http://www.collembola.org/doc/hist2004.htm - Last updated on 2007.11.06 by Frans Janssens
Checklist of the Collembola: History - 2004

Preface

This checklist has been prepared by Frans Janssens, member of the Royal Entomological Society of Antwerp, Belgium. The purpose of this check list is to resolve synonomy related problems. Typical example: given a taxon name, verify whether or not it is currently valid and if not what is then its currently valid name.

This list is updated regularly. Unfortunately, it still contains a lot of mistakes. :-(
Typical errors are a) duplication of the same species name in several genera, b) duplication of taxa at different rank (e.g., once listed at species rank and once more at subspecies rank), c) synonyms listed as valid taxa, c) incorrect spelling of the name of a taxon or author, d) incorrect publication date, e) incorrect revision authority.
Your comments, flames, remarks, suggestions, etc are welcome at Frans Janssens

Frans Janssens
2004.12.30

Status

Status of the checklist
Group # of Genera # of Species Species/Genus ratio
Actaletidae2105
Anurophorinae2841114
Arrhopalitidae1119119
Bilobellini7446
Bourletiellidae332337
Brachystomellidae161288
Caputanurininae273
Coenaletidae122
Collophoridae188
Cyphoderidae1213110
Dicyrtominae59318
Entomobryinae1844824
Frieseinae416842
Gulgastruridae111
Hypogastruridae4366915
Incerta111
Incerta17241
Isotogastruridae155
Isotominae504719
Katiannidae1718610
Lepidocyrtinae2171233
Lobellini15946
Mackenziellidae111
Microfalculidae111
Morulininae22010
Morulodini144
Neanurini1619512
Neelidae5357
Odontellidae1212410
Oncobryidae111
Oncopoduridae25125
Onychiurinae4050212
Orchesellinae1224020
Pachyotominae5183
Pachytullbergiidae331
Paleonurini1820011
Paleotullbergiidae111
Paranurini13333
Paronellidae2837413
Phylliomerini11746
Poduridae133
Proisotominae1434624
Protentomobryidae111
Pseudachorutinae484238
Ptenothricinae39531
Seirinae217788
Sensillanurini33511
Sminthurididae1014714
Sminthurinae1919510
Sphyrothecinae44010
Spinothecidae362
Sturmiidae122
Tetrodontophorinae441
Tomoceridae161368
Tullbergiidae332086
Uchidanurinae8151
Collembola626767512

Note that the species count is optimistic and includes problematic taxa such as unresolved homonyms, unresolved nomina nuda and undescribed species (e.g., 'sp.'). Including all subspecific taxa, such as subspecies, varietas and forma, the total taxa count is about 8000.

A high species/genus ratio might be an indication that it is about time to do a revision of the concerned group. Taken this into account, the following groups are candidate for revision:

History

The history of this check list has been organised on a yearly basis. This file contains the history of the current year. Previous years can be found here :
2003.12.31-2004.01.31
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on a new Tomocerina from Xinjiang, China by Ma, Chen & Christiansen (2003),
    • on a new Lepidocyrtus from China by Wang, Chen & Christiansen (2003), publications received from Dr Kenneth A. Christiansen;
    • on the history of insect paleontology by Grimaldi (2001);
    • on a soil washing system to collect Collembola from soil samples by Warlet (2000), publication received from Dr M. Dethier;
    • on the mushroom bodies of arthropodan brain systems by Strausfeld, Hansen, Li, Gomez & Ito (1998).
  • A key to the species of Tomocerina, based on Ma, Chen & Christiansen (2003), has been added.
  • A key to the subgenera of Lepidocyrtus, based on Wang, Chen & Christiansen (2003), has been added.
    2004.02.01-2004.02.29
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on Rhyniella praecursor by Engel & Grimaldi (2004), publication received from Dr Luc De Bruyn;
    • on Isotomodes bisetosus, new from Poland by Pomorski (2003),
    • on a new Protaphorura from Austria by Pomorski, Leithner & Bruckner (2003),
    • on polymorphism in Ceratophysella armata from Poland by Skarzynski (2003),
    • on Hypogastrura kelmendica, new from Poland by Skarzynski & Smolis (2003),
    • on a new Balkanura from Turkey by Smolis (2003),
    • on a new Paleonura from North Vietnam by Smolis & Deharveng (2003),
    • on a new Endonura from Israel by Smolis & Kaprus' (2003),
    • on a new Wankeliella from Poland by Smolis & Skarzynski (2003), publications received from Dr Romuald Jacek Pomorski;
    • on new Isotomiella from Brasil by de Mendonça & Fernandes (2003), publication received from Dr Maria Cleide de Mendonça;
    • on the culture of Folsomia candida by Bruse, Meyer & Schmidt (2003),
    • on culturing Collembola as food for fish and frogs kept in aquaria or terraria by Staiesse in Claude (2000), publications received from Peter De Batist;
    • on a new Protaphorura from Morocco by Arbea (2003),
    • on the effects of forestry practices on the abundance of Collembola by Klimaszewski, Bernier-Cardou & Germain (2003),
    • on Agrenia bidenticulata record from Belgium by Leleup (1948), publications received from Jos Bruers;
    • on cave Collembola records from Belgium by Dethier & Hubart (2003),
    • on cave Collembola records from Belgium by Hubart, Xhaard, Dumoulin, Monsier, Roosens & Dethier (2003),
    • on the definition of 'Superficial Underground Compartment' (Milieu Souterrain Superficiel) by Hubart (2001), publications received from Dr Michel Dethier;
    • on the germ line differentiation during embryogenesis of Allacma fusca by Klag & Swiatek (1999);
    • on Hypograstrura nivicola from Canada by Bilodeau (1998);
    • on a new Willemia from Canada and Alaska by Fjellberg (1991), publication received from Dr Gloria Ortega Munoz.
    2004.03.01-2004.04.02
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on secondary infestations by Collembola of cutaneous lesions by Amin (2004), (2003), publications brought to my attention by Sidney Presley;
    • on a revision of the Isotoma viridis complex by Fjellberg (2003), publication received from Dr Arne Fjellberg;
    • on cave fauna (including Collembola) sampling and conservation techniques by Dethier & Hubart (2000),
    • on Onychiurus records from Belgium by Gilson & Hubart (1980),
    • on Gisinea delhezi from Belgium by Hubart (1976), publications received from Dr Michel Dethier;
    • on metallothionein in Orchesella cincta by Hensbergen (1999),
    • on some alpine records of Collembola from Italy by Janssens (1997),
    • on a translation in Dutch of the original description of Podura fusca De Geer, 1743 by Janssens (1996),
    • on Hypogastruridae records from Spain by Jordana (1981),
    • on new Xenyllodes, Uzelia and Folsomia from Spain by Jordana & Ardanaz (1981),
    • on a new form of Friesea subterranea from Spain by Jordana & Asiain (1981),
    • on Hypogastrura records from Spain by Jordana (1980).
    2004.04.03-2004.05.01
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on a review of the Checklist of the Collembola of the World www.collembola.org by Leslie (2004), publication brought to my attention by Scott M. Kruse;
    • on a morphological phylogeny of the Collembola by D'Haese (2003);
    • on three species of the Isotoma viridis complex by Simonsen, Filser, Krogh & Fjellberg (1999);
    • on a new cave Pseudosinella from France by Stomp (1971),
    • on a new cave Tomocerus from Slovenia by Stomp (1969), publications received from Dr Michel Dethier;
    • on the bibliography of keys to the Collembola of Europe (1880-1963) by Göllner-Scheiding (1967),
    • on the cave Collembola catalogue by Wolf (1938), publications received from Jos Bruers;
    • on cave Collembola and a new Oncopodura from Hongary by Loksa (1961), publication received from Dr Louis Deharveng.
    2004.05.02-2004.05.31
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on Collembola from Antarctica by Carpenter (1921).
    2004.06.01-2004.07.01
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on new recent Collembola taxa based on (Feuilles d') Informations Apterygotologiques, Nos 936 - 937, (2004), e-mail received from Dr Jean-Marc Thibaud;
    • on a new Tomocerina from China by Ma, Chen & Christiansen (2004),
    • on a redescription of Tomocera baicalensis from China by Ma, Chen & Christiansen (2004),
    • on a new Homidia from China by Jia, Chen & Christiansen (2004),
    • on a survey of Pseudosinella from East Asia by Wang, Chen & Christiansen (2004), publications received from Dr Kenneth A. Christiansen;
    • on effects of fire and agricultural practices on Collembola from Mexico by Vázquez & Tun Osorio (2004),
    • on records of Collembola from a semi-dry region of Mexico by Delgadillo-Reyes & Padilla-Ramírez (2004),
    • on fossil records of Collembola in amber from Chiapas Mexico by Hernández-Meléndez, Palacios-Vargas & Cutz-Pool (2004),
    • on new Sphyrotheca from Mexico by Palacios-Vargas, Vázquez & Cuéllar (2003), publications received from Dr José G. Palacios-Vargas;
    • on the genetic variability of Gressittacantha terranova from Antarctica by Fanciulli, Summa, Dallai & Frati (2001), publication received from Dr Francesco Frati;
    • on the phylogeny of Neanurinae by Cassagnau (1983),
    • on the function of the antennal tip sensory complex of Allacma fusca by Altner & Thies (1978),
    • on the female genital orifice of Proisotoma minuta by Betsch-Pinot (1974),
    • on a revision of Maynard's species of Bourletiella by Gough (1974),
    • on the epitoky in Hypogastruridae by Bourgeois & Cassagnau in Vandel (1973),
    • on the spermatozoon of Collembola by Dallai (1969),
    • on a new Onychiurus from northern Africa by Denis (1935), publications received from Dr Nicole Balaguer ().
  • The key to the species of Pseudosinella has been actualised, based on Wang, Chen & Christiansen (2004).
  • A key to the species of Sphyrotheca from America, based on Palacios-Vargas, Vázquez & Cuéllar (2003), has been added.
    2004.07.02-2004.08.02
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on cave Collembola from Andalusia, Spain by Arbea & Baena (2004),
    • on the catalogue of Collembola from Aragon, Spain by Arbea (2004), publications received from Dr Javier I. Arbea;
    • on Collembola found in skin scrapings by Altschuler et al. (2004), publication received from Deborah Z. Altschuler;
    • on new Seira from Yemen by Barra (2004), publication received from Dr Jean-Auguste Barra;
    • on the homonymy of Adisius by Bretfeld (2004), e-mail received from Dr Gerhard Bretfeld;
    • on the survival strategy of Collembola in contaminated soil by van Straalen (2004),
    • on the genetic structure of populations of Orchesella cincta from Europe by Timmermans (2004), publications received from Victor Naveau;
    • on the pairwise culturing of seven species of Collembola by Smith (1997), unpublished thesis received from Dr Kenneth A. Christiansen;
    • on the antennal trap, a specialisation for feeding on Collembola, of the ground beetle Loricera pilicornis by Hintzpeter & Bauer (1986), publication received from François Van Kerkhoven;
    • on new Collembola from Finland by Reuter (1876), publication received from Dr Nicole Balaguer (), scanned by Veerle Jacobs.
    2004.08.03-2004.08.31
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on Collembola implicated in delusory parasitosis by Barclay (2004), publication received from Sidney Presley;
    • on records from contaminated soils from England by Fountain & Hopkin (2004),
    • on records from contaminated soils from England by Fountain & Hopkin (2004), publications received from Dr Steven P. Hopkin;
    • on neotropical records from Quintana Roo, Mexico by Vázquez & Palacios-Vargas (2004), publication received from Dr José G. Palacios-Vargas.
    2004.09.01-2004.10.01
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on new littoral Poduromorpha from Brazil by Fernandes & de Mendonça (2004), publication received from Dr Maria Cleide de Mendonça;
    • on the dorsal organ of the embryo of a dipluran compared to that of Collembola by Ikeda & Machida (2001).
    2004.10.02-2004.10.31
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on new Seira from Yemen by Barra (2004), publication received from Dr Jean-Auguste Barra;
    • on the correct authority of Entomobrya pazaristei from Europe by Jordana (2004),
    • on the key to species of Wankeliella by Jordana (2004),
    • on the biogeographic region of new Seira from Yemen by Barra (2004), emails received from Dr Rafael Jordana;
    • on Collembola in skin scrapings and delusory parasitosis by Benecke (2004), publication brought to my attention by Frauke Büttner;
    • on Collembola by Thibaud in Leraut (2003), publication received from Victor Naveau;
    • on a molecular phylogeny of the Collembola by D'Haese (2002);
    • on Collembola from New Caledonia by Yosii (1960), publication received from Dr José G. Palacios-Vargas.
  • A key to the species of Paraxenylla has been added, based on Thibaud & Weiner (1997).
  • A key to the species of Wankeliella has been added, based on Jordana (2004).
  • The taxonomic hierarchy of D'Haese (2002) has been adopted, in general.
    2004.11.01-2004.11.30
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on a new cave Onychiurinae from the Iberian Peninsula by Martínez, Baquero, Barranco, Ariño & Jordana (2004), publication received from Dr Rafael Jordana;
    • on a checklist from Poland by Skarzynski, Pomorski, Smolis, Weiner, Szeptycki, Slawska & Sterzynska (2002), publication received from Dr Romuald J. Pomorski;
    • on a combined morphological and molecular Hexapoda phylogeny [Collembola included] by Carpenter & Wheeler (1999);
    • on zoological systematics and nomenclature [Collembola included] by Kluge (1999);
    • on new Poduromorpha from New Caledonia by Najt & Weiner (1997), publication received from Dr Wanda M. Weiner;
    • on a systematic overview of apterygote Hexapoda by Palacios-Vargas (1991),
    • on new Collembola from Hungary by Stach (1930), publications received from Dr José G. Palacios-Vargas.
  • A key to the species of Spinotheca has been added, based on Greenslade (1982).
  • A key to the species of Tillieria has been added, based on Najt & Weiner (1997).
    2004.12.01-2004.12.30
  • Added the new taxa and/or corrections of the publications and/or communications:
    • on Folsomia candida by Fountain & Hopkin (2005),
    • on recent advances in Collembola systematics by Deharveng (2004),
    • on Collembola in general by Hopkin (2002),
    • on the effect of heavy metals on Folsomia candida by Fountain & Hopkin (2001),
    • on the toxicity of cupper to Folsomia fimetaria by Scott-Fordsmand, Weeks & Hopkin (2000),
    • on the toxicity of nickel to Folsomia fimetaria by Scott-Fordsmand, Krogh & Hopkin (1999),
    • on the effect of temperature on the toxicities of heavy metals on Folsomia candida by Sandifer & Hopkin (1997),
    • on effects of heavy metals on development of Eisenia fetida [Collembola references included] by Spurgeon & Hopkin (1996),
    • on heavy metals in Chilopoda due to predation on Collembola and woodlice by Hopkin & Martin (1983) publications received from Dr Steven P. Hopkin;
    • on a new Friesea from Algeria (2004), publication received from Dr Salah Hamra-Kroua;
    • on the possible relationship of Collembola with Amphipoda by Lawrence (2004), publication received from Dr Loren Benton Byrne;
    • on the effect of land-use on Collembola from Portugal by Sousa, da Gama, Pinto, Keating, Calhôa, Lemos, Castro, Luz, Leitao & Dias (2004), publication received from Dr José Paulo Sousa;
    • on the effects of model misspecification on phylogenetic inference [Collembola included] by Buckley & Cunningham (2002);
    • on the effect of glufosinate-ammonium on the demography of Paronychiurus kimi from Korea by Kang, Choi & Ryoo (2001);
    • on the ecotoxity of mercury to Folsomia candida by Lock & Janssen (2001);
    • on the phylogeny of extant hexapodans [Collembola included] by Wheeler, Whiting, Wheeler & Carpenter (2001);
    • on the molecular phylogeny of Neanuridae by Frati & Dell'Ampio (2000);
    • on the endogenous tidal rhythm of orientation in Anurida maritima by Manica, McMeechan & Foster (2000);
    • on the phylogeny of 'Apterygota' [Collembola included] by Bach de Roca, Gaju-Ricart & Compte-Sart (1999);
    • on a combined morphological and molecular phylogeny of Chilopoda [Collembola included as outgroup] by Edgecombe, Giribet & Wheeler (1999);
    • on systematics and phylogeny of Arthropoda [Collembola included] by Giribet, Edgecombe & Wheeler (1999);
    • on the automatic counting of collembolans by Krogh, Johansen & Holmstrup (1996);
    • on the ecotoxity of an anionic surfactant, linear alkylbenzene sulfonate to Folsomia fimetaria by Holmstrup & Krogh (1996);
    • on the toxicity test with the insecticide dimethoate on Folsomia candida and Folsomia fimetaria by Krogh (1995);
    • on the ultrastructure of the midgut of Tomocerus minor by Humbert (1979), publication received from Dr Nicole Balaguer ();
    • on Collembola from the island St-Helena by Lawrence (1970),
    • on Agrenia from Belgium by Collart in Cooreman (1958),
    • on Hypogastrura from Belgium by Allaer in Segers (1964),
    • on Podurae records by Fabricius (1793),
    • on Podurae records by de Geer (1778), publications received from Jos Bruers.
    The taxonomic hierarchy of Deharveng (2004:427) has been adopted, in general.