessimiani (Fernandes, LH & de Mendonça, MC, 2003:4) Palacios-Vargas, JG & Gao, Y, 2009:55
             Syn.: Furculanurida nessiamiani(sic) Palacios-Vargas, JG et Gao, Y, 2009:55 ISS
Species perplexa (Salmon, 1956) Massoud, 1967:233
Species septemoculata Palacios-Vargas, JG et Gao, Y, 2009:56
Furculanurida septemoculata from Guatemala
After Palacios JG & Gao Y, 2009 Fig.12

Species tropicalia Queiroz & Fernandes, 2011

Genus Micranurida Börner, 1901
Species agenjoi Simón, 1981
Species anini Christiansen, K et Bellinger, P, 1992:68
Species anophthalmica Stach, J, 1946:26
Species balta Fjellberg, A, 1998:77
Species bescidica Smolis, A et Skarzynski, D, 2004:1
Species candida Cassagnau, P, 1952:310
Species fluminensis Fernandes, LH et de Mendonça, MC, 2004:22
Species forsslundi Gisin, 1949, B
Species furcifera Mills, HB, 1934:27
Species grana Lee & Kim, 1984
Species hygrophila Rusek, 1973
Species intermedia Dallai, 1973
Species japonica Tamura, 1998
Species lanceolata Najt, J et Weiner, WM, 1985:35
Species meridionalis Cassagnau, P, 1952:310
Species montana Lee & Kim, 1984
Species papillosa Axelson, 1902, B
Species porcella Fjellberg, 1985
Species pygmaea Börner, 1901, t.t., B
            Syn.: Micranurida pygmaea f. hasai Kseneman, 1936
            Syn.: Micranurida meridionalis Arbea & Jordana, 1985
Micranuridae pygmaea from Italy
After Dallai R, 1973 Fig.6n
             Syn.: Micranurida pygmea(sic) Slawska, M, 1996:328 ISS
Species retezatica Gruia & Harsia, 1990
Species rostrata Babenko, AB, 2007:364
Species sensillata (Gisin, 1953) Dunger, 1974
Species sp. nov. Palacios-Vargas, 1981:89
Species sp. nov. Palacios-Vargas & Gómez-Anaya, 1993:8
Species spirillifera Hammer, 1953
             Syn.: Micranurida spirilifera(sic) Massoud, Z, 1967:236 ISS
Species tuberculata Lee & Kim, 1984
Species valiana Fjellberg, 1985
Species wladimiri Najt & Rubio, 1978

Genus Meganurida Carpenter, 1935
Species mumfordi Carpenter, 1935, t.t.

Genus Pseudachorudina Stach, 1949
Species alpina Stach, 1949
Species angelieri Cassagnau, P, 1959:76
Species berninii (Dallai, R, 1970:438) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:5

Pseudachorutes brunneus from New Zealand
After Salmon, J.T. 1941 Pl.46 Fig.101.
Species brunnea (Carpenter, GH, 1925:2) Massoud, Z, 1967:243

Species dahli (Schäffer, C, 1898:400) Yoshii, R & Suhardjono, YR, 1992:2
            Syn.: Schöttella(sic) dahlii(sic) Schäffer, C, 1898:400 IOS
            Syn.: Pseudachorudina delamarei Massoud, 1965
             Syn.: Pseudachorutes dahlii(sic) Börner, 1901 ISS
             Syn.: Schöttella(sic) dahlii(sic) Schött, 1901 ISS
             Syn.: Schöttella(sic) dahlii(sic) Stach, 1929 ISS
             Syn.: Pseudachorutes dahlii(sic) Womersley, 1937 ISS
             Syn.: Schoettella dahlii(sic) Salmon, JT, 1964:200 ISS
             Syn.: Pseudachorutes dahlii(sic) Massoud, Z, 1967:165 ISS
             Syn.: Pseudachorutes? dahlii(sic) Yayuk, RS(sic), 1989:4 ISS
             Syn.: Pseudachorudina dahlii(sic) Yoshii, R et Suhardjono, YR, 1992:2 ISS
Species evansi (Womersley, H, 1936:475) Stach, 1949
Species falteronensis (Denis, 1926) Massoud, 1967:243, t.t.
Species ignota Christiansen & Bellinger, 1980:300
Species mabirensis (Philiptschenko, 1926) Salmon, 1954
Species meridionalis (Bonet, 1929) Jordana, R, Arbea, JI, Simón, C & Luciáñez, MJ in Ramos, MA & al, 1997:777
            Syn.: Pseudachorudina bougisi Delamare Deboutteville, 1951
            Syn.: Pseudachorudina bougisi var. catalonica Massoud, 1963
            Syn.: Pseudachorudina patriciae Gough, 1976
            Syn.: Pseudachorudina najtae Simon & Pozo, 1989
Species nepalica Yosii, 1966

Brachystomella osextara from New Zealand
After Salmon, J.T. 1941 Pl.45 Fig.88.
Species osextara (Salmon, JT, 1941:303) Massoud, Z, 1967:243
Species pacifica (Womersley, H, 1936:318) Massoud, 1967:241
Species texensis Christiansen & Bellinger, 1980:301

Genus Cassagnaudina Massoud, 1967
Species coiffaiti (Cassagnau, P, 1955:145) Massoud, Z, 1967:245
            Syn.: Pseudachorudina Coiffaiti(sic) Cassagnau, P, 1955:145 IOS
Species khumbuensis Yosii, 1971
Species perisi Simón, 1981
Species tourratensis (Cassagnau, P, 1964:470)

Genus Cassagnaurida Salmon, 1964
Species dentata (Cassagnau & Rapoport, 1962) Massoud, 1967:246, t.t.

Genus Notachorudina Cassagnau & Rapoport, 1962
  Syn.: Collembola Nothachorudina(sic) Rapoport, EH et Rubio, IR, 1963:111 ISS
Species patagonica Cassagnau & Rapoport, 1962, t.t.

Genus Forsteramea Salmon, JT, 1965:645
Species megacephala (Salmon, JT, 1954:213) Salmon, JT, 1965:645, t.t.
            Syn.: Montachorutes megacephala(sic) Salmon, JT, 1954:213 IOS
Montachorutes megacephala(sic) from New Zealand
After Salmon JT, 1954 Text-Fig.1

             Syn.: Montachorutes? megacephala(sic) Rapoport, EH et Rubio, IR, 1963:113 ISS
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.


Genus Delamarellina Rapoport, EH et Rubio, IR, 1963:113
Species globulosa (Cassagnau & Rapoport, 1962) Massoud, Z, 1967:253
Species guilleni Rapoport, EH et Rubio, IR, 1963:113, t.t.
Delamarellina guilleni from Chile
After Rapoport EH & Rubio IR, 1963 Fig.62

Species ubiquata (Salmon, JT, 1944:133) Massoud, Z, 1967:251

Genus Quatacanthella Salmon, JT, 1945:68

Polyacanthella proprieta from New Zealand
After Salmon, J.T. 1941 Pl.42 Fig.57.
Species proprieta (Salmon, JT, 1941:296) Salmon, JT, 1945:68, t.t.

Genus Anurida Laboulbène, 1865 key  key
  Syn.: Aphoruridæ(sic) Aphoromma MacGillivray, AD, 1893:314
Species alpina Agrell, 1939
Species amorita Folsom, JW, 1902:90
Anurida amorita from Alaska
After Folsom JW, 1902 Pl.IV Fig.3

Species annatshagi Tshelnokov, 1988
Species ariesi Harsia & Gruia, 1992

Anurida ashbyae from the Lesser Antilles
Det. Christiansen, K.
2006 © De Silva, M.
Species ashbyae Christiansen, K et Bellinger, P, 1988:228

Species asiatica Babenko, AB, 2002:374
Species assimilis Yosii, R, 1956:28
  Subspecies persimilis Yosii, R, 1956:28
Species azurea Babenko, AB, 1997:513
Species balatovae Rusek, 1970
Species beringi Fjellberg, 1985
Species bisetosa Bagnall, 1949
Species bondarenkoae Tshelnokov, 1988
Species bonneti Cassagnau, P, 1955:147
            Syn.: Anurida Bonneti(sic) Cassagnau, P, 1955:147 IOS
Species borealis Martynova in Stebaeva, 1976, n.n.
Species callainii Dallai, R et Martinozzi, I, 1980:8
Species caprariensis Dallai, 1969
Species carpatica Babenko, AB, 1998:658
Species clavata Schäffer, 1897
Species confinis Babenko, AB, 1998:651
Species decemoculata Hammer, 1953
Species dentata Christiansen & Bellinger, 1980:326
Species desnuda Yosii, R, 1956:25
Species diabolica Yosii, R, 1956:26
Species dynkengda Babenko, AB, 1998:649
Species frigida Fjellberg, 1973
Species granaria (Nicolet, H, 1847:387) Tullberg, 1869, t.t., B
            Syn.: Anurida bagnalli Stach, J, 1949:14
            Syn.: Anurida comellinii Gisin, 1949
            Syn.: Anurida pseudogranaria Stach, J, 1949:30
Anoura granaria from the UK
After Lubbock J, 1873 Pl.49
Anurida granaria from the UK
2005 © Hopkin, S.P.
Anurida granaria from Sweden
2006 © Hall, K.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria juv. from the UK
2013.01.29 © Murray, A.
Anurida granaria from the UK
Circular post-antennal organ
2013.01.29 © Murray, A.
Anurida granaria from the UK
Granulated cuticula
2013.01.29 © Murray, A.
Anurida granaria from the UK
Antennae and mouthcone
2013.03.23 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from Holland
2014.03.14 © van Duinen, J.
Anurida granaria from Canada
Anal spines absent
2015.08.12 © Murray, A.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
Cuticle coarsely granulate
Abd.6 not bilobed
2015.09.26 © Tony.
Anurida granaria from Tasmania
Postantennal organ circular
Tapering antenna
2015.09.26 © Tony.
Anurida granaria from New Zealand
Mouth: labrum, maxillae heads, hypopharynx, split labium
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Ant.4: apical eversible trilobed sensillar vesicle
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Rudiment of furcula
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Post-antennal organ
2016.02.10 © Koken, M.

Species granulata Agrell, 1943, B
            Syn.: Micranurida sexpunctata Handschin, 1924, nec Axelson, 1902
            Syn.: Hypanurida mirabilis Denis, 1931
            Syn.: Hypanurida flavescens Kseneman, 1934
            Syn.: Micranurida handschini Gisin, 1944
            Syn.: Anurida hexophthalmica Stach, J, 1949:26
Species hammerae Christiansen, KA, 1952:135
            Syn.: Anurida hammeri(sic) Christiansen, KA, 1952:135 IOS
Anurida hammeri(sic) from Alaska
After Christiansen K, 1952 Pl.12 Fig.1-2

            Syn.: Anurida polychaetosa Tshelnokov, 1988
             Syn.: Anurida hammeri(sic) Massoud, Z, 1967:265 ISS
             Syn.: Anurida (Anurida) hammeri(sic) Christiansen, K et Bellinger, P, 1980:330 ISS
             Syn.: Anurida hammeri(sic) Christiansen, KA in Dindal, DL, 1990:978 ISS
             Syn.: Anurida hammeri(sic) Poole, RW in Poole, RW et Gentili, P, 1997:47 ISS
Species interior Fjellberg, 1985
Species iriei Yosii, 1970
Species kaya Babenko, AB, 1998:656
Species kazukoae Tamura & Zhao, 1997
             Syn.: Anurida kazokoae(sic) Yue, Q-Y et Yin, W-Y, 1999:222 ISS
Species kolymensis Tshelnokov, 1988
Species komi Babenko, AB, 1998:653
Species latosensillata Dunger, 1975
Species luciae Fjellberg, A, 1985:359
Species lvivska Babenko, AB, 1998:659
Species mandibulata Tshelnokov, 1988
Species mara Christiansen, K et Bellinger, P, 1988:230
Species marchena Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:161
             Syn.: Anurida marchana(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS
Species maritima (Guérin-Méneville, 1836) Laboulbène, 1865, t.t., B
            Syn.: Podura ambulatoria Ström in Plateau, 1890
            Syn.: Anurida bisetosa Bagnall, 1949
            Syn.: Anurida megalops Bagnall, 1949
Anurida maritima from Germany
Capitulum of maxilla
After Anonymous fig.C

             Syn.: Anurida maritime(sic) D'Haese, CA, 2003:581 ISS
Anurida maritima from France
2003 © De Bruyn, L.
Anurida maritima from the UK
2005 © Hopkin, S.P.
Anurida maritima from the UK
Post-antennal organ and eye with 5 ocelli
2005 © Hopkin, S.P.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Aggregation on tidal pool
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Circular post-antennal organ (PAO) with max 10 vesicles
2007.08.11 © Reed,D.E.
Anurida maritima from the UK
2008.01.06 © Roberts, S.
Anurida maritima from the UK
2008.03.30 © Roberts, S.
Anurida maritima from the UK
2008.06.29 © Ramone, J.
Anurida maritima from the UK
2008.07.27 © Roberts, S.
Anurida maritima from the USA
2009.02.21 © Edmonds, M.
Anurida maritima from France
2009.08.09 © Henderickx, H.
Anurida maritima from France
Egg cluster
2009.08.09 © Henderickx, H.
Anurida maritima from the USA
Aggregation on tidal pool
2013.07.21 © Kynd, F.
Anurida maritima from Portugal
Eyepatch with 5 ocelli
2016.08.06 © Almeida, J.

Species maritima Arbea JI, 2001:39, nec Guérin, 1839, si, s.i.
Species martynovae Fjellberg, 1985
Species nadezhdae Babenko, AB, 1998:654
Species narli Fjellberg, 1985
Species neanuriformis Babenko, AB, 2002:369
Species nuda Yosii, 1956
            Syn.: Anurida granaria Uchida, 1952
            Syn.: Anurida granaria Yosii, R, 1977:147
Species okamotoi Yosii, 1970
Species orientalis Martynova in Stebaeva, 1976, n.n.
Species palustris Babenko, AB, 1997:522
Species papillosa (Axelson, 1902) Fjellberg, 1985
Species papillosoides (Hammer, 1953) Massoud, Z, 1967:258
             Syn.: Anurida papillosoïdes Massoud, Z, 1967:258 ISS
Species parapapillosa Tshelnokov, 1988
Species plurichaetotica Yosii, R, 1966:544
Species polaris (Hammer, 1954) Fjellberg, 1985
Species pusilla da Gama, 1964
Species reducta Fjellberg, 1985
Species riverina Sterzynska, M, Kaprus, I et Ehrnsberger, R, 2006:255
Anurida riverina from Poland
After Sterzynska M, Kaprus I & Ehrnsberger R, 2006 Fig.1

Species sakhensis Babenko, AB, 1997:529
Species setosa Yosii, R, 1956:27
Species similis Fjellberg, 1985
Species speobia Yosii, R, 1954:64
Species stereoodorata Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49
Anurida stereoodorata from Abkhazi
After Jordana R & al, 2012 Fig.11A

             Syn.: Anurida stereoodarata(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49 ISS
Species subarctica Fjellberg, 1985
            Syn.: Anurida tajmyrica Tshelnokov, 1988
Species tamurae Yue, Q-Y et Yin, W-Y, 1999:224
            Syn.: Anurida tamurea(sic) Yue, Q-Y et Yin, W-Y, 1999:224 IOS
Species tatianae Babenko, AB, 1997:530
Species thalassophila (Bagnall, 1939) Stach, J, 1949:14
            Syn.: Anurida granaria Remy, 1928
            Syn.: Anurida remyi Denis, 1947
Species trioculata Kinoshita, 1916
Species tullbergi Schött, 1891:192
            Syn.: Anurida crassicornis Reuter L & Reuter OM, 1879-1880 =?
            Syn.: Anurida ellipsoides Stach, J, 1949:34
             Syn.: Anurida ellipsoïdes(sic) Massoud, Z, 1967:260 ISS
             Syn.: Anurida elipsoides(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS
             Syn.: Anurida ellipsoids(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:260 ISS
            Syn.: Anurida brunsvigiensis Hüther, 1964
             Syn.: Anurida brusvigiensis(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS
             Syn.: Anurida Tullbergi(sic) Schött, H, 1893:91 ISS
Anurida Tullbergii(sic) from Sweden
After Schött H, 1893 Taf.VII Fig.16


             Syn.: Anurida Tullbergi(sic) Schött, H, 1917:6 ISS
             Syn.: Anurida tulbergii(sic) Shaw, P et Benefer, CM, 2015:200 ISS
Anurida tullbergi from Belgium
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Post-antennal organ (PAO) oval
Eyepatch with 5 ocelli
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Ant.3+4 fused
2017.05.20 © Huskens, M.-L.

Species uniformis Gisin, 1953
             Syn.: Anurida uniformes(sic) Babenko, AB, 1998:654 ISS
Species vicaria Yosii, R, 1956:30
Species vontoernei Nosek, 1962
Species vulcanica Fjellberg, A, 1985:357
Species weberi Christiansen & Bellinger, 1980:335
Species wrangelensis Tshelnokov, 1988
Species zanokhae Babenko, AB, 1998:649

Genus Minotaurella Weiner, WM, 1999:373
Species banyulensis (Denis, 1947) Weiner, WM, 1999:376
Species edaphica Kaprus, IJ et Nevo, E, 2003:67
Minotaurella edaphica from Israel
After Kaprus I.J. & Nevo E, 2003 Fig.2a

Species isabellae Weiner, WM, 1999:374, t.t.
             Syn.: Minotaurella isabelle(sic) Kaprus, IJ et Nevo, E, 2003:69 ISS

Genus Anuridella Willem, 1906
  Syn.: ... Anurodes Bagnall, 1949
Anuridella sp. from the UK
2013.08.23 © Fenwick, D.

Species sp. Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:892
Species calcarata Denis, 1925:267
            Syn.: Anuridella immsiana Bagnall, 1939
Species germanica (Gisin, 1951)
Species marina Willem, V, 1906:247, t.t.
Anuridella marina
After Willem V, 1906
            Syn.: Anuridella hintoni Bagnall, 1949

Species submarina Bagnall, 1934

Genus Gastranurida Bagnall, 1949
Species denisi (Bagnall, 1939) Massoud, 1967:268, t.t.
            Syn.: Anurida trioculata Denis, 1922, nec Kinoshita, 1916
            Syn.: Paranura hibernica Bagnall, 1941

Genus Hylaeanura Arlé, R, 1966:5 key
  Syn.: Pseudachorutinae Hylaenura(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
  Syn.: Neanuridae Hyleanura(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS
Species infima (Arlé, R, 1960:163) Arlé, R, 1966:5, t.t.
Species mendoncae Zeppelini, D et Palacios-Vargas, JG, 2013:1402
Species nepalensis (Yosii, 1966) Vázquez, M, Cutz-Pool, LQ & Palacios-Vargas, JG, 1998:370
Species nohbecana Vázquez, M, Cutz-Pool, LQ et Palacios-Vargas, JG, 1998:367

Genus Anurachorutes Kuznetsova & Potapov, 1988
Species martynovae Kuznetsova & Potapov, 1988

Genus Anuritelsa Womersley, H, 1939:315
Species maritima Womersley, H, 1939:316, t.t.

Genus Gamachorutes Cassagnau, 1978
Species verrucosus Cassagnau, 1978

Genus Koreanurina Najt & Weiner, 1992
Species inexpectata Najt & Weiner, 1992
Species szeptyckii Najt & Weiner, 1992

Genus Lanzhotia Rusek, 1985
Species brachycera Rusek, 1985

Genus Paranurida Skarzynski, D et Pomorski, RJ, 1994:363
Species kopetdagi Skarzynski, D et Pomorski, RJ, 1994:364

Genus Philotella Najt, J et Weiner, WM, 1985:29
Species deharvengi Najt, J et Weiner, WM, 1985:30, t.t.
Species miracli Najt, J et Weiner, WM, 1985:32
             Syn.: Philotella miracle(sic) Babenko, AB, 2007:364 ISS
Species obesa Babenko, AB, 2007:362
Species olgae Kniss & Thibaud, 1995
Species sp. Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41
Species tertia Najt, J et Weiner, WM, 1985:34

Genus Rusekella Deharveng, 1982
Species cantabrica Deharveng, 1982
Species cassagnaui Deharveng, 1982
Species flagellata (Arbea, JI et Jordana, R, 1985:64)
Species gamae Arbea, JI et Jordana, R, 1991:98
Species nivalis Deharveng, 1982
Species peyrei (Cassagnau, P, 1955:157)
            Syn.: Micranurida Peyrei(sic) Cassagnau, P, 1955:157 IOS
Species similis Deharveng, 1982
Rusekella similis from France
2006 © Boldsystems.org


Genus Pongeia Najt & Weiner, 2002
Species asturianensis Najt & Weiner, 2002, t.t.

Genus Holacanthella Börner, C, 1906:169 key
  Syn.: Pseudachorutinae Holocanthella(sic) Greenslade, P, 2015:5 ISS
Species brevispinosa (Salmon, JT, 1942:256) Massoud, Z, 1967:222
Ceratrimeria brevispinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.1
Holacanthella brevispinosa from New Zealand
2008 Minor, M. & Robertson, A.© SoilBugs

Species duospinosa (Salmon, JT, 1942:257) Massoud, Z, 1967:222
Ceratrimeria duospinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.9
Holacanthella duospinosa from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Holacanthella duospinosa from New Zealand
2007 © Stevens, M.
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
Ant.1 & 2 with apically pointed digitations
2014.05.26 © Murray, A.
Holacanthella duospinosa from New Zealand
2016.04.03 © Sullivan, J.

Species laterospinosa (Salmon, JT, 1944:132) Massoud, Z, 1967:222
Species paucispinosa (Salmon, JT, 1941:300) Massoud, Z, 1967:222
Ceratrimeria paucispinosa from New Zealand
After Salmon JT, 1941 Pl.44 Fig.81
Holacanthella paucispinosa from New Zealand
2006 Minor, M. & Robertson, A.© SoilBugs
Holacanthella paucispinosa from New Zealand
2006 © Stevens, M.
Holacanthella paucispinosa from New Zealand
After Thibaud J-M & D'Haese C, 2010 Fig.33
2009 © D'Haese, C.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.01.28 © Thorpe, S.E.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2013.08.31 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2013.09.22 © Jewell, T.
Holacanthella paucispinosa from New Zealand
2014.mm.dd © Stevens, M.
Holacanthella paucispinosa from New Zealand
2014.02.09 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2016.05.15 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
2016.06.22 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
Head frontoventral aspect
2016.06.22 © Navest, M.P.

Species spinosa (Lubbock, 1899) Börner, C, 1906:169, t.t.
Ceratrimeria spinosa from New Zealand
After Salmon, J.T., 1941 Pl.44 Fig.77
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
Pointed mouthcone
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
Tubercles
2014.05.15 © Murray, A.


Genus Acanthanura Börner, 1906
  Syn.: ... Anoura(sic) Lubbock, 1899 partim IOS
  Syn.: ... Ceratrimeria Börner, 1906 partim
Acanthanura sp. from Australia
2007.11.19 © Midtgaard, R.

Species sp.1 Garrick, RC, Sands, CJ, Rowell, DM, Hillis, DM et Sunnucks, P, 2006:1
Acanthanura sp. from Australia
2006 © Sunnucks, P.
Acanthanura sp. from Australia
2011.05.30 © Selemba, A.

Species sp.2 Janssens F & Murray A, 2014
Acanthanura "of Henderickx" from Tasmania
2007 © Henderickx, H.
Acanthanura "of Henderickx" from Tasmania
2012.12.29 © Zosterops, K.
Acanthanura "of Henderickx" from Tasmania
4th antennal segment
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
Pointed mouth-cone
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
Post-antennal organ
2014.09.25 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2014.11.09 © Grove, S.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.

Species sp.3 Greenslade P, 2015:il
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. juv. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
Tuberculate cuticula
2014.09.13 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.10.28 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.11.14 © Murray, A.
Det.: Greenslade, P., 2015.06.01

Species dendyi (Lubbock, 1899) Börner, 1906, t.t.
            Syn.: Anoura(sic) Dendyi(sic) Lubbock, 1899 IOS
             Syn.: Neanura Dendyi(sic) Absolon, K, 1901:576 ISS
             Syn.: Acanthanura Dendyi(sic) Börner, 1906 ISS
             Syn.: Anoura(sic) Dendyi(sic) Schött, H, 1917:1 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1939 ISS
             Syn.: Acanthanura Dendyi(sic) Handschin, 1942 tt ISS
Acanthanura dendyi from Tasmania
2010.07.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010.12.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010 © Morris, R.


Genus Megalanura Ellis, WN et Bellinger, PF, 1973:30
  Syn.: ... Meganura Handschin, 1942, nec Szepligéti, 1906
Species tasmaniae (Lubbock, 1899 tt) Ellis, WN & Bellinger, PF, 1973:30
            Syn.: Anoura(sic) tasmania(sic) Lubbock, 1899 tt IOS
             Syn.: Anoura(sic) tasmaniæ(sic) Dendy, A, 1900:98 ISS
             Syn.: Neanura Tasmaniae(sic) Absolon, K, 1901:576 ISS
             Syn.: Anoura(sic) Tasmaniae(sic) Schött, H, 1917:1 ISS
Megalanura tasmaniae from Tasmania
2010 Morris, R.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Ventral aspect
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Composed eye with 8 ocelli; PAO elliptic
2014.09.26 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.


Genus Womersleymeria Stach, J, 1949:64
Womersleymeria sp. juv. from Australia
2015.07.27 © Porch, N.

Species bicornis (Womersley, H, 1940:137) Stach, J, 1949:64, t.t.
Ceratrimeria bicornis from Tasmania
After Womersley H, 1940 Fig.A
Womersleymeria bicornis from Tasmania
2010 Morris, R.
Womersleymeria bicornis from Tasmania
Aberration with fused digitations of abd.5
2014.09.22 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Pointed mouth-cone
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Post-antennal organ (PAO)
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.11.14 © Murray, A.




References