Checklist of the Collembola: History - 2009

http://www.collembola.org/doc/hist2009.htm - Last updated on 2009.12.31 by Frans Janssens

Checklist of the Collembola: History - 2009

Preface

This checklist has been prepared by Frans Janssens, member of the Royal Entomological Society of Antwerp, Belgium. The purpose of this check list is to resolve synonomy related problems. Typical example: given a taxon name, verify whether or not it is currently valid and if not what is then its currently valid name.

This list is updated regularly. Unfortunately, it still contains a lot of mistakes. :-(
Typical errors are a) duplication of the same species name in several genera, b) duplication of taxa at different rank (e.g., once listed at species rank and once more at subspecies rank), c) synonyms listed as valid taxa, c) incorrect spelling of the name of a taxon or author, d) incorrect publication date, e) incorrect revision authority.
Your comments, flames, remarks, suggestions, etc are welcome at Frans Janssens

Frans Janssens
2009.12.31

Status

Status of the checklist

Group	# of Genera	# of Species	Species/Genus ratio

Actaletidae	2	10	5
Anurophorinae	29	345	11
Arrhopalitidae	2	131	65
Bourletiellidae	36	245	6
Brachystomellidae	18	129	7
Capbryinae	2	3	1
Caputanurininae	2	8	4
Coenaletidae	1	2	2
Collophoridae	1	8	8
Cyphoderinae	12	128	10
Dicyrtominae	5	93	18
Entomobryinae	15	459	30
Frieseinae	6	190	31
Gulgastruridae	1	1	1
Hypogastruridae	39	677	17
Isotogastruridae	1	7	7
Isotominae	47	479	10
Katiannidae	111	205	1
Lepidocyrtinae	14	643	45
Lophognathellinae	2	2	1
Mackenziellidae	1	1	1
Microfalculidae	1	1	1
Morulininae	1	22	22
Neanurinae	94	735	7
Neelidae	5	33	6
Odontellidae	13	130	10
Oncobryidae	1	1	1
Oncopoduridae	2	52	26
Onychiurinae	45	555	12
Orchesellinae	13	250	19
Pachyotominae	5	28	5
Pachytullbergiidae	3	3	1
Paleotullbergiidae	1	1	1
Paronellinae	27	373	13
Poduridae	1	4	4
Praentomobryidae	2	2	1
Proisotominae	25	479	19
Protentomobryidae	1	1	1
Pseudachorutinae	50	443	8
Ptenothricinae	3	106	35
Seirinae	5	260	52
Sminthurididae	10	145	14
Sminthurinae	20	191	9
Sphyrothecinae	7	54	7
Spinothecidae	3	6	2
Sturmiidae	1	2	2
Tetrodontophorinae	3	3	1
Tomoceridae	16	147	9
Tullbergiidae	32	214	6
Uchidanurinae	8	16	2
Willowsiinae	8	57	7

Collembola	753	8080	10

Note that the species count is optimistic and includes problematic taxa such as unresolved homonyms, unresolved nomina nuda and undescribed species (e.g., 'sp.'). Including all subspecific taxa, such as subspecies, varietas and forma, the total taxa count is about 8200.

A high species/genus ratio might be an indication that it is about time to do a revision of the concerned group. Taken this into account, the following groups are candidate for revision:

Arrhopalitidae, currently being revised by Drs Douglas Zeppelini & Robert Vargovich;
Lepidocyrtinae; the revision of Lepidocyrtus s.l. of Yoshii & Yayuk (1989) has been only partly accepted up to subgenus level;
Seirinae.

History

The history of this check list has been organised on a yearly basis. This file contains the history of the current year. Previous years can be found here : 1995, 1996, 1997, 1998, 1999, 2000, 2001, 2002, 2003, 2004, 2005, 2006, 2007, 2008.

2009.01.01-2009.01.31

Added the new taxa and/or corrections of the publications and/or communications:

on corrections on the Neanurinae (2009) email received from Dr Louis Deharveng;
on the environmental correlates of species richness of Collembola from Europe by Ulrich & Fiera (2009), publication received from Cristina Fiera;
on a new Seira from Brazil by Bellini & Zeppelini (2008), publication received from Dr Bruno Bellini;
on the checklist of the Collembola from Georgia by Barjadze & Djanashvili (2008), publication received from Dr Shalva Barjadze;
on records of Collembola from the Slovak karst by Raschmanová, Kovác, & Miklisová (2008), publication received from Dr L'ubomír Kovác;
on Collembola from necrotraps from Mexico by Palacios-Vargas & Mejía-Recamier (2008), publication received from Dr José Palacios-Vargas;
on a new cave Verhoeffiella from Montenegro by Lucic, Tomic, Brajkovic & Curcic (2008);
on new Protaphorura from Russia by Kaprus & Pomorski (2008),
on a new Agraphorura from Southern Siberia by Kaprus & Stebaeva (2006),
on niche differentiation among epigeic Collembola from Ukraine by Kaprus & Sterzynska (2005),
on a new Pratanurida from Ukraine by Kaprus & Weiner (2002),
on a new Isotomiella from Poland by Sterzynska & Kaprus (2001),
on the reaction of Collembola communities to anthropogenic substitution of forests in Ukraine by Kaprus ([2000]),
on Collembola from Roztocze by Kaprus (1998),
on a new Willemia from Ukraine by Kaprus (1997),
on a new Allaphorura from Ukraine by Kaprus & Weiner ([1995]),
on a new Drepanura from Siberia by Kaprus & Szeptycki (1992), publications received from Dr Ighor J. Kaprus;
on the Collembola from the Obedska Bara Natural Reserve, Serbia by Lucic (2006);
on new Hymenaphorurini from Northern America by Pomorski & Steinmann (2004), publication received from David Steinmann;
on the ontogeny in Symphypleona by Nayrolles (1996);
on Collembola records from Hungary by Farkas (1995);
on a revision of Mackenziella from Europe by Fjellberg (1989), publication received from Dr Arne Fjellberg;
on the Collembola from Arkansas, USA by Tedder & Allen (1989);
on new Poduromorpha and Isotomidae from North Norway by Fjellberg (1988);
on new Lathriopyga from Europe by Deharveng (1985), publication received from Dr Louis Deharveng;
on Hypogastrura records from a cave in Turkey by Kosswig (1952), publication received from Dr Kadir Bogac Kunt;
on a review of Achorutes nivicola from the USA by Folsom (1902);
on a new Neelus from the USA by Folsom (1901);
on new Collembola from Kiev, Russia by Scherbakov (1898);
on the new family Neelidae from the USA by Folsom (1896);
on a new Smynthurus from the USA by Folsom (1896).

The key to the species of Protaphorura octopunctata group has been added, based on Kaprus & Pomorski (2008).

The key to the species of Agraphorura has been updated, based on Kaprus & Stebaeva (2006).

2009.02.01-2009.02.28

Added the new taxa and/or corrections of the publications and/or communications:

on new Onychiurus from Eastern Europe by Kaprus (2009), publication received from Dr Ighor J. Kaprus;
on Collembola records in compost from Mexico by Mejía-Recamier & Palacios-Vargas (2008), publication received from Dr Luc De Bruyn;
on new Pseudosinella from the America's by Simón-Benito & Palacios-Vargas (2008),
on a new Schaefferia from Mexico by Palacios-Vargas & Castaño-Meneses (2008), publications received from Dr José G. Palacios-Vargas;
on a new cave Coecobrya from Chile by Jordana & Baquero (2008) publication received from Dr Rafael Jordana;
on a rRNA phylogeny of the basal Hexapoda by Gao, Bu & Luan (2008) publication received from Dr Yan Gao;
on the global diversity of Collembola in fresh water by Deharveng, D'Haese & Bedos (2008) publication received from Dr Anne Bedos;
on new Collembola from Madagascar by Thibaud (2008) publication received from Dr Jean-Marc Thibaud;
on urbanistic Collembola records from Russia by Tarashchuk & Horban (2006);
on some xerothermic Collembola from Hungary by Traser (2002);
on the Collembola from the Aggtelek National Park of Hungary by Traser (1999);
on Collembola records from Hungary by Farkas, Szathmáry & Strenner (1998);
on the feeding biology of Folsomia and Onychiurus from Hungary by Csuták (1977);
on new Collembola from Jugoslavia by Loksa & Bogojevic (1970);
on new cave Arrhopalites from Hungary by Loksa (1969);
on new cave Collembola from Hungary by Loksa (1967);
on new Collembola from Hungary by Loksa & Rubio (1966);
on new Collembola from Hungary by Loksa (1964);
on some Collembola records from Hungary by Loksa (1960).

The key to the palearctic species group of Onychiurus obsiones has been added, based on Kaprus (2009).

The key to the species of Schaefferia from Mexico has been added, based on Palacios-Vargas & Castaño-Meneses (2008).

The key to the species of Isotogastrura has been updated, based on Thibaud (2008).

2009.03.01-2009.03.31

Added the new taxa and/or corrections of the publications and/or communications:

on an introduction to Collembola by De Ridder & De Kockere (2009), publication received from Michel Van Malderen;
on new cave Arrhopalites from Ukraine by Vargovitsh (2009), publication received from Dr Robert S. Vargovitsh;
on a new Homidia from Eastern China by Shi, Pan & Qi (2009),
on a new Pseudachorutes from China by Shi, Jiang & Pan (2008), publications received from Dr Zhi-Xiang Pan;
on Collembola populations in hayfields and pastures from Iceland by Gudleifsson & Bjarnadottir (2008);
on Collembola from a parking area in an urban nature reserve from the UK by Shaw & Reeve (2008),
on arboreal Collembola from the UK by Shaw, Ozanne, Speight & Palmer (2007),
on Collembola of pulverised fuel ash sites from the UK by Shaw (2003), publications received from Dr Peter Shaw;
on a new cave genus of Sminthuridae from Herzegovina by Curcic, Lucic, Tomic, Makarov & Karaman (2007);
on Folsomia candida living in nest cells of the cicada Graptopsaltria nigrofuscata from Japan by Maezono & Miyashita (2000);
on the types of Collembola at the Illinois Natural History Survey by Mari Mutt (1978).

The key to the genera of Arrhopallitae has been added, based on Vargovitsh (2009).

2009.04.01-2009.04.30

Added the new taxa and/or corrections of the publications and/or communications:

on Heterosminthurus novemlineatus from Holland by Berg (2009), publication received from Ab H. Baas;
on Desoria glacialis from Swiss glaciers by Duffin (2009), publication received from Dr Chris Duffin;
on a new Seira from Brazil by Bellini, Pais & Zeppelini (2009), publication received from Dr Bruno Cavalcante Bellini;
on the synonymy of Sminthurinus similitortus from the USA by Snider in Justis (2009) email received from Scott Justis;
on a new Friesea from Algeria by Hamra-Kroua, Jordana & Deharveng (2009), publication received from Dr Salah Hamra-Kroua;
on a new Hypogastrura from Ukraine by Skarzynski & Kaprus (2009), publication received from Dr Dariusz Skarzynski;
on corrections on the list of and key to the species of Arrhopalitidae (2009), (2009) emails received from Dr Robert Vargovitsh;
on the relationship between head capsule length and body length in Collembola from Svalbard by Uvarov (2004);
on a new Isotomurus from Ukraine by Potapov & Starostenko (2002).

The key to the genera of Arrhopallitae has been updated, based on Vargovitsh (2009), and (2009).

2009.05.01-2009.05.31

Added the new taxa and/or corrections of the publications and/or communications:

on new Entomobryidae from the United Arab Emirates by Barra & van Harten (2009), publication received by Dr Jean-Auguste Barra;
on Collembola from Southern Australia by Greenslade & Ireson (1986).

2009.06.01-2009.06.30

Added the new taxa and/or corrections of the publications and/or communications:

on a new Entomobrya from Algeria by Baquero, Hamra-Kroua & Jordana (2009), publication received by Salah Hamra-Kroua;
on a new Spinonychiurus and Tetracanthella from Ukraine by Kaprus' & Tsalan (2009), publication received by Dr Ighor J. Kaprus'.

2009.07.01-2009.07.31

Added the new taxa and/or corrections of the publications and/or communications:

on new Pseudachorutes from Ukraine by Kaprus' & Weiner (2009),
on new Superodontella from Ukraine by Kaprus' (2009),
on the redescription of Endonura cretensis from Israel by Smolis & Kaprus' (2009), publications received by Dr Ighor J. Kaprus';
on new records of Collembola from Mexico by Palacios-Vargas, Mejía-Recamier & Cutz-Pool (2009), publication received by Dr José G. Palacios-Vargas;
on a new name for Acanthothorax Bretfeld & Griegel nec Gaede 1832 by Bretfeld & Griegel (2006), publication received by Dr Dariusz Skarzynski;
on a new Folsomia from Siberia by Berezina & Potapov (2006);
on new Folsomia from Russia by Potapov (2006);
on Bromacanthus from Indonesia by Mitra (2003),
on Dicranocentruga from Ethiopia and the neotropical region by Mitra (2002),
on the status of Microparonella by Mitra (2002),
on the status of Paronellides, Pseudoparonellides and Micronellides by Mitra (1993),
on the status of Paronella from Africa by Mitra (1992), excerpts of publications received by Dr Saral K. Mitra;
on an accessory ocular structure in Paronellinae from Ethiopia and Neotropics by Mitra (1972), publication received by Adolf Suttrop.

The key to the species of Brachystomellides has been added.

The key to the species of Superodontella from Ukraine has been added, based on Kaprus' (2009).

The key to the species of Pseudachorutes from Ukraine has been added, based on Kaprus' & Weiner (2009).

2009.08.01-2009.08.31

Added the new taxa and/or corrections of the publications and/or communications:

on the redescription of Isotominella geophila from Algeria by Jordana, Hamra-Kroua & Baquero (2009), publication received by Salah Hamra-Kroua;
on new cave Pygmarrhopalites from the USA by Zeppelini, Taylor & Slay (2009), publication received by Dr Douglas Zeppelini;
on the emendation of Entomobrya Lönnbergi by Koerber (2009), publication received by Dr Stefan Koerber;
on a new Denisiella from Colombia by Ospina & Palacios-Vargas (2009),
on new records of Collembola from Paraguay by Palacios-Vargas & Mejía-Remarcier (2009), publications received by Dr José G. Palacios-Vargas;
on new Collembola from Indonesia by Yoshii & Suhardjono (1992), publication received by Dr Felipe Soto-Adames;
on the status of Trichorypha from Africa by Mitra (2002),
on the status of Pseudoparonella by Mitra (1992),
on Idiomerus pallidus from India by Mitra (1990), excerpts of publications received by Dr Saral K. Mitra;
on new records from Mesaphorura from Australia by Greenslade (1992);
on the chaetotaxy of Xenylla occidentalis, X. arenosa and X. littoralis by da Gama (1981);
on new Xenylla from Australia by da Gama (1980);
on a new Collembola from Australia by Womersley (1942);
on a new Isotobrya from South Australia by Womersley (1940);
on new Collembola from Australia by Womersley (1937);
on new Collembola from Australia by Womersley (1935);
on the moisture conditions of the soil in relation to the occurrence of Smynthurus(sic) viridis in Australia by Davidson (1934);
on the Bdellidae as pest control agent of Sminthurus viridis from Australia by Womersley (1933).

The key to the species of Denisiella has been added, based on Ospina & Palacios-Vargas (2009).

2009.09.01-2009.09.30

Added the new taxa and/or corrections of the publications and/or communications:

on records of in the UK introduced Symphypleona by Ardron (2009), publication received by Dr Paul A. Ardron;
on the taxonomic position of Hypogastrura montiocola from European mountains by Skarzynski (2009), publication received by Dr Dariusz Skarzynski;
on the habitat preference and dispersal ability of soil Collembola by Auclerc, Ponge, Barot & Dubs (2009), publication received by Dr Jean-François Ponge;
on new recent Collembola taxa based on (Feuilles d') Informations Apterygotologiques, Nos 1012 - 1023, (2009), e-mail received from Dr Cyrille D'Haese;
on new records of Collembola from Brazil by Zeppelini, Bellini, Creao-Duarte & Hernández (2009), publication received by Dr Douglas Zeppelini;
on new cave Proisotoma from Venezuela and Mexico by Palacios-Vargas & Arbea (2009),
on new Americanura from Mexico and Panama by Palacios-Vargas, Simón Benito & Paniagua Nucamendi (2009), publications received by Dr José G. Palacios-Vargas;
on the structure of the antifreeze protein of Hypogastrura harveyi by Lin, Graham, Campbell & Davies (2007), publication received by Dr Peter Shaw;
on a new Ceratrimeria from Tasmania by Womersley (1940);
on new Collembola from Australia by Womersley (1936);
on Collembola from Germany by Heymons & Heymons (1909).

The key to the species of Americanura has been added, based on Palacios-Vargas, Simón Benito & Paniagua Nucamendi (2009).

The key to the species-groups of Hypogastrura has been added, based on Skarzynski (2009).

The key to the genera of Katiannidae has been added, based on Bretfeld (1999) and Betsch (1980).

2009.10.01-2009.10.31

Added the new taxa and/or corrections of the publications and/or communications:

on a new Pseudosinella from Palma de Mallorca by Palacios-Vargas & Simón-Benito (2009), publication received by Dr José G. Palacios-Vargas;
on a new Subisotoma from China by Gao, Xiong & Potapov (2009),
on a new Folsomia from Russia by Potapov & Chimitova (2009),
on a new Folsomia from Russia by Potapov (2009),
on a new Orchesella from the Caucasus by Potapov & Kremenitsa (2008), publications received by Dr Mikhail B. Potapov;
on the number of Collembola species from the World by Chapman (2009), publication received by Dr Arthur D. Chapman;
on the predation of Collembola by Pselaphinae by Schomann, Afflerbach & Betz (2008), publication received by Anthony Daley;
on Podura nivalis by Vallot (1836);
on Podura from Germany by Herbst (1787);
on Podura by de Bomare (1780) (1780) (1780);
on Podura from France by Geoffroy (1764), (1762);
on Podura from Sweden by Linnaeus (1761);
on Podura by Linnaeus (1740).

2009.11.01-2009.11.30

Added the new taxa and/or corrections of the publications and/or communications:

on a new Psammisotoma from Brazil by Abrantes & de Mendonça (2009),
on Poduromorpha communities from Brazil by Fernandes, Nessimian & de Mendonça (2009), publications received by Dr Eduardo Assis Abrantes;
on sexual dimorphism in Collembola by Palacios-Vargas & Castaño-Meneses (2009),
on Collembola from Mexico by García-Gó:mez, Castaño-Meneses & Palacios-Vargas (2009),
on Collembola from Mexico by Rojas, Castaño-Meneses, Palacios-Vargas & García-Calderón (2009), publications received by Dr José G. Palacios-Vargas;
on the aberrant spermatogenesis in Symphypleona by Dallai, Fanciulli & Frati (2004), publication received by Dr Luc De Bruyn;
on Collembola from Indonesia by Yoshii & Suhardjono (1992),
on Collembola from Malaysia by Yoshii (1982),
on Collembola from South-India by Handschin (1929), publications received by Dr Zhi-Xiang Pan;
on the status of Paronella from Africa by Mitra (1992), publication received by Dr Saral K. Mitra;
on puce d'eau noire by Ledermüller (1768).

A field key to the families of Collembola from France has been added.

The key to the species of Denisiella has been corrected, based on an erratum of Ospina (2009).

The key to the species of Psammisotoma has been added, derived from Abrantes & de Mendonça (2009).

2009.12.01-2009.12.31

Added the new taxa and/or corrections of the publications and/or communications:

on the key to Holarctic Oligaphorurini by Kaprus and Weiner (2009), email received by Dr Ighor J. Kaprus;
on new Friesea from New Caledonia and Vanuatu by Weiner, Bedos & Deharveng (2009), publication received by Dr Wanda M. Weiner;
on a revision of Subisotoma and a new isotomid genus by Potapov, Babenko, Fjellberg & Greenslade (2009), publication received by Dr Mikhail Potapov;
on a new Sinella from China by Zhang & Deharveng (2009),
on new Coecobrya from China by Zhang, Deharveng & Chen (2009),
on new Acanthocyrtus from Australia by Zhang, Deharveng, Greenslade & Chen (2009),
on a new Seira from India by Zhang, Chatterjee & Chen (2009), publications received by Dr Feng Zhang;
on a phylogeny of Palmanura by Palacios-Vargas, Garcí-Barros & Simón-Benito (2009), publication received by Dr José G. Palacios-Vargas;
on a revision of Agrenia by Fjellberg & Bernard (2009),
on new psammophilic Collembola from Norway by Fjellberg (2009), publications received by Dr Arne Fjellberg;
on Campylothorax by Mitra & Dallai (1980), excerpt received by Dr Saral K. Mitra.

The key to the species of Americanura has been corrected, based on a new key and an erratum of Palacios-Vargas (2009), (2009).

The key to the species of Agrenia has been updated, based on (2009).

The key to the species of Xenyllodes has been added, mainly based on (2009).

The key to the European species of Halisotoma has been added, based on (2009).

The key to the species of Coecobrya has been updated, based on (2009).

The key to the species of Acanthocyrtus has been added, based on (2009).

The key to the species of Subisotoma has been added, based on (2009).

The key to the species of Isotopenola has been added, based on (2009).

The key to the genera of Oligaphorurini has been added, based on (2009).

The key to the species of Archaphorura has been added, based on (2009).

The key to the species of Dimorphaphorura has been added, based on (2009).

The key to the species of Oligaphorura has been added, based on (2009).