http://www.collembola.org/key/pseudosi.htm - Last updated on 2024.10.31 by Frans Janssens

Checklist of the Collembola: Key to the species of Pseudosinella

This key is still under construction. Note that missing figures will be provided as soon as possible. Currently, the key is in the feasibility study phase to find out how to integrate in the checklist in a modular way a key that has been generated with DELTA.

Kenneth A. Christiansen, Department of Biology, Grinnell College, PO Box V3, Grinnell, IA 50112-0806, USA
Peter F. Bellinger (), Department of Biology, California State University, Northridge, CA 91330, USA
Rafael Jordana, Department of Zoology and Ecology, Faculty of Science, University of Navarra, 31080 Pamplona, Spain
Frans Janssens, Department of Biology, University of Antwerp, Antwerp, B-2020, Belgium

Preamble

Rationale

Key

1(0). Habitat cave.............................................. 2 Habitat surface......................................... 122 Habitat both cave and surface........................... 202
2(1). Ungual wing tooth (fig.1) absent.......................... 3 Ungual wing tooth (fig.1) present........................ 74
Fig.1.
3(2). Dorsal cephalic macrochaeta t (fig.2) absent.............. 4 Dorsal cephalic macrochaeta t (fig.2) present............ 41
Fig.2.
4(3). Unquiculus shape (fig.1) acuminate........................ 5 Unquiculus shape (fig.1) clavate......................... 16 Unquiculus shape (fig.1) basally swollen................. 18
5(4). Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0............................................... 6 Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1.............................................. 13
Fig.3.
6(5). P (fig.4) absent.......................................... 7 P (fig.4) present......................................... 9
Fig.4.
7(6). Distance distal unpaired ungual tooth from base/total unguis up to 15............................................. nata Distance distal unpaired ungual tooth from base/total unguis 41 to 50........................................ martelii 17 Distance distal unpaired ungual tooth from base/total unguis 58 to 65............................................. lava Distance distal unpaired ungual tooth from base/total unguis 66 to 74................................................ 8
8(7). B (fig.4) smooth macrochaeta; e (fig.5) smooth macrochaeta; r (fig.5) ciliated macrochaeta; m1 (fig.5) smooth macrochaeta; m2 (fig.5) smooth macrochaeta.......... testa B (fig.4) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; r (fig.5) smooth microchaeta; m1 (fig.) absent; m2 (fig.5) ciliated macrochaeta......... palaciosi
Fig.5.
9(6). Maximum length 1.4 to 1.59........................ martelii 17 Maximum length 2.2 to 2.49.............................. ops Maximum length 2.5 to 2.99............................... 10 Maximum length 3 to 3.49................................. 11 Maximum length 4 to 4.99................................. 12
10(9). Apical organ of third antennal segment (fig.6) peg or rod-like.......................................... georgia Apical organ of third antennal segment (fig.6) expanded..... .................................................... bona Apical organ of third antennal segment (fig.) paddle-shaped....................................... fonsa
Fig.6.
11(9). Q1 (fig.4) smooth microchaeta; median(M) 4th abd dorsal macrochaetae (fig.3) 2; tenent hair shape (fig.) acuminate; m2 (fig.5) smooth macrochaeta; apical organ of third antennal segment (fig.6) expanded............. pecki Q1 (fig.4) absent; median(M) 4th abd dorsal macrochaetae (fig.3) 3; tenent hair shape (fig.1) clavate; m2 (fig.) smooth macrochaeta with supplementary seta; apical organ of third antennal segment (fig.6) peg or rod-like... dubia
12(9). B (fig.4) ciliated microchaeta; m2 (fig.5) smooth macrochaeta with supplementary seta; antennal/cephalic diagonal 6 or more; cephalic seta r1 (fig.2) present; cephalic seta r2 (fig.2) present............ christianseni B (fig.4) smooth macrochaeta; m2 (fig.5) smooth macrochaeta; antennal/cephalic diagonal 4 to 5.99; cephalic seta r1 (fig.2) absent; cephalic seta r2 (fig.2) absent... spinosa
13(5). Maximum length 1 to 1.19............................. espana Maximum length 1.4 to 1.59........................ martelii 17 Maximum length 1.6 to 1.79................ duodecimpunctata 21 Maximum length 1.8 to 2.19........................... oromii Maximum length 2.5 to 2.99............................ leoni Maximum length 3 to 3.49................................. 14
14(13). A (fig.4) smooth microchaeta; tenent hair shape (fig.) clavate................................................ 15 A (fig.4) smooth macrochaeta; tenent hair shape (fig.) acuminate......................................... voylesi A (fig.4) ciliated macrochaeta; tenent hair shape (fig.) truncate........................................... crypta
15(14). B (fig.4) smooth macrochaeta; r (fig.5) absent; m1 (fig.) smooth macrochaeta; region located Mexico, Central America, West Indies; apical organ of third antennal segment (fig.6) peg or rod-like.................. reddelli B (fig.4) ciliated macrochaeta; r (fig.5) smooth microchaeta; m1 (fig.5) ciliated macrochaeta; region located Europe and North Africa; apical organ of third antennal segment (fig.6) paddle-shaped............ riojana
16(4). Maximum length 1 to 1.19................................. 17 Maximum length 1.4 to 1.59...................... brevicornis Maximum length 1.8 to 2.19........................ mucronata
17(16). Tenent hair shape (fig.1) acuminate; number of teeth of inner unguis (fig.1) 2; unquiculus wing tooth (fig.) fully developed; number of eyes per side 0; region located North America...................................... espana Tenent hair shape (fig.1) clavate; number of teeth of inner unguis (fig.1) 3; unquiculus wing tooth (fig.1) absent; number of eyes per side 5; region located Europe and North Africa............................................ joupani
18(4). Unquiculus wing tooth (fig.1) absent..................... 19 Unquiculus wing tooth (fig.1) minute..................... 34 Unquiculus wing tooth (fig.1) weak.................... maros Unquiculus wing tooth (fig.1) fully developed........ espana
19(18). Region located Europe and North Africa................... 20 Region located North America............................. 32 Region located Mexico, Central America, West Indies.. bonita
20(19). Maximum length 1.2 to 1.39............................... 21 Maximum length 1.4 to 1.59............................... 22 Maximum length 1.6 to 1.79............................... 25 Maximum length 1.8 to 2.19............................... 26 Maximum length 2.2 to 2.49............................... 31 Maximum length 2.5 to 2.99......................... truncata
21(20). A (fig.4) smooth microchaeta; p (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0; median(M) 4th abd dorsal macrochaetae (fig.3) 3........... ............................................ octophthalma A (fig.4) ciliated macrochaeta; p (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 2; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1; median(M) 4th abd dorsal macrochaetae (fig.3) 2. impediens
22(20). Tenent hair shape (fig.1) acuminate.............. christiani Tenent hair shape (fig.1) clavate........................ 23 Tenent hair shape (fig.1) truncate................... stompi
23(22). A (fig.4) smooth microchaeta; p (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0; supplementary seta (fig.7) absent.................................. ksenemani 2 A (fig.4) ciliated macrochaeta; p (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; supplementary seta (fig.7) present........................................ 24
Fig.7.
24(23). L2 (fig.5) smooth macrochaeta; m1 (fig.5) smooth macrochaeta...................................... alfonsii L2 (fig.5) ciliated macrochaeta; m1 (fig.5) ciliated macrochaeta.................................. subefficiens
25(20). Antennal/cephalic diagonal 1.3 to 1.49....... dodecophthalma Antennal/cephalic diagonal 1.5 to 1.59.......... dodecopsis 19 Antennal/cephalic diagonal 2.6 to 2.69.......... carthusiana
26(20). Number of eyes per side 0................................ 27 Number of eyes per side 1....................... astronomica Number of eyes per side 6..................... kwartirnikovi
27(26). Antennal/cephalic diagonal 1.6 to 1.89................... 28 Antennal/cephalic diagonal 1.9 to 2.49................... 29
28(27). Number of teeth of inner unguis (fig.1) 2.......... aspinata Number of teeth of inner unguis (fig.1) 3......... plachteri
29(27). Tenent hair shape (fig.1) acuminate...................... 30 Tenent hair shape (fig.1) clavate................ propinqua 18
30(29). Posterior thoracic seg 2 macrochaeta number (fig.3) 3; median(M) 4th abd dorsal macrochaetae (fig.3) 2; m2 (fig.5) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; apical organ of third antennal segment (fig.6) peg or rod-like............................. aueri Posterior thoracic seg 2 macrochaeta number (fig.3) 4; median(M) 4th abd dorsal macrochaetae (fig.3) 3; m2 (fig.5) smooth macrochaeta; e (fig.5) smooth macrochaeta; apical organ of third antennal segment (fig.) paddle-shaped............................... salisburgiana
31(20). Supplementary seta (fig.7) absent; inner setae manubrial plaque (fig.8) 2............................... cassagnaui Supplementary seta (fig.7) present; inner setae manubrial plaque (fig.8) 3............................. superoculata
Fig.8.
32(19). Antennal/cephalic diagonal 3.5 to 3.99............... gisini Antennal/cephalic diagonal 6 or more..................... 33
33(32). Apical organ of third antennal segment (fig.6) expanded..... ........................................... christianseni Apical organ of third antennal segment (fig.) paddle-shaped..................................... boneti 23
34(18). M1 (fig.5) smooth macrochaeta............................ 35 M1 (fig.5) ciliated macrochaeta.......................... 39
35(34). Apical organ of third antennal segment (fig.6) peg or rod-like............................................... 36 Apical organ of third antennal segment (fig.) paddle-shaped.......................................... 37
36(35). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; antennal/cephalic diagonal 1.5 to 1.59............ vandeli 3 Posterior thoracic seg 2 macrochaeta number (fig.3) 1; antennal/cephalic diagonal 1.6 to 1.89......... infernalis
37(35). Number of eyes per side 0; inner setae manubrial plaque (fig.8) 2.............................................. 38 Number of eyes per side 1; inner setae manubrial plaque (fig.8) 3.......................................... alpina
38(37). Maximum length 2.2 to 2.49.......................... dallaii Maximum length 3 to 3.49............................. concii
39(34). Apical organ of third antennal segment (fig.6) peg or rod-like.......................................... vandeli 4 Apical organ of third antennal segment (fig.) paddle-shaped.......................................... 40
40(39). Antennal/cephalic diagonal 1.5 to 1.59.............. vandeli 5 Antennal/cephalic diagonal 1.6 to 1.89............ insubrica
41(3). Region located Europe and North Africa................... 42 Region located North America............................. 68 Region located Mexico, Central America, West Indies...... 72 Region located South America.................... brevicornis
42(41). Number of teeth of inner unguis (fig.1) 2................ 43 Number of teeth of inner unguis (fig.1) 3................ 45 Number of teeth of inner unguis (fig.1) 4................ 65 Number of teeth of inner unguis (fig.1) 0................ 67
43(42). Maximum length 1.4 to 1.59........................ martelii 17 Maximum length 1.8 to 2.19......................... lamperti 1 Maximum length 2.2 to 2.49...................... oxybarensis Maximum length 2.5 to 2.99............................... 44
44(43). Q1 (fig.4) smooth microchaeta; unquiculus wing tooth (fig.) minute; median(M) 4th abd dorsal macrochaetae (fig.3) 3; supplementary seta (fig.7) present............... sollaudi Q1 (fig.4) ciliated macrochaeta; unquiculus wing tooth (fig.1) absent; median(M) 4th abd dorsal macrochaetae (fig.3) 2; supplementary seta (fig.7) absent... cavernarum
45(42). Tenent hair shape (fig.1) acuminate...................... 46 Tenent hair shape (fig.1) clavate........................ 52 Tenent hair shape (fig.1) truncate....................... 63
46(45). Distance distal unpaired ungual tooth from base/total unguis up to 15...................................... carthusiana Distance distal unpaired ungual tooth from base/total unguis 16 to 20............................................... 47 Distance distal unpaired ungual tooth from base/total unguis 21 to 30...................................... unguiculata Distance distal unpaired ungual tooth from base/total unguis 41 to 50............................................... 48 Distance distal unpaired ungual tooth from base/total unguis 51 to 57............................................... 51 Distance distal unpaired ungual tooth from base/total unguis 58 to 65.......................................... inflata
47(46). Posterior thoracic seg 2 macrochaeta number (fig.3) 2....... ............................................. jeanpierrei Posterior thoracic seg 2 macrochaeta number (fig.3) 3....... ........................................... unguilonginea Posterior thoracic seg 2 macrochaeta number (fig.3) 4....... ............................................... passaueri
48(46). Unquiculus shape (fig.1) acuminate....................... 49 Unquiculus shape (fig.1) clavate................. intemerata Unquiculus shape (fig.1) basally swollen................. 50
49(48). Maximum length 1.4 to 1.59; antennal/cephalic diagonal 1.01 to 1.29......................................... martelii 17 Maximum length 2.2 to 2.49; antennal/cephalic diagonal 1.9 to 2.49...................................... arrasatensis
50(48). A (fig.4) smooth microchaeta; p (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0; supplementary seta (fig.7) absent........................... duodecimocellata A (fig.4) ciliated macrochaeta; p (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; supplementary seta (fig.7) present.................................... gineti
51(46). E (fig.5) smooth macrochaeta; m2 (fig.5) smooth macrochaeta; maximum length 2.5 to 2.99; L1 (fig.5) smooth macrochaeta; L2 (fig.5) smooth macrochaeta................ heteromurina E (fig.5) ciliated macrochaeta; m2 (fig.5) ciliated macrochaeta; maximum length 2.2 to 2.49; L1 (fig.) ciliated macrochaeta; L2 (fig.5) ciliated macrochaeta..... .............................................. racovitzai
52(45). Unquiculus shape (fig.1) acuminate....................... 53 Unquiculus shape (fig.1) clavate......................... 55 Unquiculus shape (fig.1) basally swollen................. 61
53(52). Distance distal unpaired ungual tooth from base/total unguis 21 to 30........................................... duprei Distance distal unpaired ungual tooth from base/total unguis 41 to 50............................................... 54 Distance distal unpaired ungual tooth from base/total unguis 58 to 65....................................... binoculata Distance distal unpaired ungual tooth from base/total unguis 66 to 74..................................... subilliciens
54(53). Maximum length 1.4 to 1.59; antennal/cephalic diagonal 1.01 to 1.29......................................... martelii 17 Maximum length 2.2 to 2.49; antennal/cephalic diagonal 1.9 to 2.49...................................... arrasatensis
55(52). Supplementary seta (fig.7) absent........................ 56 Supplementary seta (fig.7) present....................... 60
56(55). Distance distal unpaired ungual tooth from base/total unguis 58 to 65............................................... 57 Distance distal unpaired ungual tooth from base/total unguis 66 to 74............................................... 58
57(56). L2 (fig.5) smooth macrochaeta; e (fig.5) smooth macrochaeta; r (fig.5) smooth microchaeta; L1 (fig.5) smooth macrochaeta; m2 (fig.5) smooth macrochaeta........ azorica L2 (fig.5) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; r (fig.5) ciliated macrochaeta; L1 (fig.) ciliated macrochaeta; m2 (fig.5) ciliated macrochaeta..... ............................................ subduodecima
58(56). Antennal/cephalic diagonal 1.01 to 1.29............. joupani Antennal/cephalic diagonal 1.3 to 1.49................... 59 Antennal/cephalic diagonal 1.5 to 1.59......... subduodecima Antennal/cephalic diagonal 1.6 to 1.89......... subduodecima
59(58). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0; r (fig.) ciliated macrochaeta; a (fig.4) smooth microchaeta........ ............................................ subduodecima Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; r (fig.) smooth microchaeta; a (fig.4) ciliated macrochaeta........ ............................................ subilliciens
60(55). Maximum length 1 to 1.19............................ joupani Maximum length 1.6 to 1.79.......................... decepta Maximum length 1.8 to 2.19....................... lleidensis Maximum length 2.5 to 2.99................... superduodecima
61(52). Antennal/cephalic diagonal 1.3 to 1.49......... subilliciens Antennal/cephalic diagonal 1.5 to 1.59............. encrusae Antennal/cephalic diagonal 1.6 to 1.89................... 62 Antennal/cephalic diagonal 1.9 to 2.49........... propinqua 18
62(61). Q1 (fig.4) smooth microchaeta; supplementary seta (fig.) present; e (fig.5) smooth macrochaeta; r (fig.5) ciliated macrochaeta; number of eyes per side 6......... huescensis Q1 (fig.4) ciliated macrochaeta; supplementary seta (fig.) absent; e (fig.5) ciliated macrochaeta; r (fig.5) smooth microchaeta; number of eyes per side 5........... encrusae
63(45). Maximum length 1.4 to 1.59........................ martelii 17 Maximum length 2.2 to 2.49............................... 64 Maximum length 3 to 3.49....................... theodoridesi
64(63). L2 (fig.5) smooth macrochaeta; a (fig.4) ciliated macrochaeta; p (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2............................. arrasatensis L2 (fig.5) ciliated macrochaeta; a (fig.4) smooth microchaeta; p (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0................................ substygia
65(42). Maximum length 0.61 to 0.79........................... decui Maximum length 1.4 to 1.59............................... 66 Maximum length 1.6 to 1.79.................... bidenticulata 6 Maximum length 1.8 to 2.19.......................... bessoni
66(65). Unquiculus shape (fig.1) acuminate; distance distal unpaired ungual tooth from base/total unguis 41 to 50; antennal/cephalic diagonal 1.01 to 1.29......... martelii 17 Unquiculus shape (fig.1) clavate; distance distal unpaired ungual tooth from base/total unguis 75 to 80; antennal/cephalic diagonal 1.5 to 1.59.................... ........................................... bidenticulata 7
67(42). Unquiculus shape (fig.1) acuminate; maximum length 1.4 to 1.59; distance distal unpaired ungual tooth from base/total unguis 41 to 50...................... martelii 17 Unquiculus shape (fig.1) basally swollen; maximum length 1.6 to 1.79; distance distal unpaired ungual tooth from base/total unguis up to 15..................... cabidochei
68(41). Unquiculus shape (fig.1) acuminate....................... 69 Unquiculus shape (fig.1) basally swollen................. 70
69(68). P (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 3; r (fig.5) absent; maximum length 1.8 to 2.19; distance distal unpaired ungual tooth from base/total unguis 31 to 35........................... orba P (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 4; r (fig.5) ciliated microchaeta; maximum length 1.6 to 1.79; distance distal unpaired ungual tooth from base/total unguis 66 to 74...................... vita
70(68). Unquiculus wing tooth (fig.1) absent..................... 71 Unquiculus wing tooth (fig.1) fully developed.. parattenuata
71(70). Number of teeth of inner unguis (fig.1) 2........... boneti 23 Number of teeth of inner unguis (fig.1) 3...... meganporteri
72(41). A (fig.4) smooth microchaeta........................... yuca A (fig.4) ciliated microchaeta............. kenchristianseni A (fig.4) smooth macrochaeta............................. 73 A (fig.4) ciliated macrochaeta......................... cava
73(72). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; antennal/cephalic diagonal up to 1................. bonita Posterior thoracic seg 2 macrochaeta number (fig.3) 1; antennal/cephalic diagonal 1.9 to 2.49.............. volca Posterior thoracic seg 2 macrochaeta number (fig.3) 2; antennal/cephalic diagonal 3 to 3.49.............. huautla
74(2). Region located Europe and North Africa................... 75 Region located North America............................ 115 Region located Mexico, Central America, West Indies..... 118 Region located Asia.......................... chiangdaoensis
75(74). Tenent hair shape (fig.1) acuminate...................... 76 Tenent hair shape (fig.1) clavate....................... 107 Tenent hair shape (fig.1) truncate...................... 114
76(75). Maximum length 1.4 to 1.59............................... 77 Maximum length 1.6 to 1.79............................... 81 Maximum length 1.8 to 2.19............................... 84 Maximum length 2.2 to 2.49............................... 97 Maximum length 2.5 to 2.99.............................. 104 Maximum length 3.5 to 3.99...................... subterranea 8
77(76). Unquiculus shape (fig.1) acuminate....................... 78 Unquiculus shape (fig.1) clavate......................... 80 Unquiculus shape (fig.1) basally swollen........ feneriensis
78(77). Antennal/cephalic diagonal 1.01 to 1.29........... martelii 17 Antennal/cephalic diagonal 1.9 to 2.49................... 79
79(78). Distance distal unpaired ungual tooth from base/total unguis 36 to 40......................................... styriaca Distance distal unpaired ungual tooth from base/total unguis 41 to 50......................................... styriaca Distance distal unpaired ungual tooth from base/total unguis 66 to 74....................................... inaequalis
80(77). L1 (fig.5) smooth macrochaeta; a (fig.4) ciliated macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; supplementary seta (fig.7) present................. dobati L1 (fig.5) ciliated macrochaeta; a (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0; supplementary seta (fig.7) absent............... bulgarica
81(76). Posterior thoracic seg 2 macrochaeta number (fig.3) 0....... ........................................ vertamicoriensis Posterior thoracic seg 2 macrochaeta number (fig.3) 1....... .................................................. stygia Posterior thoracic seg 2 macrochaeta number (fig.3) 2....... .............................................. eskualduna Posterior thoracic seg 2 macrochaeta number (fig.3) 3.... 82
82(81). L1 (fig.5) smooth macrochaeta; e (fig.5) smooth macrochaeta; thoracic seg 3 macrochaetae number (fig.3) 1; m2 (fig.) smooth macrochaeta; number of teeth of inner unguis (fig.1) 2.................................... subcentralis L1 (fig.5) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; thoracic seg 3 macrochaetae number (fig.3) 2; m2 (fig.5) ciliated macrochaeta; number of teeth of inner unguis (fig.1) 3....................................... 83
83(82). Supplementary seta (fig.7) absent; dorsal cephalic macrochaeta t (fig.2) present; unquiculus shape (fig.) basally swollen; apical organ of third antennal segment (fig.6) paddle-shaped............................. aelleni Supplementary seta (fig.7) present; dorsal cephalic macrochaeta t (fig.2) absent; unquiculus shape (fig.) clavate; apical organ of third antennal segment (fig.) peg or rod-like.................................... curtii
84(76). Unquiculus wing tooth (fig.1) absent..................... 85 Unquiculus wing tooth (fig.1) minute..................... 94 Unquiculus wing tooth (fig.1) weak....................... 96
85(84). Unquiculus shape (fig.1) acuminate....................... 86 Unquiculus shape (fig.1) clavate......................... 87 Unquiculus shape (fig.1) basally swollen................. 89
86(85). Antennal/cephalic diagonal 1.9 to 2.49....... aggtelekiensis Antennal/cephalic diagonal 2.6 to 2.69............... boneti
87(85). L2 (fig.5) smooth macrochaeta; m2 (fig.5) smooth macrochaeta; dorsal cephalic macrochaeta t (fig.) present; e (fig.5) smooth macrochaeta; L1 (fig.5) smooth macrochaeta............................................ 88 L2 (fig.5) ciliated macrochaeta; m2 (fig.5) ciliated macrochaeta; dorsal cephalic macrochaeta t (fig.2) absent; e (fig.5) ciliated macrochaeta; L1 (fig.5) ciliated macrochaeta.................................. vornatscheri
88(87). Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; median(M) 4th abd dorsal macrochaetae (fig.3) 2; m1 (fig.5) ciliated macrochaeta..................................... subdobati Posterior thoracic seg 2 macrochaeta number (fig.3) 4; thoracic seg 3 macrochaetae number (fig.3) 3; median(M) 4th abd dorsal macrochaetae (fig.3) 3; m1 (fig.5) smooth macrochaeta...................................... balazuci
89(85). Number of eyes per side 0................................ 90 Number of eyes per side 1.............................. lesi Number of eyes per side 6............................ goughi Number of eyes per side 7............................ goughi
90(89). Dorsal cephalic macrochaeta t (fig.2) absent............. 91 Dorsal cephalic macrochaeta t (fig.2) present............ 93
91(90). Antennal/cephalic diagonal 1.6 to 1.89............ centralis Antennal/cephalic diagonal 1.9 to 2.49................... 92
92(91). Distance distal unpaired ungual tooth from base/total unguis 16 to 20................................... aggtelekiensis Distance distal unpaired ungual tooth from base/total unguis 31 to 35.................................... caladairensis
93(90). Distance distal unpaired ungual tooth from base/total unguis up to 15..................................... kersmaekersi Distance distal unpaired ungual tooth from base/total unguis 16 to 20................................... aggtelekiensis
94(84). L1 (fig.5) smooth macrochaeta; supplementary seta (fig.) present; e (fig.5) smooth macrochaeta; dorsal cephalic macrochaeta s (fig.2) present; m1 (fig.5) smooth macrochaeta............................................ 95 L1 (fig.5) ciliated macrochaeta; supplementary seta (fig.) absent; e (fig.5) ciliated macrochaeta; dorsal cephalic macrochaeta s (fig.2) absent; m1 (fig.5) ciliated macrochaeta...................................... lamperti 9
95(94). Q1 (fig.4) smooth microchaeta; q2 (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 4; median(M) 4th abd dorsal macrochaetae (fig.) 3; m2 (fig.5) smooth macrochaeta................... denisi Q1 (fig.4) ciliated microchaeta; q2 (fig.4) ciliated microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 3; median(M) 4th abd dorsal macrochaetae (fig.) 2; m2 (fig.5) ciliated macrochaeta................. jesusi
96(84). Number of teeth of inner unguis (fig.1) 2; unquiculus shape (fig.1) acuminate; antennal/cephalic diagonal 1.9 to 2.49. ............................................... strinatii Number of teeth of inner unguis (fig.1) 3; unquiculus shape (fig.1) basally swollen; antennal/cephalic diagonal 1.6 to 1.89............................................... paclti
97(76). Posterior thoracic seg 2 macrochaeta number (fig.3) 0....... ............................................. ashmoleorum Posterior thoracic seg 2 macrochaeta number (fig.3) 3.... 98 Posterior thoracic seg 2 macrochaeta number (fig.3) 4... 103
98(97). Dorsal cephalic macrochaeta s (fig.2) absent............. 99 Dorsal cephalic macrochaeta s (fig.2) present........... 101
99(98). L2 (fig.5) smooth macrochaeta; a (fig.4) ciliated macrochaeta; p (fig.4) present; e (fig.5) smooth macrochaeta; supplementary seta (fig.7) present....... 100 L2 (fig.5) ciliated macrochaeta; a (fig.4) smooth microchaeta; p (fig.4) absent; e (fig.5) ciliated macrochaeta; supplementary seta (fig.7) absent... chapmani
100(99). Distance distal unpaired ungual tooth from base/total unguis 31 to 35........................................... virei 10 Distance distal unpaired ungual tooth from base/total unguis 41 to 50......................................... subvirei
101(98). Distance distal unpaired ungual tooth from base/total unguis 21 to 30....................................... melatensis Distance distal unpaired ungual tooth from base/total unguis 31 to 35...................................... subterranea Distance distal unpaired ungual tooth from base/total unguis 41 to 50.............................................. 102
102(101). Antennal/cephalic diagonal 1.6 to 1.89........... subinflata Antennal/cephalic diagonal 1.9 to 2.49. tarraconensis 11 Antennal/cephalic diagonal 2.6 to 2.69. tarraconensis 12 Antennal/cephalic diagonal 2.7 to 2.79. tarraconensis 12 Antennal/cephalic diagonal 2.8 to 2.99. tarraconensis 12
103(97). Thoracic seg 3 macrochaetae number (fig.3) 2; median(M) 4th abd dorsal macrochaetae (fig.3) 2; number of teeth of inner unguis (fig.1) 3; dorsal cephalic macrochaeta t (fig.2) present; unquiculus shape (fig.1) clavate......... ........................................... barcelonensis Thoracic seg 3 macrochaetae number (fig.3) 3; median(M) 4th abd dorsal macrochaetae (fig.3) 3; number of teeth of inner unguis (fig.1) 2; dorsal cephalic macrochaeta t (fig.2) absent; unquiculus shape (fig.1) basally swollen.. ................................................. delhezi
104(76). Number of teeth of inner unguis (fig.1) 2............... 105 Number of teeth of inner unguis (fig.1) 3............... 106
105(104). Posterior thoracic seg 2 macrochaeta number (fig.3) 3; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1; median(M) 4th abd dorsal macrochaetae (fig.) 2; supplementary seta (fig.7) present; cephalic seta r3 (fig.2) present.................................. pyrenaea Posterior thoracic seg 2 macrochaeta number (fig.3) 4; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0; median(M) 4th abd dorsal macrochaetae (fig.) 3; supplementary seta (fig.7) absent; cephalic seta r3 (fig.2) absent................................... chauvini
106(104). Median(M) 4th abd dorsal macrochaetae (fig.3) 2; distance distal unpaired ungual tooth from base/total unguis 36 to 40.............................................. pieltaini Median(M) 4th abd dorsal macrochaetae (fig.3) 3; distance distal unpaired ungual tooth from base/total unguis 21 to 30.............................................. antennata
107(75). Unquiculus shape (fig.1) acuminate...................... 108 Unquiculus shape (fig.1) clavate........................ 110 Unquiculus shape (fig.1) basally swollen................ 111
108(107). Distance distal unpaired ungual tooth from base/total unguis 21 to 30........................................... duprei Distance distal unpaired ungual tooth from base/total unguis 41 to 50........................................ martelii 17 Distance distal unpaired ungual tooth from base/total unguis 58 to 65.............................................. 109 Distance distal unpaired ungual tooth from base/total unguis 66 to 74..................................... subilliciens Distance distal unpaired ungual tooth from base/total unguis 75 to 80.................................. duodecimoculata
109(108). A (fig.4) smooth microchaeta; r (fig.5) ciliated microchaeta; maximum length 2.5 to 2.99............ luquei A (fig.4) ciliated microchaeta; r (fig.5) ciliated macrochaeta; maximum length 2.2 to 2.49.......... beruetei A (fig.4) ciliated macrochaeta; r (fig.5) smooth microchaeta; maximum length 1.4 to 1.59........ binoculata
110(107). A (fig.4) smooth microchaeta; q1 (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; r (fig.) ciliated macrochaeta; thoracic seg 3 macrochaetae number (fig.3) 0...................................... lleidensis A (fig.4) ciliated macrochaeta; q1 (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 3; r (fig.5) smooth microchaeta; thoracic seg 3 macrochaetae number (fig.3) 2................ subilliciens
111(107). Number of teeth of inner unguis (fig.1) 2........... lunaris Number of teeth of inner unguis (fig.1) 3............... 112
112(111). Antennal/cephalic diagonal 1.3 to 1.49.................. 113 Antennal/cephalic diagonal 1.5 to 1.59............. manuelae Antennal/cephalic diagonal 1.9 to 2.49.......... handschini 20
113(112). Posterior thoracic seg 2 macrochaeta number (fig.3) 1; thoracic seg 3 macrochaetae number (fig.3) 0; supplementary seta (fig.7) present; m2 (fig.5) smooth macrochaeta; e (fig.5) smooth macrochaeta......... seyleri Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; supplementary seta (fig.7) absent; m2 (fig.5) ciliated macrochaeta; e (fig.5) ciliated macrochaeta.. subilliciens
114(75). Number of eyes per side 0; maximum length 1.4 to 1.59; distance distal unpaired ungual tooth from base/total unguis 41 to 50................................. martelii 17 Number of eyes per side 4; maximum length 1.6 to 1.79; distance distal unpaired ungual tooth from base/total unguis 58 to 65................................ aramendiai
115(74). Maximum length 1.2 to 1.39.............................. 116 Maximum length 1.4 to 1.59.............................. 117 Maximum length 1.6 to 1.79........................... granda Maximum length 1.8 to 2.19......................... espanita Maximum length 2.2 to 2.49.......................... vespera Maximum length 2.5 to 2.99............................ fonsa Maximum length 3 to 3.49............................. flatua
116(115). A (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 1; number of teeth of inner unguis (fig.1) 2; unquiculus wing tooth (fig.1) fully developed; m2 (fig.5) ciliated macrochaeta.......... certa A (fig.4) ciliated microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 2; number of teeth of inner unguis (fig.1) 3; unquiculus wing tooth (fig.1) absent; m2 (fig.5) smooth macrochaeta........................ erehwon
117(115). Posterior thoracic seg 2 macrochaeta number (fig.3) 1; unquiculus wing tooth (fig.1) weak; m2 (fig.5) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; L1 (fig.) ciliated macrochaeta............................ josemarii Posterior thoracic seg 2 macrochaeta number (fig.3) 2; unquiculus wing tooth (fig.1) absent; m2 (fig.5) smooth macrochaeta; e (fig.5) smooth macrochaeta; L1 (fig.) smooth macrochaeta.................................. extra
118(74). Posterior thoracic seg 2 macrochaeta number (fig.3) 0....... ................................................ rabonica Posterior thoracic seg 2 macrochaeta number (fig.3) 1... 119 Posterior thoracic seg 2 macrochaeta number (fig.3) 2. finca
119(118). Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0............................................. 120 Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1............................................. 121
120(119). R (fig.5) smooth microchaeta; tenent hair shape (fig.) acuminate; maximum length 1.8 to 2.19; distance distal unpaired ungual tooth from base/total unguis up to 15; inner setae manubrial plaque (fig.8) 3.............. volca R (fig.5) absent; tenent hair shape (fig.1) clavate; maximum length 1.6 to 1.79; distance distal unpaired ungual tooth from base/total unguis 75 to 80; inner setae manubrial plaque (fig.8) 2........................... petrustrinatii
121(119). E (fig.5) smooth macrochaeta; L1 (fig.5) smooth macrochaeta; a (fig.4) smooth macrochaeta; b (fig.4) smooth macrochaeta; m2 (fig.5) smooth macrochaeta........... vera E (fig.5) ciliated macrochaeta; L1 (fig.5) ciliated macrochaeta; a (fig.4) ciliated macrochaeta; b (fig.) ciliated macrochaeta; m2 (fig.5) ciliated macrochaeta..... .................................................... cava
122(1). Region located Europe and North Africa.................. 123 Region located North America............................ 182 Region located Mexico, Central America, West Indies..... 183 Region located South America............................ 184 Region located Sub-Saharan Africa....................... 187 Region located Asia..................................... 188 Region located Australia................................ 196 Region located Oceania.................................. 198 Region located New Zealand.............................. 201
123(122). Number of teeth of inner unguis (fig.1) 2............... 124 Number of teeth of inner unguis (fig.1) 3............... 125 Number of teeth of inner unguis (fig.1) 4............... 171
124(123). Q1 (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 1; tenent hair shape (fig.) clavate; ungual wing tooth (fig.1) absent; unquiculus shape (fig.1) acuminate....................... lafargensis Q1 (fig.4) ciliated macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 0; tenent hair shape (fig.) acuminate; ungual wing tooth (fig.1) present; unquiculus shape (fig.1) clavate........................... ghirkhani
125(123). Unquiculus wing tooth (fig.1) absent.................... 126 Unquiculus wing tooth (fig.1) minute.................... 162 Unquiculus wing tooth (fig.1) weak.............. tietarensis Unquiculus wing tooth (fig.1) fully developed........... 165
126(125). Dorsal cephalic macrochaeta s (fig.2) absent............ 127 Dorsal cephalic macrochaeta s (fig.2) present........... 153
127(126). Tenent hair shape (fig.1) acuminate..................... 128 Tenent hair shape (fig.1) clavate....................... 132
128(127). Number of eyes per side 0............................... 129 Number of eyes per side 1.......................... jordanai Number of eyes per side 2............................... 130 Number of eyes per side 3........................... serrana Number of eyes per side 4...................... luenganensis
129(128). Maximum length 1.2 to 1.39....................... jacetanica Maximum length 1.6 to 1.79......................... maritima
130(128). Maximum length 0.8 to 0.99......................... absoloni Maximum length 1 to 1.19............................ angelae Maximum length 1.2 to 1.39.............................. 131
131(130). A (fig.4) smooth microchaeta; p (fig.4) absent; posterior thoracic seg 2 macrochaeta number (fig.3) 0; supplementary seta (fig.7) absent; m1 (fig.5) ciliated macrochaeta...... ................................................ thibaudi A (fig.4) ciliated macrochaeta; p (fig.4) present; posterior thoracic seg 2 macrochaeta number (fig.3) 1; supplementary seta (fig.7) present; m1 (fig.5) smooth macrochaeta....... ................................................ leclerci
132(127). P (fig.4) absent........................................ 133 P (fig.4) present....................................... 139
133(132). Number of eyes per side 0....................... handschini 20 Number of eyes per side 2........................ turolensis Number of eyes per side 3............................... 134 Number of eyes per side 4............................... 137 Number of eyes per side 5............................... 138 Number of eyes per side 6...................... cansiliensis
134(133). Posterior thoracic seg 2 macrochaeta number (fig.3) 0... 135 Posterior thoracic seg 2 macrochaeta number (fig.3) 1... 136 Posterior thoracic seg 2 macrochaeta number (fig.3) 3....... ............................................... marcuzzii
135(134). M2 (fig.5) smooth macrochaeta; distance distal unpaired ungual tooth from base/total unguis 66 to 74; no. outer setae manubrial plaque (fig.8) 3..................... edax M2 (fig.5) ciliated macrochaeta; distance distal unpaired ungual tooth from base/total unguis 58 to 65; no. outer setae manubrial plaque (fig.8) 1................. hermanni
136(134). Supplementary seta (fig.7) absent; unquiculus shape (fig.) acuminate; apical organ of third antennal segment (fig.) expanded; antennal/cephalic diagonal 1.5 to 1.59; differentiated inner seta on hind tibiotarsus (fig.) unclear or absent.............................. zaragozana Supplementary seta (fig.7) present; unquiculus shape (fig.) clavate; apical organ of third antennal segment (fig.) peg or rod-like; antennal/cephalic diagonal 1.3 to 1.49; differentiated inner seta on hind tibiotarsus (fig.) clear, acuminate................................ templadoi
Fig.9.
137(133). Q1 (fig.4) smooth microchaeta; ungual wing tooth (fig.) present; unquiculus shape (fig.1) acuminate; r (fig.) ciliated microchaeta; dorsal cephalic macrochaeta t (fig.2) absent............................... luenganensis Q1 (fig.4) ciliated macrochaeta; ungual wing tooth (fig.) absent; unquiculus shape (fig.1) clavate; r (fig.5) smooth microchaeta; dorsal cephalic macrochaeta t (fig.) present........................................... xabieri
138(133). L1 (fig.5) smooth macrochaeta; q1 (fig.4) smooth microchaeta; e (fig.5) smooth macrochaeta; ungual wing tooth (fig.1) absent; m2 (fig.5) smooth macrochaeta....... ................................................ huetheri L1 (fig.5) ciliated macrochaeta; q1 (fig.4) ciliated macrochaeta; e (fig.5) ciliated macrochaeta; ungual wing tooth (fig.1) present; m2 (fig.5) ciliated macrochaeta.... ........................................... turiasonensis
139(132). Supplementary seta (fig.7) absent....................... 140 Supplementary seta (fig.7) present...................... 149
140(139). Ungual wing tooth (fig.1) absent........................ 141 Ungual wing tooth (fig.1) present....................... 147
141(140). A (fig.4) smooth microchaeta............................ 142 A (fig.4) ciliated macrochaeta.......................... 145
142(141). Number of eyes per side 4............................... 143 Number of eyes per side 5............................... 144
143(142). Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0; apical organ of third antennal segment (fig.) peg or rod-like; maximum length 0.8 to 0.99... canariensis Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1; apical organ of third antennal segment (fig.) expanded; maximum length 1.6 to 1.79................ picta
144(142). Unquiculus shape (fig.1) acuminate............... variabilis Unquiculus shape (fig.1) clavate................. fjellbergi
145(141). L1 (fig.5) smooth macrochaeta; m2 (fig.5) smooth macrochaeta........................................ fallax L1 (fig.5) ciliated macrochaeta; m2 (fig.5) ciliated macrochaeta........................................... 146
146(145). Posterior thoracic seg 2 macrochaeta number (fig.3) 1; thoracic seg 3 macrochaetae number (fig.3) 0; distance distal unpaired ungual tooth from base/total unguis 58 to 65............................................... apuanica Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; distance distal unpaired ungual tooth from base/total unguis 66 to 74........................................... subilliciens
147(140). Antennal/cephalic diagonal 1.01 to 1.29............. pallida Antennal/cephalic diagonal 1.3 to 1.49.................. 148 Antennal/cephalic diagonal 1.6 to 1.89............... horaki Antennal/cephalic diagonal 1.9 to 2.49.......... handschini 20
148(147). Posterior thoracic seg 2 macrochaeta number (fig.3) 0....... ............................................... hercynica Posterior thoracic seg 2 macrochaeta number (fig.3) 1....... ................................................ bohemica Posterior thoracic seg 2 macrochaeta number (fig.3) 3....... ............................................ subilliciens
149(139). Number of eyes per side 0............................... 150 Number of eyes per side 3........................ annemariae Number of eyes per side 5............................. mauli Number of eyes per side 6............................... 152
150(149). Ungual wing tooth (fig.1) absent........................ 151 Ungual wing tooth (fig.1) present............... handschini 20
151(150). Posterior thoracic seg 2 macrochaeta number (fig.3) 2; thoracic seg 3 macrochaetae number (fig.3) 0.............. ........................................... pseudodecepta Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 1.... illiciens
152(149). Ungual wing tooth (fig.1) absent; unquiculus shape (fig.) basally swollen; a (fig.4) smooth microchaeta; r (fig.) smooth microchaeta; dorsal cephalic macrochaeta t (fig.) absent.......................................... terricola Ungual wing tooth (fig.1) present; unquiculus shape (fig.) acuminate; a (fig.4) ciliated macrochaeta; r (fig.) ciliated microchaeta; dorsal cephalic macrochaeta t (fig.2) present............................... navarrensis
153(126). Unquiculus shape (fig.1) acuminate...................... 154 Unquiculus shape (fig.1) clavate........................ 156 Unquiculus shape (fig.1) basally swollen................ 159
154(153). Number of eyes per side 0.......................... maritima Number of eyes per side 1.............................. pori Number of eyes per side 2........................ biocellata Number of eyes per side 3............................... 155 Number of eyes per side 5........................... pygmaea
155(154). Distance distal unpaired ungual tooth from base/total unguis 51 to 57; antennal/cephalic diagonal 6 or more............ ............................................ burgalensis 22 Distance distal unpaired ungual tooth from base/total unguis 66 to 74; antennal/cephalic diagonal 1.5 to 1.59.......... ............................................ occidentalis Distance distal unpaired ungual tooth from base/total unguis 75 to 80; antennal/cephalic diagonal 3 to 3.49............ ........................................... montiscarmeli
156(153). Maximum length 0.8 to 0.99.............................. 157 Maximum length 1.2 to 1.39........................... albida Maximum length 1.4 to 1.59................... imparipunctata Maximum length 1.8 to 2.19.......................... charoae
157(156). Thoracic seg 3 macrochaetae number (fig.3) 0; ungual wing tooth (fig.1) absent; antennal/cephalic diagonal 1.3 to 1.49.................................................. 158 Thoracic seg 3 macrochaetae number (fig.3) 1; ungual wing tooth (fig.1) present; antennal/cephalic diagonal 1.5 to 1.59............................................. astarini
158(157). Q1 (fig.4) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 1; supplementary seta (fig.) absent; dorsal cephalic macrochaeta t (fig.2) present; distance distal unpaired ungual tooth from base/total unguis 66 to 74................................ ksenemani 13 Q1 (fig.4) ciliated macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 0; supplementary seta (fig.) present; dorsal cephalic macrochaeta t (fig.2) absent; distance distal unpaired ungual tooth from base/total unguis 75 to 80................................... ciliata
159(153). Number of eyes per side 0............................... 160 Number of eyes per side 3........................... hauseri Number of eyes per side 4............................... 161
160(159). Ungual wing tooth (fig.1) absent.................. recipiens Ungual wing tooth (fig.1) present............... handschini 20
161(159). Q1 (fig.4) smooth microchaeta; supplementary seta (fig.) present; maximum length 1 to 1.19; distance distal unpaired ungual tooth from base/total unguis 66 to 74..... ........................................... azerbaidzhani Q1 (fig.4) ciliated macrochaeta; supplementary seta (fig.) absent; maximum length 1.2 to 1.39; distance distal unpaired ungual tooth from base/total unguis 58 to 65..... ............................................... lenkorani
162(125). Number of eyes per side 0............................... 163 Number of eyes per side 3............................ simoni Number of eyes per side 5........................ difficilis
163(162). Dorsal cephalic macrochaeta t (fig.2) absent; maximum length 1.6 to 1.79........................................ pongei Dorsal cephalic macrochaeta t (fig.2) present; maximum length 1.2 to 1.39.................................... 164
164(163). Posterior thoracic seg 2 macrochaeta number (fig.3) 2; supplementary seta (fig.7) present; distance distal unpaired ungual tooth from base/total unguis 66 to 74; antennal/cephalic diagonal 1.3 to 1.49........ praecipiens Posterior thoracic seg 2 macrochaeta number (fig.3) 3; supplementary seta (fig.7) absent; distance distal unpaired ungual tooth from base/total unguis 58 to 65; antennal/cephalic diagonal 1.5 to 1.59............. noseki
165(125). Tenent hair shape (fig.1) acuminate..................... 166 Tenent hair shape (fig.1) clavate....................... 167
166(165). Maximum length 1.2 to 1.39; antennal/cephalic diagonal 1.5 to 1.59......................................... anderseni Maximum length 1.4 to 1.59; antennal/cephalic diagonal 1.9 to 2.49......................................... halophila
167(165). Posterior thoracic seg 2 macrochaeta number (fig.3) 0... 168 Posterior thoracic seg 2 macrochaeta number (fig.3) 1... 169 Posterior thoracic seg 2 macrochaeta number (fig.3) 2... 170 Posterior thoracic seg 2 macrochaeta number (fig.3) 3....... .................................................. bachae
168(167). L1 (fig.5) smooth macrochaeta; a (fig.4) ciliated macrochaeta; e (fig.5) smooth macrochaeta; r (fig.) smooth microchaeta; m1 (fig.5) smooth macrochaeta... gamae L1 (fig.5) ciliated macrochaeta; a (fig.4) ciliated microchaeta; e (fig.5) ciliated macrochaeta; r (fig.) ciliated macrochaeta; m1 (fig.5) ciliated macrochaeta..... ........................................ pseudopetterseni
169(167). Supplementary seta (fig.7) absent; ungual wing tooth (fig.) present; number of eyes per side 2; distance distal unpaired ungual tooth from base/total unguis 58 to 65..... ................................................ espanoli Supplementary seta (fig.7) present; ungual wing tooth (fig.1) absent; number of eyes per side 3; distance distal unpaired ungual tooth from base/total unguis 75 to 80..... ................................................. arretzi
170(167). Thoracic seg 3 macrochaetae number (fig.3) 0; unquiculus shape (fig.1) acuminate; antennal/cephalic diagonal 1.6 to 1.89............................................ soriensis Thoracic seg 3 macrochaetae number (fig.3) 2; unquiculus shape (fig.1) basally swollen; antennal/cephalic diagonal 1.5 to 1.59.................................... petterseni
171(123). Maximum length up to 0.6.......................... paprivata Maximum length 0.8 to 0.99.............................. 172 Maximum length 1 to 1.19................................ 173 Maximum length 1.2 to 1.39.............................. 174 Maximum length 1.4 to 1.59.............................. 178 Maximum length 1.6 to 1.79.............................. 179 Maximum length 1.8 to 2.19.............................. 180
172(171). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; antennal/cephalic diagonal 1.3 to 1.49............ ciliata Posterior thoracic seg 2 macrochaeta number (fig.3) 1; antennal/cephalic diagonal 1.5 to 1.59........ cordobensis Posterior thoracic seg 2 macrochaeta number (fig.3) 2; antennal/cephalic diagonal 1.6 to 1.89........ cementensis
173(171). Distance distal unpaired ungual tooth from base/total unguis 58 to 65.................................... torcuatoensis Distance distal unpaired ungual tooth from base/total unguis 81 to 90.......................................... cobosae Distance distal unpaired ungual tooth from base/total unguis 91 or more........................................ aidamar
174(171). Distance distal unpaired ungual tooth from base/total unguis 41 to 50..................................... logrognensis Distance distal unpaired ungual tooth from base/total unguis 51 to 57....................................... leivaensis Distance distal unpaired ungual tooth from base/total unguis 58 to 65.............................................. 175 Distance distal unpaired ungual tooth from base/total unguis 66 to 74.............................................. 176 Distance distal unpaired ungual tooth from base/total unguis 75 to 80.............................................. 177
175(174). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; thoracic seg 3 macrochaetae number (fig.3) 0; ungual wing tooth (fig.1) present; unquiculus shape (fig.1) acuminate; r (fig.5) ciliated microchaeta................ burgalensis Posterior thoracic seg 2 macrochaeta number (fig.3) 1; thoracic seg 3 macrochaetae number (fig.3) 1; ungual wing tooth (fig.1) absent; unquiculus shape (fig.1) basally swollen; r (fig.5) smooth microchaeta.............. selgae
176(174). E (fig.5) smooth macrochaeta; L1 (fig.5) smooth macrochaeta; a (fig.4) ciliated microchaeta; q1 (fig.4) ciliated macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 0..................................... gutierrezae E (fig.5) ciliated macrochaeta; L1 (fig.5) ciliated macrochaeta; a (fig.4) ciliated macrochaeta; q1 (fig.) smooth microchaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 3.................................. aeolica
177(174). E (fig.5) smooth macrochaeta; L1 (fig.5) smooth macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 1; thoracic seg 3 macrochaetae number (fig.3) 0; ungual wing tooth (fig.1) absent............................. tyrrhena E (fig.5) ciliated macrochaeta; L1 (fig.5) ciliated macrochaeta; posterior thoracic seg 2 macrochaeta number (fig.3) 2; thoracic seg 3 macrochaetae number (fig.3) 1; ungual wing tooth (fig.1) present................. helenae
178(171). Number of eyes per side 0; r (fig.5) absent.......... hrabei Number of eyes per side 3; r (fig.5) ciliated microchaeta... ............................................... moldavica Number of eyes per side 5; r (fig.5) ciliated macrochaeta... ............................................. sandelsorum
179(171). A (fig.4) smooth microchaeta; q1 (fig.4) smooth microchaeta; tenent hair shape (fig.1) clavate; unquiculus wing tooth (fig.1) absent; unquiculus shape (fig.1) clavate.... picta A (fig.4) ciliated macrochaeta; q1 (fig.4) ciliated macrochaeta; tenent hair shape (fig.1) acuminate; unquiculus wing tooth (fig.1) minute; unquiculus shape (fig.1) acuminate.................................... ioni
180(171). Antennal/cephalic diagonal 1.5 to 1.59................ codri Antennal/cephalic diagonal 1.6 to 1.89.................. 181 Antennal/cephalic diagonal 1.9 to 2.49............... gruiae
181(180). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; unquiculus shape (fig.1) basally swollen; number of eyes per side 0; dorsal cephalic macrochaeta s (fig.2) absent; dorsal cephalic macrochaeta t (fig.2) absent...... larisae Posterior thoracic seg 2 macrochaeta number (fig.3) 1; unquiculus shape (fig.1) acuminate; number of eyes per side 3; dorsal cephalic macrochaeta s (fig.2) present; dorsal cephalic macrochaeta t (fig.2) present... simpatica
182(122). Maximum length 0.8 to 0.99........................ sanguinea Maximum length 1.2 to 1.39............................. sera Maximum length 1.8 to 2.19........................... rolfsi Maximum length 2.5 to 2.99.......................... folsomi
183(122). Q1 (fig.4) smooth microchaeta; q2 (fig.4) smooth microchaeta................................. biunguiculata Q1 (fig.4) ciliated microchaeta; q2 (fig.4) ciliated microchaeta.................................. lahainaensis Q1 (fig.4) absent; q2 (fig.4) absent................ violeta
184(122). Distance distal unpaired ungual tooth from base/total unguis 36 to 40........................................ peruensis Distance distal unpaired ungual tooth from base/total unguis 41 to 50.............................................. 185 Distance distal unpaired ungual tooth from base/total unguis 66 to 74........................................ rapoporti Distance distal unpaired ungual tooth from base/total unguis 75 to 80....................................... intermixta Distance distal unpaired ungual tooth from base/total unguis 81 to 90.............................................. 186
185(184). M2 (fig.5) smooth macrochaeta; q2 (fig.4) smooth microchaeta; tenent hair shape (fig.1) acuminate; number of teeth of inner unguis (fig.1) 3; unquiculus wing tooth (fig.1) fully developed......................... federicoi M2 (fig.5) ciliated macrochaeta; q2 (fig.4) absent; tenent hair shape (fig.1) clavate; number of teeth of inner unguis (fig.1) 2; unquiculus wing tooth (fig.1) absent.... ............................................... metallica
186(184). Number of eyes per side 0; antennal/cephalic diagonal 1.01 to 1.29.......................................... gleycola Number of eyes per side 5; antennal/cephalic diagonal 1.3 to 1.49.............................................. confusa
187(122). Number of eyes per side 0....................... praecipiens Number of eyes per side 1......................... biguttata Number of eyes per side 2.................... imparipunctata
188(122). Number of eyes per side 0............................... 189 Number of eyes per side 2............................... 191 Number of eyes per side 3...................... sensillifera Number of eyes per side 4............................... 192 Number of eyes per side 5.................. pseudolanuginosa Number of eyes per side 6............................... 195
189(188). Number of teeth of inner unguis (fig.1) 2............ montis Number of teeth of inner unguis (fig.1) 3............... 190 Number of teeth of inner unguis (fig.1) 4........ bellingeri
190(189). Unquiculus wing tooth (fig.1) absent; unquiculus shape (fig.1) clavate; antennal/cephalic diagonal 1.6 to 1.89... ................................................ yosiiana Unquiculus wing tooth (fig.1) fully developed; unquiculus shape (fig.1) basally swollen; antennal/cephalic diagonal 1.5 to 1.59.................................... petterseni
191(188). Maximum length 0.61 to 0.79...................... baghdadica Maximum length 0.8 to 0.99..................... genghizkhani
192(188). Maximum length up to 0.6................................ 193 Maximum length 0.61 to 0.79............................. 194 Maximum length 1 to 1.19............................. vallis Maximum length 1.8 to 2.19............................. paxi
193(192). R (fig.5) smooth microchaeta; a (fig.4) ciliated macrochaeta; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1; dorsal cephalic macrochaeta s (fig.2) present; dorsal cephalic macrochaeta t (fig.) present......................................... mutabilis R (fig.5) absent; a (fig.4) ciliated microchaeta; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0; dorsal cephalic macrochaeta s (fig.2) absent; dorsal cephalic macrochaeta t (fig.2) absent.............. tumula
194(192). A (fig.4) ciliated microchaeta; q1 (fig.4) smooth microchaeta; antennal/cephalic diagonal 1.9 to 2.49; b (fig.4) smooth microchaeta................... tridentifera A (fig.4) ciliated macrochaeta; q1 (fig.4) ciliated macrochaeta; antennal/cephalic diagonal 1.3 to 1.49; b (fig.4) ciliated macrochaeta....................... staryi A (fig.4) absent; q1 (fig.4) ciliated microchaeta; antennal/cephalic diagonal 1.01 to 1.29; b (fig.) ciliated microchaeta........................... grinnellia
195(188). Number of teeth of inner unguis (fig.1) 2................... ........................................... sandakanicus 24 Number of teeth of inner unguis (fig.1) 3........ segamanus 24 Number of teeth of inner unguis (fig.1) 4................... ........................................... sepilokensis 24
196(122). Number of eyes per side 1; maximum length 5 or more; unquiculus shape (fig.1) acuminate............. unioculata Number of eyes per side 6; maximum length 1.4 to 1.59; unquiculus shape (fig.1) clavate...................... 197
197(196). Distance distal unpaired ungual tooth from base/total unguis up to 15......................................... spelunca Distance distal unpaired ungual tooth from base/total unguis 66 to 74......................................... fasciata
198(122). Q2 (fig.4) smooth microchaeta........................... 199 Q2 (fig.4) ciliated microchaeta......................... 200 Q2 (fig.4) absent.................................. fujiokai
199(198). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; tenent hair shape (fig.1) truncate; unquiculus shape (fig.1) clavate; number of eyes per side 4; maximum length 1.2 to 1.39.................................. kalalauensis Posterior thoracic seg 2 macrochaeta number (fig.3) 1; tenent hair shape (fig.1) clavate; unquiculus shape (fig.1) acuminate; number of eyes per side 6; maximum length 1.6 to 1.79............................... ashmolae
200(198). Posterior thoracic seg 2 macrochaeta number (fig.3) 0; tenent hair shape (fig.1) acuminate; number of teeth of inner unguis (fig.1) 2; unquiculus wing tooth (fig.) fully developed; dorsal cephalic macrochaeta t (fig.) absent....................................... lahainaensis Posterior thoracic seg 2 macrochaeta number (fig.3) 1; tenent hair shape (fig.1) clavate; number of teeth of inner unguis (fig.1) 3; unquiculus wing tooth (fig.) absent; dorsal cephalic macrochaeta t (fig.2) present..... ................................................. miratio
201(122). Maximum length 0.8 to 0.99; distance distal unpaired ungual tooth from base/total unguis 41 to 50; antennal/cephalic diagonal 1.01 to 1.29........................ dispadentata Maximum length 1 to 1.19; distance distal unpaired ungual tooth from base/total unguis 91 or more; antennal/cephalic diagonal 1.5 to 1.59......................... insoloculata Maximum length 1.2 to 1.39; distance distal unpaired ungual tooth from base/total unguis 36 to 40; antennal/cephalic diagonal 1.6 to 1.89........................... nonoculata
202(1). Unquiculus wing tooth (fig.1) absent.................... 203 Unquiculus wing tooth (fig.1) minute.................... 223 Unquiculus wing tooth (fig.1) fully developed........... 224
203(202). Ungual wing tooth (fig.1) absent........................ 204 Ungual wing tooth (fig.1) present....................... 221
204(203). Dorsal cephalic macrochaeta t (fig.2) absent............ 205 Dorsal cephalic macrochaeta t (fig.2) present........... 208
205(204). Maximum length 1.2 to 1.39............................ alba 14 Maximum length 1.4 to 1.59........................... stachi Maximum length 1.6 to 1.79.............................. 206 Maximum length 1.8 to 2.19........................ efficiens Maximum length 2.5 to 2.99.............................. 207 Maximum length 3.5 to 3.99.......................... hirsuta
206(205). E (fig.5) smooth macrochaeta; L1 (fig.5) smooth macrochaeta; q1 (fig.4) smooth macrochaeta; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1; m2 (fig.5) smooth macrochaeta................................... sexoculata 14 E (fig.5) ciliated macrochaeta; L1 (fig.5) ciliated macrochaeta; q1 (fig.4) smooth microchaeta; anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0; m2 (fig.5) ciliated macrochaeta................... collina
207(205). B (fig.4) smooth microchaeta; distance distal unpaired ungual tooth from base/total unguis 66 to 74; antennal/cephalic diagonal 1.5 to 1.59; apical organ of third antennal segment (fig.6) peg or rod-like; differentiated inner seta on hind tibiotarsus (fig.) clear, acuminate..................................... aera B (fig.4) smooth macrochaeta; distance distal unpaired ungual tooth from base/total unguis 58 to 65; antennal/cephalic diagonal 1.6 to 1.89; apical organ of third antennal segment (fig.6) expanded; differentiated inner seta on hind tibiotarsus (fig.9) unclear or absent. ................................................ argentea
208(204). Tenent hair shape (fig.1) acuminate............... decipiens Tenent hair shape (fig.1) clavate....................... 209
209(208). Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 0............................................. 210 Anterior lateral(P) fourth abdominal dorsal macrochaetae (fig.3) 1............................................. 211
210(209). Number of eyes per side 0; maximum length 0.61 to 0.79...... .............................................. infrequens Number of eyes per side 2; maximum length up to 0.6......... ............................................... insularum Number of eyes per side 3; maximum length 0.8 to 0.99....... .............................................. trioculata Number of eyes per side 4; maximum length 1.4 to 1.59....... .................................................. stachi
211(209). Supplementary seta (fig.7) absent....................... 212 Supplementary seta (fig.7) present...................... 220
212(211). Dorsal cephalic macrochaeta s (fig.2) absent............ 213 Dorsal cephalic macrochaeta s (fig.2) present........... 219
213(212). Cephalic seta r0 (fig.2) absent......................... 214 Cephalic seta r0 (fig.2) present........................ 217
214(213). Thoracic seg 3 macrochaetae number (fig.3) 0............ 215 Thoracic seg 3 macrochaetae number (fig.3) 1......... stachi Thoracic seg 3 macrochaetae number (fig.3) 2............ 216 Thoracic seg 3 macrochaetae number (fig.3) 3......... stachi
215(214). Number of eyes per side 4; maximum length 1.4 to 1.59....... .................................................. stachi Number of eyes per side 5; maximum length 1 to 1.19......... ............................................... wahlgreni
216(214). Region located Europe and North Africa; distance distal unpaired ungual tooth from base/total unguis 66 to 74..... ............................................ subilliciens Region located Asia; distance distal unpaired ungual tooth from base/total unguis 58 to 65.................... stachi
217(213). Region located Europe and North Africa.................. 218 Region located Asia.................................. stachi
218(217). Posterior thoracic seg 2 macrochaeta number (fig.3) 1; thoracic seg 3 macrochaetae number (fig.3) 0; a (fig.) smooth microchaeta.............................. zygophora Posterior thoracic seg 2 macrochaeta number (fig.3) 3; thoracic seg 3 macrochaetae number (fig.3) 2; a (fig.) ciliated macrochaeta......................... subilliciens
219(212). Distance distal unpaired ungual tooth from base/total unguis up to 15; differentiated inner seta on hind tibiotarsus (fig.9) clear, truncate or clavate......... octopunctata 14 Distance distal unpaired ungual tooth from base/total unguis 58 to 65; differentiated inner seta on hind tibiotarsus (fig.9) unclear or absent......................... stachi Distance distal unpaired ungual tooth from base/total unguis 75 to 80; differentiated inner seta on hind tibiotarsus (fig.9) clear, acuminate................... octopunctata 15
220(211). Unquiculus shape (fig.1) acuminate; number of eyes per side 4.................................................. stachi Unquiculus shape (fig.1) clavate; number of eyes per side 2. ................................................... alba 15 Unquiculus shape (fig.1) basally swollen; number of eyes per side 3........................................ sexoculata 15
221(203). Antennal/cephalic diagonal up to 1...................... hui Antennal/cephalic diagonal 1.3 to 1.49......... subilliciens Antennal/cephalic diagonal 1.5 to 1.59........... immaculata Antennal/cephalic diagonal 1.9 to 2.49.................. 222
222(221). Unquiculus shape (fig.1) clavate; number of eyes per side 6. .............................................. suboculata Unquiculus shape (fig.1) basally swollen; number of eyes per side 0........................................ handschini 20
223(202). Q1 (fig.4) smooth microchaeta; dorsal cephalic macrochaeta t (fig.2) absent; unquiculus shape (fig.1) basally swollen; number of eyes per side 0; maximum length 2.2 to 2.49..... ................................................ vandeli 16 Q1 (fig.4) absent; dorsal cephalic macrochaeta t (fig.) present; unquiculus shape (fig.1) clavate; number of eyes per side 6; maximum length 1.8 to 2.19....... problematica
224(202). Q1 (fig.4) ciliated microchaeta; q2 (fig.4) ciliated microchaeta; unquiculus shape (fig.1) clavate; maximum length 1 to 1.19; m2 (fig.5) absent.................. caoi Q1 (fig.4) smooth macrochaeta; q2 (fig.4) smooth microchaeta; unquiculus shape (fig.1) acuminate; maximum length 1.8 to 2.19; m2 (fig.5) smooth macrochaeta......... ................................................ violenta

Endnotes

¹. As lamperti lamperti.

². Sensu Gisin - da Gama.

³. As vandeli meridionalis.

⁴. As vandeli relicta.

⁵. As vandeli praealpina.

⁶. As bidenticulata crenelata.

⁷. As bidenticulata bidenticulata.

⁸. As subterranea baztanensis.

⁹. As lamperti germanica.

¹⁰. With the current characters and states, Pseudosinella subvirei can only be distinguished from virei in the relative position of the distal tooth on the claw.

¹¹. As tarraconensis tarraconensis.

¹². As tarraconensis longicornis.

¹³. Sensu Hüther.

¹⁴. From America.

¹⁵. From Europe.

¹⁶. As vandeli vandeli.

¹⁷. martelii has been sunk to synonymy with immaculata by Schött (1902 cfr Handschin, 1929).

¹⁸. propinqua has been sunk to synonymy with decipiens by Goto & Lawrence in Kloet & Hincks (1964).

¹⁹. As dodecopsis caerulea.

²⁰. handschini has been sunk to synonymy with decipiens by Goto & Lawrence in Kloet & Hincks (1964).

²¹. duodecimpunctata has been sunk to synonymy with duodecimoculata by Dallai & Malatesta (1983).

²². ????

²³. Of Christiansen, K. 1960:24 nec Bagnall 1941.

²⁴. Currently in Lepidocyrtus.

References