http://www.collembola.org/taxa/pseuinae.htm - Last updated on 2024.08.28 by Frans Janssens
Checklist of the Collembola: Pseudachorutinae

Subfamilia Pseudachorutinae Börner C, 1906:160

Genus Number
of species
Acanthanura6
Aethiopella23
Aethiopellina4
Anurachorutes1
Anurida89
Anuridella5
Anuritelsa1
Arlesia7
Arlesiella6
Brasilimeria4
Cassagnaudina4
Cassagnaurida1
Cephalachorutes16
Ceratrimeria12
Delamarellina2
Forsteramea1
Furculanurida17
Gamachorutes1
Gastranurida1
Grananurida5
Halachorutes2
Handschinurida3
Holacanthella5
Hylaeanura5
Intermediurida1
Israelimeria2
Kenyura8
Koreanurina4
Lanzhotia1
Linnaniemia2
Megalanura1
Meganurida1
Micranurida29
Minotaurella3
Najtafrica1
Neotropiella19
Notachorudina1
Oudemansia10
Paranurida1
Philotella17
Platanurida3
Pongeia1
Pratanurida7
Protachorutes2
Pseudachorudina15
Pseudachorutella16
Pseudachorutes131
PseudachorutinaeGennov1
PseudachorutinaeIncertaeSedis2
Pseudanurida6
Quatacanthella1
Rusekella7
Sernatropiella2
Simonachorutes3
Stachorutes19
Tasmanura1
Tijucameria3
Venezuelida1
Womersleymeria2
Pseudachorutinae545
Genera with number of species

  Syn.: Collembola Lipuridæ(sic) Schött, H, 1891:192
  Syn.: ... Achorutinae Börner, 1907 partim
  Syn.: Hypogastruridae Acharutinae(sic) Womersley, H, 1936:318 ISS
  Syn.: Collembola Neanurini(sic) Becker, E, 1905:72
  Syn.: Collembola Anuridae(sic) Stach, J, 1949:1
  Syn.: ... Aphorommidae Yosii, 1961
  Syn.: Poduromorpha Brachystomellidae Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:14 partim lapsus


  Syn.: Neanuridae Pseudachorutine(sic) Skarzynski, D et Pomorski, RJ, 1994:364 ISS
  Syn.: Neanuridae Psuedachorutinae(sic) Alijani, M, Shayanmehr, M, Skarzynski, D, Piwnik, A et Smolis, A, 2015:268 ISS
Pseudachorutinae from Australia
2017.03.04 © Steve & Alison.
Pseudachorutinae from the USA
2017.10.16 © Johnson, S.T.

  Syn.: Neanuridae Psuedachorutinae(sic) Smolis, A et Pasnik, G, 2020:1 ISS
Pseudachorutinae from Singapore
2021.11.29 © Bay, N.
Pseudachorutinae from Taiwan
Pointed mouthcone
2022.04.09 © Cheng, H.-J.




Genus PseudachorutinaeIncertaeSedis Bellini, BC, Weiner, WM, Queiroz, GC et Paz, RV, 2020:3

Pseudachorutinae from Australia
2023.07.16 © Watts, Br.

Species denisi (Arlé, 1939) Bellini, BC, Weiner, WM, Queiroz, GC & Paz, RV, 2020:3, i.s.

Species mirabilis (Handschin, 1929) Bellini, BC, Weiner, WM, Queiroz, GC & Paz, RV, 2020:3, i.s.


Genus Cephalachorutes Bedos, A et Deharveng, L, 1991:145

Species sp.1 Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178

            Syn.: Cephalachorutes minimus Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178 nr

Species anneae Queiroz, GC et Mendonça, MC, 2016:47

Cephalachorutes anneae from Brazil
After Queiroz GC & de Mendonça MC, 2016 Fig.8A

Species asiaticus Bedos, A et Deharveng, L, 1991:152, t.t.

Species barthae Bedos, A et Deharveng, L, 1991:149

Species caecus Bedos, A et Deharveng, L, 1991:149

Species centurionis Bedos, A et Deharveng, L, 1991:149

Species deharvengi Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41

Species delamarei (Murphy, 1965) Bedos, A & Deharveng, L, 1991:147

Species judithae Shveenkova, YB, 2017:82

            Syn.: Cephalachorutes sp.n. Shveenkova, 2011
Cephalachorutes judithae from Vietnam
After Shveenkova YB, 2017 Fig.4A



Species masi Thibaud, J-M, 2008:507

Remark by Janssens, F. 2009.02.06: Given the name is referring to the surname "Masy", strictly spoken the correct latinisation "of Masy" is "masyi".

Species microphthalmus (Barra, 1969) Bedos, A & Deharveng, L, 1991:147

Species minimus (Massoud, 1963) Bedos, A & Deharveng, L, 1991:147

Species murphyi Bedos, A et Deharveng, L, 1991:151

Species nakaoi (Yosii, 1966) Bedos, A & Deharveng, L, 1991:152

Species pestilentiae Bedos, A et Deharveng, L, 1991:151

Species pillaii (Prabhoo, 1971) Bedos, A & Deharveng, L, 1991:147


Genus Pseudachorutes Tullberg, 1871 key

  Syn.: Poduridæ(sic) Brachysius MacGillivray, AD, 1893:317
  Syn.: Poduridæ(sic) Gnathocephalus MacGillivray, AD, 1893:318
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Sphragiphora Houlbert, 1924
  Syn.: ... Sphagiphorura(sic) Pomorski, RJ et Skarzynski, D, 1992:57 ISS
  Syn.: ... Logacanura Wray, 1958
  Syn.: ... Metachorudina Massoud, 1963
  Syn.: ... Neoafricella Salmon, 1964


Pseudachorutes sp. from Germany
Mandibula
After Anonymous Fig.B

  Syn.: Collembola Pseudachotures(sic) Verlinden, 1977 ISS
Pseudachorutes sp. from the USA
2010.01.17 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Ocular area
2010.01.18 © Ten Eyck, C.
Pseudachorutes sp. from the USA
Furcula
2010.02.12 © Ten Eyck, C.
Pseudachorutes sp. from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutes sp. from Japan
2012.04.18 © Miyazaki, Y.
Pseudachorutes sp. from the USA
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Pointed buccal cone
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from the USA
Furcula
2012.09.28 © Kouri, J.D.
Pseudachorutes sp. from Portugal
2012.10.10 © Murray, A.
Pseudachorutes sp. from New Zealand
2014.mm.dd © Murray, A.
Pseudachorutes sp. from the USA
2014.05.22 © Murray, T.
Pseudachorutes sp. from Holland
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Holland
Pointed mouthcone
2014.10.27 © van Duinen, J.
Pseudachorutes sp. from Germany
Large round post-antennal organ (PAO)
2015.02.20 © Mattew, M.
Pseudachorutes sp. from the UK
2015.02.26 © Murray, A.
Pseudachorutes sp. from the UK
Pointed mouth-cone
2015.02.26 © Murray, A.
Pseudachorutes sp. from France
2016.03.03 © Garcelon, P.
Pseudachorutes sp. from Australia
2016.04.25 © Murray, A.

  Syn.: Neanuridae Pseudahorutes(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:241 ISS
Pseudachorutes sp. from France
2018.01.06 © Picard, J.
Pseudachorutes sp. from France
Unguiculus reduced (*)
2018.01.06 © Picard, J.
Pseudachorutes sp. from France
Anal spines absent (*)
2018.01.06 © Picard, J.
Pseudachorutes sp. from the USA
2018.03.20 © Montgomery, G.
Pseudachorutes sp. from Tenerife
2018.03.30 © Huskens, M.-L.
Pseudachorutes sp. from Tenerife
Large ring-shaped PAO with many vesicles
2018.03.30 © Huskens, M.-L.
Pseudachorutes sp. from the USA
2018.12.09 © Pearson, R.R.
Pseudachorutes sp. from the USA
Pointed mouthcone (*)
2018.12.09 © Pearson, R.R.
Pseudachorutes sp. from the USA
Post-antennal organ roundish (PAO)
2018.12.09 © Pearson, R.R.
Pseudachorutes sp. juv. from the USA
2018.12.14 © Pearson, R.R.
Pseudachorutes sp. from the UK
Large cauliform post-antennal organ (PAO)
2019.02.25 © Smart, J.
Pseudachorutes sp. from Belgium
Furcula with broad manubrium and short dentes
2019.05.25 © Huskens, M.-L.
Pseudachorutes sp. subadult from the USA
2021.01.09 © Parker, H.S.
Pseudachorutes sp. subadult from the USA
Moulting
2021.01.09 © Parker, H.S.
Pseudachorutes sp. from the USA
With phoretic Nematoda
2021.01.19 © Shahan, T.
Pseudachorutes sp. from the USA
With phoretic Nematoda
2021.01.19 © Shahan, T.
Pseudachorutes sp. from Belgium
2021.02.23 © Huskens, M.-L.
Pseudachorutes sp. from Belgium
Pointed mouthcone
Eye with 8 ocelli
2021.02.23 © Huskens, M.-L.
Pseudachorutes sp. from Hungary
2021.03.07 © Ferenc, S.
Pseudachorutes sp. from Tasmania
2021.07.05 © Ellingsen, K.
Pseudachorutes sp. from Tasmania
Intersegmental tergite
2021.07.05 © Ellingsen, K.
Pseudachorutes sp. from Tasmania
Eye with 8 ocelli
2021.07.05 © Ellingsen, K.
Pseudachorutes sp. from the USA
Pointed mouthcone (*)
Post-antennal organ round (PAO)
2022.01.12 © Parker, H.S.
Pseudachorutes sp. from Australia
Ventral aspect
2023.06.08 © Watts, Br.
Pseudachorutes sp. from Australia
Feeding from plasmodium
Mouthcone distinctly pointed
2023.06.11 © Watts, Br.
Pseudachorutes sp. from Australia
Yellow colour form
2023.07.02 © Watts, Br.
Pseudachorutes sp. from Australia
Blue colour form
2023.07.14 © Watts, Br.
Pseudachorutes sp. from Australia
White colour form
Eyes lacking
2023.07.16 © Watts, Br.
Pseudachorutes sp. from the USA
"Nudging" behaviour
2023.07.24 © Watts, Br.
Pseudachorutes sp. from Australia
Ant.3+4 fused
2023.07.25 © Watts, Br.
Pseudachorutes sp. from Australia
Pink colour form
2023.08.25 © Watts, Br.
Pseudachorutes sp. from Australia
2023.08.25 © Watts, Br.
Pseudachorutes sp. from Australia
Eye with 8 ocelli
2023.08.25 © Watts, Br.
Pseudachorutes sp. from Thailand
2023.11.28 © van der Ende, T.
Pseudachorutes sp. from the USA
Post-antennal organ with many lobes
2024.07.13 © Critchfield, E.





Species sp.1 Janssens F sensu Denis, 1924

            Syn.: Pseudachorutes sp.A Denis, J-R, 1924:223

Species sp.2 Janssens F sensu Denis, 1924

            Syn.: Pseudachorutes sp.B Denis, J-R, 1924:223

Species sp.3 Janssens F sensu Delamare Deboutteville, 1953

            Syn.: Hypogastrura sp.n. Delamare Deboutteville, C, 1953:823

Species sp.4 Janssens F sensu Palacios-Vargas & Gómez-Anaya, 1995

            Syn.: Pseudachorutes sp.3 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8

Species sp.5 Janssens F sensu Palacios-Vargas & Gómez-Anaya, 1995

            Syn.: Pseudachorutes sp.4 Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8

Species sp.6 Janssens F sensu Christiansen & Bellinger, 1992

            Syn.: Pseudachorutes sp.A Christiansen, K et Bellinger, P, 1980:297bis

Species sp.7 Janssens F sensu Christiansen & Bellinger, 1992

            Syn.: Pseudachorutes sp.C Christiansen, K et Bellinger, P, 1980:297bis

Species sp.8 Janssens F sensu Christiansen & Bellinger, 1992

            Syn.: Pseudachorutes sp.D Christiansen, K et Bellinger, P, 1980:297bis

Species sp.9 Janssens F sensu Soto-Adames & Taylor, 2013

            Syn.: Pseudachorutes sp.undescr. Soto-Adames, FN et Taylor, SJ, 2013:153

Species sp.10 Janssens F sensu Queiroz & Zeppelini, 2017

            Syn.: Pseudachorutes sp. Queiroz, GC et Zeppelini, D, 2017:132

Species sp.11 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Pseudachorutes sp.nov.1 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:9

Species sp.12 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Pseudachorutes sp.nov.2 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:9

Species aleksandrae Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species algidensis Carpenter, GH, 1925:4

             Syn.: Pseudachorutes aldgidenssi(sic) Womersley, H, 1936:327 ISS

Species alluaudi Delamare Deboutteville, 1945

             Syn.: Pseudachorutes allauaudi(sic) Massoud, 1967 ISS

Species americanus Stach, 1949

            Syn.: Pseudachorutes boerneri Nearctic records
            Syn.: Pseudachorutes corticolus Mills, 1934
            Syn.: Pseudachorutes corticicolus f. globulisetosa Gisin, 1944, Nearctic records

Species amucronatus Díaz & Najt, 1995

Species andrei Weiner, WM et Najt, J, 1985:260

            Syn.: Pseudachorutes sibiricus Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351 =?


Species anomalus Imms, 1912

Species aphysus Christiansen, K et Bellinger, P, 1980:304

Pseudachorutes aphysus from the USA
Post-antennal organ with 16 vesicles
2023.01.03 © Wang, G.
Pseudachorutes aphysus from the USA
Mucro with long curved apex and basal swelling
2023.01.03 © Wang, G.
Pseudachorutes aphysus from the USA
Third foot
With reduced unguiculus and inner tooth (*)
2023.01.03 © Wang, G.
Pseudachorutes aphysus from the USA
2023.01.21 © Wang, G.
Pseudachorutes aphysus from the USA
2023.01.21 © Wang, G.
Pseudachorutes aphysus from the USA
With distinctly visible post-antennal organ
2023.12.16 © Wang, G.
Pseudachorutes aphysus from the USA
Post-antennal organ with 16 vesicles
2023.12.16 © Wang, G.
Pseudachorutes aphysus from the USA
Mucro with basal swelling
2023.12.16 © Wang, G.
Pseudachorutes aphysus from the USA
Dorsal setae blunt
2023.12.16 © Wang, G.
Pseudachorutes aphysus from the USA
Mandibula apex
2023.12.16 © Wang, G.
Pseudachorutes aphysus from the USA
Maxilla apex
2023.12.16 © Wang, G.

Species armatus Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species aureofasciatus (Mac Gillivray, 1893) Mills, HB, 1934:23

            Syn.: Pseudachorutes aureofasciatus f. millsi Maynard, EA, 1951:58
Pseudachorutes aureofasciatus from the USA
Non-banded juvenile
2007.02.22 © Bernard, E.C.

            Syn.: Pseudachorutes (Pseudachorutes) bifidus Christiansen & Bellinger, 1980



Pseudachorutes aureofasciatus
1980 © Norton, R.A. in Dindal.
Pseudachorutes aureofasciatus from the USA
Banded juvenile
2007.02.22 © Bernard, E.C.
Pseudachorutes aureofasciatus from the USA
"Butts up colony"
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
Badly smelling defensive secretion
2009.09.13 © Hodge, K.
Pseudachorutes aureofasciatus from the USA
2012.05.01 © Murray, T.
Pseudachorutes aureofasciatus from the USA
2018.05.08 © Caterino, M.
Pseudachorutes aureofasciatus from the USA
2018.06.04 © Caterino, M.

Species auricularius Cassagnau & Rapoport, 1962

Species beta Najt, J et Weiner, WM, 1991:106

Species bifasciatus Oliveira & Deharveng, 1994

Species bifidus Christiansen & Bellinger, 1980:307

Species bobeitio Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32

Species boerneri Schött, 1902, B

            Syn.: Pseudachorutes corticicolus Mills, 1934
            Syn.: Pseudachorutes forsslundi Agrell, 1943
            Syn.: Pseudachorutes corticicola f. globulisetosa Gisin, 1944
            Syn.: Pseudachorutes boerneri ssp. americanus Stach, 1949


             Syn.: Pseudachorutes börneri(sic) Wahlgren, E, 1906:4 ISS
Pratanurida boerneri from Mexico
1997 © Palacios-Vargas, J.G.



Species cacaoiensis Najt, Thibaud & Weiner, 1990

Species caerulescens Schött, H, 1893:24

            Syn.: Pseudachorutes cærulescens(sic) Schött, H, 1893:24 IOS
             Syn.: Pseudachorutes coerulescens(sic) Wahlgren, E, 1906:4 ISS



Species caramel Kaprus, IJ, Weiner, WM et Nevo, E, [2005]:263

Pseudachorutes caramel from Israel
After Kaprus, I.J., Weiner, W.M. & Nevo, E., [2005] Fig.1

Species chazeaui Najt, J et Weiner, WM, 1991:100

Species cheni Shi, S, Jiang, J et Pan, Z-X, 2008:57

Pseudachorutes cheni from China
After Shi S et al., 2008 Fig.1-2

Species columbicus Rusek, J, 1991:120

Species complexus (MacGillivray, AD, 1893:318) Folsom, 1916

Pseudachorutes complexus from the USA
2021.11.03 © Nelson, D.
Pseudachorutes complexus from the USA
2021.11.03 © Nelson, D.
Pseudachorutes complexus from the USA
Mouthcone long elongate
2021.11.03 © Nelson, D.
Pseudachorutes complexus from the USA
Anal spines absent
2021.11.03 © Nelson, D.
Pseudachorutes complexus from the USA
Dens 1.5 times as long as mucro
2021.11.03 © Nelson, D.

Species concinnus Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species conicus Lee & Kim, 1984

Species conspicuatus Salmon, JT, 1944:137

            Syn.: Pseudachorutes conspicuatus ssp. principalis Massoud, Z, 1967:156


Pseudachorutes conspicuatus juv. from New Zealand
2014.02.dd © Murray, A.
Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.
Pseudachorutes conspicuatus from New Zealand
2014.03.03 © Murray, A.

  Subspecies flavus Salmon, JT, 1944:139

  Subspecies lineatus Salmon, JT, 1944:138

Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2009 © Te Papa Museum
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from Tasmania
2014.03.09 © Murray, A.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
2016.05.17 © McQuillan, Br.
Pseudachorutes conspicuatus ssp. lineatus from New Zealand
About 5mm
2020.08.26 © Maurice.

  Subspecies maximus Salmon, JT, 1944:138

Pseudachorutes conspicuatus ssp. maximus from New Zealand
2020.08.18 © Maurice.
Pseudachorutes conspicuatus ssp. maximus from New Zealand
About 5mm
2020.08.18 © Maurice.

Species corticicolus (Schäffer, 1896) Krausbauer, 1898

             Syn.: Pseudachorutes börneri(sic) Schött, 1902 ISS
            Syn.: Pseudachorutes corticicolus f. globulisetosa Gisin, 1944 partim


             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1898 ISS
             Syn.: Pseudachorutes corticicola(sic) Krausbauer, 1901 ISS
             Syn.: Pseudachorutes corticicola(sic) Handschin, E in Dahl, F, 1929:34 ISS
             Syn.: Pseudachorutes corticola(sic) Kloet, GS et Hincks, WD, 1945:4 ISS
             Syn.: Pseudachorutes (Pseudachorutes) corticicola(sic) Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:83 ISS
             Syn.: Pseudachorutes corticolus(sic) Snider, RJ, 1967:192 ISS
             Syn.: Pseudachorutes corticicola(sic) Loksa, I, 1978:55 ISS
             Syn.: Pseudachorutes corticicollus(sic) Kuznetsova, NA, 1984:257 ISS
             Syn.: Pseudachorutes corticolus(sic) Palacios-Vargas, JG, 1990:19 ISS
             Syn.: Pseudachorutes corticolus(sic) Palacios-Vargas, JG, 1990:19 nr ISS
             Syn.: Pseudachorutes corticolus(sic) Szeptycki, A et Weiner, WM in Razowski J, 1990:20 ISS
             Syn.: Pseudachorutes corticolus(sic) Barra, J-A, 1993:71 ISS
             Syn.: Pseudachorutes corticicola(sic) Farkas, J, 1995:46 ISS
             Syn.: Pseudachorutes corticolus(sic) Kopeszki, H et Meyer, E, 1996:224 ISS
             Syn.: Pseudachorutes corticicola(sic) Arbea, JI et Jordana, R in Ramos, MA et al, 1997:323 ISS
Pseudachorutes corticicolus from France
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Pointed mouthcone
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Ant.3+4 fused
2018.06.16 © Tiky, S.
Pseudachorutes corticicolus from France
Abd.4 chaetotaxy
2018.06.16 © Tiky, S.



Species crassus da Gama, 1964

Species curtus Christiansen & Bellinger, 1980:232

Species dalensi Weiner, WM et Najt, J, 1985:258

Species decussus (Salmon, JT, 1941:298)

Micranurida decussa from New Zealand
After Salmon JT, 1941 Pl.43 Fig.72

Species deficiens Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:558

Pseudachorutes deficiens from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.1A,E

Species deserticus Palacios-Vargas, 1990:11

Species difficilis Denis, 1931

Species dilatatus (MacGillivray, AD, 1893:317) Christiansen & Bellinger, 1980

Species dubius Krausbauer, 1898

            Syn.: Pseudachorutes lapponicus Ågren, H, 1904:8
            Syn.: Schöttella(sic) poppei Schäffer, 1897


             Syn.: Pseudahorutes(sic) dibius(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:241 ISS
             Syn.: Pseudachorutes dibius(sic) Özata, MA, Sevgili, H et Kaprus, IJ, 2017:242 ISS

Species galapagoensis Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:157

Species gilvus Oliveira & Deharveng, 1995

Species geronensis (Massoud, 1963) Massoud, 1967:158

Species gilvus Oliveira & Deharveng, 1994

Species guyanensis Najt, Thibaud & Weiner, 1990

Species herberti Arlé & Rufino, 1976

Species hitakamiensis Tamura, 2001

Species hodeberti Thibaud, J-M, 2008:508

Pseudachorutes hodeberti from Madagascar
After Cipola NG in Cipola NG & al, 2022 Fig.9.17.C

Species hyrcanicus Shayanmehr, M, Kaprus, IJ, Ghobari, H, Yahyapour, E, Samani, KM, Lafooraki, EY et Sepanlou, MG, 2022:?

Species indiana Christiansen, K et Bellinger, P, 1980:312

Species innominatus Denis, 1931

Species insularis Yosii, R, 1965:8

Pseudachorutes-insularis from Japan
Modified after Yosii R, 1965 Fig.5A

Species isawaensis Tamura, 2001

Species janstachi Kaprus, IJ et Weiner, WM, 2009:9

Pseudachorutes janstachi from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.32

Species japonicus Kinoshita, 1916

Species javanicus (Handschin, 1920, i.s.) Handschin, 1920

Species jeonjuensis Lee & Kim, 1984

Species jianxiucheni Gao, Yin & Palacios-Vargas, 2008

Species kangchenjungae Yosii, 1966

Species kurdistanicus Shayanmehr, M, Kaprus, IJ, Ghobari, H, Yahyapour, E, Samani, KM, Lafooraki, EY et Sepanlou, MG, 2022:?

Species labiatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:565

Pseudachorutes labiatus from Algeria
After Zoughailech A, Hamra-Kroua S & Deharveng L, 2016 Fig.5A

Species laricis Arbea, JI et Jordana, R, 1989:161

            Syn.: Pseudachorutes parvulus Arbea & Jordana, 1985 partim


Species legrisi Thibaud & Massoud, 1983

Species libanensis (Cassagnau, P et Delamare Deboutteville, C, 1955:372)

Species libanensis Ellis, WN, 1976:269

Remark by Kaprus, I.J., Weiner, W.M. & Nevo, E. [2005]:263: P. libanensis was described from Lebanon as Aetiopella(sic) libanensis by CASSAGNAU and DELAMARE (1955) and subsequently redescribed by ELLIS (1976) on the basis of the material from Central Crete. The comparison of both descriptions shows, that they concern two separate species.

Species lishanensis Gao, Yin & Palacios-Vargas, 2008

Species longisetis Yosii, 1961

            Syn.: Pseudachorutes conicus Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351 =?


             Syn.: Pseudachorutes longisetus(sic) Cassagnau, P, 1974:309 ISS

Species lunatus Folsom, 1916

            Syn.: Pseudachorutes (Pseudachorudina) ignotus Christiansen, K et Bellinger, P, 1980:301 =?


Species macrochaetus Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:158

Species massoudi Arlé, R, 1966:7

Species medjugorjensis Barra, 1993:69

Species minimus Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species minor Peja, 1985

Species morulifer Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species multidentatus Najt, Thibaud & Weiner, 1990

Species murphyi Massoud, 1965

             Syn.: Pseudachorutes murphy(sic) Peck, SB et Jacquemart, J, 2011:5,cf ISS

Species namphoensis Weiner, WM et Najt, J, 1985:262

Species nica Palacios-Vargas, JG et Mejía-Madrid, HH, 2012:64

Species niloticus Wahlgren, E, 1906:3

Species octosensillatus Zoughailech, A, Hamra-Kroua, S et Deharveng, L, 2016:561

Species orghidani Massoud, Z et Gruia, M, 1973:329

Species orientalis Loring, 1984

Species ouatilouensis Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:32

Species palmiensis Börner, 1903

            Syn.: Pseudachorutes palmiensis f. secunda(sic) Denis, J-R, 1947:1 IOS
             Syn.: Pseudachorutes palmiensis f. secundo(sic) Gisin, H, 1960:86 ISS
             Syn.: Pseudachorutes palmiensis f. secundo(sic) Massoud, Z, 1967:162 ISS
            Syn.: Pseudachorudina conjungens Stach, 1949
            Syn.: Pseudachorutes cantabricus Simón & Pozo, 1979


             Syn.: Pseudachorutes palmatus(sic) Wahlgren, E, 1906:4 lapsus ISS
Pseudachorutes palmiensis from France
2006 © Boldsystems.org
Pseudachorutes palmiensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.42
Pseudachorutes palmiensis from Spain
2016.04.24 © Tizon, J.

             Syn.: Pseudachorutes palminensis(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:3 ISS
             Syn.: Pseudachorutes palminensis(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:16 ISS
Pseudachorutes palmiensis from Portugal
After Arbea JI & Almeida J, 2022 Fig.10





Species parvulus Börner, C, 1901:701, B

            Syn.: Pseudachorutes parvulus f. infuscata Yosii, 1954
            Syn.: Pseudachorutes corticicolus Selga, 1971
            Syn.: Pseudachorutes subcrassus Selga, 1971, partim Arbea & Jordana in Ramos, 1997


Pseudachorutes parvulus from Spain
Modified after Arbea JI & Jordana R, 1997 Fig.144A
Pseudachorutes parvulus from Spain
Axis (G-A) of eyepatch directed to the side
Modified after Arbea JI & Jordana R, 1997 Fig.144A
Pseudachorutes parvulus from Spain
In state of anhydrobiosis
Arbea JI & Blasco-Zumeta J, 2001 Fig.2
Pseudachorutes parvulus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.52
Pseudachorutes parvulus from Belgium
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Post-antennal organ (PAO) with 6 vesicles
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus ♂ from Belgium
Genital plate with star-shaped orifice
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Axis (G-A) of eyepatch directed to the side
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Post-antennal organ (PAO) with 6 vesicles
2019.09.01 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Pointed mouthcone (*)
2019.09.14 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Furcula with short dentes and mucrones
2019.09.14 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
8 ocelli (A-H); post-antennal organ (PAO)
2020.02.26 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Preparing for a jump
Anal sacs exposed
2020.02.26 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Anal sacs exposed
2020.02.26 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
2020.07.11 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
2020.07.11 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Tuberculate cuticula
2020.07.11 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Ant.3+4 dorsally fused
2020.07.11 © Huskens, M.-L.
Pseudachorutes parvulus from Belgium
Ant.3 sensory organ (A3O)
Distinctly pointed mouthcone (*)
2020.07.11 © Huskens, M.-L.

Species pasurus (Wray, 1958) Christiansen & Bellinger, 1980

Species pauliani Delamare Deboutteville, C in Paulian, R, 1952:63

            Syn.: Pseudachorutes Pauliani(sic) Delamare Deboutteville, C in Paulian, R, 1952:63 IOS
Pseudachorutes Pauliani(sic) from Ivory Coast
After Delamare Deboutteville C, 1952 Fig.5



Species periyarensis Prabhoo, 1971

Species plurichaetosus Arbea, JI et Jordana, R, 1991:78

Species poahi Christiansen, K et Bellinger, P, 1992:65

Species polychaetosus Gao, Y et Palacios-Vargas, J, 2008:54

Species ponmudiensis (Prabhoo, 1971, i.s.) Prabhoo, 1971

Species pratensis Rusek, 1973

Pseudachorutes pratensis from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.65

Species prosimplex Weiner, WM et Najt, J, 1985:256

Species pseudoniloticus Massoud, 1963

Species puniceus Salmon, JT, 1944:139

Species reductus Thibaud & Massoud, 1983

Species rhaeticus (Carl, J, 1899:298) Handschin, E, 1924:96

            Syn.: Schöttella(sic) rhätica(sic) Carl, J, 1899:298 IOS
            Syn.: Schöttella(sic) rhætica(sic) Carl, J, 1899:298 IOS

Species rugatus Wray, 1952

Species saxatilis Macnamara, 1920

Species scythicus Kaprus, IJ et Weiner, WM, 2009:3

Pseudachorutes scythicus from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.1

Species sexsetosus Denis, 1934

Species shiragamiensis Hisamatsu & Tamura, 1998

Species sibiricus Reuter, 1891

Species sibiricus (Rusek J, 1991:123) Rusek, J, 1991:123

Species simplex Maynard, EA, 1951:59

Pseudachorutes simplex from the USA
2006 © Bernard, E.C.
Pseudachorutes simplex from the USA
2007.02.22 © Bernard, E.C.

Species solaris da Silveira, TC et de Mendonça, MC, 2018:2

Pseudachorutes solaris from Brazil
After da Silveira TC & de Mendonça MC, 2018 Fig.1i

Species subabdominalis Steiner, 1958

Species subcrassoides Mills, HB, 1934:24

             Syn.: Pseudachorutes subcrassoïdes Massoud, 1967 ISS

Species subcrassus Tullberg, 1871, t.t., B

            Syn.: Pseudachorutes tuberculatus Schött, 1902


Pseudachorutes subcrassus from Russia
After Scherbakov AM, 1898 Table I Fig.4
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from the Azores
2009.09.15 © E.D.E.N.
Pseudachorutes subcrassus from Holland
2015.02.21 © Baas, A.H.

Species subcrassus Schäffer, 1897, nec Tullberg, 1871

Species tamajonensis Simón, 1986

Species tasmaniensis Womersley, H, 1936:475

Species tavignani Poinsot-Balaguer, 1978

Species tillieri Najt, J et Weiner, WM, 1991:104

Species unidentatus Salmon, 1956, i.s.

Species uniformis Thibaud & Massoud, 1983

Species univesicatus Weiner & Najt, 1991

             Syn.: Pseudachorutes universicatus(sic) Greenslade, P, 1997-:www0312 ISS

Species variabilis Babenko, A, Kuznetsova, N, Nakamori, T et Shveenkova, Y, 2021:351

Species vasylii Kaprus, IJ et Weiner, WM, 2009:7

Pseudachorutes vasylii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.22

Species vitalii Kaprus, IJ et Weiner, WM, 2009:5

Pseudachorutes vitalii from Ukraine
After Kaprus IJ & Weiner WM, 2009 Fig.11

Species wandae Gao, Yin & Palacios-Vargas, 2008

Species yoshiii Najt, J et Weiner, WM, 1991:102

            Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM, 1991:102 IOS
             Syn.: Pseudachorutes yoshii(sic) Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36 ISS
             Syn.: Pseudachorutes yoshii(sic) Gao, Y et Palacios-Vargas, J, 2008:55 ISS
             Syn.: Pseudachorutes yoshii(sic) da Silveira, TC et de Mendonça, MC, 2018:4 ISS




Genus Simonachorutes Skarzynski, D, Arbea, JI et Piwnik, A, 2016:172

Species agrensis (Arbea, JI & Jordana, R, 1989:164) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:175

Simonachorutes agrensis from Spain
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.18

Species romeroi (Simón, 1986) Skarzynski, D, Arbea, JI & Piwnik, A, 2016:172, t.t.

Simonachorutes romeroi from Spain
After Skarzynski D, Arbea JI & Piwnik A, 2016 Fig.1-2

Species weinerae Skarzynski, D, Arbea, JI et Piwnik, A, 2016:178


Genus Pratanurida Rusek, 1973

  Syn.: ... Pratanuria(sic) Rusek, 1973 IOS
Pratanurida sp. from Spain
After Arbea JI, 2013 Fig.8



Species cassagnaui (Rusek, 1973) Kaprus, IJ & Weiner, WM, 2002:283, t.t.

Species foxi Fjellberg, 1985

Species harti Christiansen & Bellinger, 1980:340

Species minuta (Selga, 1966) Arbea, JI & Jordana, R, 1990:189

  Subspecies menorquina Arbea, JI et Jordana, R, 1990:190

Species mucronata (Ellis, WN, 1976:272) Kaprus, IJ & Weiner, WM, 2002:283

Species podolica Kaprus, IJ et Weiner, WM, 2002:283

Pratanurida podolica from Ukraine
After Kaprus, I.J. & Weiner, W.M., 2002 Fig.1

Species tananensis Fjellberg, 1985


Genus Pseudachorutella Stach, J, 1949:79 key

  Syn.: ... Pseudachorutes Tullberg, 1871 partim
  Syn.: Pseudachorutidae Montachorutes Stach, J, 1949:55
  Syn.: ... Notachoridina Cassagnau & Rapoport, 1962 partim


Pseudachorutella gigas from Chile
Ant.3+4 dorsally fused, ventrally separate (*)
2016.09.02 © Invertebrados, C.

Species africana Weiner & Najt, 1991

Species asigillata (Börner, C, 1901:36) Stach, J, 1949:55, t.t.

            Syn.: Pseudachorutes clavatus Börner, C, 1901:36
            Syn.: Pseudachorutes silvaticus Ågren, 1903


             Syn.: Pseudachorutella assigilata(sic) Jucevica, E, Grina, V et Melecis, V, 2023:258 ISS

Species balcanica Cassagnau, P et Peja, N, 1979:205

Species bescidica Smolis & Skarzynski, 2007

Species boudinoti Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:39

Species castrii (Rapoport, EH & Rubio, IR, 1963:111) Massoud, Z, 1967:170

Notachorudina castrii from Chile
After Rapoport EH & Rubio IR, 1963 Fig.49

Species circassiana Babenko, A et Kuznetsova, N, 2024:189

Pseudachorutella circassiana from Russia
After Babenko A & Kuznetsova N, 2024 Fig.1-2

Species clavata (Börner, C, 1901:36)

Remark by Dányi, L. & Traser, G. 2008:18: According to da Gama (1964) the species is a junior synonym of Pseudachorutella asigillata (Börner), but it is a valid, separate taxon in our opinion.

Species ellisi Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:338

            Syn.: Pseudachorutella sp. Gwiazdowicz & al, 2022
            Syn.: Pseudachorutella sp. Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:333


Species gigas (Rapoport, EH & Rubio, IR, 1963:112) Massoud, Z, 1967:169

Pseudachorutella gigas from Chile
2016.09.02 © Invertebrados, C.
Pseudachorutella gigas from Chile
Unguis with lateral apical teeth (*)
2016.09.02 © Invertebrados, C.

Species intricata Stryczek, B et Smolis, A, 2024:?

Species maxima Najt, Thibaud & Weiner, 1990

Species plurichaetosa Babenko, A et Kuznetsova, N, 2024:192

Pseudachorutella plurichaetosa from Russia
After Babenko A & Kuznetsova N, 2024 Fig.3-4

Species remyi (Denis, 1933) Massoud, Z, 1967:170, t.t.

            Syn.: Pseudachorutes Remyi(sic) Denis, 1933 IOS
             Syn.: Pseudachorutes Remyi(sic) Stach, J, 1949:56,tt ISS

  Subspecies alpina Stach, J, 1949:65





Species stachi Massoud, 1965

Species stebaevae Babenko, A et Kuznetsova, N, 2024:195

Pseudachorutella stebaevae from Russia
After Babenko A & Kuznetsova N, 2024 Fig.20


Genus Pseudanurida Schött, 1901 key

  Syn.: ... Neachorutes Womersley, 1933
  Syn.: Neanurini(sic) Pseudachorutides Becker, E, 1905:72
  Syn.: Neanurini(sic) Pseudochorutides(sic) Becker, E, 1905:72 ISS
  Syn.: Neanurini(sic) Pseudochorutides(sic) Becker, E, 1906:72 ISS
  Syn.: ... Pseudachorutoides(sic) Denis, 1938 ISS
  Syn.: Pseudachorutini Pseudachorutoides(sic) Womersley, H, 1939:315 ISS
  Syn.: Collembola Pseudachorutoides(sic) Ellis, WN et Bellinger, PF, 1973:45 ISS


Pseudanurida sp. from Singapore
2011.01.04 © Koh, J.

Species billitonensis Schött, 1901, t.t.

Pseudanurida billitonensis from Tokara Island
After Yosii R, 1955 Abb.1A
Pseudanurida billitonensis from Japan
After Uchida H, 1962 Fig.A
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.
Pseudanurida billitonensis from Singapore
2007.06.07 © Ng, M.

Species bogoyawlenskyi (Becker, E, 1905:72) Salmon, 1951

            Syn.: Pseudachorutides Bogoyawlensky(sic) Becker, E, 1905:72 IOS
            Syn.: Pseudachorutides clysmae Jackson, 1927
            Syn.: Pseudachorutoides(sic) dollfusi Denis, 1938


             Syn.: Pseudochorutides(sic) Bogoyawlensky(sic) Becker, E, 1905:72 ISS
             Syn.: Pseudachorutides Bogoyawlensky(sic) Becker, E, 1906:72 ISS
             Syn.: Pseudochorutides(sic) Bogoyawlensky(sic) Becker, E, 1906:72 ISS
             Syn.: Pseudachorutoides(sic) bogoyawlensky(sic) Womersley, H, 1939:315 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Stach, J, 1949:54 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Gisin, H, 1960:69 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Salmon, JT, 1964:277 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Massoud, Z, 1967:172 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Massoud, Z, 1967:174 ISS
             Syn.: Pseudachorutides bogoyawlensky(sic) Massoud, Z, 1967:175 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Massoud, Z, 1967:175 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:49 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:50 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:53 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:55 ISS
             Syn.: Pseudachorutes bogoyawlensky(sic) Murphy, DH, 1971:55 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:56 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, DH, 1971:57 ISS
             Syn.: Pseudanurida bogoyawlensky(sic) Queiroz, GC et Zeppelini, D, 2017:150 ISS
Pseudanurida bogoyawlenskyi from Pakistan
After Aslam S, Arbea JI & Siddiqui G, 2018 Fig.3
Pseudanurida bogoyawlenskyi from Pakistan
With orange eggs
After Aslam S, Arbea JI & Siddiqui G, 2018 Fig.10





Species glauerti (Womersley, 1933) Murphy, DH, 1971:56

            Syn.: Pseudanurida australica Salmon, 1955
            Syn.: Pseudanurida billitonensis Womersley, 1939


Species longisetosa Lee, B-H et Kim, J-T, 1994:208

Species sawayana Schuster, 1965

             Syn.: Pseudanurida bogoyawlensky(sic) Murphy, 1965 cf ISS


             Syn.: Pseudanurida sawayai(sic) Palacios-Vargas, JG et Bu, Y, 2020:? ISS

Species yini Murphy, DH, 1971:62


Genus Halachorutes Arlé, R, 1967:368

  Syn.: ... Holachorutes(sic) Jacquemart, S et Jacques, J-M, 1980:14 ISS

Species quadrisetosus (Snider, RJ, 1981:244) Queiroz, GC & Zeppelini, D, 2017:151

Species schusteri Arlé, R, 1967:370, t.t.

Halachorutes schusteri from Brazil
After Queiroz GC & Zeppelini D, 2017 Fig.17C
Halachorutes schusteri from Brazil
After Queiroz GC & Zeppelini D, 2017 Fig.17B


Genus Oudemansia Schött, 1893 key

Species barnardi (Womersley, H, 1934:447) Uchida, H, 1965:87

            Syn.: Oudemansia dubia Paclt, 1959


Species chenorum Palacios-Vargas, JG et Bu, Y, 2020:?

Species coerulea Schött, 1893, t.t.

            Syn.: Oudemansia subcoerulea Yoshii & Sawada, 1997
            Syn.: Polyacanthella oceanica Uchida, 1955 si
            Syn.: Oudemansia hirashimai Uchida, H, 1965:86
            Syn.: Oudemansia hirashimai Yosii, R, 1977:146


             Syn.: Oudemansia cærulea(sic) MacGillivray, AD, 1893:318 ISS
             Syn.: Friesea coerula(sic) Salmon, 1964 ISS

Species dhritiae Mandal, GP, Mandal, P, Suman, KK et Bhattacharya, KK, 2024:26

Oudemansia dhritiae from India
After Mandal GP, 2024 & al Fig.1
Oudemansia dhritiae from India
After Mandal GP, 2024 & al Fig.2
Oudemansia dhritiae from India
With anal spines
With pseudocelli
After Mandal GP, 2024 & al Fig.21
             Syn.: Oudemansia dhririae(sic) Mandal, GP, Mandal, P, Suman, KK et Bhattacharya, KK, 2024:32 ISS


Species dubia Denis, 1947

Species esakii (Kinoshita, 1932) Yosii, 1958

             Syn.: Oudemansia esaki(sic) Park, K-H et Lee, B-H, 2000:228 ISS
             Syn.: Oudemansia esaki(sic) Park, K-H, Kim, K-G, Park, S-J, Kim, B-J et Lee, B-H, 2000:415 ISS

Species georgia Christiansen, K et Bellinger, P, 1980:295

Species petiti Delamare Deboutteville, C et Massoud, Z, 1964:387

Species shotti Denis, 1948

             Syn.: Friesea schoetti(sic) Salmon, JT, 1964:267 ISS
             Syn.: Oudemansia shötti(sic) Massoud, Z, 1967:176 ISS
             Syn.: Oudemansia schoetti(sic) Mandal, GP, Mandal, P, Suman, KK et Bhattacharya, KK, 2024:30 ISS
             Syn.: Oudemansia schoetti(sic) Mandal, GP, Mandal, P, Suman, KK et Bhattacharya, KK, 2024:32 ISS

Species subcoerulea Denis, 1948

             Syn.: Oudemansia subcoerula(sic) Jacquemart, S et Jacques, J-M, 1980:14 ISS
             Syn.: Oudemansia subcoerula(sic) Jacquemart, S et Jacques, J-M, 1980:18 ISS



Genus Tasmanura Womersley, H, 1937:154

Species evansi Womersley, H, 1937:154, t.t.

Tasmanura evansi from Tasmania
After Womersley H, 1937 Fig.1a
Tasmanura evansi from Tasmania
After Womersley H, 1939 Fig.44M

             Syn.: Ceratrimeria evansii(sic) Greenslade, P, 1974:258,cf ISS
             Syn.: Ceratrimeria evansii(sic) Greenslade, P, 1974:260,cf ISS
Tasmanura evansi from Tasmania
2014.04.dd © Murray, A.




Genus Ceratrimeria Börner, C, 1906:167

  Syn.: ... Platanurida Carpenter, 1925
  Syn.: ... Tasmanura Womersley, 1937
  Syn.: Pseudachorutidae Zealandmeria Stach, J, 1949:59
  Syn.: Pseudachorutinae Schöttella(sic) Babenko, AB, Nakamori, T, Ohira, A et Potapov, MB, 2021:19 IOS


  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1924:220 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1924:222 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:446 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:447 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:448 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:449 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:450 ISS
  Syn.: Collembola Ceratimeria(sic) Handschin, E, 1926:451 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1929:172 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1929:173 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1929:174 ISS
  Syn.: ... Ceratimeria(sic) Denis, 1931 ISS
  Syn.: Collembola Ceratimeria(sic) Denis, J-R, 1933:318 ISS
  Syn.: Pseudachorutinae Caratrimeria(sic) Yosii, R, 1977:147 ISS
Ceratrimeria sp. from New Zealand
2006 © Minor, M. & Robertson, A.
Ceratrimeria sp. from Tasmania
2012.10.28 © Zosterops, K.
Ceratrimeria sp. from Australia
2015.07.27 © Porch, N.





Species aurea Salmon, JT, 1944:133

Species crassa (Oudemans, JT in Weber, M, 1890:74) Yayuk, 1989:5

Species harrisi Salmon, JT, 1942:258

Ceratrimeria harrisi from New Zealand
After Salmon J.T., 1942 Pl.43 Fig.4
Ceratrimeria harrisi from New Zealand
2015 © TePapa Museum

Species indica (Handschin, E, 1929:231) Massoud, Z, 1967:180

Species longicornis (Handschin, E, 1926:451) Massoud, Z, 1967:180

Ceratimeria(sic) longicornis from Java
After Handschin E, 1926 Fig.2a
Ceratimeria(sic) longicornis from Java
After Handschin E, 1926 Fig.2b

Species lydiae Najt, J et Weiner, WM, 1991:108

Species maxima (Schött, 1901) Börner, C, 1906:167, t.t.

Ceratrimeria maxima from Australia
After Schött H, 1917 Fig.4

Species novaezealandiae (Womersley, H, 1936:318) Womersley, 1937

            Syn.: Pseudachorutes novae-zealandiae(sic) Womersley, H, 1936:318 IOS
             Syn.: Ceratrimeria novae-zealandiae(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1941:302 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Salmon, JT, 1942:254 ISS
             Syn.: Zealandmeria novae-zealandiae(sic) Stach, J, 1949:59,si ISS
             Syn.: Aoteareria novae-zealandiae(sic) Salmon, 1964 ISS
             Syn.: Ceratrimeria novae-zealandiae(sic) Massoud, Z, 1967:182 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Zealandmeria novazealandiae(sic) Greenslade, 2012 ISS
             Syn.: Pseudachorutes novazealandiae(sic) Greenslade, P, 2015:6 ISS
             Syn.: Ceratrimeria novazealandiae(sic) Greenslade, P, 2015:6 ISS
Ceratrimeria novaezealandiae from New Zealand
2017.12.01 © Maurice
Ceratrimeria novaezealandiae from New Zealand
Eyepatch with 8 ocelli
2017.12.01 © Maurice





Species pulchella Handschin, 1926

Species takaoensis (Kinoshita, 1916) Yosii, R, 1977:147

Ceratrimeria takaoensis from Japan
After Babenko AB & al, 2021 Fig.1
Ceratrimeria takaoensis from Japan
After Babenko AB & al, 2021 Fig.11-12

Species termiticola Delamare Deboutteville, 1951

            Syn.: Ceratrimeria termiticola Delamare Deboutteville, C, 1948:309 nn
            Syn.: Ceratrimeria termiticola Delamare Deboutteville, C, 1948:422 nn


Species yasumatsui (Uchida, 1940) Yosii, R, 1977:147

Ceratrimeria yasumatsui from Japan
After Uchida, 1965 in Babenko AB & al, 2021 Fig.4
Ceratrimeria yasumatsui from Japan
Type specimen
After Babenko AB & al, 2021 Fig.3
Ceratrimeria yasumatsui from Japan
Dorsal aspect
After Babenko AB & al, 2021 Fig.5
Ceratrimeria yasumatsui from Japan
Ventral aspect
After Babenko AB & al, 2021 Fig.6
Ceratrimeria yasumatsui from Japan
After Babenko AB & al, 2021 Fig.7-10


Genus PseudachorutinaeGennov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105

Species spnov Garrick, RC, Rowell, DM, Simmons, CS, Hillis, DM et Sunnucks, P, 2008:1105

Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.
Pseudachorutinae sp. nov.from Australia
2011.03.11 © Keszei, A.


Genus Aethiopellina Delamare Deboutteville, C, 1951:284

Species bimaculata Delamare Deboutteville, C, 1951:284

Species hebrardi Poinsot-Balaguer, 1978

Species proboscina Delamare Deboutteville, C, 1951:284, t.t.

Species sphaeroculata Massoud, 1963


Genus Aethiopella Handschin, 1942 key

  Syn.: Arthropleona Certrimeria(sic) Womersley, H, 1934:443 ISS
  Syn.: Protachorutidae Ruwenzoriella Salmon, JT, 1964:269


  Syn.: Neanuridae Aethiopiella(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS

Species africana (Delamare Deboutteville, 1945) Massoud, Z, 1967:188

  Subspecies nigra Delamare Deboutteville, 1945

Species ariana Najt, Thibaud & Weiner, 1990

Species basilewskyi (Salmon, 1956) Massoud, Z, 1967:188

Species caraibensis Thibaud & Massoud, 1983

            Syn.: Aethiopella caraïbensis(sic) Thibaud & Massoud, 1983 IOS
             Syn.: Aethiopella caraïbensis(sic) Mari Mutt & Bellinger, 1990 ISS



Species condei Massoud, 1963

Species delamarei Arlé, R, 1960:164

Species flavoantennata (Philiptschenko, 1926 tt) Handschin, 1942

            Syn.: Certrimeria(sic) flavoantennatus(sic) var. capensis Womersley, H, 1934:448


             Syn.: Ceratimeria(sic) flavoantennatus(sic) Denis, J-R, 1933:320 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, H, 1934:448 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria flavoantennatus(sic) Womersley, 1939 ISS

Species guineensis Handschin, 1942

Species handschini (Denis, J-R, 1924:221) Handschin, 1942

            Syn.: Pseudachorutes Handschini(sic) Denis, J-R, 1924:221 IOS
Pseudachorutes Handschini(sic) from Ethiopia
After Denis J-R, 1924 Fig.31

            Syn.: Pseudachorutes Handschini(sic) Denis, J-R, 1929:174 IOS


Species jeanneli Massoud, Z, 1963:60

Species kuolo Christiansen, K et Bellinger, P, 1992:63

Species littoralis Fernandes, LH et de Mendonça, MC, 2003:1

Species machadoi Massoud, 1963

Species maculata (Delamare Deboutteville, 1945) Massoud, Z, 1967:188

Species mandibulata (Delamare Deboutteville, 1945) Salmon, 1956

Species mangenoti Massoud, 1963

Species pedifalx (Salmon, 1956) Massoud, Z, 1967:190

Species pilarandresae Palacios-Vargas, JG et Montejo-Cruz, M, 2014:1039

Species ricardoi da Paz, RV, Queiroz, GC et Bellini, BC, 2019:41

Aethiopella ricardoi from Brazil
After da Paz et al, 2019 Fig.1

Species silvestris Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:36

Species tournieri Delamare Deboutteville, 1950

            Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, 1950 IOS


             Syn.: Aethiopella Tournieri(sic) Delamare Deboutteville, C, 1951:282 ISS



Species ugandensis Salmon, 1954

Species villiersi Massoud, 1963


Genus Protachorutes Cassagnau, P, 1955:154

Species bicolor (Christiansen & Bellinger, 1980:343) Christiansen, KA in Dindal, DL, 1990:979

Species pyrenaeus Cassagnau, P, 1955:154, t.t.

Protachorutes pyrenaeus
Modified after Deharveng L, 1983 Fig.2E


Genus Arlesiella Delamare Deboutteville, C, 1951:285

Species amazonica Arlé, R, 1966:8

Species camerounensis Massoud, 1963

Species delamarei Massoud, Z, 1963:64

Species monodi (Delamare Deboutteville, C in Paulian, R, 1952:65) Massoud, Z, 1967:198

            Syn.: Ceratrimeria Monodi(sic) Delamare Deboutteville, C in Paulian, R, 1952:65 IOS
Ceratrimeria Monodi (sic) from Ivory Coast
After Delamare Deboutteville C, 1952 Fig.16

            Syn.: Ceratrimeria monodi Delamare Deboutteville, 1951 nn




Species saprophila Delamare Deboutteville, C, 1951:285, t.t.

Species simulans Massoud, Z, 1963:67


Genus Linnaniemia Philiptschenko, 1926

Species bicolor (Delamare Deboutteville, 1945) Massoud, Z, 1967:202

Species gigas Philiptschenko, 1926:202, t.t.


Genus Neotropiella Handschin, 1942 key

  Syn.: ... Americotrimeria Stach, 1949


  Syn.: Brachystomellidae(lapsus) Neotropiella Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:14

Species sp.1 Janssens F sensu Bellini & Zeppelini, 2004

            Syn.: Neotropiella sp.undescr. Bellini, BC et Zeppelini, D, 2004:588

Species arlei Najt, Thibaud & Weiner, 1990

Species arretada Paz RV, Bellini BC et Queiroz GC in Bellini, BC, Weiner, WM, Queiroz, GC et Paz, RV, 2020:3

Neotropiella arretada from Brazil
After Bellini BC & al, 2020 Fig.2H
Neotropiella arretada from Brazil
Tentative designation of ocelli of right eye
Ocelli ABCD fused to large ocellus
Modified after Bellini BC & al, 2020 Fig.2H

Species barbatae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:43

Species carli (Denis, J-R, 1924:222) Arlé, 1960

            Syn.: Pseudachorutes Carli(sic) Denis, J-R, 1924:222 IOS
            Syn.: Ceratrimeria pentoculata Womersley, 1930
            Syn.: Neotropiella araguaensis Rapoport & Maño, 1969



            Syn.: Pseudachorutes Carli(sic) Denis, J-R, 1924:223 IOS
            Syn.: Ceratimeria(sic) Carli(sic) Denis, J-R, 1929:172 IOS
            Syn.: Pseudachorutes Carli(sic) Denis, J-R, 1929:174 IOS
            Syn.: Ceratimeria(sic) Carli(sic) Denis, J-R, 1929:174 IOS
            Syn.: Ceratimeria(sic) Carli(sic) Denis, J-R, 1933:320 IOS
Neotropiella carli from French Guyana
Pointed mouthcone
After Thibaud J-M & D'Haese C, 2010 Fig.2
2009 © D'Haese, C.
Neotropiella carli from Ecuador
Eyepatch with 5 ocelli; PAO present
2014.01.31 © McClarin, J.
Neotropiella carli from Ecuador
Body segments trilobed
2014.01.31 © McClarin, J.



Species digitomucronata Massoud & Thibaud, 1982

Species duranti Díaz & Najt, 1995

Species gordae Díaz & Najt, 1995

Species insularis Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:47

Species macunaimae Queiroz, GC, da Silveira, TC et de Mendonça, MC, 2013:45

Species meridionalis (Arlé, 1939) Stach, 1949

Neotropiella meridionalis from Brazil
After Cipola NG & al, 2019 Fig.14

Species minima Thibaud & Oliveira, 2010

Species murphyi Massoud, 1965

             Syn.: Neotropiella murphy(sic) Bellini, BC, Weiner, WM, Queiroz, GC et Paz, RV, 2020:10 ISS
?Neotropiella murphyi? from Java
2020.05.12 © Naufal, U.



Species pedisensilla Najt, Thibaud & Weiner, 1990

Species peruana Palacios-Vargas, JG et Callohuari, YT, 2020:4

Neotropiella peruana from Peru
After Palacios-Vargas JG & Callohuari YT, 2020 Fig.2a
Neotropiella peruana from Peru
Tentative designation of ocelli of left eye
Ocelli ABCD fused to large ocellus
Modified after Palacios-Vargas JG & Callohuari YT, 2020 Fig.8

Species plurichaetosa Thibaud & Oliveira, 2010

Species quinqueoculata (Denis, 1931) Arlé, 1962

            Syn.: Pseudachorutes 5-oculatus(sic) Denis, 1931 IOS
            Syn.: Pseudachorutes 5-oculatus(sic) Denis, J-R, 1933:318 IOS
             Syn.: Americotrimeria 5-oculata(sic) Stach, 1949 ISS
             Syn.: Neotropiella quinque-oculata(sic) Arlé, R, 1966:13 ISS
             Syn.: Neotropiella 5-oculata(sic) Arlé, R, 1966:15 ISS
             Syn.: Neotropiella pentoculata(sic) Massoud, 1967 ISS
Neotropiella quinqueoculata from Panama
After Palacios-Vargas JG, 2021 Fig.1A
Neotropiella quinqueoculata from Mexico
After Palacios-Vargas JG, 2021 Fig.1B





Species silvestrii (Denis, J-R, 1929:172) Massoud, Z, 1967:208, t.t.

            Syn.: Ceratimeria(sic) Silvestrii(sic) Denis, J-R, 1929:172 IOS
            Syn.: Ceratimeria(sic) Silvestrii(sic) Denis, J-R, 1929:173 IOS
Ceratimeria(sic) Silvestrii(sic) from Cuba
After Denis J-R, 1929 Fig.IV.1



Species vanderdrifti Massoud, 1963


Genus Sernatropiella Palacios-Vargas, JG, 2019:374

Species malkini (Arlé, 1981) Palacios-Vargas, JG, 2019:379

Species pinzonae Palacios-Vargas, JG, 2019:374, t.t.

Sernatropiella pinzonae from Colombia
After Palacios-Vargas JG, 2019 Fig.1
Sernatropiella pinzonae from Colombia
After Palacios-Vargas JG, 2019 Fig.2
Sernatropiella pinzonae from Colombia
With cephalic pseudocelli (*)
After Palacios-Vargas JG, 2019 Fig.5


Genus Grananurida Yosii, 1954

  Syn.: Neanuridae Agranurida(sic) Kim, J-T et Lee, B-H, 2000:180 ISS
Remark by Janssens F. & Bernard, E.C. 2007:in litt.: Agranurida(sic) Kim, J.T. & Lee.-H., 2000:180 is an incorrect subsequent spelling of Grananurida.


Species alba (Yosii, 1966) Weiner, WM & Najt, J, 1985:281, i.s.

Species baicalica Rusek, J, 1991:126

Species koreana Lee, B-H et Kim, J-T, 1984:21

Species minuta Kim, J-T et Lee, B-H, 2000:180

Species tuberculata Yosii, 1954, t.t.


Genus Najtafrica Barra, J-A, 2002:178

Species riebi (Barra, J-A, 1994:184) Barra, J-A, 2002:178

Najtafrica riebi from South Africa
After Barra J-A, 2002 Fig.1C


Genus Stachorutes Dallai, 1973 key

  Syn.: Neanuridae Chihuahuachorutes Palacios-Vargas, JG, 1990:7


Species arlei (Thibaud & Massoud, 1980) Weiner, WM & Najt, J, 1998:225

Species cabagnerensis Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150

Species cuihuaensis Gao & Yin, 2007

Species dallaii Weiner, WM et Najt, J, 1998:226

             Syn.: Stachorutes dallai(sic) Kovác, L, 1999:135 ISS
             Syn.: Stachorutes dallai(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS
             Syn.: Stachorutes dallai(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS

Species dematteisi Dallai, 1973, t.t.

             Syn.: Stachorutes dematteisae(sic) Palacios-Vargas, 1990 ISS
             Syn.: Stachorutes demattiesi(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Stachorutes dematteisae(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:108 ISS

Species escobarae (Palacios-Vargas, JG, 1990:8) Thibaud, J-M & Palacios-Vargas, JG, 2000:107

Species gracilis Smolis, A et Shvejonkova, JB, 2006:44

Species jizuensis Tamura & Zhao, 1997

Species longirostris Deharveng & Lienhard, 1983

Species maya Thibaud, J-M et Palacios-Vargas, JG, 2000:108

Species najtae Fanciulli, PP, Dallai, R, Frati, F et Carapelli, A, 2017:32

Stachorutes najtae from Italy
After Fanciulli PP & al, 2017 Fig.3A

Species navajellus Fjellberg, A, 1984:198

Stachorutes navajellus from the USA
After Fjellberg A, 1984 Fig.3AB
             Syn.: Stachorutes navajelus(sic) Slawska, M, 1996:329 ISS
             Syn.: Stachorutes novajellus(sic) Thibaud, J-M et Palacios-Vargas, JG, 2000:110 ISS


Species ruseki Kovác, L, 1999:135

Species scherae Deharveng & Lienhard, 1983

Species sphagnophilus Slawska, M, 1996:325

             Syn.: Stachorutes sphagnphilus(sic) Slawska, M, 1996:327 ISS
Stachorutes sphagnphilus(sic) from Poland
After Slawska M, 1996 Fig.8
Stachorutes sphagnphilus(sic) from Poland
Abdominal ventral aspect
Furcal area at anterior margin of abd.4
After Slawska M, 1996 Fig.9
Stachorutes sphagnphilus(sic) from Poland
Retinaculum fully developed
Furca reduced; mucro absent
After Slawska M, 1996 Fig.10

             Syn.: Stachorutes sphagnophillus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:156 ISS


Species tatricus Smolis, A et Skarzynski, D, [2002]:407

             Syn.: Stachorutes tatricusa(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS
             Syn.: Stachorutes tatricusb(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:153 ISS

Species tieni Pomorski, RJ et Smolis, A, 1999:?

Species triocellatus Pomorski, RJ et Smolis, A, 1999:?

             Syn.: Stachorutes triocelatus(sic) Simón-Benito, JC, Espantaleón, D et García-Barros, E,, 2005:150 ISS

Species valdeaibarensis Arbea, JI et Jordana, R, 1991:80

             Syn.: Stachorutes valdeibarensis(sic) Slawska, M, 1996:329 ISS



Genus Platanurida Carpenter, GH, 1925:5

Platanurida sp. from New Zealand
2016.03.18 © Murray, A.
Platanurida sp. from New Zealand
2016.03.18 © Murray, A.
Platanurida sp. from New Zealand
2017 © Murray, A.

Species lata Carpenter, GH, 1925:5, t.t.

Ceratrimeria lata from New Zealand
After Salmon JT, 1941 Pl.44 Fig.73

Species marplesi (Salmon, JT, 1941:301) Massoud, Z, 1967:212

Ceratrimeria marplesi from New Zealand
After Salmon JT, 1941 Pl.44 Fig.79
Platanurida marplesi from New Zealand
2014 © TePapa Museum
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.
Platanurida marplesi from New Zealand
2014.05.21 © Murray, A.

Species marplesoides Massoud, Z, 1967:210

            Syn.: Platanurida marplesoïdes(sic) Massoud, Z, 1967:210 IOS
            Syn.: Ceratrimeria marplesi Salmon, 1942, nec Salmon, 1941





Genus Israelimeria Weiner, WM et Kaprus, IJ, 2005:49

Species persica Alijani, M, Shayanmehr, M, Skarzynski, D, Piwnik, A et Smolis, A, 2015:269

Israelimeria persica from Iran
After Alijani M & al, 2015 Fig.1

Species poliakovae Weiner, WM et Kaprus, IJ, 2005:50, t.t.

Israelimeria poliakovae from Israel
After Weiner W.M. & Kaprus, I.J, 2005 Fig.1


Genus Arlesia Handschin, 1942

  Syn.: ... Arlésia(sic) Handschin, 1942 IOS
  Syn.: Neanuridae Portachorutes Wray, DL, 1953:147
  Syn.: ... Pseudarlesia Salmon, 1964
  Syn.: Pseudachorutinae Handschinia Massoud, Z, 1967:214




Species sp.1 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Arlesia sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:8

Species albipes (Folsom, 1927) Stach, J, 1949:57, t.t.

            Syn.: Portachorutes mambatus Wray, DL, 1953:147
            Syn.: Arlesia albipes f. decorata Mari Mutt, JA et Bellinger, PF, 1990:41


             Syn.: Arlesia albibes(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:45 ISS

Species arleana de Mendonça, MC et Fernandes, LH, 1999:1196

            Syn.: Arlesia albipes f. decorata Arlé & Rufino, 1976


Species cochabambensis Cassagnau & Rapoport, 1962

Species innominata (Salmon, JT, 1964:276) Bellinger, 1996 il

            Syn.: Ceratrimeria albipes Denis, 1931
            Syn.: Ceratrimeria n.sp. innominata Denis, 1931


Species intermedia Fernandes, LH et de Mendonça, MC, 2004:21

Species variabilis Thibaud & Massoud, 1983


Genus Intermediurida Najt, Thibaud & Weiner, 1990

Species anophthalma Najt, Thibaud & Weiner, 1990, t.t.

             Syn.: Intermediurida anophtalma(sic) Queiroz, GC et Zeppelini, D, 2017:152,tt ISS
             Syn.: Intermediurida anophtalma(sic) Queiroz, GC et Zeppelini, D, 2017:153 ISS



Genus Venezuelida Díaz & Najt, 1995

Species polylepiana Díaz & Najt, 1995, t.t.


Genus Handschinurida Queiroz, GC, 2015:499

  Syn.: Pseudachorutidae Handschinia Stach, J, 1949:56 nec Lallemand, 1935
  Syn.: Collembola Handschinia Ellis, WN et Bellinger, PF, 1973:22 nec Lallemand, 1935
  Syn.: Pseudachorutinae Handschinia Queiroz, GC et de Mendonça MC, 2014:171 nec Lallemand, 1935


Species fluminensis (Arlé, 1939 tt) Queiroz, GC, 2015:499, t.t.

             Syn.: Arlesia flaminensis(sic) Palacios-Vargas, JG in Alvarez-Sánchez, J et Naranjo-García, E, 2003:221,ca ISS
Handschinia fluminensis from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.15



Species proxima (Arlé, 1939) Queiroz, GC, 2015:499

Handschinia proxima from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.19

Species rauli (Queiroz, GC & de Mendonça MC, 2014:172) Queiroz, GC, 2015:499

Handschinia rauli from Brazil
After Queiroz GC & de Mendonça MC, 2014 Fig.23


Genus Brasilimeria Stach, J, 1949:56 key

Species sp. Queiroz, GC et Zeppelini, D, 2017:133

Species anura (Arlé, 1939) Stach, J, 1949:56, t.t.

Brasilimeria anura from Brazil
Dorsal aspect
Hemolymphatic defense bleeding (arrow)
After Zeppelini D et al, 2019 Fig.1a
Brasilimeria anura from Brazil
Ventral aspect
Hemolymphatic defense bleeding (arrows)
After Zeppelini D et al, 2019 Fig.1b

Species assua Zeppelini, D, Queiroz, GC, Lopes, NP et Mendonça-Junior, FJB, 2019:5

            Syn.: Brasilimeria assu(sic) Zeppelini, D, Queiroz, GC, Lopes, NP et Mendonça-Junior, FJB, 2019:5 IOS

Species wygodzinskyi (Arlé, R, 1943:110) Arlé, 1959

            Syn.: Ceratrimeria Wygodzinskyi(sic) Arlé, R, 1943:110 IOS
Brasilimeria wygodzinskyi from Brazil
After Massoud Z, 1967 Fig.69G




Genus Tijucameria de Mendonça, MC et Fernandes, LH, 2005:699

  Syn.: Pseudachorutinae Tijucameira(sic) Queiroz, GC et Zeppelini, D, 2017:131 ISS

Species sp. Queiroz, GC et Zeppelini, D, 2017:133

            Syn.: Neotropiella denisi Arlé, 1981


Species gabrieli de Mendonça & Silveira, 2012

Species mame de Mendonça, MC et Fernandes, LH, 2005:699, t.t.


Genus Kenyura Salmon, 1954

  Syn.: Neanuridae Paranurella Arlé, R, 1960:162
  Syn.: Pseudachorutini Pseudoparanura Massoud, Z, 1963:70 =?


Species sp. Bendjaballah, M, Zoughailech, A, Brahim-Bounab, H, Hamra-Kroua, S, Bedos, A et Deharveng, L, 2018:399

Kenyura sp. from Algeria
After Bendjaballah M & al, 2018 Fig.10

Species africana (Massoud, Z, 1963:70) Massoud, Z, 1967:230

Species delicata Arlé, R, 1966:4

Species monoculata (Winter, C, 1963:484) Mari Mutt, JA & Bellinger, PF, 1990:43

            Syn.: Paranurella monoculata Winter, C, 1963:484 nn
Remark by Winter, C. 1963:395: 37 von mir neu aufgestellte Arten bzw. Formen sind in der vorliegende Arbeit mit n.sp. bzw. n.f. (ohne Autorennamen) bezeichnet und müssen bis zu ihrer Beschreibung (voraussichtlich im kommenden Jahr) als Nomina nuda betrachtet werden.


Species multisensa Salmon, 1954, t.t.

Species nicaraguensis Palacios-Vargas & Deharveng, 2010

Species porcula (Arlé, R, 1960:163) Arlé, R, 1966:2

            Syn.: Paranurella porculus(sic) Arlé, R, 1960:163 IOS


             Syn.: Kenyura porculus(sic) Arlé, R, 1966:2 ISS
             Syn.: Kenyura porculus(sic) Massoud, Z, 1967:229 ISS
             Syn.: Kenyura porculus(sic) Arlé, 1981 ISS
             Syn.: Kenyura porculus(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
             Syn.: Kenyura porculus(sic) Culik, MP et Zeppelini, DF, 2003:1124 ISS



Species xinguensis Arlé, R, 1966:3


Genus Furculanurida Massoud, Z, 1967:231 key  key

Species africana (Massoud, Z, 1963:68) Massoud, Z, 1967:231, t.t.

Species arawakensis Thibaud & Massoud, 1983

Species ashrafi (Yosii, 1966) Thibaud, J-M & Palacios-Vargas, JG, 2000:112

Species belemensis Arlé & Rufino, 1976

Species bistribusata Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2020:3

            Syn.: Furculanurida bistribus(sic) Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2020:3 IOS
Furculanurida bistribus(sic) from Puerto Rica
After Ospina-Sánchez CM & al, 2020 Fig.1
Furculanurida bistribus(sic) from Puerto Rica
Dorsal chaetotaxy
After Ospina-Sanchez CM & al, 2020 Fig.2



Species boiunae da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:4

            Syn.: Furculanurida boiuna(sic) da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:4 IOS
            Syn.: Furculanurida (Furculanirida) boiuna(sic) da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:6 IOS
            Syn.: Furculanurida boiuna(sic) Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2020:8 IOS
            Syn.: Furculanurida boiuna(sic) Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2020:9 IOS

Species duodecimoculata Thibaud & Massoud, 1980

Species emucronata Zon, Tano & Deharveng, 2014

Species furculata (Salmon, 1956) Massoud, Z, 1967:233

Species goeldiana Arlé & Rufino, 1976

Species grandcolasorum Weiner, WM et Najt, J, 1998:223

Furculanurida grandcolasorum from Tanzania
Bicolour form
After Weiner W.M. & Najt J, 1998 Fig.17

Species guatemalensis Palacios-Vargas, JG et Gao, Y, 2009:55

Furculanurida guatemalensis from Guatemala
After Palacios JG & Gao Y, 2009 Fig.1
             Syn.: Furculanurida guatemaltesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS
             Syn.: Furculanurida guatemalesis(sic) Palacios-Vargas, JG et Gao, Y, 2009:56 ISS


Species langdoni Bernard, 2007

Species longisensillata Najt, Thibaud & Weiner, 1990

Species nessimiani (Fernandes, LH & de Mendonça, MC, 2003:4) Palacios-Vargas, JG & Gao, Y, 2009:55

             Syn.: Furculanurida nessiamiani(sic) Palacios-Vargas, JG et Gao, Y, 2009:55 ISS

Species septemoculata Palacios-Vargas, JG et Gao, Y, 2009:56

Furculanurida septemoculata from Guatemala
After Palacios JG & Gao Y, 2009 Fig.12

Species tropicalia Queiroz & Fernandes, 2011


Genus Micranurida Börner, C, 1901:702

  Syn.: Pseudachorutinae Lanzhotia Smolis, A, Deharveng, L et Bedos, A, 2012:154 =?


  Syn.: Collembola Micranurisa(sic) Haybach, G, 1959:36 ISS
  Syn.: Collembola Microanurida(sic) Sterzynska, M et Kuznetsova, N, 1995:147 ISS
  Syn.: Collembola Microanurida(sic) Sterzynska, M et Kuznetsova, N, 1995:151 ISS
  Syn.: Collembola Microanurida(sic) Kopeszki, H et Meyer, E, 1996:230 ISS
  Syn.: Collembola Microannurida(sic) Kopeszki, H et Meyer, E, 1996:232 ISS
  Syn.: Collembola Microanurida(sic) Kopeszki, H et Meyer, E, 1996:233 ISS
  Syn.: Collembola Microanurida(sic) Kopeszki, H et Meyer, E, 1996:234 ISS
  Syn.: Pseudachorutini Microanurida(sic) Mendoza Arviso, S, Villalobos, FJ, Ruíz Montoya, L et Castro, AER, 1999:90 ISS
  Syn.: Pseudachorutini Microanurida(sic) Mendoza Arviso, S, Villalobos, FJ, Ruíz Montoya, L et Castro, AER, 1999:96 ISS
Micranurida sp. from Far East Russia
After Babenko A & al, 2022 cover



Species sp.1 Janssens F sensu Palacios-Vargas, 1981

            Syn.: Micranurida sp.nov. Palacios-Vargas, JG, 1981:89


Species sp.2 Janssens F sensu Palacios-Vargas & Gómez-Anaya, 1995

            Syn.: Micranurida sp.nov. Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8


Species agenjoi Simón, 1981

Species anini Christiansen, K et Bellinger, P, 1992:68

Species anophthalmica Stach, J, 1946:26

Species balta Fjellberg, A, 1998:77

Species bescidica Smolis, A et Skarzynski, D, 2004:1

Species candida Cassagnau, P, 1952:310

Micranurida candida from Algeria
After Bendjaballah M & al, 2018 Fig.11

Species distincta Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species fluminensis Fernandes, LH et de Mendonça, MC, 2004:22

Species forsslundi Gisin, 1949, B

Species furcifera Mills, HB, 1934:27

Species hygrophila Rusek, 1973

Species intermedia Dallai, 1973

Species lanceolata Najt, J et Weiner, WM, 1985:35

Species meridionalis Cassagnau, P, 1952:310

Species montana Lee & Kim, 1984

Species papillosa Axelson, 1902, B

Species potapovi Smolis, A, Deharveng, L et Bedos, A, 2012:153

Micranurida potapovi from Far East Russia
After Smolis A, Deharveng L & Bedos A, 2012 Fig.2a+e

Species pygmaea Börner, C, 1901:702, t.t., B

Micranurida pygmaea from Germany
After Börner, 1901 Fig.6

            Syn.: Micranurida pygmaea f. hasai Kseneman, 1936
            Syn.: Micranurida meridionalis Arbea & Jordana, 1985



             Syn.: Micranurida pygmea(sic) Winter, C, 1963:475 ISS
Micranurida pygmaea from Italy
After Dallai R, 1973 Fig.6n

             Syn.: Micranurida pygmea(sic) Slawska, M, 1996:328 ISS
             Syn.: Micranurida pygmea(sic) Smolis, A, Deharveng, L et Bedos, A, 2012:149 ISS


Species retezatica Gruia & Harsia, 1990

Species rostrata Babenko, AB, 2007:364

Species russica Smolis, A, Deharveng, L et Bedos, A, 2012:150

Micranurida russica from Far East Russia
After Smolis A, Deharveng L & Bedos A, 2012 Fig.1a+i

Species sensillata (Gisin, 1953) Dunger, 1974

Species sectilis Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species spirillifera Hammer, 1953

             Syn.: Micranurida spirilifera(sic) Massoud, Z, 1967:236 ISS

Species tuberculata Lee & Kim, 1984

Species valiana Fjellberg, 1985

Species wladimiri Najt & Rubio, 1978

  Subspecies caribena Ospina-Sánchez, 2019

            Syn.: Micranurida wladimiri ssp. caribeña(sic) Ospina-Sánchez, 2019 IOS

            Syn.: Micranurida wladimiri ssp. caribeña(sic) Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:9 IOS


Genus Meganurida Carpenter, 1935

Species mumfordi Carpenter, 1935, t.t.


Genus Pseudachorudina Stach, 1949

Species alpina Stach, 1949 tt, t.t.

             Syn.: Pseudachorudina alpine(sic) Budha, PB et Shrestha, P, 2021:28 ISS

Species angelieri Cassagnau, P, 1959:76

            Syn.: Pseudachorutes subcrassus Selga, 1971 partim Arbea & Jordana in Ramos, 1997
            Syn.: Pseudachorudina santanderensis Simon & Pozo, 1989


Species berninii (Dallai, R, 1970:438) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:5


Pseudachorutes brunneus from New Zealand
After Salmon, J.T. 1941 Pl.46 Fig.101.
Species brunnea (Carpenter, GH, 1925:2) Massoud, Z, 1967:243


Species dahli (Schäffer, C, 1898:400) Yoshii, R & Suhardjno, YR, 1992:2

            Syn.: Schöttella(sic) dahlii(sic) Schäffer, C, 1898:400 IOS
            Syn.: Pseudachorudina delamarei Massoud, 1965



            Syn.: Pseudachorutes dahlii(sic) Börner, 1901 IOS
            Syn.: Schöttella(sic) dahlii(sic) Schött, 1901 IOS
            Syn.: Schöttella(sic) dahlii(sic) Stach, 1929 IOS
            Syn.: Pseudachorutes dahlii(sic) Womersley, 1937 IOS
            Syn.: Schoettella dahlii(sic) Salmon, JT, 1964:200 IOS
            Syn.: Pseudachorutes dahlii(sic) Yosii, R, 1965:8 IOS
            Syn.: Pseudachorutes dahlii(sic) Massoud, Z, 1967:165 IOS
            Syn.: Pseudachorutes? dahlii(sic) Yayuk, RS(sic), 1989:4 IOS
            Syn.: Pseudachorudina dahlii(sic) Yoshii, R et Suhardjno, YR, 1992:2 IOS
Pseudachoridina dahlii(sic) from Indonesia
After Yoshii R & Suhardjno YR, 1992 Fig.1A



Species evansi (Womersley, H, 1936:475) Stach, 1949

Species falteronensis (Denis, 1926) Massoud, 1967:243, t.t.

Species ignota (Christiansen, K & Bellinger, P, 1980:300) Palacios-Vargas, JG, 2020:1

             Syn.: Pseudachorudina ignotus(sic) Palacios-Vargas, JG, 2020:1 ISS
             Syn.: Pseudachorudina ignotus(sic) Palacios-Vargas, JG, 2020:4 ISS
Pseudachorudina ignota from the USA
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Ant.4 apical bulb
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Postantennal organ with 11 vesicles
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Eye with 8 ocelli (A-H)
Postantennal organ (PAO) subequal to ocellus A
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Maxilla head with serrate lamella
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Mandibula apically with 5 small and 2 large teeth
2024.02.12 © Wang, G.
Pseudachorudina ignota from the USA
Terminal body segments
with capitate macrosetae
2024.02.12 © Wang, G.



Species mabirensis (Philiptschenko, 1926) Salmon, 1954

Species meridionalis (Bonet, 1929) Jordana, R, Arbea, JI, Simón, C & Luciáñez, MJ in Ramos, MA & al, 1997:777

            Syn.: Pseudachorudina bougisi Delamare Deboutteville, 1951
            Syn.: Pseudachorudina bougisi var. catalonica Massoud, 1963
            Syn.: Pseudachorudina patriciae Gough, 1976
            Syn.: Pseudachorudina najtae Simon & Pozo, 1989


Species navarretei Palacios-Vargas, JG, 2020:2

Pseudachorudina navarretei from Mexico
After Palacios-Vargas JG, 2020 Fig.1

Species nepalica Yosii, 1966


Brachystomella osextara from New Zealand
After Salmon, J.T. 1941 Pl.45 Fig.88.
Species osextara (Salmon, JT, 1941:303) Massoud, Z, 1967:243

Species pacifica (Womersley, H, 1936:318) Massoud, Z, 1967:241

             Syn.: Pseudachorudina pacificus(sic) Massoud, Z, 1967:241 ISS

Species texensis (Christiansen & Bellinger, 1980:301) Palacios-Vargas, JG, 2020:1


Genus Cassagnaudina Massoud, 1967

Species coiffaiti (Cassagnau, P, 1955:145) Massoud, Z, 1967:245

            Syn.: Pseudachorudina Coiffaiti(sic) Cassagnau, P, 1955:145 IOS

Species khumbuensis Yosii, 1971

Species perisi Simón, 1981

Species tourratensis (Cassagnau, P, 1964:470)


Genus Cassagnaurida Salmon, 1964

Species dentata (Cassagnau & Rapoport, 1962) Massoud, Z, 1967:246, t.t.

Cassagnaurida dentata from Chile
Abdominal dorsal chaetotaxy
After Palacios-Vargas JG, 2021 Fig.8
Cassagnaurida dentata from Chile
Cephalic + thoracic dorsal chaetotaxy
After Palacios-Vargas JG, 2021 Fig.7


Genus Notachorudina Cassagnau & Rapoport, 1962

  Syn.: Collembola Nothachorudina(sic) Rapoport, EH et Rubio, IR, 1963:111 ISS

Species patagonica Cassagnau & Rapoport, 1962, t.t.

            Syn.: Notachorudina rapoporti Queiroz, GC et Zeppelini, D, 2017:138
Notachorudina rapoporti from Chile
With everted anal sacs
After Queiroz GC & Zeppelini D, 2017 Fig.11B

            Syn.: Notachorudina rapoporti Palacios-Vargas, JG, 2021:12



Notachorudina patagonica from Chile
Head
After Palacios-Vargas JG, 2021 Fig.40
Notachorudina patagonica from Chile
Body dorsal chaetotaxy
After Palacios-Vargas JG, 2021 Fig.41


Genus Forsteramea Salmon, JT, 1965:645

Species megacephala (Salmon, JT, 1954:213) Salmon, JT, 1965:645, t.t.

            Syn.: Montachorutes megacephala(sic) Salmon, JT, 1954:213 IOS
Montachorutes megacephala(sic) from New Zealand
After Salmon JT, 1954 Text-Fig.1

             Syn.: Montachorutes? megacephala(sic) Rapoport, EH et Rubio, IR, 1963:113 ISS
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.
Forsteramea megacephala from New Zealand
2014.05.31 © Murray, A.






Genus Delamarellina Rapoport, EH et Rubio, IR, 1963:113

  Syn.: Pseudachorutinae Dellamarelina Queiroz, GC et Zeppelini, D, 2017:129 ISS

Species globulosa (Cassagnau & Rapoport, 1962) Massoud, Z, 1967:253

            Syn.: Delamarellina guilleni Rapoport, EH et Rubio, IR, 1963:113,tt
Delamarellina guilleni from Chile
After Rapoport EH & Rubio IR, 1963 Fig.62

            Syn.: Delamarellina guilleni Palacios-Vargas, JG, 2021:6



Delamarellina globulosa from Chile
After Palacios-Vargas JG, 2021 Fig.14
Delamarellina globulosa juv. from Chile
After Palacios-Vargas JG, 2021 Fig.32

Species ubiquata (Salmon, JT, 1944:133) Massoud, Z, 1967:251


Genus Quatacanthella Salmon, JT, 1945:68


Polyacanthella proprieta from New Zealand
After Salmon, J.T. 1941 Pl.42 Fig.57.
Species proprieta (Salmon, JT, 1941:296) Salmon, JT, 1945:68, t.t.


Genus Anurida Laboulbène, 1865 key  key  key  key

  Syn.: ... Anoura(sic) Gervais, 1844
  Syn.: Lipuridæ(sic) Anoura(sic) Lubbock, J, 1862:601 partim ISS
  Syn.: Aphoruridæ(sic) Aphoromma MacGillivray, AD, 1893:314
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Hypanurida Denis, 1931
  Syn.: Pseudachorutini Anuridella Mills, HB, 1934:26
  Syn.: Neanurinae Anuridella Gisin, H, 1960:87
  Syn.: Neanurinae Anuridella Poole, RW in Poole, RW et Gentili, P, 1997:46
  Syn.: Neanurinae Gastranurida Gisin, H, 1960:87


Anurida sp. from Holland
2018.09.30 © van Leeuwen, J.



Species abashiriensis Babenko, A et Nakamori, T, 2018:377

Anurida abashiriensis from Japan
After Babenko A & Nakamori T, 2018 Fig.12-13

Species alpina Agrell, 1939

            Syn.: Anurida tundricola Tshelnokov, 1988
            Syn.: Anurida tundricola Babenko, AB, 1997:512


Species amorita Folsom, JW, 1902:90

Anurida amorita from Alaska
After Folsom JW, 1902 Pl.IV Fig.3

Species annatshagi Tshelnokov, 1988

            Syn.: Anurida balagannakhi Tshelnokov, 1988
            Syn.: Anurida balagannakhi Babenko, AB, 1997:512


Species ariesi Harsia & Gruia, 1992

Species armifera Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:134


Anurida ashbyae from the Lesser Antilles
Det. Christiansen, K.
2006 © De Silva, M.
Species ashbyae Christiansen, K et Bellinger, P, 1988:228


Species asiatica Babenko, AB, 2002:374

Species aspera Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:144

Species assimilis Yosii, R, 1956:28

  Subspecies persimilis Yosii, R, 1956:28

Species azurea Babenko, AB, 1997:513

Species balatovae Rusek, 1970

Species bisetosa Bagnall, 1949

Species bondarenkoae Tshelnokov, 1988

            Syn.: Anurida jakutica Tshelnokov, 1988
            Syn.: Anurida jakutica Babenko, AB, 1997:517 =?
            Syn.: Anurida jakutica Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:146


Species bonneti Cassagnau, P, 1955:147

            Syn.: Anurida Bonneti(sic) Cassagnau, P, 1955:147 IOS

Species borealis Martynova in Stebaeva, 1976 nn

Species callainii Dallai, R et Martinozzi, I, 1980:8

Species caprariensis Dallai, 1969

Species carpatica Babenko, AB, 1998:658

Species clavata Schäffer, 1897

Species confinis Babenko, AB, 1998:651

Species decemoculata Hammer, 1953

Species decipiens Yosii, R, 1966:546

Species dentata Christiansen & Bellinger, 1980:326

            Syn.: Anurida (Anurida) decemoculata Christiansen & Bellinger, 1980,si


Species desnuda Yosii, R, 1956:25

Species diabolica Yosii, R, 1956:26

Species dynkengda Babenko, AB, 1998:649

Species elegans Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:140

Species ellipsoides Stach, J, 1949:34

             Syn.: Anurida ellipsoïdes(sic) Massoud, Z, 1967:260 ISS
             Syn.: Anurida elipsoides(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS
             Syn.: Anurida ellipsoids(sic) Falahati HosseinAbad, A, Mehr, MS et Khyroodin, A, 2013:260 ISS

Species granaria (Nicolet, H, 1847:387) Tullberg, 1869, t.t., B

            Syn.: Anurida bagnalli Stach, J, 1949:14
            Syn.: Anurida comellinii Gisin, 1949
            Syn.: Anurida pseudogranaria Stach, J, 1949:30
            Syn.: Megaphorura arctica Shaw, P et Trewhella, S, 2019:14 lapsus
Megaphorura arctica (lapsus) from the UK
After Shaw P & Trewhella S, 2019 Fig.20
= Anurida granaria Janssens F, 2021.02.17




Anoura granaria from the UK
After Lubbock J, 1873 Pl.49
Anurida granaria from the UK
2005 © Hopkin, S.P.
Anurida granaria from Sweden
2006 © Hall, K.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2012.11.22 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria from the UK
2013.01.28 © Murray, A.
Anurida granaria juv. from the UK
2013.01.29 © Murray, A.
Anurida granaria from the UK
Circular post-antennal organ
2013.01.29 © Murray, A.
Anurida granaria from the UK
Granulated cuticula
2013.01.29 © Murray, A.
Anurida granaria from the UK
Antennae and mouthcone
2013.03.23 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from New Zealand
2014.02.17 © Murray, A.
Anurida granaria from Holland
2014.03.14 © van Duinen, J.
Anurida nr granaria from Canada
Anal spines absent
2015.08.12 © Murray, A.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
2015.09.26 © Tony.
Anurida granaria from Tasmania
Cuticle coarsely granulate
Abd.6 not bilobed
2015.09.26 © Tony.
Anurida granaria from Tasmania
Postantennal organ circular
Tapering antenna
2015.09.26 © Tony.
Anurida granaria from New Zealand
Mouth: labrum, maxillae heads, hypopharynx, split labium
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Ant.4: apical eversible trilobed sensillar vesicle
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Rudiment of furcula
2016.02.10 © Koken, M.
Anurida granaria from New Zealand
Post-antennal organ
2016.02.10 © Koken, M.
Anurida nr granaria from Canada
2021.01.14 © Rennert, M.
Anurida granaria from Belgium
2021.11.19 © Huskens, M.-L.
Anurida granaria from Belgium
2021.11.19 © Huskens, M.-L.
Anurida granaria from Belgium
Post-antennal organ (PAO) circular
2021.11.19 © Huskens, M.-L.
Anurida granaria from Belgium
Post-antennal organ (PAO) circular
2021.11.23 © Huskens, M.-L.
Anurida granaria from Belgium
Post-antennal organ (PAO) circular
Enhanced contrast
2021.11.23 © Huskens, M.-L.
Anurida nr granaria from the USA
2023.10.28 © Wang, G.
Anurida nr granaria from the USA
2023.10.28 © Wang, G.
Anurida nr granaria from the USA
Ant.4 sensilla stalked at base
2023.10.28 © Wang, G.
Anurida nr granaria from the USA
Post-antennal organ
2023.10.28 © Wang, G.
Anurida nr granaria ♀ from the USA
2023.11.19 © Wang, G.
Anurida nr granaria ♀ from the USA
2023.11.19 © Wang, G.
Anurida nr granaria ♀ from the USA
Mouthcone appressed
2023.11.19 © Wang, G.
Anurida nr granaria ♀ from the USA
Genital area
2023.11.19 © Wang, G.
Anurida nr granaria ♀ from the USA
Unguiculus absent
2023.11.19 © Wang, G.

Species granulata Agrell, 1943, B

            Syn.: Micranurida sexpunctata Handschin, 1924, nec Axelson, 1902
            Syn.: Hypanurida mirabilis Denis, 1931
            Syn.: Hypanurida flavescens Kseneman, 1934
            Syn.: Micranurida handschini Gisin, 1944
            Syn.: Anurida hexophthalmica Stach, J, 1949:26


Species interior Fjellberg, 1985

Species iriei Yosii, R, 1970:4

Species kaya Babenko, AB, 1998:656

Species kazukoae Tamura & Zhao, 1997

             Syn.: Anurida kazokoae(sic) Yue, Q-Y et Yin, W-Y, 1999:222 ISS

Species kolymensis Tshelnokov, 1988

Species komi Babenko, AB, 1998:653

Species kyshyensis Babenko, A et Nakamori, T, 2018:575

Anurida kyshyensis from Russia
After Babenko A & Nakamori T, 2018 Fig.2-5

Species latosensillata Dunger, 1975

Species longilamellata Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:141

Species lvivska Babenko, AB, 1998:659

Species mara Christiansen, K et Bellinger, P, 1988:230

Species marchena Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:161

             Syn.: Anurida marchana(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 ISS

Species maritima (Guérin-Méneville, 1836 tt) Laboulbène, 1865, t.t., B

            Syn.: Podura ambulatoria Ström in Plateau, 1890
            Syn.: Anurida bisetosa Goto & Delamare-Deboutteville, 1953
            Syn.: Anurida bisetosa Gisin, H, 1960:89
            Syn.: Anurida bisetosa Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:5
            Syn.: Anurida bisetosa Massoud, Z, 1967:256
            Syn.: Anurida bisetosa Timmermans, MJTN, Arbea, JI, Campbell, G, King, MC, Prins, A et Kett, S, 2021:2
            Syn.: Anurida megalops Bagnall, 1949


Anurida maritima from Germany
Capitulum of maxilla
After Anonymous fig.C

             Syn.: Anurida maritime(sic) D'Haese, CA, 2003:581 ISS
Anurida maritima from France
2003 © De Bruyn, L.
Anurida maritima from the UK
2005 © Hopkin, S.P.
Anurida maritima from the UK
Post-antennal organ and eye with 5 ocelli
2005 © Hopkin, S.P.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.06 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Aggregation on tidal pool
2007.08.08 © Reed, D.E.
Anurida maritima from the USA
Circular post-antennal organ (PAO) with max 10 vesicles
2007.08.11 © Reed,D.E.
Anurida maritima from the UK
2008.01.06 © Roberts, S.
Anurida maritima from the UK
2008.03.30 © Roberts, S.
Anurida maritima from the UK
2008.06.29 © Ramone, J.
Anurida maritima from the UK
2008.07.27 © Roberts, S.
Anurida maritima from the USA
2009.02.21 © Edmonds, M.
Anurida maritima from France
2009.08.09 © Henderickx, H.
Anurida maritima from France
Egg cluster
2009.08.09 © Henderickx, H.
Anurida maritima from the USA
Aggregation on tidal pool
2013.07.21 © Kynd, F.

             Syn.: Anurida maritime(sic) Hensel, R, Neinhuis, C et Werner, C, 2015:7 ISS
Anurida maritima from the USA
Aggregation on tidal pool
2015.08.18 © Hasselman, K.
Anurida maritima from Portugal
Eyepatch with 5 ocelli
2016.08.06 © Almeida, J.
Anurida maritima from Holland
Mouthcone moderately pointed
2017.07.21 © van Duinen, J.
Anurida maritima from Holland
2017.08.04 © Wieringa, P.
Anurida maritima from Holland
With protruded mouthcone
2017.08.04 © Wieringa, P.
Anurida maritima from France
2019.08.31 © Tiky, S.
Anurida maritima from France
With protruded mouthcone
2019.08.31 © Tiky, S.
Anurida maritima from France
Anal spines absent
2019.08.31 © Tiky, S.
Anurida maritima from France
Unguiculus absent (*)
2019.08.31 © Tiky, S.
Anurida maritima
Jumping on water using tension of surfacefilm
2020.07.01 © Life Science Studios



Species maritima2 Arbea, JI, 2001:39

Species martynovae Fjellberg, 1985

Species multisensillata Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:135

Species nadezhdae Babenko, AB, 1998:654

Species neanuriformis Babenko, AB, 2002:369

Species nuda Yosii, 1956

            Syn.: Anurida granaria Uchida, 1952
            Syn.: Anurida granaria Yosii, R, 1977:147


Species olaensis Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:137

            Syn.: Anurida (weberi-group) olae(sic) Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:137 IOS
Remark by Janssens, F. 2021.02.22: According to the etymology the new species name is a toponym refering to the type locality, the Ola river. However, the genitive 'olae' is not a toponym but a patronym that refers to a female person with name 'Ola'. This meaning does not match the etymology. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). 'olaensis' is a toponym that refers to a place called 'Ola', which is in line with the etymology.

            Syn.: Anurida (weberi-group) olae(sic) Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:130 IOS
            Syn.: Anurida (weberi-group) olae(sic) Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:140 IOS
            Syn.: Anurida (weberi-group) olae(sic) Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:146 IOS

Species orientalis Martynova in Stebaeva, 1976 nn

Species pallens Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:131

Species palustris Babenko, AB, 1997:522

Species papillosa (Axelson, 1902) Fjellberg, 1985

Species papillosoides (Hammer, 1953) Massoud, Z, 1967:258

            Syn.: Micranurida papillosa Hammer, 1953 =?
            Syn.: Anurida papillosa Massoud, Z, 1967:265 =?
            Syn.: Anurida arctica Tshelnokov, 1988
            Syn.: Anurida arctica Babenko, AB, 1997:526 =?
            Syn.: Anurida arctica Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:146
            Syn.: Anurida violacea Tshelnokov in Babenko AB nn


             Syn.: Anurida papillosoïdes(sic) Massoud, Z, 1967:258 ISS

Species parapapillosa Tshelnokov, 1988

Species perplexa (Salmon, 1956) Janssens F, 2020

Species plurichaetotica Yosii, R, 1966:544

Anurida (Aphoromma) plurichaetotica from Korea
After Yosii R, 1966 Fig.2A

            Syn.: Anurida (Aphoromma) plurichaetotica ssp. decipiens Yosii, R, 1966:548



Species polaris (Hammer, 1954) Fjellberg, 1985

            Syn.: Anurida frigida Fjellberg, 1973
            Syn.: Anurida frigida Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:146
            Syn.: Anurida wrangelensis Tshelnokov, 1988
            Syn.: Anurida wrangelensis Babenko, AB, 1997:528


Species pusilla da Gama, 1964

Species riverina Sterzynska, M, Kaprus, I et Ehrnsberger, R, 2006:255

Anurida riverina from Poland
After Sterzynska M, Kaprus I & Ehrnsberger R, 2006 Fig.1

Species sakhensis Babenko, AB, 1997:529

Species setosa Yosii, R, 1956:27

Species similis Fjellberg, 1985

Species speobia Yosii, R, 1954:64

Species stereoodorata Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49

Anurida stereoodorata from Abkhazi
After Jordana R & al, 2012 Fig.11A

             Syn.: Anurida stereoodarata(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:49 ISS
Anurida stereoodorata from Europe
After Vargovitsh R in Fiera C & al, 2021 Fig.1a



Species subarctica Fjellberg, 1985

            Syn.: Anurida tajmyrica Tshelnokov, 1988
            Syn.: Anurida tajmyrica Babenko, AB, 1997:530


Species tamurae Yue, Q-Y et Yin, W-Y, 1999:224

            Syn.: Anurida tamurea(sic) Yue, Q-Y et Yin, W-Y, 1999:224 IOS

Species tatianae Babenko, AB, 1997:530

Species thalassophila (Bagnall, 1939) Stach, J, 1949:14

            Syn.: Anurida granaria Remy, 1928
            Syn.: Anurida remyi Denis, 1947


Species troglodyta Lee & Park, 2016

Species tullbergi Schött, H, 1891:192

            Syn.: Anurida Tullbergi(sic) Schött, H, 1891:192 IOS
            Syn.: Anurida crassicornis Reuter L & Reuter OM, 1879-1880 =?
            Syn.: Anurida ellipsoides Jordana, R, Arbea, JI, Simón, C et Luciáñez, MJ in Ramos, MA et al, 1997:778
            Syn.: Anurida ellipsoides Babenko, AB, 2021:il =e?
            Syn.: Anurida ellipsoides Jordana, R, 2021:il =e
            Syn.: Anurida brunsvigiensis Hüther, 1964
             Syn.: Anurida brusvigiensis(sic) Pomorski, RJ et Skarzynski, D, 1994:367 ISS



            Syn.: Anurida Tullbergi(sic) Schött, H, 1893:91 IOS
Anurida Tullbergii(sic) from Sweden
After Schött H, 1893 Taf.VII Fig.16


            Syn.: Anurida Tullbergi(sic) Reuter, OM, 1895:5 IOS
            Syn.: Anurida Tullbergi(sic) Reuter, OM, 1895:33 IOS
            Syn.: Anurida Tullbergi(sic) Reuter, OM, 1901:130 IOS
            Syn.: Anurida Tullbergi(sic) Schött, H, 1917:6 IOS
Anurida tullbergi from Spain
2009.04.21 © Pato, J.R.
Det.: Arbea, J.

             Syn.: Anurida tulbergii(sic) Shaw, P et Benefer, CM, 2015:200 ISS
Anurida tullbergi from Belgium
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Post-antennal organ (PAO) oval
Eyepatch with 5 ocelli
2017.05.20 © Huskens, M.-L.
Anurida tullbergi from Belgium
Ant.3+4 fused
2017.05.20 © Huskens, M.-L.





Species uniformis Gisin, 1953

             Syn.: Anurida uniformes(sic) Babenko, AB, 1998:654 ISS

Species vicaria Yosii, R, 1956:30

Species vontoernei Nosek, 1962

Species weberi Christiansen & Bellinger, 1980:335

            Syn.: Micranurida papillosa Hammer, 1953 nec Axelson, 1902
            Syn.: Micranurida papillosoides Hammer, 1953 =?


Species zanokhae Babenko, AB, 1998:649


Species-group hammerae
            Syn.: Anurida hammerae Fjellberg, 1985 gr

Species sp.1 Janssens F sensu Babenko & Nakamori, 2021

            Syn.: Anurida (hammerae-group) sp. gr. Babenko, A et Nakamori, T, 2021:371
Anurida sp. gr hammerae from Russia
After Babenko A & Nakamori T, 2021 Fig.11

            Syn.: Anurida (hammerae-group) sp. juv. undescribed Babenko, A et Nakamori, T, 2021:372



Species sp.2 Janssens F sensu Babenko & Nakamori, 2021

            Syn.: Anurida (hammerae-group?) trioculata Babenko, A et Nakamori, T, 2021:371,cf
Anurida cf. trioculata from Japan
After Babenko A & Nakamori T, 2021 Fig.12-13




Species beringi Fjellberg, 1985

            Syn.: Anurida tullbergi Hammer, 1953
            Syn.: Anurida (Anurida) tullbergi Christiansen, K et Bellinger, P, 1980:323
            Syn.: Anurida nerensis Tshelnokov, 1988
            Syn.: Anurida nerensis Babenko, AB, 1997:517
             Syn.: Anurida neriensis(sic) Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:145 ISS


Species hammerae Christiansen, KA, 1952:135

            Syn.: Anurida hammeri(sic) Christiansen, KA, 1952:135 IOS
Anurida hammeri(sic) from Alaska
After Christiansen K, 1952 Pl.12 Fig.1-2

            Syn.: Anurida mandibulata Tshelnokov, 1988
            Syn.: Anurida mandibulata Babenko, AB, Shveenkova, YB et Kuznetsova, NA, 2019:145
            Syn.: Anurida polychaetosa Tshelnokov, 1988
            Syn.: Anurida polychaetosa Babenko, AB, 1997:519 =?



             Syn.: Anurida hammeri(sic) Massoud, Z, 1967:265 ISS
             Syn.: Anurida (Anurida) hammeri(sic) Christiansen, K et Bellinger, P, 1980:330 ISS
             Syn.: Anurida hammeri(sic) Christiansen, KA in Dindal, DL, 1990:978 ISS
             Syn.: Anurida hammeri(sic) Poole, RW in Poole, RW et Gentili, P, 1997:47 ISS



Species hirsuta Babenko, A et Nakamori, T, 2021:368

Anurida hirsuta from Russia
After Babenko A & Nakamori T, 2021 Fig.8

Species luciae Fjellberg, A, 1985:359

            Syn.: Anurida macrochaetosa Tshelnokov, 1988
            Syn.: Anurida macrochaetosa Babenko, AB, 1997:520


Species narli Fjellberg, 1985

            Syn.: Anurida tshuktshorum Tshelnokov, 1988
            Syn.: Anurida tshuktshorum Babenko, AB, 1997:520


Species okamotoi Yosii, R, 1970:2

Species reducta Fjellberg, 1985

Species trioculata Kinoshita, 1916

Species vulcanica Fjellberg, A, 1985:357


Genus Minotaurella Weiner, WM, 1999:373

Species banyulensis (Denis, 1947) Weiner, WM, 1999:376

Species edaphica Kaprus, IJ et Nevo, E, 2003:67

Minotaurella edaphica from Israel
After Kaprus I.J. & Nevo E, 2003 Fig.2a

Species isabellae Weiner, WM, 1999:374, t.t.

             Syn.: Minotaurella isabelle(sic) Kaprus, IJ et Nevo, E, 2003:69 ISS



Genus Anuridella Willem, 1906

  Syn.: ... Anurodes Bagnall, 1949
Anuridella sp. from the UK
2013.08.23 © Fenwick, D.



Species sp. Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:892

Species calcarata Denis, 1925:267

            Syn.: Anuridella immsiana Bagnall, 1939


Species germanica (Gisin, 1951)

Species marina Willem, V, 1906:247, t.t.

Anuridella marina
After Willem V, 1906
            Syn.: Anuridella hintoni Bagnall, 1949



Species submarina Bagnall, 1934


Genus Gastranurida Bagnall, 1949

Species denisi (Bagnall, 1939) Massoud, Z, 1967:268, t.t.

            Syn.: Anurida trioculata Denis, 1922, nec Kinoshita, 1916
            Syn.: Paranura hibernica Bagnall, 1941
             Syn.: Gastranurida gisini(sic) Babenko, A et Nakamori, T, 2018:578 lapsus ISS



Genus Hylaeanura Arlé, R, 1966:5 key  key

  Syn.: Pseudachorutinae Hylaenura(sic) Mari Mutt, JA et Bellinger, PF, 1990:43 ISS
  Syn.: Neanuridae Hyleanura(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS

Species emiliae Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2020:33

Hylaeanura emiliae from Puerto Rico
After Ospina-Sánchez CM, Palacios-Vargas JG & González G, 2020 Fig.1

            Syn.: Hylaeanura sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:9



Species infima (Arlé, R, 1960:163) Arlé, R, 1966:5, t.t.

Species mendoncae Zeppelini, D et Palacios-Vargas, JG, 2013:1402

Species nepalensis (Yosii, 1966) Vázquez, M, Cutz-Pool, LQ & Palacios-Vargas, JG, 1998:370

Species nohbecana Vázquez, M, Cutz-Pool, LQ et Palacios-Vargas, JG, 1998:367


Genus Anurachorutes Kuznetsova & Potapov, 1988

Species martynovae Kuznetsova & Potapov, 1988


Genus Anuritelsa Womersley, H, 1939:315

Species maritima Womersley, H, 1939:316, t.t.


Genus Gamachorutes Cassagnau, 1978

Species verrucosus Cassagnau, 1978


Genus Koreanurina Najt & Weiner, 1992

Species alba Wu, Bedos & Deharveng, 2014

Species inexpectata Najt & Weiner, 1992

Species szeptyckii Najt & Weiner, 1992

Species weinerae Wu, Bedos & Deharveng, 2014


Genus Lanzhotia Rusek, 1985

Species brachycera Rusek, 1985


Genus Paranurida Skarzynski, D et Pomorski, RJ, 1994:363

Species kopetdagi Skarzynski, D et Pomorski, RJ, 1994:364


Genus Philotella Najt, J et Weiner, WM, 1985:29 key

Species sp.1 Janssens F sensu Najt & Weiner, 1997

            Syn.: Philotella sp.n. Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:41


Species alba Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species aliniensis Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species amurica Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species caeca Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species cedrina Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species deharvengi Najt, J et Weiner, WM, 1985:30, t.t.

Species grana (Lee & Kim, 1984) Babenko, A, Shveenkova, Y & Potapov, M, 2022:?

Species japonica (Tamura, 1998) Babenko, A, Shveenkova, Y & Potapov, M, 2022:?

Species miracli Najt, J et Weiner, WM, 1985:32

             Syn.: Philotella miracle(sic) Babenko, AB, 2007:364 ISS

Species obesa Babenko, AB, 2007:362

Species olgae Kniss & Thibaud, 1995

Species oligotricha Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species pallida Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species pax Babenko, A, Shveenkova, Y et Potapov, M, 2022:?

Species porcella (Fjellberg, 1985) Babenko, A, Shveenkova, Y & Potapov, M, 2022:?

Species tertia Najt, J et Weiner, WM, 1985:34


Genus Rusekella Deharveng, 1982

Species cantabrica Deharveng, 1982

Species cassagnaui Deharveng, 1982

Species flagellata (Arbea, JI & Jordana, R, 1985:64) Arbea, JI & Jordana, R, 1991:99

Species gamae Arbea, JI et Jordana, R, 1991:98

Rusekella gamae from Portugal
After Salinas JA in Arbea JI & Jordana R, 1991 Fig.1A
Rusekella gamae from Portugal
Ocellus H missing
Modified after Salinas JA in Arbea JI & Jordana R, 1991 Fig.1A

Species nivalis Deharveng, 1982

Species peyrei (Cassagnau, P, 1955:157)

            Syn.: Micranurida Peyrei(sic) Cassagnau, P, 1955:157 IOS

Species similis Deharveng, 1982

Rusekella similis from France
2006 © Boldsystems.org


Genus Pongeia Najt & Weiner, 2002

Species asturianensis Najt & Weiner, 2002, t.t.


Genus Holacanthella Börner, C, 1906:169 key

Holacanthella spinosa and duospinosa from New Zealand
2014 © Murray, A.
  Syn.: Pseudachorutinae Holocanthella(sic) Greenslade, P, 2015:5 ISS


Species brevispinosa (Salmon, JT, 1942:256) Massoud, Z, 1967:222

Ceratrimeria brevispinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.1
Holacanthella brevispinosa from New Zealand
2008 Minor, M. & Robertson, A.© SoilBugs
Holacanthella brevispinosa from New Zealand
2014 © Murray, A.
Holacanthella brevispinosa from New Zealand
2014 © Murray, A.
Holacanthella sp. from New Zealand
With longer purple dorsal digitations
After Ashwood F & Janssens F, 2024 Fig.1 top
2024 © Ashwood, F.
Holacanthella sp. juv. from New Zealand
With longer purple dorsal digitations
After Ashwood F & Janssens F, 2024 Fig.1 middle
2024 © Ashwood, F.
Holacanthella sp. from New Zealand
With longer purple dorsal digitations
After Ashwood F & Janssens F, 2024 Fig.1 bottom
2024 © Ashwood, F.
Holacanthella sp. from New Zealand
With longer purple dorsal digitations
After Ashwood F & Janssens F, 2024 Fig.2 top
2024 © Ashwood, F.

Species duospinosa (Salmon, JT, 1942:257) Massoud, Z, 1967:222

Ceratrimeria duospinosa from New Zealand
After Salmon JT, 1942 Pl.43 Fig.9
Holacanthella duospinosa from New Zealand
2007 © Stevens, M.
Holacanthella duospinosa from New Zealand
2014 © Murray, A.
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
2014.03.15 © McQuillan., B
Holacanthella duospinosa from New Zealand
Ant.1 & 2 with apically pointed digitations
2014.05.26 © Murray, A.
Holacanthella duospinosa from New Zealand
2016.04.03 © Sullivan, J.
Holacanthella duospinosa from New Zealand
2017.06.13 © Maurice.
Holacanthella duospinosa from New Zealand
2017.12.13 © Maurice.
Holacanthella duospinosa from New Zealand
2020.07.31 © Maurice.
Holacanthella duospinosa from New Zealand
2020.07.31 © Maurice.
Holacanthella duospinosa from New Zealand
In defensive 'hedgehog' posture
2020.07.31 © Maurice.
Holacanthella duospinosa from New Zealand
2020.12.30 © Huang, D.

Species laterospinosa (Salmon, JT, 1944:132) Massoud, Z, 1967:222

Holacanthella laterospinosa from New Zealand
2014 © Murray, A.
Holacanthella laterospinosa juv. from New Zealand
2014 © Murray, A.

Species paucispinosa (Salmon, JT, 1941:300) Massoud, Z, 1967:222

Ceratrimeria paucispinosa from New Zealand
After Salmon JT, 1941 Pl.44 Fig.81
Holacanthella paucispinosa from New Zealand
2006 Minor, M. & Robertson, A.© SoilBugs
Holacanthella paucispinosa from New Zealand
2006 © Stevens, M.
Holacanthella paucispinosa from New Zealand
After Thibaud J-M & D'Haese C, 2010 Fig.33
2009 © D'Haese, C.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.01.28 © Thorpe, S.E.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
Reduced furcula
2010.08.14 © Winter, D.
Holacanthella paucispinosa from New Zealand
2013.08.31 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2013.09.22 © Jewell, T.
Holacanthella paucispinosa juv. from New Zealand
2014 © Murray, A.
Holacanthella paucispinosa from New Zealand
2014.mm.dd © Stevens, M.
Holacanthella paucispinosa from New Zealand
2014.02.09 © Kerr, S.
Holacanthella paucispinosa from New Zealand
2016.05.15 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
2016.06.22 © Navest, M.P.
Holacanthella paucispinosa from New Zealand
Head frontoventral aspect
2016.06.22 © Navest, M.P.

Species spinosa (Lubbock, 1899) Börner, C, 1906:169, t.t.

Ceratrimeria spinosa from New Zealand
After Salmon, J.T., 1941 Pl.44 Fig.77
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2007.11.21 © Pselaphid.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa from New Zealand
2009.11.08 © Ronny & Raviv.
Holacanthella spinosa juv. from New Zealand
Freshly hatched
2014 © Murray, A.
Holacanthella spinosa juv. from New Zealand
2014 © Murray, A.
Holacanthella spinosa subadult from New Zealand
2014 © Murray, A.
Holacanthella spinosa from New Zealand
2014.05.15 © Murray, A.
Holacanthella spinosa juv. from New Zealand
Pointed mouthcone
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
Tubercles
2014.05.15 © Murray, A.
Holacanthella spinosa from New Zealand
2020.07.29 © Knox, C.


Genus Acanthanura Börner, 1906

  Syn.: ... Anoura(sic) Lubbock, 1899 partim IOS
  Syn.: ... Ceratrimeria Börner, 1906 partim


Acanthanura sp. from Australia
2007.11.19 © Midtgaard, R.
Acanthanura sp. from Tasmania
Defreezing
2023.09.09 © Ellingsen, K.



Species sp.1 Garrick, RC, Sands, CJ, Rowell, DM, Hillis, DM et Sunnucks, P, 2006:1

Acanthanura sp. from Australia
2006 © Sunnucks, P.

Species sp.2 Janssens F & Murray A, 2014

Acanthanura "of Henderickx" from Tasmania
2007 © Henderickx, H.
Acanthanura "of Henderickx" from Tasmania
2012.12.29 © Zosterops, K.
Acanthanura "of Henderickx" from Australia
4th antennal segment
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Australia
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Australia
Pointed mouth-cone
2014.03.10 © Murray, A.
Acanthanura "of Henderickx" from Australia
Ocular tubercle with 7 ocelli in circular formation
Post-antennal organ
2014.09.25 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2014.11.09 © Grove, S.
Acanthanura "of Henderickx" from Australia
2015 © Murray, A.
Acanthanura "of Henderickx" from Australia
Pointed mouthcone
2015 © Murray, A.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.
Acanthanura "of Henderickx" from Tasmania
2017.02.10 © Satler, J.
Acanthanura "of Henderickx" from Tasmania
2019.12.21 © Grove, S.
Acanthanura "of Henderickx" from Tasmania
Ocular tubercle with 7 ocelli in circular formation
2020.01.01 © Grove, S.
Acanthanura "of Henderickx" from Tasmania
2020.05.04 © Rogers, J.
Acanthanura "of Henderickx" from Tasmania
2022.11.30 © Grove, S.

Species sp.3 Greenslade P, 2015:il

Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. juv. from Tasmania
2014.03.29 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.10.28 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2014.11.14 © Murray, A.
Det.: Greenslade, P., 2015.06.01
Acanthanura sp. from Tasmania
2015 © Murray, A.
Acanthanura sp. from Tasmania
In defensive posture
2015 © Murray, A.
Acanthanura sp. from Tasmania
2020.07.09 © Ellingsen, K.
Acanthanura sp. from Tasmania
2020.07.09 © Ellingsen, K.
Acanthanura sp. from Tasmania
Cephalic digitations not pointed
2020.07.09 © Ellingsen, K.
Acanthanura sp. from Tasmania
Cephalic digitations not pointed
2020.07.09 © Ellingsen, K.

Species sp.4 Janssens F & Huang D, 2019

Acanthanura sp. from Tasmania
2017.11.30 © Huang, D.
Acanthanura sp. from Tasmania
2017.11.30 © Huang, D.
Acanthanura sp. from Tasmania
Head and thorax
2017.11.30 © Huang, D.

Species sp.5 Janssens F & Murray A, 2021

Acanthanura sp. from Australia
2015 © Murray, A.

Species dendyi (Lubbock, 1899) Börner, 1906, t.t.

            Syn.: Anoura(sic) Dendyi(sic) Lubbock, 1899 IOS
             Syn.: Neanura Dendyi(sic) Absolon, K, 1901:576 ISS
             Syn.: Acanthanura Dendyi(sic) Börner, 1906 ISS
             Syn.: Anoura(sic) Dendyi(sic) Schött, H, 1917:1 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1937 ISS
             Syn.: Ceratrimeria Dendyi(sic) Womersley, 1939 ISS
             Syn.: Acanthanura Dendyi(sic) Handschin, 1942 tt ISS
Acanthanura dendyi from Tasmania
2010.07.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010.12.dd © LLoyd, S.
Acanthanura dendyi from Tasmania
2010 © Morris, R.
Acanthanura dendyi from Tasmania
Tuberculate cuticula
2014.09.14© Murray, A.
Det.: Murray, A.






Genus Megalanura Ellis, WN et Bellinger, PF, 1973:30

  Syn.: ... Meganura Handschin, 1942, nec Szepligéti, 1906

Species tasmaniae (Lubbock, 1899 tt) Ellis, WN & Bellinger, PF, 1973:30

            Syn.: Anoura(sic) tasmania(sic) Lubbock, 1899 tt IOS
             Syn.: Anoura(sic) tasmaniæ(sic) Dendy, A, 1900:98 ISS
             Syn.: Neanura Tasmaniae(sic) Absolon, K, 1901:576 ISS
             Syn.: Anoura(sic) Tasmaniae(sic) Schött, H, 1917:1 ISS
Megalanura tasmaniae from Tasmania
2010 Morris, R.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Ventral aspect
2014.04.06 © Murray, A.
Megalanura tasmaniae from Tasmania
Composed eye with 8 ocelli; PAO elliptic
2014.09.26 © Murray, A.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.
Megalanura tasmaniae from Tasmania
2014.11.09 © Grove, S.
Megalanura tasmaniae from Tasmania
2015 © Murray, A.
Megalanura tasmaniae from Tasmania
2015 © Murray, A.
Megalanura tasmaniae from Tasmania
2020.01.17 © Grove, S.






Genus Womersleymeria Stach, J, 1949:64

Womersleymeria sp. from Australia
2011.05.30 © Selemba, A.

Species sp.1 Janssens F & Reiner, 2020

Womersleymeria sp. from Australia
2015 © Murray, A.
Womersleymeria sp. from Australia
2015.07.27 © Porch, N.
Womersleymeria sp. from Australia
2019.09.22 © Invertophiles.
Womersleymeria sp. from Australia
2020.06.07 © Reiner.
Womersleymeria sp. from Australia
2020.06.07 © Reiner.
Womersleymeria sp. from Australia
2020.06.07 © Reiner.

Species bicornis (Womersley, H, 1940:137) Stach, J, 1949:64, t.t.

Ceratrimeria bicornis from Tasmania
After Womersley H, 1940 Fig.A
Womersleymeria bicornis from Tasmania
2010 Morris, R.
Womersleymeria bicornis from Tasmania
Aberration with fused digitations of abd.5
2014.09.22 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Pointed mouth-cone
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
Post-antennal organ (PAO)
2014.10.25 Murray, A.
Womersleymeria bicornis from Tasmania
2014.11.14 © Murray, A.
Womersleymeria bicornis from Tasmania
2015 Murray, A.
Womersleymeria bicornis from Tasmania
2015 Murray, A.
Womersleymeria bicornis from Tasmania
Caudal aspect
Producing faecal pellet
2015 Murray, A.
Womersleymeria bicornis from Tasmania
2015 Murray, A.
Womersleymeria bicornis from Australia
Pointed mouthcone
2017 Murray, A.
Womersleymeria bicornis from Tasmania
2020.02.01 © Porch, N.

References