http://www.collembola.org/taxa/smi1idae.htm - Last updated on 2017.04.28 by Frans Janssens
Checklist of the Collembola: Sminthurididae

Familia Sminthurididae Börner, 1906:163 sensu Betsch J-M & Massoud Z, 1970:199
Genus Number
of species
Boernerides1
Debouttevillea1
Denisiella12
Pedonides1
Pseudosminthurides1
Pygicornides2
Sinnamarides1
Sminthurides58
Sminthuridia1
Sphaeridia69
Stenacidia3
Yosiides2
Sminthurididae152
Genera with number of species
  Syn.: Collembola Sminthurides(sic) Willem, V, 1900:55 ISS
  Syn.: Symphypleona Smynthuridae(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
Sminthurides aquaticus from Belgium
2000 © Hopkin, S.P.
Sphaeridia serrata from the USA
2006 © Maddison, D.R.


Genus Pseudosminthurides Sánchez-García, A et Engel, MS, 2016:4, -
Species stoechus Sánchez-García, A et Engel, MS, 2016:4, -
Pseudosminthurides stoechus in Cretaceous amber from Spain
After Sánchez-García A & Engel MS, 2016 Fig.2


Tribus Sphaeridiini Richards WR, 1968:18

Genus Sphaeridia Linnaniemi, WM, 1912:248
  Syn.: Symphypleona Sphaerida(sic) Russell, DJ, 2008:126 ISS
Species aserrata Mari Mutt, JA, 1987:374
Sphaeridia aserrata ♂ from Colombia
After Mari Mutt JA, 1987 Fig.42
Sphaeridia aserrata ♀ from Colombia
After Mari Mutt JA, 1987 Fig.45

Species asiatica Rusek, 1971
Species aspinosa Bretfeld & Trinklein, 2000
Species betschi Arlé, 1984
Species biclava Bretfeld & Trinklein, 2000
Species biniserrata (Salmon, 1951) Massoud, 1964
Sphaeridia biniserrata juv. from India
After Bano R, 2016 p.64

Species bivirgata Bretfeld, 2002
Species boettgeri Bretfeld & Gauer, 1994
Species brevipila (Murphy, 1960) Murphy, 1966
Species cardosi Arlé, 1984
Species carioca Arlé, 1984
Species catapulta Bretfeld & Gauer, 1994
Species cerastes Bretfeld & Gauer, 1994
Species chisacae Bretfeld & Gauer, 1994
Species clara Bretfeld & Gauer, 1994
Species cornuta Murphy, 1966
Species coronata Bretfeld & Gauer, 1994
Species decemdigitata Bretfeld & Schulz, 2012
Species delamarei Bretfeld, 1997
Species denisi Massoud & Delamare Deboutteville, 1964
Species duckei Bretfeld, 2002
Species fernandoi Murphy, 1966
Species fibulifera Bretfeld & Gauer, 1994
Species fluminensis Arlé, 1984
Species foliata Bretfeld, 1997
Species franklinae Bretfeld & Gauer, 1994
Species furcata Dunger, W et Bretfeld, G, 1989:7
Species gladiolifer Delamare Deboutteville, C et Massoud, Z, 1964:58
Species heloisae Arlé, 1984
Sphaeridia heloisea from Peru
2009.01-03.dd © Contreras-Valdez, A.

Species indica Prabhoo, 1971
Species inflata Bretfeld, 1997
Species irmleri Bretfeld & Gauer, 1994
Species jaliensis Bretfeld, 1997
Species leutrensis Dunger, W et Bretfeld, G, 1989:3
Species lobata Bretfeld & Gauer, 1994
Species mandibulata Bretfeld & Gauer, 1994
Species martii Bretfeld & Gauer, 1994
Species massoudi Murphy, 1966
Species minimus (Schött, H, 1893:7)
Species murphyi Yosii, 1966
Species neopumilis Bretfeld & Gauer, 1994
Species obtusa Murphy, 1966
Species panguanae Bretfeld & Schulz, 2012
Species paroara Arlé, 1984
Species peruensis Bretfeld & Schulz, 2012
Species pilleata Bretfeld & Gauer, 1994
Species pippetti Murphy, 1966
Species posterospina Murphy, 1966
Species proxima Murphy, 1960
Species pumilis (Krausbauer, 1898) Agrell, 1934, t.t., B
            Syn.: Sphaeridia minima Scherbakov, 1898
            Syn.: Sminthurides globosus Axelson, 1902
            Syn.: Sphaeridia tunicata Yosii, 1954
            Syn.: Sminthurides (Stenacidia) violacea Salmon, 1956
            Syn.: Sphaeridia pumilis f. spinosa Bretfeld, G, 1995:18
             Syn.: Sminthurides pumilio(sic) Börner, 1901 ISS
             Syn.: Sminthurides pumilio(sic) Axelson, 1905 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Linnaniemi, 1912 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Handschin, 1929 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Denis, 1931 ISS
             Syn.: Sminthurides (Sphaeridia) pumilio(sic) Kseneman, M, 1932:22 ISS
             Syn.: Sphaeridia pumilio(sic) Agrell, 1934 ISS
             Syn.: Sminthurides pumilio(sic) Obenberger, 1955 ISS
             Syn.: Sphaeridia pumilio(sic) Delamare Deboutteville & Massoud, 1963 ISS
Neozygites sp. from Denmark. Bar=10µm
Conidia infecting Sphaeridia pumilis
After Dromph & al, 2001 Fig.3
Sphaeridia pumilis from the UK
Male antenna
2004 © Hopkin, S.P.
Sphaeridia pumilis from France
2007.08.13 © Lebeaux, P.
Sphaeridia pumilis from Belgium
2007.12.18
© Salehian, S., Beladjal, L. & Mertens, J.
Sphaeridia pumilis female from Belgium
2007.12.18
© Salehian, S., Beladjal, L. & Mertens, J.
Sphaeridia pumilis juv. from the UK
2012.06.04 © Murray, A.
 
Sphaeridia pumilis ♂ from the UK
2012.06.04 © Murray, A.
 
Sphaeridia pumilis ♂ from the UK
Male clasping antenna
2012.06.04 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.06.07 © Murray, A.
 
Sphaeridia pumilis ♀ from the UK
2012.06.07 © Murray, A.
 
Sphaeridia pumilis ♀ from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis 1st instar from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis juv. from the UK
2012.06.07 © Murray, A.
Sphaeridia pumilis juv. from the UK
2012.06.11 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.06.12 © Murray, A.
Sphaeridia pumilis ♂ from the UK
With clasping antennae
2012.07.08 © Nurcombe, E.
Sphaeridia pumilis ♂ from the UK
Male grasping antenna
2012.08.19 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.08.24 © Nurcombe, E.
Sphaeridia pumilis ♀ from Holland
2012.10.04 © van Duinen, J.
Sphaeridia pumilis ♀ from Portugal
2012.10.13 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2012.10.15 © Nurcombe, E.
Sphaeridia pumilis ♀ from the UK
2012.10.15 © Nurcombe, E.
Sphaeridia pumilis ♀ from the UK
2012.12.08 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2013.01.13 © Murray, A.
Sphaeridia pumilis ♀ from the UK
2013.09.24 © Nurcombe, E.
Sphaeridia pumilis ♀ from Holland
2013.10.23 © van Duinen, J.
Sphaeridia pumilis ♂ from Holland
With clasping antennae
2013.10.23 © van Duinen, J.
Sphaeridia pumilis ♀ from the UK
2013.11.09 © Murray, A.
Sphaeridia pumilis ♀ from Holland
2014.10.03 © van Duinen, J.
Sphaeridia pumilis ♀ from Holland
2014.11.07 © van Duinen, J.
Sphaeridia pumilis ♀ & ♂ from Holland
End of copulation
2014.11.11 © van Duinen, J.
Sphaeridia pumilis ♀ from Germany
2014.11.15 © Beck, K.
Sphaeridia pumilis ♀ from Germany
2014.11.16 © Beck, K.

Species robusta Bretfeld & Gauer, 1994
Species salmoni Massoud & Delamare Deboutteville, 1964
Species schalleri Massoud & Delamare Deboutteville, 1964
Species serrata Folsom & Mills, 1938
             Syn.: Sphaeridia serratus(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS
Sphaeridia serrata from the USA
2006 © Maddison, D.R.
Sphaeridia serrata ♀ & ♂ from the USA
2009 © Haney, J.

Species sphaera (Salmon, JT, 1946:58) Massoud, 1964
Species spinifer (da Gama, 1964)
Species spinifrons Murphy, 1966
Species spira Bretfeld & Gauer, 1994
Species squamifera Bretfeld & Gauer, 1994
Species sturmi Bretfeld & Gauer, 1994
Species superstes Bretfeld in Bretfeld, Poliakov & Broza, 2000
Species torifera Bretfeld & Schulz, 2012
Species tropica Bretfeld & Schulz, 2012
Species tschirnhausi Bretfeld & Schulz, 2012
Species tunicata Yosii, 1954
Species vampyra Bretfeld & Schulz, 2012
Species vanharteni Bretfeld, 2000
Species winteri Massoud & Delamare Deboutteville, 1964
Species zaheri Yosii, 1966

Tribus Sminthuridini Börner, 1913 sensu Richards WR, 1968:18

Genus Sminthurides Börner, 1900
  Syn.: Sminthuri(sic) Smynthurus(sic) Folsom, JW, 1896:446 partim
  Syn.: Sminthuridae Sminthurus Oudemans, 1890
  Syn.: Sminthuridæ(sic) Sminthurus Schött, H, 1891:7 partim
  Syn.: Sminthuridae Sminthurus Levander, KM, 1894:3 partim
  Syn.: Sminthurus Pilosi Schött, H, 1891:11 partim
  Syn.: Sminthurides(sic) Prosminthurus Willem, V, 1900:55
  Syn.: Sminthurides Spinosella Palissa, 1961, nec Vosmaer, 1885
  Syn.: Sminthurides Spinosellina Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:240
  Syn.: ... Spinsellina(sic) Pomorski, RJ et Skarzynski, D, 1992:71 ISS
Remark by ICZN. 2007:759: Placed on the Official List of Names in Zoology. Opinion 291.

  Syn.: Sminthuridae Sminthurus(sic) Glime, JM, 2015:5 lapsus ISS
Sminthurides sp. from India
After Bano R, 2016 p.63
Det.: Janssens F

Species sp.1 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:895
Species sp.2 Janssens F & Dell T, 2008
Sminthurides nr pseudassimilis from Australia
2002.11.05 © Dell T.
Sminthurides nr pseudassimilis from Australia
Furca with lamellate mucrones
2002.11.05 © Dell T.
Sminthurides nr pseudassimilis from Australia
Female specimen
Antenna with annulated 4th antennomere
2002.11.05 © Dell T.

Species sp.3 Soto-Adames, FN et Taylor, SJ, 2013:163
Species acanthus Hüther, 1967
Species afer Marlier, 1944:5
Species annulicornis Axelson, WM, 1905:793
             Syn.: Sminthurides ann.(sic) Bretfeld, G et Griegel, A, 1999:223 ISS
Species antennatus Baijal & Verma, 1986
Species appendiculatus Imms, 1912
Species aquaticus (Bourlet, 1842) Isa, 1963, t.t., B
            Syn.: Sminthurus apicalis Reuter, 1880
             Syn.: Sminthurus apiculis(sic) Schött, H, 1893:36 ISS
Sminthurus apicalis from Finnland
Mucro with 3 lamellae: inner serrate, outer + median smooth
After Levander KM, 1894:10
Sminthurus apicalis from Germany
Male grasping antenna
After Anonymous Fig.G

            Syn.: Sminthurides aquaticus f. principalis Schött
            Syn.: Sminthurus aquaticus var. levanderi Schött, H, 1893:37
             Syn.: Sminthurides aquaticus var. levanderis(sic) Isa, 1963 ISS
Sminthurides aquaticus var. levanderi from France
2008.03.17 © Lebeaux, P.
 
Sminthurides aquaticus var. levanderi from France
2008.03.17 © Lebeaux, P.
 
Sminthurides aquaticus var. levanderi from France, courtship
2008.03.17 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France, courtship
2008.03.18 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2008.04.02 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France, courtship
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France, courtship
2008.04.06 © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi ♂ from France
With highly modified grasping antennae
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
Courtship
2009.09.dd © Lebeaux, P.
Sminthurides aquaticus var. levanderi from France
Predation by spider
2009.09.13 © Lebeaux, P.
Sminthurides aquaticus var. levanderi ♀ from Holland
2011.04.dd © van Duinen, J.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.09.29 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.10.10 © Barton, T.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
2012.11.19 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2012.12.23 © Barton, T.
Sminthurides aquaticus var. levanderi ♂ & ♀ juv. from the UK
Male practising courtship ritual
2012.12.23 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the USA
Feeding instar with dark dorsum due to gut contents
2013.02.09 © Fleming, K.
Sminthurides aquaticus var. levanderi ♀ from the UK
2013.04.23 © Murray, A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2013.11.04 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♂ & ♀ juv. from the UK
Courtship
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Pale basal antennomeres
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Lamellate mucro
2014.10.20 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Fighting females
2014.11.21 © Murray, A.
Sminthurides aquaticus var. levanderi from the UK
2014.11.26 © Murray, A.
Sminthurides aquaticus var. levanderi from the UK
Black foot and lamellate mucro
2014.11.26 © Murray, A.
Sminthurides aquaticus var. levanderi ♂ & ♀ from the UK
Courtship ritual
2014.12.01 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2014.12.08 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
Proleg: black foot; short empodium
2014.12.10 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Ready to moult
2014.12.10 © Ardron,P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.01.11 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.02.15 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from the UK
2015.04.26 © Ardron, P.A.
Sminthurides aquaticus var. levanderi ♂ from the UK
2015.04.26 © Ardron, P.A.
Sminthurides aquaticus var. levanderi ♀ from the UK
Guarding eggs on Lemna minor
2015.05.09 © Barton, T.
Sminthurides aquaticus var. levanderi ♀ from France
2016.11.27 © Garcelon, P.
Sminthurides aquaticus var. levanderi ♀ from France
Dorsal aspect
2016.11.27 © Garcelon, P.
Sminthurides aquaticus var. levanderi ♀ from France
2017.02.28 © Picard, J.
Sminthurides aquaticus var. levanderi ♀ from France
2017.02.28 © Picard, J.
Sminthurides aquaticus var. levanderi ♂ from France
2017.02.28 © Picard, J.


            Syn.: Sminthurus aquaticus var. viridulus Schött, H, 1893:37
             Syn.: Sminthurides aquaticus var. viridula(sic) Bretfeld, G, 1999:38 ISS
Sminthurides aquaticus var. viridulus ♀ & ♂ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
Long narrow unguis and needle-like unguiculus of 1st leg
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
Composed eye with 6 large ocelli and 2 distinctly smaller ocelli
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Germany
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ & ♂ from Germany
With embryo's in eggs
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus from Germany
Embryo's in eggs embedded in epidermis of Lemna minor
2014.11.05 © Gutekunst, V.
Sminthurides aquaticus var. viridulus ♀ from Poland
2015.05.14 © Beck, K.


            Syn.: Sminthurus apicalis f. principalis Reuter, 1890
            Syn.: Smynthurus(sic) amicus Folsom, JW, 1896:447
Smynthurus(sic) amicus from the USA
Fore foot with long unguis and shorter unguiculus
After Folsom JW, 1896 Plate10 Fig.6
Smynthurus(sic) amicus from the USA
Hind foot with short unguis and longer unguiculus
After Folsom JW, 1896 Plate10 Fig.8
Smynthurus(sic) amicus from the USA
Mucro with serrate inner and smooth outer lamella
After Folsom JW, 1896 Plate10 Fig.7

Sminthurus aquaticus from Sweden
After Schött H, 1893 Taf.II Fig.22
Sminthurides aquaticus from Germany
Foot of proleg with short unguiculus
After Börner C, 1901 Fig.43a
Sminthurides aquaticus from Germany
Foot of mesoleg with long unguiculus
After Börner C, 1901 Fig.43b
Sminthurides aquaticus from Germany
Foot of metaleg with long unguiculus
and tibiotarsal organ
After Börner C, 1901 Fig.43c
Sminthurides aquaticus from Germany
After Heymons R & Heymons H, 1909 Fig.22
Sminthurus aquaticus from Germany
After Anonymous Fig.F
Sminthurides aquaticus from the UK
Broad lamellate mucro
2004 © Hopkin, S.P.
Sminthurides aquaticus from the UK
2006 © Jones, D.
Sminthurides aquaticus from Sweden
2006 © Hall, K.
Sminthurides aquaticus from the UK
2007.03.04 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.18 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.18 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.03.26 © Cornwall, N.J.
Sminthurides aquaticus from the UK
2007.04.01 © Jones, D.
Sminthurides aquaticus from Germany
2007.08.13 © Schmitt, R.
Sminthurides aquaticus from Germany
2007.08.14 © Schmitt, R.
Sminthurides aquaticus from the UK
2007.08.31 © Oakshott, A.
Sminthurides aquaticus from Germany
2007.10.12 © Schmitt, R.
Sminthurides aquaticus from Belgium
2008.04.22 © Vancoppenolle B.
Sminthurides aquaticus from the UK
2009.03.29 © Michaelson, J.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.10 © Robertson, A.
Sminthurides aquaticus from the UK
2009.05.17 © Robertson, A.
Embryo in transparant egg ready to hatch
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male with clasping antennae
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male with clasping antennae
Sminthurides aquaticus from the UK
2009.06.17 © Robertson, A.
Male advancing larger female
Sminthurides aquaticus from the UK
2009.09.30 © Robertson, A.
Bothriotricha A-D
Sminthurides aquaticus from the UK
2010.07.03 © Robertson, A.
Male clasped to female
Sminthurides aquaticus from the UK
2010.08.07 © Robertson, A.
Unsubdivided 4th antennal segment
Sminthurides aquaticus from the UK
Furca with broad lamellate mucrones
2012.04.12 © Peacock, H.
Sminthurides aquaticus from the UK
2012.05.01 © Murray, A.
Sminthurides aquaticus ♂ from the UK
2012.05.02 © Murray, A.
Sminthurides aquaticus juv. from the UK
2012.05.14 © Murray, A.
Sminthurides aquaticus from the UK
Eggs in 'nests'
2012.05.19 © Murray, A.
Sminthurides aquaticus from the UK
Embryo in transparant egg
2012.05.19 © Murray, A.
Sminthurides aquaticus from the UK
Embryo in transparant egg
2012.05.22 © Murray, A.
Sminthurides aquaticus ♂ & juv. ♀ from the UK
2012.05.28 © Murray, A.
Sminthurides aquaticus juv. from the UK
2012.06.04 © Murray, A.
Sminthurides aquaticus juv. ♀ & ♂ from the UK
2012.08.12 © Nurcombe, E.
Sminthurides aquaticus juv. & ♂ from the UK
Male grasps juvenile at wrong end...
2012.10.03 © Nurcombe, E.
Sminthurides aquaticus juv. ♀ from the UK
2013.02.11 © Nurcombe, E.
Sminthurides aquaticus ♂ & ♀ from the UK
Mucro with broad lamellae
2013.02.17 © Nurcombe, E.
Sminthurides aquaticus ♂ from Germany
Antennal clasping organ
2014.09.19 © Beck, K.
Sminthurides aquaticus ♀ from Germany
Ant.4 not subdivided
2014.10.05 © Kluck, A.
Sminthurides aquaticus ♂'s from Germany
Figthing?
2014.11.09 © Kluck, A.
Sminthurides aquaticus from the UK
2014.11.14 © Murray, A.
Sminthurides aquaticus ♀ from Germany
2015.10.31 © Kluck, A.
Sminthurides aquaticus ♂ from Germany
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ & ♂ from Germany
Male grasping antenna
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ from Germany
Antenna
2015.11.15 © Kluck, A.
Sminthurides aquaticus ♀ from Belgium
2016.07.10 © Huskens, M.-L.
Sminthurides aquaticus juv. ♂ from Belgium
With swollen ant.2-3
2016.07.12 © Huskens, M.-L.
Sminthurides aquaticus subadult ♂ from Belgium
With precursive grasping antenna
2016.07.12 © Huskens, M.-L.
Sminthurides aquaticus juv. ♀ & ♂ from Belgium
Courtship
2016.07.13 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Ant.4 not subdivided
2016.07.19 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Grooming with a drop of saliva
2016.07.25 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Chewing cup in epidermis of Lemna minor to deposit egg
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Depositing egg in epidermal cup of Lemna minor
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♀ from Belgium
Covering egg with faecal pellet
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus from Belgium
Embryo's embedded in epidermis of Lemna minor
2016.07.27 © Huskens, M.-L.
Sminthurides aquaticus ♂ from the UK
Male clasping antenna
2016.08.19 © Phillips, E.
Sminthurides aquaticus ♀ from the UK
Cleaning dorsum with drop of saliva
2016.08.23 © Phillips, E.
Sminthurides aquaticus ♀ from Belgium
Balancing on 3 right legs
2016.08.25 © Huskens, M.-L.
Sminthurides aquaticus ♀ + ♂ from Belgium
Begin of courtship ritual
2016.09.22 © Huskens, M.-L.
Sminthurides aquaticus ♂ from Belgium
Clasping antenna
2016.09.22 © Huskens, M.-L.
Sminthurides aquaticus ♂ juv. from Belgium
First instar
2016.09.30 © Huskens, M.-L.
Sminthurides aquaticus ♀ from the UK
Bothriotricha A-D
2016.10.09 © Barton, T.
Sminthurides aquaticus ♀ from Belgium
Balancing on 3 right legs
2016.10.25 © Huskens, M.-L.

Species armatus Bretfeld, 2000
Species aureolus Maynard, EA, 1951:213
Species avicenniae Murphy, 1960
Species bicuspidatus Bretfeld, 2001
Species biwae Yosii, 1970
Species bothrium Nguyen, 2001
Species condei (Delamare Deboutteville & Massoud, 1963:177) Delamare Deboutteville & Massoud, 1963
            Syn.: Sminthurides Condéi Delamare Deboutteville & Massoud, 1963 IOS
Species cruciatus Axelson, WM, 1905:792
Sminthurides cruciatus from Russia
2003.mm.dd © Potapov, M.

Species gambiae Murphy, 1960
Species globocerus Folsom & Mills, 1938
Species hessei (Poppe, 1886)
Species hospes Börner, 1907
Species hyogramme Pedigo, 1966
             Syn.: Sminthurides hygromae(sic) Palacios-Vargas, 1990 ISS
             Syn.: Sminthurides hyogramma(sic) Miranda Rangel & Palacios-Vargas, 1992 ISS
             Syn.: Sminthurides hypogramme(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurides hyogramme from Canada
2009 © Porco, D.

             Syn.: Sminthurides hygramme(sic) Cruz-Leal, JI, Escalante-Jiménez, AL, Salgado-Ortiz, J et Palacios-Vargas, JG, 2013:1666 ISS
Species inaequalis Börner, 1903
            Syn.: Sminthurides sexoculatus Unknown YYYY =?
             Syn.: Sminthurides inequalis(sic) Stach, J, 1956:19 ISS
Sminthurides inaequalis ♀ from Italy
After Dallai R, 1973 Fig.14a
Sminthurides inaequalis ♂ from Italy
After Dallai R, 1973 Fig.14a

Species lepus Mills, HB, 1934:91
Sminthurides lepus from the USA
After Mills, H.B., 1934 Plate 2 Fig.13
Sminthurides (Sminthurides) lepus from the USA
After Maynard, E.A., 1951 Pl.XXII Fig.360
Sminthurides (Sminthurides) lepus from the USA
After Maynard, E.A., 1951 Pl.XXII Fig.361
Sminthurides (Sminthurides) lepus from the USA
After Maynard, E.A., 1951 Pl.XXII Fig.362
Sminthurides (Sminthurides) lepus from the USA
After Maynard, E.A., 1951 Pl.XXII Fig.363
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA, female & male
2008.06.04 © Schoenherr, J.
Sminthurides lepus from the USA
2008.06.04 © Schoenherr, J.

Species lolelua Christiansen, K et Bellinger, P, 1992:340
Species longicornis (Joseph G, 1882) Börner, C, 1901:137
            Syn.: Arrhopalites? sp. Stach, J, 1956:239
Species macgregori Murphy, 1960
Species macnamarai Folsom & Mills, 1938
Species macroceros Arlé, R, 1961:36
Species malmgreni (Tullberg, T, 1877:30) Axelson, 1905, B
            Syn.: Sminthurus Malmgrenii(sic) Tullberg, T, 1877:30 IOS
            Syn.: ... principalis Tullberg, 1876
            Syn.: Sminthurus elegantulus Reuter, 1880
Sminthurides Malmgreni(sic) var. elegantula from Germany
After Heymons R & Heymons H, 1909 Fig.24

            Syn.: Smynthurus(sic) socialis Folsom, JW, 1896:446
Smynthurus(sic) socialis from the USA
Fore foot with long unguis
After Folsom JW, 1896 Plate10 Fig.1
Smynthurus(sic) socialis from the USA
Hind foot with short unguis
After Folsom JW, 1896 Plate10 Fig.3
Smynthurus(sic) socialis from the USA
Mucro with serrate inner and smooth outer lamella
After Folsom JW, 1896 Plate10 Fig.2
Smynthurus(sic) ♂ socialis from the USA
Modified clasping antenna
After Folsom JW, 1896 Plate10 Fig.4

            Syn.: Sminthurides malmgreni var. nigrescens Börner, C, 1901:96
Sminthurides malmgreni f. nigrescens ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♂ & ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♀ & ♂ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♂ & ♀ from the UK
2008.02.dd © Ardron, P.A.
Sminthurides malmgreni f. nigrescens ♀ from Germany
2009.12.12 © Marquard, F.
Sminthurides malmgreni f. nigrescens ♀ from Norway
2009.04.16 © Fjellberg, A.
Sminthurides malmgreni f. nigrescens ♀ from Norway
2009.12.18 © Fjellberg, A.
Sminthurides malmgreni f. nigrescens ♀ from Germany
2010.02.09 © Marquard, F.
Sminthurides malmgreni f. nigrescens ♀ from Poland
2015.05.14 © Beck, K.
Sminthurides malmgreni f. nigrescens ♀ from Poland
2015.05.14 © Beck, K.
Sminthurides malmgreni f. nigrescens ♀ from Holland
2016.02.13 © van Duinen, J.
Sminthurides malmgreni f. nigrescens ♀ from the USA
2016.12.23 © Dobbs, Br.


            Syn.: ... abrupta Ågren, 1903
            Syn.: Sminthurides maculatus Ågren, 1903
            Syn.: Sminthurides quadrilineatus Ågren, 1903
            Syn.: Sminthurides malmgreni var. immaculata Axelson, WM, 1905:792
Sminthurides malmgreni f. immaculata from France
2008.04.13 © Lebeaux, P.
Sminthurides malmgreni f. immaculata from the UK
2011 © Shaw, P.
Sminthurides malmgreni f. immaculata from Germany
2014.10.23 © Horak, T.


            Syn.: Sminthurides cruciatus Ionescu, 1915
            Syn.: Sminthurides ludovicianus Folsom & Mills, 1938
            Syn.: Sminthurides palustris Folsom & Mills, 1938
            Syn.: Sminthurides sp. Greenslade, 1987
            Syn.: Sminthurides sp. Greenslade, 1990
            Syn.: Sminthurides sp. Greenslade, 2002
             Syn.: Sminthurus Malmgrenii(sic) Schött, H, 1893:34 ISS
Sminthurus Malmgrenii(sic) from Sweden
After Schött H, 1893 Taf.II Fig.20


             Syn.: Smynthurus(sic) malmgrenii(sic) Folsom, JW, 1896:447 ISS
Sminthurides malmgreni ♀ light form
After Stach J, 1956 Pl.II Fig.4
Sminthurides malmgreni ♀ medium form
After Stach J, 1956 Pl.II Fig.5
Sminthurides malmgreni ♀ dark form
After Stach J, 1956 Pl.II Fig.6

             Syn.: Sminthurides Malmgreni(sic) Delamare Deboutteville & Massoud, 1963 ISS
Sminthurides malmgreni from Germany
After Palissa, A., 1964 Fig.2.

             Syn.: Sminthurides malmgremi(sic) Richards, WR, 1968:16 ISS
Sminthurides malmgreni ♀ light form
After Janssens F, 1995 Fig.2c


             Syn.: Sminthurides malgreni(sic) Shaw, P, 2003:5 ISS
Sminthurides malmgreni from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides malmgreni from Sweden
2006 © Hall, K.
 
Sminthurides malmgreni from the UK
2007.03.04 © Cornwall, N.J.
 
Sminthurides malmgreni from France
2008.03.02 © Lebeaux, P.
Sminthurides malmgreni from France
Male with clasping antennae
2008.03.02 © Lebeaux, P.
Sminthurides malmgreni from France
2008.04.13 © Lebeaux, P.
 
Sminthurides malmgreni ♂ juv. from France
2009.09.dd © Lebeaux, P.
Sminthurides malmgreni from Norway
2009.11.24 © Fjellberg, A.
Sminthurides malmgreni ♀ from Holland
2011.02.16 © van Duinen, J.
Sminthurides malmgreni ♂ from Holland
2011.02.16 © van Duinen, J.
Sminthurides malmgreni juv. from France
2012.01.15 © Lebeaux, P.
Sminthurides malmgreni from Holland
2012.01.25 © van Duinen, J.
Sminthurides malmgreni ♂ from the UK
2012.04.26 © Murray, A.
Sminthurides malmgreni ♂ & ♀ from the UK
Courtship
2012.05.11 © Murray, A.
Sminthurides malmgreni ♂ from the UK
2012.08.02 © Murray, A.
Sminthurides malmgreni from the UK
2013.02.15 © Nurcombe, E.
Sminthurides malmgreni juv. from the UK
2013.02.17 © Nurcombe, E.
Sminthurides malmgreni ♀ from Holland
2014.11.07 © van Duinen, J.
Sminthurides malmgreni ♂ from Holland
2014.11.07 © van Duinen, J.
Sminthurides malmgreni ♀ & Dicyrtomina saundersi ♂ from Holland
Love at first sight ;-)
2014.11.11 © van Duinen, J.
Sminthurides malmgreni ♀ from the UK
2015.01.19 © Murray, A.
Sminthurides malmgreni ♂ & ♀ juv. from Germany
Courtship ritual
2015.03.27 © Landgraf, A.
Sminthurides malmgreni ♀ from Belgium
2016.04.17 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
2016.08.11 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
2016.08.12 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Caudal aspect
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Cleaning antenna with drop of saliva
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Eyepatch with 6 large + 2 small ocelli
2016.09.23 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Grooming balancing on 3 right legs
2016.11.04 © Huskens, M.-L.
Sminthurides malmgreni ♂ & ♀ from France
Courtship
2016.12.08 © Garcelon, P.
Sminthurides malmgreni ♀ from the USA
2016.12.23 © Dobbs, Br.
Sminthurides malmgreni ♀ from Belgium
2017.02.19 © Huskens, M.-L.
Sminthurides malmgreni ♀ from Belgium
Mucro with lamellae
2017.02.19 © Huskens, M.-L.

Species melanotus Börner, 1908
Species millsi (Marlier, 1944:6) Stach, 1956, s.i.
            Syn.: Sminthurides (Sminthurides) Millsi Marlier, 1944 IOS
Species monnioti Massoud & Betsch, 1966
Species napostaensis Najt & Rapoport, 1965
Species occultus Mills, HB, 1934:91
Sminthurides occultus from Canada
2010 © Biodiv. Inst. Ontario.

Species olivieri Delamare Deboutteville & Massoud, 1963:185
            Syn.: Sminthurides Olivieri(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species parvulus (Krausbauer, 1898) Cassagnau, P, 1953:227, B
            Syn.: Sminthurides quinquelineatus Gisin, 1943
            Syn.: Sminthurides quinquearticulatus Gisin, 1944
             Syn.: Sminthurides 5-articulatus(sic) Cassagnau, P, 1953:227 ISS
            Syn.: Sminthurides alpinus Cassagnau, P, 1960:407
Sminthurides parvulus from Germany
After Heymons R & Heymons H, 1909 Fig.18
Sminthurides parvulus from the UK
Mucrones gutter-like
2004 © Hopkin, S.P.
Sminthurides parvulus ♀ from Luxembourg
Ant.4 with 5 subdivisions
After Marx MT & Weber D, 2013 Fig.4
Sminthurides parvulus ♀ from the UK
Fourth antennal segment with 5 subdivisions
2013.01.05 © Murray, A.
Sminthurides parvulus ♂ from the UK
2013.01.06 © Murray, A.
Sminthurides parvulus ♂ from the UK
Male grasping antenna
2013.01.06 © Murray, A.
Sminthurides parvulus ♀ juv. from the UK
2015.02.13 © Murray, A.
Sminthurides parvulus ♀ juv. from the UK
Ant.4 with 5 subdivisions
2015.02.13 © Murray, A.
Sminthurides parvulus ♀ juv. from Holland
2016.08.25 © van Duinen, J.

Species penicillifer (Schäffer, 1896) Börner, C, 1901:94, B
            Syn.: Sminthurides penicillifer var. incompta(sic) Börner, C, 1901:94 IOS
             Syn.: Sminthurides penicillifer var. incompta(sic) Voigts, 1903 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Heymons, R et Heymons, H in Brauer, A, 1909:12 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Linnaniemi, WM, 1912:252 ISS
             Syn.: Sminthurides penicillifer f. incompta(sic) Handschin, E in Dahl, F, 1929:108 ISS
             Syn.: Sminthurus penicillifer var. incompta(sic) Kel'schtein, 1929 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Stach, J, 1956:41 ISS
             Syn.: Sminthurides penicillifer var. incompta(sic) Grinbergs, 1960 ISS
             Syn.: Sminthurides penicillifer var. incompacta(sic) Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:241 ISS
             Syn.: Sminthurides penicillifer f. incomota(sic) Tarashchuk, 1995 ISS
Sminthurides penicillifer var. incomptus ♀ from the UK
With Podura aquatica spermatophore
2010.07.09 © Michaelson, J.
Sminthurides penicillifer var. incomptus from the UK
Footcomplex of proleg; striated mucro
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Footcomplex of mesoleg; hydrophylic outer face of unguis
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Extended mouthparts
2013.11.01 © Murray, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.15 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Feeding instar
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Ant.4 with 7 subdivisions and rings
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Mucro
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♂ juv. from Germany
Ant.2 with two bothriotricha
2014.09.17 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2014.09.17 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
Facial interantennal spot
2014.09.20 © Beck, K.
Sminthurides penicillifer var. incomptus ♀ from Germany
Mucro with shortened outer lamella
2014.11.09 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from Germany
Facial interantennal spot
2014.11.09 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides penicillifer var. incomptus ♀ from the UK
Aggressive? encounter of two females
2014.12.03 © Ardron,P.A.
Sminthurides penicillifer var. incomptus ♀ from Germany
2015.08.30 © Kluck, A.
Sminthurides penicillifer var. incomptus ♂ from Germany
2015.08.30 © Kluck, A.
Sminthurides penicillifer var. incomptus ♀ from the UK
2016.12.11 © Phillips, E.
Sminthurides penicillifer var. incomptus ♀ from the UK
Unguiculus (ung) longer than unguis (UNG)
2016.12.15 © Phillips, E.


            Syn.: Sminthurides penicillifer f. typica(sic) Börner, C, 1901:94 IOS
            Syn.: Sminthurides Pauliani(sic) Denis, 1936
            Syn.: Sminthurides malmgreni Janssens, F, 1992:99,B
Sminthurides penicillifer ♀ & ♂ from Germany
After Börner, 1901
Sminthurides penicillifer from Germany
Foreleg with long unguiculus
After Börner, 1901 Taf.II Fig.11a
Sminthurides penicillifer from Germany
Hindleg with multifaded unguiculus
After Börner, 1901 Taf.II Fig.11b

             Syn.: Sminthurides penicilifer(sic) Kseneman, M, 1932:22 ISS
Sminthurides penicillifer ♀ from Poland
After Stach J, 1956 Pl.V Fig.1
Sminthurides penicillifer ♀ from Poland
After Janssens F, 1995 Fig.1a
Sminthurides penicillifer ♀ from Belgium
After Janssens F, 1995 Fig.1b
Sminthurides penicillifer ♀ from Belgium
After Janssens F, 1995 Fig.2b
Sminthurides penicillifer ♂ from Belgium
After Janssens F, 1995 Fig.3a
Sminthurides penicillifer ♂ from Belgium
After Janssens F, 1995 Fig.3b
Sminthurides penicillifer ♂ juv. from Germany
2014.03.23 © Beck, K.
Sminthurides penicillifer ♂ & ♀ from Germany
Courtship
2014.09.12 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ from Germany
Male antennal clasping organ
2014.09.17 © Beck, K.
Sminthurides penicillifer ♂ juv. from Germany
With undeveloped pearly ant.2-3
2014.09.23 © Beck, K.
Sminthurides penicillifer ♂ & ♂ from Germany
Juvenile ♂ grabbed by adult ♂
2014.11.09 © Kluck, A.
Sminthurides penicillifer ♂ & ♀ juv. from the UK
Courtship practise
2014.12.09 © Ardron,P.A.

  Subspecies ssp.1 Janssens F & Murray A, 2012
Remark by Janssens, F. 2017.03.03: With white circumlateral band.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.21 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.04.26 © Murray, A.
Sminthurides nr penicillifer ♂ from the UK
2012.04.26 © Murray, A.
Sminthurides nr penicillifer from the UK
Courtship ritual
2012.04.26 © Murray, A.
Sminthurides nr penicillifer from the UK
2012.04.30 © Murray, A.
 
Sminthurides nr penicillifer from the UK
2012.04.30 © Murray, A.
 
Sminthurides nr penicillifer from the UK
Footcomplex with split empodium
2012.05.01 © Shaw, P.
Sminthurides nr penicillifer from the UK
2012.05.02 © Murray, A.
 
Sminthurides nr penicillifer from the UK
2012.05.02 © Murray, A.
 
Sminthurides nr penicillifer juv. from the UK
2012.05.24 © Murray, A.
Sminthurides nr penicillifer juv. ♂ from the UK
2012.05.28 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.06.02 © Murray, A.
Sminthurides nr penicillifer ♀ & ♂ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
 
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
 
Sminthurides nr penicillifer ♀ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer ♀ & ♂ from the UK
2012.06.04 © Murray, A.
Sminthurides nr penicillifer juv. ♂ from the UK
2012.06.06 © Murray, A.
 
Sminthurides nr penicillifer from the UK
Lamellate mucro
2012.06.dd © Shaw, P.
Sminthurides nr penicillifer ♀ from the UK
2013.09.24 © Murray, A.
Sminthurides nr penicillifer from the UK
2013.09.24 © Murray, A.
Sminthurides nr penicillifer from the UK
2013.10.05 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ juv. from the UK
Courtship ritual
2013.10.08 © Nurcombe, E.
Sminthurides nr penicillifer ♀ from the UK
Eyepatch with 8 ocelli: 6 large 2 small
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Courtship ritual 1. ♂ clasps to ♀
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Courtship ritual 2. ♀ lifts up ♂
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from the UK
Clasping mechanism
2014.07.07 © Nurcombe, E.
Sminthurides nr penicillifer ♂ & ♀ from Germany
Courtship
2014.09.12 © Beck, K.
Sminthurides nr penicillifer ♂ & ♀ from Germany
♂ approaches ♀ in courtship
2014.09.19 © Beck, K.
Sminthurides nr penicillifer ♂ & ♀ from Germany
♂ grasps antenna of ♀ in courtship
2014.09.19 © Beck, K.
Sminthurides nr penicillifer ♀ from Germany
Facial pigmentation; lamellate mucro
2014.09.20 © Beck, K.
Sminthurides nr penicillifer ♀ from Germany
Anal segment
2014.09.20 © Beck, K.
Sminthurides nr penicillifer ♀ & ♂ from Holland
Courtship
2015.07.02 © van Leeuwen, J.
Sminthurides nr penicillifer ♂ & ♀ from Germany
Courtship
2015.08.13 © Jäschke, J.
Sminthurides nr penicillifer ♀ from Germany
Eyepatches
2015.08.13 © Jäschke, J.

  Subspecies bifidus Mills, HB, 1934:90
            Syn.: Sminthurus penicillifer Guthrie, JE, 1903:55
Sminthurides (Sminthurides) bifidus ♀ from the USA
After Maynard EA PlateXXII Fig.366
Sminthurides (Sminthurides) bifidus ♀ from the USA
After Maynard EA PlateXXII Fig.367

             Syn.: Sminthurides (Sminthurides) bifidis(sic) Richards, 1956 ISS
             Syn.: Sminthurides (Sminthurides) bifidis(sic) Richards, WR, 1968:19 ISS
             Syn.: Sminthurides bificus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Sminthurides bifidus ♂ & ♀ from Russia
From aquarium
2005.mm.dd © Potapov, M.
Sminthurides bifidus from Germany
Eye cluster with 7 eyes
2006 © Schulz, H.-J. & Schubert H.D.
Sminthurides bifidus from the USA
2008.01.16 © Mcmillan, A.S.
 
Sminthurides bifidus ♀ from the USA
2009.02.15 © Fleming, K.
Sminthurides bifidus from the USA
2009.05.02 © Zungri, D.
Sminthurides bifidus ♀ from the USA
2012.04.23 © Kouri, J.
Sminthurides bifidus from the USA
Male approaching female
2012.04.23 © Kouri, J.
Sminthurides bifidus from the USA
Male grasped juvenile female
2012.04.23 © Kouri, J.
Sminthurides bifidus from Germany
Habitus dorsal
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Habitus lateral
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Mucro SEM 400x
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from Germany
Foot complex of leg1 or leg2
2012.05.04 © Schulz, H.-J.
Sminthurides bifidus from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus ♂ from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus ♂ & ♀ from the USA
2012.05.07 © Kouri, J.
Sminthurides bifidus from the USA
2012.05.13 © Kouri, J.
Sminthurides bifidus ♀ from the USA
2012.07.11 © Janssens, F.
Leg. Kouri, J.
Sminthurides bifidus ♀ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ & ♂ from the UK
2013.04.19 © Murray, A.
Sminthurides bifidus ♀ from the UK
Composed eye with 6 large ocelli and 2 distinctly smaller ocelli
2013.04.23 © Murray, A.
Sminthurides bifidus ♂ from the UK
2013.04.23 © Murray, A.
Sminthurides bifidus ♀ from Germany
2015.05.23 © Gutekunst, V.
Sminthurides penicillifer ssp. bifidus ♀ France
On aquarium
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ France
on aquarium; ant.4 subdivided
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ juv. & ♂ France
On aquarium; courtship
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♂ France
On aquarium; with modified grasping antennae
2017.02.08 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France
On aquarium; albino
2017.03.01 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France
On aquarium
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France
On aquarium; 8 ocelli (C+D small)
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ juv. & albino ♂ from France
On aquarium
2017.03.02 © Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France on aquarium
Ant.4 with intersubdivisional rings
2017.04.11 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France on aquarium
Foot3 with bifid unguiculus
2017.04.21 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♀ from France on aquarium
Mucro with shortened outer lamella
2017.04.21 © Janssens, F. Leg.: Picard, J.
Sminthurides penicillifer ssp. bifidus ♂ from France on aquarium
Ant.2+3 clasping mechanism
2017.04.21 © Janssens, F. Leg.: Picard, J.

Species pilleatus (Yao AL & Kluge G, 1995) Janssens F, 2007
Sminthurus pilleatus
1995 © Yao, A.-L. & Kluge, G.
Sminthurus pilleatus
2000 © Chen, S.

Species plicatus (Schött, H, 1891:13) Stach, 1956, i.s.
Sminthurus plicatus from the USA
After Schött H, 1891 Taf.II Fig.2

Species potamobius Yosii, 1970
Species pseudassimilis Stach, J, 1956:43
            Syn.: Sminthurides assimilis Linnaniemi, 1912, nec Krausbauer, 1898
            Syn.: Sminthurides assimilis Folsom & Mills, 1938, nec Krausbauer, 1898
            Syn.: Sminthurides signatus var. distincta Linnaniemi, WM, 1912:262
            Syn.: Sminthurus signatus Krausbauer, 1898
            Syn.: Sminthurides sp. Axelson, 1905
Species quinquesensillatus Hüther, 1967
Species rapoporti Delamare Deboutteville & Massoud, 1963:187
            Syn.: Sminthurides Rapoporti(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species ringueleti Delamare Deboutteville & Massoud, 1963:182
            Syn.: Sminthurides Ringueleti(sic) Delamare Deboutteville & Massoud, 1963 IOS
Species rupium Najt, 1967
Species schoetti Axelson, WM, 1903:12, B
            Syn.: Sminthurides Schötti(sic) Axelson, WM, 1903:12 IOS
            Syn.: Sminthurus signatus Krausbauer, 1898, nec Templeton, 1835
            Syn.: Sminthurides sp. Schött, 1902
            Syn.: Sminthurides Schötti(sic) var. bilineata Axelson, WM, 1903:13
            Syn.: Sminthurides Schötti(sic) var. ornata Axelson, WM, 1903:13
Sminthurides schoetti var. ornata ♀ from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornata ♀ from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornata ♀ from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornata ♀ from the UK
2012.07.08 © Nurcombe, E.
Sminthurides schoetti var. ornata ♀ from the UK
2012.07.08 © Nurcombe, E.


            Syn.: Sminthurides krausbaueri Folsom & Mills, 1938
             Syn.: Sminthurides schötti(sic) Axelson, 1904 ISS
             Syn.: Sminthurides schötti(sic) Axelson, 1905 ISS
             Syn.: Sminthurides schötti(sic) Linnaniemi, WM, 1912:267 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) Handschin, E in Dahl, F, 1929:110 ISS
             Syn.: Sminthurides (Sminthurides) schötti(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Sminthurides schötti(sic) Joosse-Van Damme, ENG et Naaktgeboren, C, 1961:34 ISS
             Syn.: Sminthurides schötti(sic) Paclt, J, 1961:235 ISS
Sminthurides schoetti from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides schoetti
2006 © Boldsystems.org
 
Sminthurides schoetti juv. from the UK
2012.06.21 © Murray, A.
 
Sminthurides schoetti juv. from the UK
2012.06.27 © Nurcombe, E.
Sminthurides schoetti juv. from the UK
2012.06.27 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
Distinctly pale basal antennal segment
2012.07.08 © Nurcombe, E.
Sminthurides schoetti from the UK
Pale colour form
2012.07.08 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
Subdivided 4th antennal segment
2012.07.08 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2012.09.07 © Nurcombe, E.
 
Sminthurides schoetti ♀ from the UK
2012.10.07 © Murray, A.
Sminthurides schoetti ♀ from the UK
2012.10.07 © Murray, A.
Sminthurides schoetti ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides schoetti ♀ & ♂ from the UK
2012.10.09 © Nurcombe, E.
Sminthurides nr schoetti ♀ juv. from Portugal
2012.10.16 © Murray, A.
Sminthurides schoetti juv. from the UK
2012.12.21 © Barton, T.
Sminthurides schoetti ♂ juv. from the UK
Swollen 2nd & 3rd antennal segment
2013.09.18 © Nurcombe, E.
Sminthurides schoetti juv. from the UK
2013.09.24 © Nurcombe, E.
Sminthurides schoetti ♀ from the UK
2013.09.24 © Nurcombe, E.
Sminthurides schoetti ♀ from Holland
2014.10.05 © van Duinen, J.
Sminthurides schoetti ♀ from Holland
2014.10.09 © van Duinen, J.
Sminthurides schoetti ♂ & ♀ from Holland
Courtship
2014.11.17 © van Duinen, J.
Sminthurides schoetti ♂ & ♀ from Holland
Courtship
2014.11.17 © van Duinen, J.
Sminthurides schoetti ♀ from Belgium
Facial chaetotaxy
2016.01.02 © Huskens, M.-L.
Sminthurides schoetti ♀ from Belgium
2017.03.08 © Huskens, M.-L.

Species sensillatus Massoud & Betsch, 1966
Species sexoculatus Betsch, J-M et Massoud, Z, 1970:213
Sminthurides sexoculatus ♀ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus ♀ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus subadult ♀ & ♂ from France
2017.03.20 © Picard, J.
Sminthurides sexoculatus ♀ from France
2017.03.22 © Picard, J.

Species sexsensillatus Hüther, 1967
Species signatus (Krausbauer, 1898) Heymons, R & Heymons, H in Brauer, A, 1909:12
            Syn.: Sminthurus assimilis Krausbauer, 1898
Sminthurides (Sminthurides) assimilis from the USA
After Maynard, E.A., 1951 Pl.XXII Fig.364.

            Syn.: Sminthurides krausbaueri Folsom & Mills, 1938
Sminthurides signatus from Germany
After Heymons R & Heymons H, 1909 Fig.17
Sminthurides signatus from the UK
Mucrones
2004 © Hopkin, S.P.
Sminthurides signatus from the UK
2012.07.22 © Murray, A.
 
Sminthurides signatus from the UK
2012.07.22 © Murray, A.
Sminthurides signatus from the UK
2012.08.14 © Murray, A.
Sminthurides signatus ♂ from the UK
2012.08.14 © Murray, A.
Sminthurides signatus juv. from the UK
2012.09.09 © Nurcombe, E.
Sminthurides signatus juv. from the UK
2012.09.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.09 © Nurcombe, E.
Sminthurides signatus from the UK
2012.10.21 © Murray, A.
Sminthurides signatus ♀ from the UK
2012.10.25 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2012.10.25 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♂ from the UK
2013.01.04 © Nurcombe, E.
Sminthurides signatus ♀ from the UK
2013.01.04 © Nurcombe, E.

Species spegazzinii Börner, 1907
             Syn.: Sminthurides Spegazzinii Murphy, 1960 ISS
             Syn.: Sminthurides Spegazzinii Delamare Deboutteville & Massoud, 1963 ISS
Species sundanus Yoshii, R et Suhardjono, YR, 1992:89
Sminthurides sundanus from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.8A

Species terrestris Maynard, EA, 1951:213
Sminthurides terrestris from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.396.

Species tropicus Delamare Deboutteville, 1952
Species velii Prabhoo, 1971
             Syn.: Sminthurides velli(sic) Bano, R, 2016:56 ISS
             Syn.: Sminthurides velli(sic) Bano, R, 2016:64 ISS
Species ventanae Najt, 1969
Species weichseli Christiansen & Bellinger, 1981
Sminthurides weichseli from the USA
Eggs deposited in epidermal cavity
2013.09.07 © Krebs, C.
Sminthurides weichseli from the USA
Pale embryo and well pigmented embryo
2013.09.07 © Krebs, C.
Sminthurides weichseli juv. from the USA
2013.09.07 © Krebs, C.
Sminthurides weichseli ♂ & ♀ from the USA
2013.09.10 © Krebs, C.
Sminthurides weichseli ♂ & ♀ from the USA
Ant.4 with weakly developed, spherical subapical knob (*)
2013.09.10 © Krebs, C.


Genus Denisiella Folsom & Mills, 1938 key  key
Species sp.1 Christiansen & Bellinger, 1981
Species sp.2 Bellini, BC et Zeppelini, D, 2004:588
Species bretfeldi Schulz, H-J et van Harten, A, 2013:15
Denisiella bretfeldi ♀ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.5
Denisiella bretfeldi ♂ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.6

Species colombiana Ospina, M et Palacios-Vargas, JG, 2009:63
Species diomedesi Palacios-Vargas, JG, 2007:64
Species lithophila Snider, RJ, 1988:124
             Syn.: Sminthurides lithophila(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Species maesorum Palacios-Vargas, JG, 1995:28
Species nayarita Palacios-Vargas, JG et Bernava, V, 1999:447
Species ramosa (Folsom, JW, 1932:72) Stach, J, 1956:21
Species seurati (Denis, 1925) Stach, J, 1956:11, t.t.
Species serroseta (Börner, C in Schultze, 1908:58) Stach, J, 1956:56
            Syn.: Sminthurides (Stenacidia) serroseta(sic) Börner, C in Schultze, 1908:58 IOS
            Syn.: Sminthurides distilacinia(sic) Crampton, 1922 IOS
             Syn.: Sminthurides (Stenacidia) serroseta(sic) Börner in Voeltzkowi, 1907 ISS
             Syn.: Sminthurides (Denisiella) serroseta(sic) Folsom & Mills, 1938 ISS
             Syn.: Sminthurides (Stenacidia) serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides (Denisiella) serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Denisiella serrosetosa(sic) Stach, J, 1956:56 ISS
             Syn.: Sminthurides (Denisiella) serroseta(sic) Paclt, 1959 ISS
Denisiella serroseta ♀ from the UAE
After Schulz H-J & van Harten A, 2013 Pl.7

Species sexpinnata (Denis, 1931) Stach, J, 1956:21
             Syn.: Denisiella sexpinnatus(sic) Snider, RJ, 1988:128 ISS
             Syn.: Denisiella sexpinnatus(sic) Palacios-Vargas, JG, 1995:33 ISS
             Syn.: Sminthurides (Denisiella) sexspinnatus(sic) Skidmore, RE, 1995:60 ISS
             Syn.: Denisiella sexpinnatus(sic) Palacios-Vargas, JG et Bernava, V, 1999:447 ISS
             Syn.: Denisiella sexpinnatus(sic) Ospina, M et Palacios-Vargas, JG, 2009:67 ISS

Genus Debouttevillea Murphy, DH, 1965:31
  Syn.: Sminthuridinae Deboutevillea(sic) Murphy, DH, 1965:31 IOS
Species marina Murphy, DH, 1965:31, t.t.

Genus Pygicornides Betsch, J-M, 1969:355
Species stagnalis (Womersley, H, 1932:16) Greenslade, P in Houston, WWK, 1994:129
Species torridus Betsch, J-M, 1969:349, t.t.

Genus Sinnamarides Betsch, J-M et Waller, A, 1991:234
Species delamarei Betsch, J-M et Waller, A, 1991:230, t.t.

Genus Sminthuridia Massoud & Betsch, 1972
Species sphaeridioides (Murphy, 1960) Massoud & Betsch, 1972

Genus Yosiides Massoud & Betsch, 1972
Species chinensis Itoh & Zhao, 1993
Species himachal (Yosii, 1966) Massoud & Betsch, 1972

Genus Stenacidia Börner, C, 1906:182
  Syn.: Sminthuridae Jeannenotia Stach, J, 1956:10
  Syn.: ... Sminthurides Jeannenot, 1955
  Syn.: ... Spinosellina Palissa, 1961 partim
Species picta Yosii, 1966
Species stachi (Jeannenot, 1955) Dallai, R, 1973:547
Species violacea (Reuter, 1881) auctt., t.t.
            Syn.: Stenacidia violacea var. variabilis Krausbauer, 1898
Sminthurides (Stenacidia) violaceus var. variabilis from Germany
After Heymons R & Heymons H, 1909 Fig.13

            Syn.: Arrhopalites adelaidica Womersley, 1933 partim
            Syn.: Arrhopalites coccineus Salmon, JT, 1941:408
Arrhopalites coccineus from New Zealand
After Salmon JT, 1941 Pl.70 Fig.484

            Syn.: Jeannenotia stachi ssp. australiensis Betsch, J-M et Massoud, Z, 1970:173
Sminthurus violaceus from Finland
After Schött H, 1893 Taf.II Fig.8

             Syn.: Sminthurides (Stenacidia) violacea(sic) Womersley, H, 1932:14 ISS
Sminthurides (Stenacidia) violaceus ♂ from Australia
After Womersley H, 1932 Fig.2b
Sminthurides violaceus from the USA
After Maynard, E.A., 1951 Pl.XXIII Fig.412.
 
Stenacidia violacea from the UK
Male antenna
2004 © Hopkin, S.P.
Stenacidia violacea from the UK
2011.08.17 © Robertson, A.
Stenacidia violacea from the UK
2011.08.17 © Robertson, A.
Stenacidia violacea ♂ juv. from the UK
With partly developed grasping antennae
2011.08.17 © Robertson, A.
Stenacidia violacea ♀ & ♂ from the UK
Courtship
2012.06.21 © Murray, A.
Stenacidia violacea ♀ & ♂ from the UK
Courtship
2012.06.21 © Murray, A.
Stenacidia violacea ♂ from the UK
Abdominal dorsal rows of setae
2012.06.21 © Murray, A.
Stenacidia violacea ♂ from the UK
Grasping antenna
2012.06.22 © Murray, A.
Stenacidia violacea ♀ from the UK
2012.06.22 © Murray, A.
Stenacidia violacea juv. from the UK
2012.07.20 © Murray, A.
Stenacidia violacea ♂ from the UK
2012.08.08 © Nurcombe, E.
Stenacidia violacea ♂ from the UK
2012.08.08 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2012.08.19 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♂ & ♀ from the UK
♂ grasps ♀ with modified antennae
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♀ & ♂ from the UK
♀ lifts up ♂ in courtship ritual
2012.09.07 © Nurcombe, E.
Stenacidia violacea ♂ from the UK
2012.10.10 © Nurcombe, E.
Stenacidia violacea ♀ from France
2013.03.dd © Laprevote, S.
Stenacidia violacea ♀ from the UK
2013.04.01 © Nurcombe, E.
Stenacidia violacea ♀ from the UK
2013.09.24 © Nurcombe, E.
Stenacidia violacea ♂ from the UK
Male clasping antenna
2013.10.24 © Nurcombe, E.
Stenacidia violacea ♂ & ♀ from the UK
Male clasped to antennae of female
2013.10.24 © Nurcombe, E.
Stenacidia violacea ♂ juv. from Holland
2014.10.03 © van Duinen, J.
Stenacidia violacea ♂ from Holland
Antennal clasping organ
2014.10.04 © van Duinen, J.
Stenacidia violacea ♀ & ♀ from Holland
Courtship
2014.10.04 © van Duinen, J.
Stenacidia violacea ♂ from Holland
Antennal clasping organ
2014.10.05 © van Duinen, J.
Stenacidia violacea ♂ from Holland
2014.10.09 © van Duinen, J.


Genus Pedonides Bretfeld, G, 2010:317
Species alcochetensis Bretfeld, G, 2010:318, t.t.
Pedonides alcochetensis from Portugal
with clasping second legs
After Bretfeld G, 2010 Fig.1,13


Genus Boernerides Bretfeld, G, 1999:32
Species hystrix (Börner, 1903) Bretfeld, G, 1999:34, t.t.

Stenacidia hystrix ♀ from Italy
After Dallai R, 1973 Fig.17a
Stenacidia hystrix ♂ from Italy
After Dallai R, 1973 Fig.17b
Stenacidia hystrix ♂ from Italy
After Dallai R, 1973 Fig.18a




References