http://www.collembola.org/taxa/willinae.htm - Last updated on 2017.01.27 by Frans Janssens
Checklist of the Collembola: Willowsiinae

Subfamilia Willowsiinae Yoshii R & Suhardjono YR, 1989:35 key  key  key
Genus Number
of species
Americabrya3
Desertia3
Drepanosira18
Epimetrura4
Hawinella2
IncertaeSedis4
Janetschekbrya2
Lepidobrya4
Lepidocyrtoides14
Lepidodens3
Lepidosinella1
Lepidosira60
Szeptyckiella3
Willowsia33
Willowsiinae154
Genera with number of species
  Syn.: Entomobryidae Willowsini(sic) Yoshii, R et Suhardjono, YR, 1989:35 ISS
  Syn.: Siraeformes Willowsini(sic) Yosii & Suhardjono, 1989 ISS
Willowsia buski from France
2007.06.09 © Lebeaux, P.
Willowsia platani from the UK
2007.06.30 © Cornwall, N.J.

  Syn.: Entomobryinae Willowsini(sic) Soto-Adames, FN, Barra, J-A, Christiansen, K et Jordana R, 2008:502 ISS
Remark by Janssens, F. 2016.12.26: Yoshii & Suhardjono, 1989 defined Willowsiini as a tribe of Entomobryidae. In this checklist, the polyphyletic scaled Entomobryinae sensu Zhang et al, 2016 are maintained at subfamily level mainly for convenience.



Genus IncertaeSedis Zhang, F, Chen, J-X et Deharveng, L, 2011:8
Species brahma (Imms, 1912) Zhang, F, Chen, J-X & Deharveng, L, 2011:8
             Syn.: Sira(sic) brahama(sic) Denis, JR, 1936:266 ISS
             Syn.: Sira(sic) brahama(sic) Yosii, R, 1942:490 ISS
Species crassicornis (Nicolet, H, 1847:370) Janssens, 2016
            Syn.: Sira(sic) intermedia Schött, 1921 =?
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species intermedia (Schött, 1921) Zhang, F, Chen, J-X & Deharveng, L, 2011:8
Species membranea (Nicolet, H, 1847:371) Janssens, 2016
            Syn.: Entomobrya multifasciata Handschin, 1924
Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

             Syn.: Degeeria membranacea(sic) Kolenati, 1858 ISS

Tribus Willowsiini Yoshii R & Suhardjono YR, 1989:35

Genus Willowsia Shoebotham, 1917 key  key
  Syn.: ... Seira Lubbock, J, 1873:143 partim
Willowsia platani from the UK
Apically pointed scales
2013.12.24 © Murray, A.
Willowsia nigromaculata from the USA
Apically pointed scales
2015.02.13 © Dean, T. & S.B.
Willowsia sp. from Tasmania
2015.09.26 © Tony.
Willowsia sp. from Tasmania
Apically pointed scales
2015.09.26 © Tony.
Willowsia nr nigromaculata from Australia
Apically pointed scales
2015.10.01 © Anonymous.
Willowsia sp. from Ireland
Apically pointed scales with long basal ribs
2015.11.26 © Gaff, K.
Willowsia sp. from Ireland
Broad apically pointed scales with long uninterrupted apically raised ribs
2015.11.26 © Gaff, K.
Willowsia nigromaculata from the UK
Apically pointed backwards directed scales
2016.01.14 © Horton, M.

Species sp.1 Zhang, F, Chen, J-X et Deharveng, L, 2011:3
Species sp.2 Katz, AD, Giordano, R et Soto-Adames, FN, 2015:8
Species abrupta (Schött, H, 1917:30) Greenslade, P in Houston, WWK, 1994:102
Sira(sic) abrupta from Australia
After Schött H, 1917 Fig.36

Species bartkei Stach, J, 1965:361
Willowsia bartkei from Vietnam
After Zhang F, 2015 Fig.26

Species buski (Lubbock, J, 1870:280) Schoebotham, 1917, B
            Syn.: Seira Buskii(sic) Lubbock, J, 1870:280 IOS
            Syn.: Degeeria pruni Nicolet, H, 1842:73
            Syn.: Degeeria cyanea Tullberg, 1871
            Syn.: Sira flava Ågren, 1903
            Syn.: Sira corticaloides Stach, 1922
             Syn.: Sira(sic) Buskii(sic) Tullberg, 1872 ISS
             Syn.: Seira Buskii(sic) Lubbock, J, 1873:145 ISS
Seira Buskii(sic) from the UK
After Lubbock J, 1873 Pl.22
Seira Buskii(sic) from the UK
Apically pointed scale with long basal ribs
After Beck J in Lubbock J, 1873 Pl.70

             Syn.: Seira Buskii(sic) Beck, J in Lubbock, J, 1873:253 ISS
             Syn.: Sira(sic) Buskii(sic) Reuter, OM, 1876:81 ISS
             Syn.: Sira(sic) buskii(sic) Tullberg, 1872 ISS
             Syn.: Seira Buski(sic) Lameere, A, 1895:593,B+ ISS
             Syn.: Sira(sic) Buski(sic) Carl, J, 1899:343 ISS
             Syn.: Sira(sic) busquii(sic) Krausbauer, 1902 ISS
             Syn.: Seira Buskii(sic) Schött, H, 1917:33 ISS
             Syn.: Sira(sic) Buski(sic) Stach, J, 1930:299 ISS
             Syn.: Willowsia buskii(sic) Salmon, JT, 1945:71 ISS
             Syn.: Seira buskii(sic) Kloet, GS et Hincks, WD, 1945:8 ISS
             Syn.: Willowsia buskii(sic) Palacios-Vargas & Gómez, 1991 ISS
             Syn.: Willowsia buskii(sic) Skidmore, 1995 ISS
Willowsia buskii(sic)
After Zhang, Palacios-Vargas & Chen, 2007 Fig.18a.

             Syn.: Willowsia buskii(sic) Fiera, C, 2007:27 ISS
             Syn.: Willowsia buskii(sic) Zhang, F, Palacios-Vargas, JG et Chen, J-X, 2007:36 ISS
Willowsia buski from Germany
2006 © Kohl, F.
Willowsia buski from the USA
2006 © McClarin, J.
Willowsia buski from Germany
2006.12.27 © Kohl, F.
Willowsia buski from Germany
2006.12.30 © Kohl, F.
Willowsia buski from the USA
2007 © McClarin, J.
Willowsia buski juvenile from the USA
2007.02.25 © Murray, T.
Willowsia buski juvenile from the USA
2007.02.25 © Murray, T.
Willowsia buski subadult from the USA
2007.02.25 © Murray, T.
Willowsia buski adult from the USA
2007.02.25 © Murray, T.
Willowsia buski from Canada
2007.04.18 © Luk, S.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from the USA
2007.04.22 © Reed, D.
Willowsia buski from Hungary
2008.10.23 © Keresztes, G.
Willowsia buski from the USA
2009.04.24 © Zungri, D.
Willowsia buski from Holland
2010.01.09 © van Duinen, J.
Willowsia buski from Czechia
2010.08.02 © Deml, M.
Willowsia buski from Czechia
2010.08.05 © Deml, M.
Willowsia buski from the USA
2011.06.07 © Nanz, S.
Willowsia buski juv. from the UK
Descaled specimen
2012.08.06 © Murray, A.
Willowsia buski juv. from the UK
2012.08.25 © Murray, A.

             Syn.: Willowsia buskii(sic) Soto-Adames, FN et Taylor, SJ, 2013:63 ISS
Willowsia buski from the USA
2014.11.10 © Cristofono, P.

             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:11 ISS
             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:15 ISS
             Syn.: Willowsia buskii(sic) Pan, Z et Zhang, F, 2016:23 ISS
             Syn.: Willowsia buskii(sic) Soto-Adames, FN, 2016:3 ISS
Willowsia buski from the USA
2017.01.04 © Dobbs, Br.

Species cassagnaui Zhang, F, 2015:490
Willowsia cassagnaui from Nepal
After Zhang F, 2015 Fig.1

Species fascia Zhang F et Pan Z in Pan, Z et Zhang, F, 2016:3
Willowsia fascia from China
After Pan Z & Zhang F, 2016 Fig.1A
Willowsia fascia from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.1B

Species formosana (Denis, 1929) Yosii, R, 1956:73
Species fuscana (Uchida, 1944)
Species guangdongensis Zhang, F, Xu, G-L et Chen, J-X, 2007:?
Species guangxiensis Shi & Chen, 2004
Species hyalina (Handschin, 1928) Yosii, 1966
Species ieti Yosii, 1971
Species jacobsoni (Börner, 1913) Stach, 1965
            Syn.: Willowsia jacobsoni var. lipostropha Börner, 1913
            Syn.: Sira(sic) tricincta Schött, H, 1917:31
Sira(sic) tricincta from Australia
After Schött H, 1917 Fig.37

            Syn.: Sira(sic) jacobsoni var. indica Handschin, E, 1929:240
            Syn.: Sira(sic) parajacobsoni Denis, 1929
            Syn.: Sira(sic) abrupta Womersley, 1934
            Syn.: Sira(sic) abrupta Womersley, 1937
            Syn.: Sira(sic) abrupta Womersley, 1942
            Syn.: Sira(sic) jacobsoni var. handschini Uchida, 1944
            Syn.: Sira(sic) jacobsoni var. africana Delamare Deboutteville, 1951
            Syn.: Willowsia mesothoraxa Nguyen, 2001
             Syn.: Sira(sic) Jacobsoni(sic) Denis, JR, 1936:266 ISS
Willowsia jacobsoni
After Zhang, Palacios-Vargas & Chen, 2007 Fig.18c.

Species japonica (Folsom, JW, 1898:55) Yosii, 1955
Seira japonica from Japan
After Folsom JW, 1898 Fig.15
            Syn.: Ptenura bimaculata Börner, C, 1909:121
             Syn.: Sira(sic) binoculata(sic) Denis, JR, 1936:265 ISS
            Syn.: Sira(sic) bimaculata Yosii, R, 1942:488
Sira(sic) bimaculata from Japan
After Yosii R, 1942 Abb.11.1
Sira(sic) bimaculata from Japan
After Yosii R, 1942 Abb.11.2

            Syn.: Willowsia kahlertae Christiansen, K et Bellinger, P, 1992:254
            Syn.: Willowsia qui Zhang, F, Chen, J-X et Deharveng, L, 2011:9 partim
            Syn.: Willowsia qui Pan, Z et Zhang, F, 2016:23
Sira(sic) japonica from Japan
After Yosii R, 1942 Abb.18.1
Willowsia japonica ♀ from Japan
After Yoshii, 1992
Willowsia japonica ♂ from Japan
After Yoshii, 1992
Willowsia japonica from Japan
2005.05.12 © Keresztes, G.
Willowsia japonica from Japan
2005.05.12 © Keresztes, G.
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10A
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10B
Willowsia japonica ♀ from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.10C
Willowsia japonica ♀ from Japan
2013.03.12 © Hara, Y.

Species kalatopensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:35
            Syn.: Willowsia kalatopii(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:35 IOS
Willowsia kalatopii(sic) from India
After Baquero E & al, 2015 Fig.22

Species mekila Christiansen, K et Bellinger, P, 1992:256
Species mexicana Zhang, F, Palacios-Vargas, JG et Chen, J-X, 2007:37
            Syn.: Willowsia nigromaculata Palacios-Vargas, 1997
Willowsia mexicana from Mexico
After Zhang, Palacios-Vargas & Chen, 2007 Fig.1

Species neocaledonica Zhang, F, Bedos, A et Deharveng, L, 2014:382
Willowsia neocaledonica from New Caledonia
After Zhang F & al, 2014 Fig.1

Species nigra Zhang, F, Bedos, A et Deharveng, L, 2014:384
Species nigromaculata (Lubbock, J, 1873:146) Shoebotham, 1917, t.t., B
            Syn.: Sira(sic) elongata Nicolet
            Syn.: Seira mimica Harvey, 1894
            Syn.: Lepidocyrtus americanus Marlatt, 1896
Lepidocyrtus americanus from North America
Dorsal aspect
After Howard LO, 1901 Fig.246 left
Lepidocyrtus americanus from North America
Ventral aspect
After Howard LO, 1901 Fig.246 right
Lepidocyrtus americanus from North America
Lateral aspect
After Howard LO, 1901 Fig.247

            Syn.: Lepidocyrtus americanus Christiansen, K et Bellinger, P, 1980:920 =?
            Syn.: Lepidocyrtus americanus Christiansen, K et Bellinger, P, 1998:1020 =?
            Syn.: Degeeria platani Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:9
            Syn.: Willowsia bimaculata Yosii, R, 1977:161 =?
Willowsia nigromaculata from Europe
After Gisin H, 1960 Abb.433
Willowsia nigromaculata from the USA
2006 © Bernard, E.C.
Willowsia nigromaculata from Germany
2006.03.27 © Rodeland, J.
Willowsia nigromaculata from Germany
2006.08.24 © Kohl, F.
Willowsia nigromaculata from Germany
2006.08.24 © Kohl, F.
Willowsia nigromaculata from the USA
2007.03.23 © Bradford, A.
Willowsia nigromaculata from Belgium
2007.05.10 © Janssens, F.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the UK, juvenile
2007.06.09 © Barton, T.
Willowsia nigromaculata from the USA
caught by an ant...
2007.06.13 © DiTerlizzi, T.
Willowsia nigromaculata from the UK
2007.08.22 © Valentine., B
Willowsia nigromaculata from the UK
2007.09.17 © Valentine., B
Willowsia nigromaculata from Hungary
2008.01.12 © Keresztes, G.
Willowsia nigromaculata from Hungary
2008.03.04 © Keresztes, G.
Willowsia nigromaculata from Hungary
2008.03.16 © Keresztes, G.
Willowsia nigromaculata from the USA
2008.03.23 © Tortuga, I.
Willowsia nigromaculata from the USA
2008.04.05 © Justis, P.S.
Willowsia nigromaculata from the USA
2008.09.11 © Bradford, A.
Willowsia nigromaculata from Hungary
2008.10.19 © Keresztes, G.
Willowsia nigromaculata from Russia
2008.10.21 © Macroclub.ru
Willowsia nigromaculata from the USA
2008.12.17 © Bradford, A.
Willowsia nigromaculata from Canada
2009.01.07 © Luk, S.
Willowsia nigromaculata from the USA
2009.05.04 © Murray, T.
Willowsia nigromaculata from the USA
2009.07.23 © Bradford, A.
Willowsia nigromaculata from Czechia
2009.09.14 © Deml, M.
Willowsia nigromaculata from Belgium
2009.12.16 © De Bruyn, L.
Willowsia nigromaculata
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11B
Willowsia nigromaculata
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11C
Willowsia nr nigromaculata from Australia
2011.04.21 © Flyman.
Willowsia nigromaculata from the UK
2011.10.05 © Shaw, P.
Willowsia nigromaculata from the USA
2011.10.13 © Schmidt, M.L.
Willowsia nigromaculata juv. from the USA
2012.01.08 © Allen, K.
Willowsia nr nigromaculata from Australia
2014.12.08 © Flyman.
Willowsia nr nigromaculata from Australia
Dorsal aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Lateral aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Ventral aspect
2015.10.01 © Anonymous.
Willowsia nr nigromaculata from Australia
Head
2015.10.01 © Anonymous.
Willowsia nigromaculata from the UK
2016.01.14 © Horton, M.
Willowsia nigromaculata from the UK
Caudal aspect cephalic mimicry
2016.01.14 © Horton, M.
Willowsia nigromaculata from the USA
2017.01.04 © Dobbs, Br.

Species nivalis Yosii, 1971
            Syn.: Willowsia brahmides Yosii, 1966
Species platani (Nicolet, H, 1842:72) Palissa, 1959, B
Degeeria platani from Switzerland
After Nicolet H, 1842 Pl.8 Fig.4

            Syn.: Podura argenteo cincta Bourlet, 1842
            Syn.: Degeeria platani var. var. Nicolet, H, 1842:73
            Syn.: Sira(sic) elongata Tullberg, 1871, nec Nicolet, 1842
            Syn.: Seira mimica Harvey, 1894
            Syn.: Sira(sic) flava Ågren, 1903
            Syn.: Sira(sic) corticaloides Stach, 1922
            Syn.: Sira(sic) platani ab. corticalis Stach, J, 1930:300
            Syn.: Lepidocyrtus americanus Mills, HB, 1934:71
            Syn.: Lepidocyrtus americanus Denis, 1941
            Syn.: Lepidocyrtus americanus Salmon, JT, 1964:487
            Syn.: Sira(sic) nigromaculata Mills, HB, 1934:71
            Syn.: Willowsia japonica Yosii, R, 1977:161 =?
             Syn.: Sira(sic) plantani(sic) Bryk, F, 1955:1823 ISS
             Syn.: Willowsia planati(sic) Gisin, H, 1960:234 ISS
Willowsia platani from Europe
Modified after Gisin H, 1960 Abb.434
Willowsia platani from Europe
Modified after Gisin H, 1960 Abb.434

Willowsia platani from the UK
Leaf-shaped scales
2005 © Hopkin, S.P.
Willowsia platani from Germany
2006 © Kohl, F.
Willowsia platani from Germany, juvenile
2007 © Peters, J.
Willowsia platani from Germany, juvenile
2007.03.14 © Peters, J.
Willowsia platani from Czechia
2007.07.22 © Krásenský, P.
Willowsia platani from Germany, juvenile
2008.01.11 © Peters, J.
Willowsia platani from the UK, juvenile
2008.04.23 © Barton, T.
Willowsia platani from the UK
2008.05.24 © Barton, T.
Willowsia platani from France
2008.07.09 © Lavoué., B
Willowsia platani from the UK
2009.07.04 © Talbot, M.E.
Willowsia cf. platani from the USA
2009.07.12 © Zungri, D.
Willowsia platani from the UK
2010.03.24 © Barton, T.
Willowsia platani from Europe
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.11A
Willowsia platani from the UK
2011.06.30 © Shaw, P.
Willowsia platani from the UK
2011.06.30 © Shaw, P.
Willowsia platani juv. from the UK
2011.12.11 © Barton, T.
Willowsia platani from Portugal
2012.04.20 © Coelho, J.
Willowsia platani from Spain
2012.09.23 © Campo., B
Willowsia platani juv. from the UK
2012.12.15 © Barton, T.
Willowsia platani from Spain
2013.04.25 © Baquero, E.
Willowsia platani from Holland
2014.01.24 © van Duinen, J.
Willowsia platani from Holland
2014.01.24 © van Duinen, J.
Willowsia platani from Czechia
2014.05.01 © Miroslav, D.
Willowsia platani from the UK
2015.02.09 © Barton, T.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani from Holland
2015.02.27 © Baas, A.H.
Willowsia platani juv. from Holland
2015.02.27 © Baas, A.H.

Species potapovi Zhang, F, Chen, J-X et Deharveng, L, 2011:13
Willowsia potapovi from Azerbaijan
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.8A
Willowsia potapovi from Azerbaijan
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.8B

Species pseudobuski Pan, Z et Zhang, F, 2016:11
            Syn.: Willowsia pseudobuskii(sic) Pan, Z et Zhang, F, 2016:11 IOS
Willowsia pseudobuskii(sic) from China
After Pan Z & Zhang F, 2016 Fig.7A
Willowsia pseudobuskii(sic) from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.7E

             Syn.: Willowsia pseudobuskii(sic) Pan, Z et Zhang, F, 2016:23 ISS
Species pseudoplatani Zhang F et Pan Z in Pan, Z et Zhang, F, 2016:7
Willowsia pseudoplatani from China
After Pan Z & Zhang F, 2016 Fig.4A
Willowsia pseudoplatani from China
Body scales with long basal ribs
After Pan Z & Zhang F, 2016 Fig.4B

Species pseudosocia Stach, J, 1965:363
Species qui Zhang, F, Chen, J-X et Deharveng, L, 2011:9 partim
Willowsia qui from China
After Zhang F, Chen J-X & Deharveng L, 2011 Fig.6A
Willowsia qui from China
After Pan Z & Zhang F, 2016 Fig.13A
Willowsia qui from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.13B

Species samarkandica Martynova, 1972
Species shiae Pan, J-L, Zhang, F et Chen, J-X, 2006:261
Willowsia shiae from China
After Pan J-L, Zhang F & Chen J-X, 2006 Fig.1

Species similis Pan, Z et Zhang, F, 2016:15
Willowsia similis from China
After Pan Z & Zhang F, 2016 Fig.10A
Willowsia similis from China
Body scale with long basal ribs
After Pan Z & Zhang F, 2016 Fig.10D

Species yamashitai Uchida, 1969
Species yiningensis Zhang, F, Chen, J-X et Deharveng, L, 2011:12

Genus Lepidodens Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:600
Species hainanicus Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:610
Lepidodens hainanicus from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3C

             Syn.: Lepidodens hananicus(sic) Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:602 ISS
Species nigrofasciatus Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:602, t.t.
Lepidodens nigrofasciatus from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3A

Species similis Zhang F et Pan Z-X in Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:607
Lepidodens similis from South China
After Zhang F & Pan Z-X in Zhang F & al, 2016 Fig.3B


Genus Janetschekbrya Yosii, 1971 key
  Syn.: Xenyllodes pallescens Scott, HG, 1960:186 partim
  Syn.: Entomobryidae Janetschekbra(sic) Mari Mutt, JA et Palacios-Vargas, JG, 1987:100 ISS
Species brahamides (Denis, JR, 1936:265) Yosii, 1971
Sira(sic) brahamides from India
After Denis JR, 1936 Fig.13

Species himalica Yosii, 1971, t.t.

Genus Americabrya Mari Mutt, JA et Palacios-Vargas, JG, 1987:99 key
  Syn.: Entomobryidae Janetschekbrya Mari Mutt & Palacios-Vargas, 1987 partim
Species arida (Christiansen, K & Bellinger, P, 1980:918) Mari Mutt, JA & Palacios-Vargas, JG, 1987:100, t.t.
            Syn.: Janetschekbrya sp. Palacios-Vargas, 1979
            Syn.: Janetschekbrya sp. Palacios-Vargas, 1981
Americabrya arida from the USA
2015.03.17 © Landerman, J.
Americabrya arida from the USA
2015.03.17 © Landerman, J.
Americabrya arida from the USA
2015.03.23 © Christopherson, S.

Species epiphyta (Loring, 1984) Mari Mutt, JA & Palacios-Vargas, JG, 1987:107
Species matthewsi (Snider, RJ, 1981:39) Mari Mutt, JA & Palacios-Vargas, JG, 1987:107

Genus Drepanosira Bonet, F, 1942:57
Species albescens Stach, J, 1960:538
Species altimontana Stach, 1963
Species frigida (Imms, 1912)
Species gisini Nosek, 1964
Species grisescens Stach, J, 1960:532
Species hussi Neuherz, 1976
Drepanosira hussi from India
After Baquero E & al, 2015 Fig.17

Species lineata Stach, J, 1960:533
Species ombrophila Stach, J, 1960:535
Species ornata (Bonet, 1930) Bonet, F, 1942:58, t.t.
            Syn.: Parasira subornata Denis, 1936
Species ornata (Stach, 1963 nec Bonet, 1930) Baquero, E, Mandal, G & Jordana, R, 2015:34
Species pallida (Brown, JM, 1926:209)
Species pulchra Stach, J, 1960:534
Species rara Stach, J, 1960:537
Species raviensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:32
            Syn.: Drepanosira ravi(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:32 IOS
Drepanosira ravi(sic) from India
After Baquero E & al, 2015 Fig.18
Drepanosira ravi(sic) from India
After Baquero E & al, 2015 Fig.19

Species shimlaensis Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:34
            Syn.: Drepanosira shimlai(sic) Baquero E et Jordana R in Baquero, E, Mandal, G et Jordana, R, 2015:34 IOS
Drepanosira shimlai(sic) from India
After Baquero E & al, 2015 Fig.20
Drepanosira shimlai(sic) from India
After Baquero E & al, 2015 Fig.21

Species stachi Nosek, 1964
Species subornata (Denis, JR, 1936:267) Bonet, F, 1942:58
Parasira subornata from India
After Denis JR, 1936 Fig.21
Parasira subornata from India
After Denis JR, 1936 Fig.22

Species villosa (Börner, 1903)

Genus Desertia Tshelnokov, 1979
Species kaplini Tshelnokov, 1979, t.t.
Species pulchra (Handschin, 1928)
             Syn.: Seira pulcher(sic) Mari Mutt, JA et Bellinger, PF, 1990:106 ISS
             Syn.: Seira pulcher(sic) Culik, MP et Zeppelini, DF, 2003:1129 ISS
Species semicolorata (Handschin, 1928)

Genus Hawinella Bellinger & Christiansen, 1974
Species kuaola Christiansen, K et Bellinger, P, 1992:260
Species lava Bellinger, PF et Christiansen, KA, 1974:36, t.t.

Genus Szeptyckiella Zhang, F, Bedos, A et Deharveng, L, 2014:2 key
Species boulouparica Zhang, F, Bedos, A et Deharveng, L, 2014:4, t.t.
Szeptyckiella boulouparica from New Caledonia
After Zhang F, Bedos A & Deharveng L, 2014 Fig.1A

Species lii Zhang, F, Bedos, A et Deharveng, L, 2014:12
Szeptyckiella lii from South China
After Zhang F, Bedos A & Deharveng L, 2014 Fig.7A

Species sinelloides Zhang, F, Bedos, A et Deharveng, L, 2014:8
Szeptyckiella sinelloides from New Caledonia
After Zhang F, Bedos A & Deharveng L, 2014 Fig.4A


Genus Lepidosinella Handschin, 1920
Species armata Handschin, 1920, t.t.

Tribus Lepidosirini Yoshii R & Suhardjono YR, 1989:36 sensu Zhang F & al, 2017:534

Genus Lepidobrya Womersley, 1937
Species mawsoni (Tillyard, 1920) Greenslade P, 1990, t.t.
            Syn.: Lepidobrya aurantiaca Salmon, JT, 1949:46
             Syn.: Lepidobrya aurantica(sic) Greenslade P, 1990 ISS
             Syn.: Lepidobrya aurantica(sic) Greenslade, P in Houston, WWK, 1994:93 ISS
Lepidobrya mawsoni from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.76.
Lepidobrya mawsoni from Macquarie Island
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.1
Lepidobrya mawsoni from Macquarie Island
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.2
Lepidobrya mawsoni from Macquarie Island
Narrow pointed body scales
After Zhang F, Greenslade P & Stevens MI, 2017 Fig.5

Species mitchelli (Womersley, 1934) Greenslade, P in Houston, WWK, 1994:93
Entomobrya mitchelli from Australia
After Womersley H, 1939 Fig.58M

Species thalassarchia Salmon, JT, 1949:44
Species violacea Salmon, JT, 1949:46

Genus Lepidosira Schött, 1925
  Syn.: Entomobryini Urewera Salmon, JT, 1938:349
  Syn.: Entomobryinae Urewara(sic) Womersley, H, 1939:322 ISS
Lepidosira sp. from Australia
2008.12.10 © Bert
Lepidosira? sp. from Australia
2015.02.28 © Murray, A.

Species sp.1 Yosii, R, 1940:116
Species sp.2 Yayuk, 1989:10
            Syn.: Lepidocyrtus variabilis Oudemans, JT in Weber, M, 1890:74
Species alba (Nguyen, TT, 2005:8) Zhang, F, Chen, J-X & Deharveng, L, 2011:5
Species angulata (Schött, H, 1917:46) Howard, GW, 1969:189
Lepidocyrtoides angulatus from Australia
After Schött H, 1917 Taf.3 Fig.5

             Syn.: Lepidocyrtus (Lepidocyrtoides) angustatus(sic) Schött, 1925 ISS
Species anomala Salmon, JT, 1944:164
Species arborea Salmon, JT, 1944:163
Species australica (Schött, H, 1917:40) Greenslade P & Majer in Dindal, 1980
Lepidocyrtus australicus from Australia
After Schött H, 1917 Taf.3 Fig.1

  Subspecies tamburinensis Schött, H, 1917:41
Lepidocyrtus australicus var. tamburinensis from Australia
After Schött H, 1917 Taf.3 Fig.2

             Syn.: Lepidosira tambourinensis(sic) Yoshii, R et Greenslade, P, 1994:18 ISS
Species bidentata Salmon, JT, 1938:358
Species bifasciata (Salmon, JT, 1944:157)
Species bisecta (Salmon, JT, 1944:162) Greenslade, 2012
Species brunnea (Womersley, H, 1935:217) Greenslade, P in Houston, WWK, 1994:98
Species calolepis (Börner, 1913) Yayuk, 1989:10
            Syn.: Lepidosira montana Schött, 1925
Species congoia Salmon, 1956
Species dorsalis (Salmon, JT, 1941:383) Greenslade, 2012
Urewera flava
subsp. dorsalis from New Zealand
After Salmon JT, 1941 Pl.64 Fig.401

Species faaroana (Carpenter, 1934) Greenslade, 2012
Species fallaciosa Yoshii, 1989
Species flava (Salmon, JT, 1938:353) Greenslade, 2012
Urewera flava from New Zealand
After Salmon JT, 1938 Pl.37 Fig.38
Urewera flava from New Zealand
After Salmon JT, 1941 Pl.64 Fig.400

Species fuchsiata (Salmon, JT, 1938:350) Greenslade, 2012, t.t.
Urewera fuchsiata from New Zealand
After Salmon JT, 1938 Pl.35 Fig.1
Urewera fuchsiata from New Zealand
After Salmon JT, 1941 Pl.63 Fig.390

Species fuscata Womersley, 1930
Species gigantea (Börner, C, 1909:122) Yosii, R, 1956:73
Species glebosa Salmon, JT, 1941:394
Species gupta Howard, GW, 1969:189
Lepidosira gupta from Australia
After Howard GW, 1969 Fig.1

Species ianthina (Salmon, JT, 1941:378) Greenslade, 2012
Species inconstans (Salmon, JT, 1938:353) Greenslade, 2012
Urewera inconstans from New Zealand
After Salmon JT, 1941 Pl.64 Fig.394

Species indistincta Salmon, JT, 1938:359
Species javana (Börner, 1913) Yayuk, RS (sic), 1989:10
Species laboriosa Greenslade, P in Houston, WWK, 1994:98
            Syn.: Lepidocyrtoides cinctus Schött, H, 1917:45 nec Schäffer, 1898
Species lichenata (Salmon, JT, 1938:352) Greenslade, 2012
Species longicornis (Schött, H, 1917:38) Yoshii, R & Greenslade, P, 1994:14
Lepidocyrtoides longicornis from Australia
After Schött H, 1917 Taf.3 Fig.6

Species magna (Salmon, JT, 1937:355) Greenslade, 2012
Pseudosinella magna from New Zealand
After Salmon JT, 1937 Pl.50 Fig.e
Urewera tridentifera from New Zealand
After Salmon JT, 1938 Pl.35 Fig.8
Urewera magna from New Zealand
After Salmon JT, 1941 Pl.64 Fig.397

Species minima Salmon, JT, 1938:356
Lepidosira minima from New Zealand
After Salmon JT, 1938 Pl.36 Fig.15
Lepidosira minima from New Zealand
After Salmon JT, 1941 Pl.65 Fig.416

Species minuta Salmon, JT, 1938:356
Lepidosira minuta from New Zealand
After Salmon JT, 1938 Pl.36 Fig.22

Species nigrocephala (Womersley, H, 1936:480) Yosii, 1960
Mesira nigrocephala from Australia
After Womersley H, 1936 Fig.4a

Species nigropunctata (Nguyen, TT, 2005:8) Zhang, F, Chen, J-X & Deharveng, L, 2011:5
Species nilgiri (Denis, JR, 1936:266) Yosii, 1955
Sira(sic) nilgiri from India
After Denis JR, 1936 Fig.17
             Syn.: Lepidosira nilghiri(sic) Yosii, 1955 ISS
Species obscura (Salmon, JT, 1944:160) Greenslade, 2012
Species okarita Salmon, JT, 1938:360
Species omniofusca Salmon, JT, 1941:391
Species pallida (Ritter, 1911) Salmon, JT, 1964:507
Species parva (Salmon, JT, 1941:379) Greenslade, 2012
Species pigmenta Salmon, JT, 1944:164
Species punctata Yosii, 1960
Species purpurea (Salmon, JT, 1938:354) Greenslade, 2012
Species quadradentata (Salmon, JT, 1941:382) Greenslade, 2012
Urewera quadradentata from New Zealand
After Salmon JT, 1941 Pl.63 Fig.387

Species reducta (Salmon, JT, 1938:355) Greenslade, 2012
Species rotorua Salmon, JT, 1938:357
Lepidosira rotorua from New Zealand
After Salmon, J.T., 1938 Pl.36 Fig.24.

Species sagmaria (Schött, H, 1917:43) Schött, 1925
Lepidocyrtoides sagmarius from Australia
After Schött H, 1917 Fig.45
             Syn.: Lepidosira (Lepidocyrtoides) sagmarius(sic) Womersley, 1930 ISS
             Syn.: Lepidosira sagmarius(sic) Womersley, H, 1936:328 ISS
             Syn.: Lepidosira sagmarius(sic) Salmon, JT, 1941:390 ISS
Lepidosira sagmarius(sic) from New Zealand
After Salmon JT, 1941 Pl.66 Fig.430

Species sexmacula Salmon, JT, 1938:358
Lepidosira sexmacula from New Zealand
After Salmon JT, 1938 Pl.37 Fig.29
Lepidosira sexmacula from New Zealand
After Salmon JT, 1941 Pl.65 Fig.412

Species splendida (Salmon, JT, 1941:380) Greenslade, 2012
Urewera splendida from New Zealand
After Salmon JT, 1941 Pl.63 Fig.382

Species sundana Yoshii, R et Suhardjono, YR, 1989:38
Species tapuia Arlé & Guimaraes, 1981
Species terraereginae (Ellis, WN & Bellinger, PF, 1973:28) Howard, GW, 1969:189, t.t.
            Syn.: Lepidocyrtoides coeruleus Schött, H, 1917:45 nec Ritter, 1911
             Syn.: Lepidosira coerulus(sic) Howard, GW, 1969:189 ISS
Species unguserrata Salmon, 1970
Species variabilis (Oudemans, 1890)
Species variabilis (Schäffer, 1897) Christiansen & Bellinger, 2000:42, nec Oudemans, 1890
  Varietas albella Börner, 1906
  Varietas albiceps Börner, 1906
  Varietas bimaculata Börner, 1906
Species vicina Yoshii, 1989
Species villasboasi Arlé & Guimaraes, 1981
Species violacea (Salmon, JT, 1938:352) Greenslade, 2012
Species violaceapallipes (Denis, 1931) Christiansen, K & Bellinger, P, 2000:42
            Syn.: Lepidocyrtinus domesticus f. violacea-pallipes(sic) Denis, 1931 IOS
Species womersleyi Greenslade P in Houston, 1994:99

Genus Epimetrura Schött, 1925
Species caudata (Carpenter, 1917) Denis, 1948
Lepidocyrtus caudatus from India
After Mandal GP & Hazra AK, 2009 Fig.31a

Species mirabilis Schött, 1925, t.t.
Species rostrata Greenslade, P et Sutrisno, 1994:53
Species tonkinensis Denis, 1948

Genus Lepidocyrtoides Schött, 1917
  Syn.: Entomobryinae Promesira Womersley, H, 1942:29
  Syn.: Entomobryidae Najtsira Yoshii, 1989
  Syn.: Entomobryidae Sepidocyrtoides(sic) Zhang, F, Bedos, A et Deharveng, L, 2016:19 ISS
Lepidocyrtoides sp. from India
2016.03.15 © Mandal G.P.

Species bicolor Yoshii, R et Suhardjono, YR, 1992:17bis
Species cheesmani Womersley, 1937
Species cucullaris Schött, H, 1917:39, t.t.
Lepidocyrtoides cucullaris from Australia
After Schött H, 1917 Taf.3 Fig.3

Species fasciatus (Womersley, 1934) Greenslade, P in Houston, WWK, 1994:94
Species flavocinctus Schött, H, 1917:42
Lepidocyrtoides flavocinctus from Australia
After Schött H, 1917 Fig.44

  Subspecies unicolor Womersley, 1934
Species matahari Yoshii, R et Suhardjono, YR, 1992:21
Species meraukeae Yoshii, R et Suhardjono, YR, 1992:19
Species najtae (Yoshii, 1989) Yoshii, R & Greenslade, P, 1994:11
Species novacaledoniae Yosii, 1960
             Syn.: Lepidocyrtoidea(sic) novaecaledoniae(sic) Deharveng, L et Najt, J in Tillier, S, 1988:18 ISS
Species oliveri Liu, L, Chen, J-X et Greenslade, P, 2008:?
Species quatuordecimocellata Prabhoo, 1971
Species striatus (Schött, 1901)
Species timorensis Yoshii, R et Suhardjono, YR, 1992:80
Lepidocyrtoides timorensis from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.3A

Species yasumatsui Uchida, 1944



References