http://www.collembola.org/taxa/pteninae.htm - Last updated on 2018.12.31 by Frans Janssens
Checklist of the Collembola: Ptenothricinae

Subfamilia Ptenothricinae Richards WR, 1968:25 sensu Bretfeld, G, 1999:128

Genus Number
of species
Bothriovulsus4
Papirioides12
Ptenothrix89
Ptenothricinae105
Genera with number of species
  Syn.: Dicyrtomidae Pterothricinae(sic) Deharveng, L, 2004:427 ISS
Ptenothrix sp. from Canada
Anterior dorsal spinelike setae
2008.12.11 © McCann, S.
Ptenothrix sp. from the USA
Two thick unpaired facial setae
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from New Zealand
Abd.6 dorsal macrosetae pointing upwards (*)
2016.02.04 © Murray, A.




Genus Ptenothrix Börner, C, 1906:184

F
  Syn.: ... Papirius Lubbock, 1862 partim
  Syn.: Dicyrtoma fusca Yosii, 1940
  Syn.: Dicyrtoma fusca Yosii, R, 1977:166
  Syn.: Dicyrtomidae Dicyrtoma Greenslade, P, Boyer, S, Shields, MW et Wratten, SD, 2016:5 partim


  Syn.: Sminthuridae Ptenothryx(sic) Yosii, R, 1956:101 ISS
  Syn.: ... Ptenothryx(sic) Yosii, 1961 ISS
Ptenothrix sp. from the USA
Long tube-like everted vesicles of collophore
2009.03.30 © Bradford, A.
Ptenothrix sp. from the USA
Apical part of 3rd antennal segment annulated
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from Canada
Ocelli with transparent crystal lenses
2011.10.26 © Hochhalter, R.
Ptenothrix sp. from Canada
"Eyebrow" and composed eye
2012.03.24 © Hochhalter, R.
Ptenothrix sp. from Canada
Footcomplex with long empodium
2012.05.07 © Hochhalter, R.
Ptenothrix sp. from Canada
Two thick unpaired midfacial macrosetae (*)
2015.10.30 © Barbin, T.
Ptenothrix maculosa from the USA
Abd.6 with spinelike macrosetae : 3M+2N
2016.03.08 © Dobbs, Br.



Species sp.3 Janssens F & Moorehead C, 2008

            Syn.: Ptenothrix sp.3 f.1 Janssens F & Moorehead C, 2015
Remark by Janssens, F. 2015.01.dd: Middorsal posterior patch split in 2 patches.
Ptenothrix sp. from the USA
2007.12.25 © Moorehead, C.
Ptenothrix sp. from the USA
2009.01.12 © Davis, J.
Ptenothrix sp. from the USA
2009.01.12 © Davis, J.
Ptenothrix sp. from the USA
2010.02.05 © Schimming, L.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. from the USA
Split butt patch
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
Split butt patch
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.12.10 © Dobbs, Br.
Ptenothrix sp. from the USA
Tubelike bipartite collophore vesicle:
warted apical part + smooth basal part
2018.01.15 © Dobbs, Br.

            Syn.: Ptenothrix sp.3 f.2 Janssens F & Leppin C, 2015
Remark by Janssens, F. 2015.01.dd: Middorsal posterior patch reduced to middorsal stripe.
Ptenothrix sp. from the USA
2009.01.01 © Leppin, M.
Ptenothrix sp. from the USA
2009.01.01 © Leppin, M.

            Syn.: Ptenothrix sp.3 f.3 Janssens F & Kyron TF, 2016
Remark by Janssens, F. 2016.03.14: Middorsal posterior patch fused with lateral pigmentation.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. ♀ from Canada
2017.01.21 © Barbin, T.

            Syn.: Ptenothrix sp.3 f.4 Janssens F & Dobbs Br, 2016
Remark by Janssens, F. 2016.12.27: Middorsal posterior patch with many cross bars.
Ptenothrix sp. from the USA
2016.11.25 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.25 © Dobbs, Br.

            Syn.: Ptenothrix sp.3 f.5 Janssens F & Dobbs Br, 2018
Remark by Janssens, F. 2018.01.11: With broad pale longitudinal middorsal band.
Ptenothrix sp. from Canada
2010.01.04 © McCann, S.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.10 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.11 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.11 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2018.01.08 © Dobbs, Br.




Remark by Janssens, F. 2018.01.11: Principal form with pale middorsal anterior stripe and dark middorsal posterior patch.
Ptenothrix sp. from the USA
2006.12.03 © Moorehead, C.
Ptenothrix sp. from the USA
2006.12.10 © Moorehead, C.
Ptenothrix sp. from the USA
2006.12.10 © Moorehead, C.
Ptenothrix sp. from the USA
2007.03.08 © Moorehead, C.
Ptenothrix sp. from the USA
2007.03.08 © Moorehead, C.
Ptenothrix sp. from the USA
2007.03.13 © Moorehead, C.
Ptenothrix sp. from the USA
2007.11.26 © Moorehead, C.
Ptenothrix sp. from the USA
2007.12.09 © Moorehead, C.
Ptenothrix sp. from the USA
2007.12.25 © Moorehead, C.
Ptenothrix sp. from the USA
2008.02.03 © Moorehead, C.
Ptenothrix sp. from the USA
2008.02.03 © Moorehead, C.
Ptenothrix sp. from the USA
2008.02.03 © Moorehead, C.
Ptenothrix sp. from Canada
2008.11.27 © McCann, S.
Ptenothrix sp. from Canada
2008.12.11 © McCann, S.
Ptenothrix sp. from Canada
Two unpaired facial setae
2008.12.11 © McCann, S.
Ptenothrix sp. from the USA
2009.01.04 © Leppin, M.
Ptenothrix sp. from the USA
2009.01.04 © Leppin, M.
Ptenothrix sp. from the USA
2009.02.06 © Hall, K.
Ptenothrix sp. from the USA
2009.02.20 © Hall, K.
Ptenothrix sp. from the USA
2009.02.20 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.31 © LeMons, J.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. juv. from Canada
2013.01.17 © Kyron, T.F.
Ptenothrix sp. from the USA
2015.01.20 © Faberha, J.
Ptenothrix sp. from the USA
2015.02.15 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.01.21 © Dobbs, Br.
Ptenothrix sp. ♀ from the USA
2016.01.21 © Dobbs, Br.
Ptenothrix sp. from the USA
Two unpaired facial setae
2016.01.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Juvenile and adult
2016.02.22 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.02.23 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.02.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.02 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. ♂ from the USA
Raised male genital papilla (*)
2016.11.06 © Johnson, S.T.
Ptenothrix sp. ♂ from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix sp. from Canada
2018.01.12 © Barbin, T.

Species sp.4 Janssens F & Moorehead C, 2008

Ptenothrix sp. from the USA
2007.12.24 © Moorehead, C.
Ptenothrix sp. from Canada
2008.12.01 © McCann, S.
Ptenothrix sp. from Canada
2008.12.02 © McCann, S.
Ptenothrix sp. from the USA
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.19 © Haley, R.
Ptenothrix sp. from the USA
2016.03.12 © Necturus.
Ptenothrix sp. from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Dark form
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Cephalic dorsal pattern
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Collophore vesicles extended
2017.10.26 © Dobbs, Br.
Ptenothrix sp. from the USA
Caudal aspect
2017.10.26 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.26 © Dobbs, Br.

Species sp.5 Janssens F & LeMons J, 2008

Ptenothrix sp. from the USA
2007.03.05 © LeMons, J.
Ptenothrix sp. from the USA
2007.03.05 © LeMons, J.
Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from Canada
2016.02.09 © Barbin, T.
Ptenothrix sp. from the USA
With pale femur and dark tibiotarsus
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.12.03 © Johnson, S.T.

Species sp.6 Janssens F & Ten Eyck C, 2010

Ptenothrix sp. from the USA
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from the USA
Two thick unpaired facial setae
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from Canada
2011.03.21 © Hochhalter, R.
Ptenothrix sp. from Canada
2011.03.26 © Hochhalter, R.
Ptenothrix sp. from Canada
2011.03.26 © Hochhalter, R.
Ptenothrix sp. from Canada
2016.02.08 © Barbin, T.
Ptenothrix sp. from Canada
2016.02.08 © Barbin, T.
Ptenothrix sp. from the USA
2016.11.18 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.18 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.11.30 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.11.29 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.01.02 © Dobbs, Br.

Species sp.7 Janssens F & Alexander C, 2012

Ptenothrix sp. from the USA
2012.12.27 © Alexander, C.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. from the USA
2017.11.27 © Dobbs, Br.

Species sp.8 Janssens F & Ten Eyck, 2008

Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
Unguis with outer tooth and 2 lateral teeth
2010.01.10 © Ten Eyck, C.

Species sp.9 Janssens F & Kyron TF, 2013

Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.

Species sp.10 Janssens F & Dobbs B, 2016

Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
Unguiculus of 3rd leg with long knobbed filament
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Abd.6 dorsally with 3M+2N spinelike setae
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Eyepatch with reddish background
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Two unpair midfacial setae
2016.11.27 © Dobbs, Br.

Species sp.11 Janssens F & Dobbs B, 2017

Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
Antennae
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Antenna pigmentation
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Abd.6 with dark dorsal patch
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.02.22 © Dobbs, Br.

Species sp.12 Janssens F & Dobbs B, 2017

Ptenothrix sp. from the USA
Dark uniform body colour
2017.11.06 © Dobbs, Br.
Ptenothrix sp. from the USA
Pale ant.1+2; pale tibiotarsi
2017.11.06 © Dobbs, Br.
Ptenothrix sp. from the USA
Pale mouth region
2017.11.06 © Dobbs, Br.

Species albosignata (Parona, 1888) Stach, J, 1957:90

Species annulata Lin & Xia, 1985, nec Becker

Species argentina Delamare Deboutteville & Massoud, 1963:281

Ptenothrix argentina from Peru
2009.01-03.dd © Contreras Valdez, A.

Species atra (Linnæus, C, 1758:608) Börner, 1906, t.t., B

            Syn.: Podura antennis longioribus Hill, 1752 i
            Syn.: Podura polypoda Linnæus, 1761
             Syn.: Podura Polypoda(sic) Walckenær, CA, 1802:4 ISS
Papirius polypodus from the UK
After Lubbock, J., 1873 Pl.11.

            Syn.: Podura globosa fusca de Geer, C, 1778:35
            Syn.: Papirius chloropus Stach, J, 1957:84 =?,si
            Syn.: Papirius annulatus Becker, 1902
            Syn.: Dicyrtoma setosa-ceratoleucaspis Börner, 1903
            Syn.: Ptenothrix setosa Denis, 1931



            Syn.: Podura atra Linnæus, 1746 i


Ptenothrix atra from the UK
2004 © Hopkin, S.P.
Ptenothrix atra from the UK
2006.06.22 © Botting, J.
Ptenothrix atra from Estonia
2006.09.17 © Tartes, U.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from France
2009.09.dd © Lebeaux, P.
Ptenothrix atra juv. from the UK
2010.01.16 © Henton-Jones, D.
Ptenothrix atra juv. from Holland
2011.12.02 © van Duinen, J.
Ptenothrix atra from the UK
Trunk dorsal anterior macrosetae strongly spinelike
2013.02.10 © Prince, M.
Ptenothrix atra from the UK
2015.02.17 © Murray, A.
Ptenothrix atra from the UK
Ant.4 short and white
2015.02.17 © Murray, A.
Ptenothrix atra juv. from the UK
2015.02.17 © Murray, A.
Ptenothrix atra from the UK
2015.02.18 © Murray, A.
Ptenothrix atra from the UK
2015.02.18 © Murray, A.
Ptenothrix atra from the UK
Posterior dorsal great abdomen
2015.02.18 © Murray, A.
Ptenothrix atra from Sweden
2016.07.26 © Al Abbasi, S.
Ptenothrix atra from the UK
First instar 0.4mm
2016.12.03 © Holla, D.

Species australis Snider, RJ, 1990:247

Ptenothrix (Ptenothrix) australis from the USA
After Snider, R., 1990 Fig.1.
Ptenothrix (Ptenothrix) australis from the USA
After Snider, R., 1990 Fig.2.




             Syn.: Ptenothrix (Ptenothrix) austalis(sic) Snider, RJ, 1990:247 ISS

Species beta (Christiansen, K & Bellinger, P, 1981:1233) Christiansen, K & Bellinger, P, 1998:1407

Ptenothrix (Ptenothrix) beta from the USA
After Christiansen K & Bellinger P, 1998 Fig.1035A
Ptenothrix beta from the USA
2004 © Maddison, D.R.
Ptenothrix beta from the USA
2004 © Maddison, D.R.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.10.28 © Gross, J.
Ptenothrix beta from the USA
2007.10.28 © Gross, J.
Ptenothrix beta from the USA
2007.11.07 © Ten Eyck, C.
Ptenothrix beta from the USA
2007.12.25 © Moorehead, C.
Ptenothrix beta from the USA
2007.12.25 © Moorehead, C.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.12.09 © Bradford, A.
Ptenothrix beta from the USA
2009.03.30 © Bradford, A.
Ptenothrix beta from the USA
2009.04.01 © Bradford, A.
Ptenothrix beta from the USA
2009.04.08 © Zungri, D.
Ptenothrix beta from the USA
2009.10.22 © Ten Eyck, C.
Ptenothrix beta from the USA
2009.10.22 © Ten Eyck, C.
Ptenothrix beta from the USA
2010.01.02 © LeMons, J.
Ptenothrix beta from the USA
2010.01.03 © Zungri, D.
Ptenothrix beta from the USA
2010.01.07 © LeMons, J.
Ptenothrix beta from the USA
2010.01.21 © Bradford, A.
Ptenothrix beta from the USA
2010.06.11 © Matthews, E.
Ptenothrix beta from the USA
2012.02.26 © Elliot.
Ptenothrix beta from the USA
Pale colour form
2012.07.11 © Goff, M.
Ptenothrix beta from the USA
2012.10.06 © Baker, A.
Ptenothrix beta from the USA
2016.02.16 © Lee, C.
Ptenothrix beta juv. from the USA
Preparing to moult
2017.01.02 © Abela, A.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from Canada
2017.05.17 © Barbin, T.
Ptenothrix beta from Canada
2017.05.17 © Barbin, T.
Ptenothrix beta from Canada
2017.05.17 © Barbin, T.
Ptenothrix beta from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix beta from the USA
2018.03.18 © Schneider, K.
Ptenothrix beta from the USA
2018.03.18 © Schneider, K.

Species blidana Bretfeld, 2001

Species borincana Soto-Adames, FN, 1988:65

Ptenothrix borincana from Puerto Rico
After Soto-Adames FN, 1988 Fig.34

Species brasiliensis Delamare Deboutteville & Massoud, 1963:285

Species brouquissei Nayrolles, 1989

Species californica (Christiansen, K & Bellinger, P, 1981:1234) Christiansen, K & Bellinger, P, 1998:1408

            Syn.: ... marmorata Scott & Yosii, 1972


Ptenothrix (Ptenothrix) californica from the USA
After Christiansen K & Bellinger P, 1998 Fig.1036A
Ptenothrix cf. californica from the USA
2005 © Leonard, W.
Ptenothrix californica from the USA
2009.12.31 © Zungri, D.

Species castanea Snider, RJ, 1985:582

Dicyrtoma (Ptenothrix) castanea from the USA
After Snider RJ, 1985 Fig.1
Dicyrtoma (Ptenothrix) castanea from the USA
After Snider RJ, 1985 Fig.2
Ptenothrix castanea from Canada
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.14 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.14 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.24 © Hochhalter, R.
Ptenothrix castanea from Canada
Annulated 3rd and 4th antennal segments
2011.07.26 © Hochhalter, R.
Ptenothrix castanea from the USA
2014.04.28 © Peterson, M.A.

Species catenata (Schött, 1921) Richards, 1956

Species cavicola Cassagnau, P et Delamare Deboutteville, C, 1955:392

Species ciliata Stach, J, 1957:102

Ptenothrix ciliata from Poland
After Stach J, 1957 Pl.VIII fig.1
            Syn.: Ptenothrix atra Berg, M, 2008:50
Ptenothrix atra from Holland
After Berg, M, 2008 Fig.1b
Ptenothrix atra from Holland
After Berg, M, 2008 Fig.1a




Ptenothrix ciliata from Holland
After Berg, M, 2009 Fig.1

Species ciliophora Yosii, R et Lee, C-E, 1963:17

Ptenothrix ciliophora from Korea
After Yosii R & Lee C-E, 1963 Fig.9A

Species corynophora Börner, C, 1909:130

            Syn.: Ptenothrix corynophora var. cincta Börner, C, 1909:130
            Syn.: Ptenothrix corynephora(sic) var. cincta Yosii, R, 1977:166
            Syn.: Ptenothrix corynophora var. sellata Börner, C, 1909:130
            Syn.: Ptenothrix corynephora(sic) var. sellata Yosii, R, 1977:166
            Syn.: Ptenothrix corynephora(sic) f. shibanaii Yosii, 1955
            Syn.: Ptenothrix corynephora(sic) f. shibanaii Yosii, R, 1977:166
            Syn.: Ptenothrix tokarensis Yosii, 1965


             Syn.: Ptenothrix corynephora(sic) Yosii, 1954 ISS
             Syn.: Ptenothrix corynephora(sic) Stach, J, 1957:78 ISS
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4A
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4B
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4C

             Syn.: Ptenothrix corynephora(sic) Yosii, R et Lee, C-E, 1963:26 ISS
Ptenothrix corynephora(sic) from Japan
After Yosii R & Lee E-C, 1963 Fig.14A

             Syn.: Ptenothrix corynephora(sic) Yosii, R, 1977:166 ISS


Species curvilineata (Wray, 1949) Snider, RJ, 1990:256

Ptenothrix (Ptenothrix) curvilineata from the USA
After Snider, R., 1990 Fig.76.
Ptenothrix (Ptenothrix) curvilineata from the USA
After Snider, R., 1990 Fig.77.
Ptenothrix curvilineata from the USA
2012.01.30 © DeSantis, T.
Ptenothrix curvilineata from the USA
2018.01.20 © Abbott, J.C.

Species deharvengi Nayrolles, 1989

Species delongi (Christiansen & Bellinger, 1981) Christiansen, K & Bellinger, P, 1998:1409

Ptenothrix (Ptenothrix) delongi from the USA
After Christiansen K & Bellinger P, 1998 Fig.1037A

Species denticulata (Folsom, JW, 1899:268) Börner, C, 1909:131

Papirius denticulatus from Japan
After Folsom JW, 1899 Pl.3 Fig.33
            Syn.: Ptenothrix denticulata var. catenata Börner, C, 1909:131



Ptenothrix denticulata from Japan
After Yosii R & Lee E-C, 1963 Fig.13A
Ptenothrix denticulata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata from Japan
2014.08.29 © Hara, Y.
Ptenothrix denticulata from Japan
2014.08.29 © Hara, Y.
Ptenothrix denticulata from Japan
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
Modified rough setae on inner side of 3rd tibiotarsus
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
2015.11.07 © Murray, A.
Ptenothrix denticulata from Japan
2016.07.01 © Hara, Y.
Ptenothrix denticulata from Japan
Cleaning dorsum with drop of saliva
2016.07.04 © Hara, Y.
Ptenothrix denticulata from Japan
Just moulted
2018.10.31 © Hara, Y.
Ptenothrix denticulata from Japan
Ant.3 apically annulated (*)
2018.10.31 © Hara, Y.

Species denticulata Yosii, 1954 nec Folsom

Species dinghuensis Lin & Xia, 1985

             Syn.: Ptenothrix diinghuensis(sic) Chen & Christiansen, 1996 ISS

Species dominicana Mari Mutt, JA, 1977:285

Species dorsalis (Reuter, OM, 1876:85) Linnaniemi, 1912

            Syn.: Papirius ater var. albosignatus Parona, 1888
            Syn.: Dicyrtoma setosa var. ceratoleucaspis Börner, 1903 =?


Remark by Bretfeld, G. 1999:158: The colour variety dorsalis Reuter, 1876 of atra is incompletely known and may represent a separate species.

Species fiscellata Handschin, 1925

Species flavescens (Axelson, WM, 1905:794) Axelson, 1907

Remark by Bretfeld, G. 1999:158: The colour variety flavescens Axelson, 1905 of atra is incompletely known and may represent a separate species.

Species fuscamaculata Bretfeld & Trinklein, 2000

Species gaoligongshanensis Itoh & Zhao, 2000

Species gibbosa Börner, C, 1906:185

Species gracilicornis (Schäffer, C, 1898:423) Börner, C, 1906:185

Species hawaiiensis Snider, RJ, 1990:74

Species higashihirajii Yosii, 1965

Species higumai Yosii, 1965

Species himalayensis Yosii, 1966

Species huangshanensis Chen, J-X et Christiansen, K, 1996:587

Ptenothrix huangshanensis from China
After Chen & Christiansen, 1996 Fig.1A.
             Syn.: Ptenothrix huanshanensis(sic) Chen & Christiansen, 1996 ISS


Species iriomotensis Uchida, H, 1965:92

Species italica Dallai, R, 1973:570

Ptenothrix italica from Italy
After Dallai R, 1973 Fig.25a
Ptenothrix italica from Italy
Ant.2 after Dallai R, 1973 Fig.25d
Compared with specimen from France, 2015.11.01 © Gros, P.
Ptenothrix italica from Italy
After Dallai R & al, 2000 Fig.3
Ptenothrix italica from Portugal
2006 © Almeida, J.
Ptenothrix italica from France
2010.12.24 © Nicolosi., B
Ptenothrix italica juv. from Portugal
2011.02.25 © Almeida, J.
Ptenothrix italica from La Palma
Antenna with white rings
2012.12.01 © Barton, T.
Ptenothrix italica from La Palma
2012.12.01 © Barton, T.
Ptenothrix italica from La Palma
2012.12.04 © Barton, T.
Ptenothrix italica from La Palma
More than 2 unpaired facial setae
2012.12.04 © Barton, T.
Ptenothrix italica from Spain
2013.11.04 © Encarna, Q.V.
Ptenothrix italica from France
Marbled body
2015.11.01 © Gros, P.
Ptenothrix italica from France
Thick mid-facial setae: 1,1,3,1,1,1
2015.11.01 © Gros, P.
Ptenothrix italica from France
Antennae with 3 broad black and white rings
2015.11.01 © Gros, P.
Ptenothrix italica from France
2015.11.01 © Gros, P.
Ptenothrix italica from France
Bicoloured eyepatch
2015.11.01 © Gros, P.
Ptenothrix italica from France
Abd.6 with backwards directed dorsal macrosetae (*)
2015.11.01 © Gros, P.
Ptenothrix italica from France
Lateral aspect
2015.11.14 © Gros, P.
Ptenothrix italica from France
Caudal aspect
2015.11.14 © Gros, P.
Ptenothrix italica from France
Ant.3 and 4
2015.11.14 © Gros, P.
Ptenothrix italica from France
Short dorsal spines
2015.11.14 © Gros, P.
Ptenothrix italica ♀ from France
Female gonopore (*)
2015.11.17 © Gros, P.
Remark by Janssens, F. 2015.11.17: Absent strong macrosetae on ant.2, absent dorsal spiky macrosetae in the anterior part of the great abdomen, and presence of backwards directed macrosetae on abd.6 are atypical for Ptenothrix and more characteristic to Dicyrtoma.

Ptenothrix italica from Italy
2017.02.05 © Rinaldo, R.
Ptenothrix italica from Italy
Midfacial setae: 1+1+2+1+1
2017.02.05 © Rinaldo, R.
Ptenothrix italica from Italy
Antenna
2017.02.05 © Rinaldo, R.
Ptenothrix italica from France
2017.10.02 © Tiky, S.
Ptenothrix italica from Portugal
2018.02.18 © Almeida, J.
Ptenothrix italica from France
2018.10.25 © Tiky, S.
Ptenothrix italica from France
More than 2 unpaired midfacial macrosetae suggest Dicyrtoma...
2018.10.25 © Tiky, S.
Ptenothrix italica from France
2018.11.01 © Gros, P.
Ptenothrix italica from France
2018.11.01 © Gros, P.
Ptenothrix italica from France
Caudal aspect
2018.11.01 © Gros, P.

Species janthina Börner, C, 1909:132

            Syn.: Papirius unicolor Börner, C, 1909:132 =?
            Syn.: Ptenothrix setosa var. picta Börner, C, 1909:131
             Syn.: Ptenothrix setosa var. picata(sic) Yosii, R, 1977:166 ISS
            Syn.: Ptenothrix atra var. setosa Uchida, 1952
            Syn.: Ptenothrix atra var. setosa Uchida, 1959
            Syn.: Ptenothrix atra ssp. setosa Salmon, JT, 1964:641
            Syn.: Ptenothrix setosa f. principalis Yosii, R, 1977:166


Ptenothrix janthina from Japan
2014.01.13 © Komatsu, T.
Ptenothrix janthina juv. from Japan
2015.01.16 © Jiku.
Ptenothrix janthina juv. from Japan
2015.01.16 © Jiku.
Ptenothrix janthina juv. from Japan
Pre-jump posture
2015.01.16 © Jiku.

Species keralae Prabhoo, 1971

Species kuraschvilii Djanaschvili, 1970

Species lactea Uchida, 1953

Species leucostrigata Stach, J, 1957:98

Ptenothrix leucostrigata from Poland
After Stach J, 1957 Pl.VIII fig.6
            Syn.: Ptenothrix leucostrigata var. simplificata Stach, J, 1957:101



Ptenothrix leucostrigata from Estonia
2006.09.17 © Tartes, U.
Ptenothrix leucostrigata from Estonia
2006.10.01 © Tartes, U.
Ptenothrix leucostrigata from Denmark
2015.08.05 © Jäschke, J.
Ptenothrix leucostrigata from Germany
2016.07.02 © Jäschke, J.
Ptenothrix leucostrigata from Germany
Antenna colouration pattern
2016.07.02 © Jäschke, J.
Ptenothrix leucostrigata from Poland
2017.09.29 © Ryszard.
Ptenothrix leucostrigata from Poland
Dark form
2017.10.18 © Ryszard.
Ptenothrix leucostrigata from Poland
Dark form with asymmetric dorsal pattern
2017.10.18 © Ryszard.

Species macomba Wray, 1967

Ptenothrix (Ptenothrix) macomba from the USA
After Snider, R., 1990 Fig.74.
Ptenothrix (Ptenothrix) macomba from the USA
After Snider, R., 1990 Fig.75.
Ptenothrix macomba from the USA, juvenile
2007.06.22 © Montgomery, G.
Det. Snider, R.
Ptenothrix macomba from the USA
2007.07.07 © Montgomery, G.
Det. Snider, R.
Ptenothrix cf. macomba from New Zealand
2007.12.17 © van de Vyver, L.

Species maculosa (Schött, H, 1891:14) Matsumura & Ishida, 1931

Papirius maculosus from the USA
After Schött H, 1891 Taf.III Fig.1
Papirius maculosus from the USA
Mucro
After Schött H, 1891 Taf.III Fig.2
Papirius maculosus from the USA
Foot of first leg
After Schött H, 1891 Taf.III Fig.3
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1981:1218 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1998:1411 ISS



             Syn.: Ptenothrix maculosus(sic) Richards, WR, 1968:26 ISS
Dicyrtoma (Ptenothrix) maculosa from the USA
After Christiansen K & Bellinger P, 1981 Fig.1038A
Dicyrtoma (Ptenothrix) maculosa from the USA
After Christiansen K & Bellinger P, 1981 Fig.1038B
Dicyrtoma (Ptenothrix) maculosa from the USA
After Christiansen K & Bellinger P, 1981 Fig.1038C

             Syn.: Dicyrtoma maculosus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix maculosa from the USA
2017.01.19 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix maculosa from the USA
Dark colour form
2017.01.27 © Dobbs, Br.
Ptenothrix maculosa from the USA
Very dark colour form
2017.01.29 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.02.12 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern weak
2018.01.13 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern intermediate
2018.01.13 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern strong
2018.01.13 © Dobbs, Br.

  Subspecies olympia (MacGillivray AD, 1894:110) sensu Janssens F, 2017

            Syn.: Papirius olympius MacGillivray, AD, 1894:110
            Syn.: Ptenothrix maculosa ssp. olympia f.1 Janssens F & Moorehead C, 2008 sensu Janssens F, 2017
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.01.14 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.01.14 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.02.15 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.02.16 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2010.01.02 © LeMons, J.
Ptenothrix sp. f.1 ♂ from the USA
Dorsum pale
2010.01.02 © LeMons, J.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2010.01.03 © LeMons, J.
Ptenothrix sp. f.1 from Canada
Dorsum pale
2015.02.14 © Gilmore, S.
Ptenothrix sp. f.1 from Canada
Dorsum pale
2015.02.14 © Gilmore, S.
Ptenothrix maculosa f.1 from the USA
Dorsum pale
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa f.1 from the USA
2016.03.14 © Dobbs, Br.
Ptenothrix maculosa f.1 from the USA
Facial pigmentation
2016.03.14 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.02.27 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Owl, Bo.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Owl, Bo.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Owl, Bo.

            Syn.: Ptenothrix maculosa ssp. olympia f.2 Janssens F & Moorehead C, 2008 sensu Janssens F, 2017
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.11 © Moorehead, C.
Ptenothrix maculosa f.2 from the USA
Anterior dark + posterior pale
2016.02.04 © Dobbs, Br.
Ptenothrix maculosa f.2 from the USA
Anterior dark + posterior pale
2016.02.04 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.2 from the USA
2018.02.07 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.2 from the USA
2018.02.07 © Dobbs, Br.

            Syn.: Ptenothrix maculosa ssp. olympia f.3 Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix sp. from the USA
Only pigmented middorsally
2008.01.12 © Moorehead, C.
Ptenothrix sp. from the USA
Only pigmented middorsally
2008.01.12 © Moorehead, C.
Ptenothrix maculosa from the USA
2009.01.31 © Leppin, M.
Ptenothrix maculosa f.3 from the USA
Midfacial setae : 1+1+2+2
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from the USA
Pigmented middorsally, unpigmented laterally
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from the USA
Caudal aspect
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from Canada
2016.11.11 © Barbin, T.

            Syn.: Ptenothrix maculosa ssp. olympia f.4 Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from Canada
2016.11.11 © Barbin, T.
Ptenothrix maculosa f.4 from Canada
1+1+2+2+1+1 midfacial setae
2017.01.19 © Barbin, T.

            Syn.: Ptenothrix maculosa ssp. olympia f. subadult Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix maculosa subadult from the USA
2015.09.07 © Brown, M.H.
Ptenothrix maculosa subadult from the USA
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa subadult from the USA
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.11 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.11 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
Male genital papilla (*)
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
Roundish male genital papilla (*)
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult from the USA
With pale dorsum
2018.12.03 © Dobbs, Br.
Ptenothrix maculosa subadult from the USA
With pale dorsum
2018.12.03 © Dobbs, Br.




             Syn.: Ptenothrix olympius(sic) Mills & Rolfs, 1933 ISS
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.02.23 © Peden, S.
Ptenothrix maculosa from the USA
2007.03.19 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.12.06 © Ten Eyck, C.
Ptenothrix maculosa from the USA
2007.12.29 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.12.29 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.01.14 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.01.14 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.01.14 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.30 © McDonald, G.
Ptenothrix maculosa from the USA
2008.02.09 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.02.10 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.02.16 © Moorehead, C.
Ptenothrix maculosa from Canada
2008.12.08 © Turner, E.
Ptenothrix maculosa from Canada
2008.12.08 © Turner, E.
Ptenothrix maculosa from the USA
2009.01.01 © Schneider, K.
Ptenothrix maculosa from the USA
2009.01.01 © Schneider, K.
Ptenothrix maculosa from the USA
2009.02.22 © Schimming, L.
Ptenothrix maculosa from the USA
2010.01.07 © LeMons, J.
Ptenothrix maculosa from the USA
2010.01.22 © LeMons, J.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2012.02.07 © Hochhalter, R.
Ptenothrix maculosa from Canada
Grooming with mouth secreted drop
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.02.12 © Dobbs, Br.
Ptenothrix maculosa from the USA
Mid-facial setae : 1+1+2+2+1+1
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa from the USA
Unguis 1st leg slender + unguiculus with long filament
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa ♂ from the USA
Raised genital papilla (*)
2016.03.08 © Dobbs, Br.
Ptenothrix maculosa ♂ from the USA
Dark lateral body
2016.03.08 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
Transvers split genital papilla (*)
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa from Canada
2016.03.20 © Gatten, J.
Ptenothrix maculosa from Canada
2016.04.06 © Barbin, T.
Ptenothrix maculosa from Canada
2016.04.06 © Barbin, T.
Ptenothrix maculosa from Canada
Dorsal macrosetae AA+BB indistinctly spinelike
2016.09.26 © Barbin, T.
Ptenothrix maculosa from Canada
Caudal aspect pigmentation
2016.09.26 © Barbin, T.
Ptenothrix maculosa juv. from the USA
With extended tubelike collophore vesicles
2016.10.17 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.11.24 © Gross, J.
Ptenothrix maculosa from the USA
2016.11.24 © Gross, J.
Ptenothrix maculosa from the USA
Greenish colour variant
2016.11.29 © Dobbs, Br.
Ptenothrix maculosa from the USA
Midfacial setae: 1+1+2+2+1+1
2016.11.29 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
Caudal aspect
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.10.07 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia ♂ from the USA
2017.10.16 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia ♂ from the USA
2017.10.16 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.21 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.21 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.22 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2018.02.06 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2018.02.06 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2018.12.29 © Schneider, K.



Species marmorata (Packard, AS, 1873:42) Richards, WR, 1968:26

            Syn.: Papirius testudineatus Folsom, 1896
             Syn.: Papirius Testudineatus(sic) Jackson, AD, 1907:173 ISS
             Syn.: Dicyrtoma testudineatus(sic) James, 1933 ISS
             Syn.: Ptenothrix testudinatus(sic) Richards, WR, 1968:26 ISS
            Syn.: Papirius maculosus Guthrie, JE, 1903:58 =?
Papirius maculosus from the USA
Ant.3 apically with 8 subdivisions
After Guthrie JE, 1903 Plate I Fig.1
Papirius maculosus from the USA
Foot of 3rd leg
After Guthrie JE, 1903 Plate I Fig.2
Papirius maculosus from the USA
Foot of 1st leg
After Guthrie JE, 1903 Plate I Fig.3
Papirius maculosus from the USA
Right eyepatch
Modified after Guthrie JE, 1903 Plate I Fig.4
Papirius maculosus from the USA
Mucro: dorsal (a), lateral (c), notched seta (b)
After Guthrie JE, 1903 Plate I Fig.5

            Syn.: Papirius maculosus James, 1933 =?
            Syn.: Ptenothrix oswegatchiensis Maynard, EA, 1951:283
Ptenothrix oswegatchiensis from the USA
After Maynard EA, 1951 Plate XXIV Fig.435




             Syn.: Ptenothrix marmoratus(sic) Folsom, 1928 ISS
             Syn.: Ptenothrix marmoratus(sic) Brimley, 1938 ISS
Ptenothrix marmorata from the USA
After Maynard EA, 1951 Plate XXIV Fig.434

             Syn.: Ptenothrix marmolatus(sic) Terrón-Sierra & Palacios-Vargas, 1991 ISS
             Syn.: Ptenothrix marmoratus(sic) Palacios-Vargas & Gómez-Anaya, 1993 ISS
             Syn.: Dicyrtoma marmoratus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix (Ptenothrix) marmorata from the USA
After Christiansen K & Bellinger P, 1998 Fig.1039A
Ptenothrix (Ptenothrix) marmorata from the USA
After Christiansen K & Bellinger P, 1998 Fig.1039B
Ptenothrix (Ptenothrix) marmorata from the USA
After Christiansen K & Bellinger P, 1998 Fig.1039C

             Syn.: Ptenothrix marmoratus(sic) Palacios-Vargas, JG et Castaño-Meneses, G, 2002:399 ISS
Ptenothrix marmorata from the USA
2006 © Murray, T.
Ptenothrix marmorata from the USA
2006 © Murray, T.
Ptenothrix marmorata from the USA
2006 © Murray, T.
Ptenothrix marmorata from the USA
2006 © Murray, T.
Ptenothrix marmorata from the USA
2006 © Cresswell, S.
Ptenothrix marmorata from the USA
2006 © Cresswell, S.
Ptenothrix marmorata from the USA
Climbing on smooth glass wall
Everting its collophore vesicles at high speed
2007 © Bernard, E.C.
Ptenothrix marmorata from the USA
2007.03.11 © Montgomery, G.
Ptenothrix marmorata from the USA, juvenile
2007.06.07 © Murray, T.
Ptenothrix marmorata from the USA, juvenile
2007.06.07 © Murray, T.
Ptenothrix marmorata from the USA, juvenile
2007.06.07 © Murray, T.
Ptenothrix marmorata from the USA
2007.07.04 © McClarin, J.
Ptenothrix marmorata from the USA
2007.08.02 © Murray, T.
Ptenothrix marmorata from Canada
2007.09.20 © Pilon, C.
Ptenothrix marmorata from the USA
2008.02.07 © Lowery, J.M.
Ptenothrix marmorata from the USA
2008.10.04 © Murray, T.
Ptenothrix marmorata from the USA
2008.10.25 © Murray, T.
Ptenothrix marmorata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata from the USA
2016.08.13 © Schulz, K.
Ptenothrix marmorata from the USA
2016.08.13 © Schulz, K.
Ptenothrix marmorata subadult from the USA
2017.10.23 © Dobbs, Br.



Species mongolica Yosii, 1954

            Syn.: Ptenothrix atra Yosii, 1949


Ptenothrix mongolica from Mongolia
Antenna
After Yosii R & Lee C-E, 1963 Fig.17A

Species monochroma Yosii, R et Lee, C-E, 1963:20

Ptenothrix monochroma from Japan
2013.05.30 © Hara, Y.
Ptenothrix monochroma from Japan
1,1,2,2,1,1 thick midfacial setae
2013.05.30 © Hara, Y.
Ptenothrix monochroma from Japan
Grooming
2013.05.30 © Hara, Y.
Ptenothrix monochroma from China
2014.08.05 © Anonymous
Ptenothrix monochroma from Taiwan
2014.10.04 © Huang, D.

Species narumii Uchida, 1940

Species nigricornis (Ionesco, 1916) Stach, J, 1957:91

Remark by Bretfeld, G. 1999:158: The colour variety nigricornis Ionesco, 1916 of atra is incompletely known and may represent a separate species.

Species ocellata (Becker, 1902)

Species palawanensis Yosii, 1969

Species palmata (Folsom, JW, 1902:102) Stach, J, 1957:77

             Syn.: Ptenothrix palmatus(sic) Richards, WR, 1968:26 ISS
             Syn.: Dicyrtoma palmatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS


Ptenothrix palmata from the USA
2008.11.08 © Fleming, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from Canada
2011.07.24 © Hochhalter, R.
Ptenothrix palmata from Canada
2011.10.24 © Hochhalter, R.
Ptenothrix palmata from the USA
2016.06.22 © Dobbs, Br.
Ptenothrix palmata from the USA
2016.06.22 © Dobbs, Br.
Ptenothrix palmata from the USA
2017.07.15 © Dobbs, Br.
Ptenothrix palmata from the USA
2017.07.15 © Dobbs, Br.

Species palmisetacea Lin & Feng, 1985

            Syn.: Ptenothrix palmisetaceus(sic) Lin & Feng, 1985 IOS


             Syn.: Ptenothrix palmisetaceus(sic) Chen & Christiansen, 1996 ISS



Species plumiseta Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:235

Ptenothrix plumiseta from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.1A
             Syn.: Ptenothrix pluniseta(sic) Park, K-H et Kang, J-S, 2007:A106 ISS


Species quadrangularis (Mills, HB, 1934:106)

Species renateae (Snider, RJ, 1985:561) Snider, 1990

Dicyrtoma (Ptenothrix) renateae from the USA
After Snider RJ, 1985 Fig.1
Dicyrtoma (Ptenothrix) renateae from the USA
After Snider RJ, 1985 Fig.2

             Syn.: Ptenothrix (Ptenothrix) renatae(sic) Christiansen, K et Bellinger, P, 1998:1404 ISS
Ptenothrix renateae from the USA
2006 © Bernard, E.C.
Ptenothrix renateae from the USA
2006.12.06 © Fleming, K.
Ptenothrix renateae from the USA
2008.02.28 © Bradford, A.
Ptenothrix renateae from the USA
2008.12.05 © Bradford, A.
Ptenothrix renateae from the USA
2008.12.09 © Bradford, A.
Ptenothrix renateae from the USA
2009.01.16 © Kermit, J.
Ptenothrix renateae from the USA
2009.02.01 © Zimlich, R.L.
Ptenothrix renateae from the USA
2009.02.24 © Bradford, A.
Ptenothrix renateae from the USA
2009.03.26 © Burishkin, J.
Ptenothrix renateae from the USA
Drinking from the watersurfacefilm with everted vesicles of collophore
2009.03.29 © Bradford, A.
Ptenothrix renateae from the USA
2009.03.30 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.07 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.07 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.13 © Hart, J.
Ptenothrix renateae ♂ from the USA
2010.01.21 © Bradford, A.
Ptenothrix renateae from the USA
2010.01.21 © Bradford, A.
Ptenothrix renateae from the USA
2010.03.09 © Zimlich, R.L.
Ptenothrix renateae from the USA
2011.12.30 © Anderson, J.
Ptenothrix renateae from the USA
2013.02.11 © Anderson, J.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2016.01.01 © Gallagher, J.
Ptenothrix renateae from the USA
2016.01.01 © Gallagher, J.
Ptenothrix renateae from the USA
2016.01.07 © Fleming, K.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
Antenna
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.03.08 © Childs, K.
Ptenothrix renateae juv. from the USA
2016.12.06 © Fleming, K.
Ptenothrix renateae from the USA
4 unpaired midfacial setae
2017.01.03 © Fleming, K.
Ptenothrix renateae from the USA
2018.01.12 © Fleming, K.



Species reticulata Stach, J, 1957:105

Ptenothrix reticulata from Poland
After Stach J., 1957 Pl.IX fig.1.

Species rubra Bretfeld & Trinklein, 2000

Species ryoheii Uchida, 1953

Species saxatilis Yosii, R et Lee, C-E, 1963:19

Ptenothrix saxatilis from Korea
After Yosii R & Lee C-E, 1963 Fig.10A

Species setosa (Krausbauer, 1898:24) Handschin, 1929:125, B

            Syn.: Papirius palmatus Folsom, 1902


Species simplificata Stach, 1957

Species sinensis Lin & Feng, 1985

Species sp. BB Palacios-Vargas & Gómez-Anaya, 1995:29

Species sp. DD Palacios-Vargas & Gómez-Anaya, 1995:29

Species texensis (Packard, AS, 1873:42) Stach, J, 1957:80

            Syn.: Papirius Texensis(sic) Packard, AS, 1873:42 IOS

Species tranvansoii Yoshii, 1994

Species tricycla Uchida, 1953

Species tsutsuii Yosii, 1955

Species unicolor (Harvey, 1893) Folsom, 1928

            Syn.: Papirius pini Folsom, 1896
            Syn.: Ptenothrix atra ssp. apigmentata Stach, 1966
            Syn.: Ptenothrix atra ssp. apigmentata Christiansen, K et Bellinger, P, 1998:1405
            Syn.: Ptenothrix atra ssp. apigmentata Bretfeld, G, 1999:158
            Syn.: Dicyrtoma purpurata Maynard, EA, 1951:280
Dicyrtoma purpurata from the USA
After Maynard EA PlateXXII Fig.377b

            Syn.: Dicyrtoma purpurata Christiansen, K et Bellinger, P, 1981:1231
            Syn.: Dicyrtoma purpurata Christiansen, K et Bellinger, P, 1998:1405



Papirius unicolor from the USA
After Jackson, A.D., 1907 Pl.XI.
Ptenothrix unicolor from the USA
After Maynard EA, 1951 Plate XXIV Fig.430
Ptenothrix unicolor from the USA
After Maynard EA, 1951 Plate XXXVII Fig.663
Ptenothrix unicolor from the USA
2007.05.17 © Murray, T.
Ptenothrix unicolor from the USA
2007.05.17 © Murray, T.
Ptenothrix unicolor from the USA
2007.05.17 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.09 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.09 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.23 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.23 © McClarin, J.
Ptenothrix unicolor from the USA
2007.11.02 © Murray, T.
Ptenothrix unicolor from the USA
2007.11.02 © Murray, T.
Ptenothrix unicolor from the USA
2008.08.20 © Nanz, S.
Ptenothrix unicolor from the USA
2008.08.20 © Nanz, S.
Ptenothrix unicolor from the USA
2008.08.20 © Nanz, S.
Ptenothrix unicolor from the USA
2008.10.07 © Murray, T.
Ptenothrix unicolor from the USA
2008.10.15 © Murray, T.
Ptenothrix unicolor from the USA
2008.11.09 © Justis, S.
Ptenothrix unicolor from the USA
2008.11.09 © Justis, S.
Ptenothrix unicolor from the USA
2008.12.20 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.03.21 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.03.21 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.05.29 © Fleming, K.R.
Ptenothrix unicolor from the USA
2011.07.04 © Nanz, S.
Ptenothrix unicolor from the USA
2011.07.04 © Nanz, S.
Ptenothrix unicolor from the USA
2011.11.13 © Justis, S.
Ptenothrix unicolor from the USA
2011.12.24 © Justis, S.
Ptenothrix unicolor from the USA
Very short 4th antennal segment
2011.12.24 © Justis, S.
Ptenothrix unicolor from the USA
2012.06.03 © Murray, T.
Ptenothrix unicolor from the USA
2012.10.25 © Pugh, D.E.
Ptenothrix unicolor from the USA
2014.07.03 © Dankowicz, E.
Ptenothrix unicolor from the USA
2018.11.01 © Webb, D.

Species unicolor Bueker, 1939 nec Harvey, 1893

Species utingae Arlé & Guimaraes, 1976

Species vinnula Uchida, H, 1957:26

Ptenothrix vinnula from Japan
After Uchida H, 1957 Fig.3A
Ptenothrix vinnula from Japan
2008.04.05 © Tsukiji, T.
Ptenothrix vinnula from Japan
2008.04.05 © Tsukiji, T.
Ptenothrix vinnula from Japan
2014.01.13 © Komatsu, T.
Ptenothrix vinnula from Japan
2015.01.12 © Jiku.
Ptenothrix vinnula from Japan
2015.01.12 © Jiku.
Ptenothrix vinnula from Japan
Frontal spines
2015.01.12 © Jiku.

Species violacea Handschin, 1929

Species violaceopa Hüther, 1967

            Syn.: Ptenothrix violaceopus(sic) Hüther, 1967 IOS

Species vittata (Folsom, 1896) Womersley, H, 1932:40

            Syn.: Ptenothrix fasciatus(sic) Womersley, H, 1932:40 IOS
             Syn.: Ptenothrix fasciatus(sic) Womersley, H, 1939:261 ISS
Ptenothrix fasciata from Australia
After Womersley H, 1939 Fig.75N

             Syn.: Ptenothrix fasciatus(sic) Yosii, R et Lee, C-E, 1963:30 ISS
             Syn.: Ptenothrix fasciatus(sic) Salmon, JT, 1964:642 ISS
             Syn.: Ptenothrix fasciatus(sic) Greenslade, P in Houston, WWK, 1994:138 ISS



Ptenothrix vittata from Japan
After Yosii R & Lee C-E, 1963 Fig.16A
Ptenothrix vittata from Japan
Modified after Yosii R & Lee C-E, 1963 Fig.16B
Dicyrtoma (Ptenothrix) vittata from the USA
After Folsom in Christiansen K & Bellinger P, 1981 Fig.1043C
Dicyrtoma (Ptenothrix) vittata from the USA
After Folsom in Christiansen K & Bellinger P, 1981 Fig.1043D
             Syn.: Dicyrtoma vittatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix nr vittata from Japan
2015.11.02 © Murray, A.
Ptenothrix nr vittata from Japan
2015.11.03 © Murray, A.





Species yakushimana Yosii, 1965

Species yunnana Itoh & Zhao, 2000

            Syn.: Ptenothrix yunnanus(sic) Itoh & Zhao, 2000 IOS





Genus Papirioides Folsom, 1924


  Syn.: Ptenothricini Papiriodes(sic) Richards, WR, 1968:25 ISS
  Syn.: Sminthuridae Papiriodes(sic) Zeppelini Filho, D, 1996:17 ISS
  Syn.: Dicyrtominae Papiriodes(sic) Hopkin, SP, 1997:219 ISS

Species aequituberculatus Stach, J, 1965:369

            Syn.: Papirioides aequituberculata(sic) Stach, J, 1965:369 IOS

Species caishihiensis Wu & Chen, 1996

             Syn.: Papirioides caishijiensis(sic) Li, Z-H, Chen, J-X et Li, J, 2007:61 ISS

Species dubius (Folsom, JW, 1932:74)

Species jacobsoni Folsom, 1924, t.t.

Papiriodes jacobsoni from Japan
After Yosii R & Lee C-E, 1963 Fig.18A
Papiriodes jacobsoni from Japan
After Yosii R & Lee C-E, 1963 Fig.18B
Papirioides jacobsoni from Indonesia
2013.07.13 © Lam, P.

Species kauaiensis (Snider, RJ, 1990:75)

Species mirabilis (Denis, 1929)

Species serratus (Snider, RJ, 1990:79)

Species suzhouensis (Guo & Chen, 1996)

Species tonsori (Greenslade, P, 1994:48)

Species uenoi Uchida, 1957

Species yunnanus Itoh, R et Zhao, L-J, 1993:847

Species zhejiangensis Li, Z-H, Chen, J-X et Li, J, 2007:?


Genus Bothriovulsus Richards, WR, 1968:26


Bothriovulsus sp. from the USA
2009.10.06 © Houston, S.
Bothriovulsus sp. from the USA
3rd & 4th antennal segment
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.04.18 © Kouri, J.
Bothriovulsus sp. from the USA
2012.05.16 © Kouri, J.
Bothriovulsus sp. from the USA
2013.01.28 © Fleming, K.
Bothriovulsus sp. from the USA
2013.12.13 © Fleming, K.
Bothriovulvus sp. from the USA
2015.05.04 © Anderson, J.
Bothriovulsus sp. from the USA
2016.09.13 © Fleming, K.
Bothriovulsus sp. from the USA
2018.02.03 © Fleming, K.

Species sp. Christiansen, K et Bellinger, P, 1981:1219

Species kymgangensis Weiner & Betsch, 1992

Species pineolae (Wray, 1946 tt) Richards, WR, 1968:26

             Syn.: Ptenothrix pineola(sic) Richards, WR, 1968:26 ISS
             Syn.: Bothriovulsus pineolus(sic) Richards, WR, 1968:26 ISS
Dicyrtoma (Bothriovulsus) pineolae from the USA
After Christiansen K & Bellinger P, 1981 Fig.1025A
Dicyrtoma (Bothriovulsus) pineolae from the USA
After Christiansen K & Bellinger P, 1981 Fig.1025A



Species sohungensis Weiner & Betsch, 1992


References