Checklist of the Collembola: Nomina Collembola Belgica

http://www.collembola.org/publicat/belgium/collemb/nomina.htm - Last updated on 2022.06.29 by Frans Janssens

Checklist of the Collembola: Nomina Collembola Belgica, iteratio 20

Frans Janssens, Department of Biology, University of Antwerp, Antwerp, B-2020, Belgium

Abstract

Samenvatting

Deze voorlopige uitgave van de naamlijst van de Collembola (Hexapoda) van België is gebaseerd op literatuur onderzoek. De lijst omvat de referenties naar de originele en latere vermeldingen voor alle 217 soorten van ons land en koppelingen naar 10*10 UTM verspreidingskaarten.

Résumé

Une édition provisoire de la liste des Collembola (Hexapoda) de Belgique basée sur la littérature est fournie. Cette liste inclut les références aux citations originales et à certaines citations subséquentes des 217 espèces de notre pays ainsi qu'un lien à leurs cartes de distribution sur un quadrillage UTM 10*10 km.

Keywords: Collembola, checklist, distribution maps, Belgium

Preamble

This article is a translation and a revision of
Janssens, F. 1996. Nomina Collembola Belgica. Antwerpse Vereniging voor Entomologie, Entomo-Info, Jaargang 7, 1996, nr 3, p.79-90.
Since this first edition, 87 new species have been added to the list.

Contineously being revised.

100 years of Belgian Springtails research

This provisional edition is based on a literature survey and does not yet take into account all our own observations.
The following publications give a short overview of the history of 100 year of Belgian Collembola research :

Lameere (1895) : 45 species
Marlier (1942) : 84 species (+ 5 subspecific forms)
Berbiers & Mertens (1989) : 53 species
Janssens (1996) : 113 species

The current provisional list contains 217 species.

Systematic species list

A provisional edition of the checklist of Collembola (Hexapoda) from Belgium based on a literature survey is provided. The list includes the references to the original and several subsequent quotations for all 217 species from our country and a link to their distribution maps on a 10*10 km UTM grid. The checklist is a systematic list of valid taxon names with reference to the original authority, followed by the revision authority, if applicable. Each name follows the original orthography and mentions the original genus. The valid taxon names are ordered phylogenetically up to the rank of genus. The species valid names are ordered alphabethically within each genus. However, the references to the unidentified species are listed first. In each species, the references to the quotations are ordered chronologically, according to the traditional presentation of zoological catalogs (i.e. the name in this original spelling and combination; the cases of original invalid use of homonyms or incorrect spellings are indicated respectively by nec and sic). Synonyms, homonyms (cf. 'nec'), incorrect spellings (cf. 'sic') are ordered chronologically for each species. Whenever the original publication was verified, the page reference is mentioned following the publication date. (cf. 'yyyy:pp'; e.g. '1902:8'). In case the genus was not explicitly mentioned in the referenced publication, it is indicated by '...'. Publication dates are not always mentioned in such case.

Abbreviations used in the species list

?	tentative species determination
cf	confer species
e	erroneous species determination
gr	group species
h	homonym
il	in litteris
ios	incorrect original spelling
ip	in print
iss	incorrect subsequent spelling
sp	undefined species
un	unrecorded specimens

A preliminary distribution map is provided for each species. The maps record the presence of all species in a 10 x 10 km UTM grid. Records published before 1950 are indicated with open white circle symbols, records published after 1950 are indicated with solid black circle symbols.
The maps are produced with the mapping module of the modular software package Inventar 1.3, intended to keep track of data collected by nature monitoring projects and data of collections of natural objects. Inventar is developed by Jan de Boer and it is distributed by Mistletoe Information, Maastricht, the Netherlands.
Click on the species name to open the associated species distribution map. Note that the available distribution data is rather sparse for most species and that several records can be added still.
Click on the Collembola taxon to open a map that gives an overview of all record localities.

Classis Collembola Lubbock J, 1870:295

Ordo Poduromorpha Börner C, 1913:318, sensu D'Haese CA, 2002:1148

Familia Poduridae Latreille, 1804, i.s.

Genus Podura Linnæus, C, 1758:608

Species 1. aquatica Linnæus, C, 1758:609

Podura aquatica Lameere, A, 1895:594
Podura aquatica Willem, V, 1900:9
Podura aquatica Börner, C, 1901:131
Podura aquatica Lameere, A, 1940:8
Podura aquatica Marlier, GJ, 1942:8
Podura aquatica Geyskens, J, 1945:15
Podura aquatica Janssens, A, 1945:10
Podura aquatica Janssens, F, 1993:67

Familia Hypogastruridae Börner, 1906

Genus Hypogastrura Bourlet, C, 1839:404

Hypogastruridae Hypogastrura Allaer, L in Segers, C, 1964:Collembola included:66 ^sp
Hypogastruridae Hypogastrura Dethier, M et Hubart, J-M, 2000:27 ^sp
Hypogastruridae Hypogastrura Janssens, F et De Bruyn, L, 2005:ip ^sp
Hypogastruridae Hypogastrura Janssens, F et Dethier, M, 2005:150

Species 2. burkilli (Bagnall, RS, 1940:167) Weidner, 1953

Hypogastrura burkilli Dethier, M, 1998:55
Hypogastrura burkilli Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Hypogastrura burkilli Dethier, M et Hubart, J-M, 2000:27
Hypogastrura burkilli Dethier, M et Hubart, J-M, 2000:27 ^cf

Species 3. purpurescens (Lubbock, J, 1870:302) Wolf, B, 1938:109

Achorutes purpurascens(sic) var. pallens Willem, V, 1902:278
Hypogastrura (Hypogastrura) purpurascens(sic) var. pallens Bonet, F, 1931:374
Hypogastrura (Hypogastrura) purpurascens(sic) var. pallescens(sic) Wolf, B, 1938:109 laps
Hypogastrura (Hypogastrura) purpurascens(sic) var. pallescens(sic) Leruth, R, 1939:196 laps
Hypogastrura purpurascens(sic) var. pallescens(sic) Marlier, GJ, 1942:8 laps
Hypogastrura purpurascens(sic) Marlier, GJ, 1942:8 ^iss
Hypogastrura purpurascens(sic) Delhez, F et Houssa, M, 1969:207 ^iss
Hypogastrura purpurescens Delhez, F et Kersmaekers, M, 1973:8
Hypogastrura purpurescens Heungens, A et Van Daele, E, 1973:138
Hypogastrura purpurescens Thibaud, J-M et Massoud, Z, 1977:6
Hypogastrura purpurascens(sic) Hubart, J-M, 1982:343 ^iss
Hypogastrura purpurescens Dethier, M, 1998:55
Hypogastrura purpurescens Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Hypogastrura purpurescens Dethier, M et Hubart, J-M, 2000:27
Hypogastrura purpurescens Dethier, M et Hubart, J-M, 2003:52
Hypogastrura purpurescens Janssens, F et Dethier, M, 2005:151

Species 4. viatica (Tullberg, 1872:50) Carpenter, 1909

Hypogastrura viatica f. inermis Marlier, GJ, 1942:8
Achorutes viaticus Lameere, A, 1895:594
Achorutes viaticus Willem, V, 1900:19
Achorutes (Euachorutes) viaticus Börner, C, 1901:131
Achorutes viaticus Lameere, A, 1940:8
Hypogastrura viatica Marlier, GJ, 1942:8
Achorutes viaticus Geyskens, J, 1945:15
Hypogastrura viatica Heungens, A et Van Daele, E, 1973:140
Hypogastrura viatica Mertens, J et Bourgoignie, R, 1975:369
Hypogastrura viatica Mertens, J et Bourgoignie, R, 1977:41
Hypogastrura viatica Mertens, J, Coessens, R et Blancquaert, J-P, 1983:567
Hypogastrura viatica Berbiers, P et Mertens J, 1989:236
Hypogastrura viatica Janssens, F, Jacobs, W et De Boey, B, 1998:47
Hypogastrura viatica Lock, K, Janssens, F et Janssen, CR, 2003:27

Species 5. assimilis (Krausbauer, 1898) Handschin, E in Dahl, F, 1929:25

Hypogastrura manubrialis ssp. assimilis Delamare Deboutteville, C, 1948:136
Hypogastrura manubrialis ssp. assimilis Marlier, G in Pasteels, M, 1948:59

Species 6. manubrialis (Tullberg, 1869:9) Linnaniemi, 1912

Hypogastrura manubrialis var. neglecta Marlier, GJ, 1942:8
Hypogastrura manubrialis Marlier, GJ, 1942:8
Hypogastrura manubrialis Heungens, A et Van Daele, E, 1973:138
Hypogastrura manubrialis Grégoire-Wibo, C, 1981:632
Hypogastrura manubrialis Janssens, F et Witters, J, 2007:ip

Species 7. vernalis (Carl, J, 1901:249) Handschin, E, 1924:91

Hypogastrura vernalis Heungens, A et Van Daele, E, 1973:136

Species 8. sahlbergi (Reuter, OM, 1895:31) Handschin, E, 1924:91

Hypogastrura sahlbergi Marlier, GJ, 1942:8
Hypogastrura sahlbergi Lock, K, Janssens, F et Janssen, CR, 2003:27 ^cf

Genus Willemia Börner, 1901

Species 9. anophthalma Börner, 1901

Willemia anophtalma(sic) Marlier, 1942:4 ^iss
Willemia anophthalma Ponge, J-F, 2000:510
Willemia anophthalma Gillet, S et Ponge, J-F, 2004:223

Species 10. denisi Mills, HB, 1932:263

Willemia aspinata Ponge, J-F, 2000:510
Willemia denisi Gillet, S et Ponge, J-F, 2004:223

Genus Mesachorutes Absolon, K, 1900:266

Species 11. quadriocellatus Absolon, K, 1900:267

Mesachorutes marlieri Delamare Deboutteville, 1947
Hypogastrura (Mesachorutes) quadriocellata Leleup, N, 1947:322
Hypogastrura (Mesachorutes) quadriocellata Marlier, G in Brien, P, 1947:158
Hypogastrura (Mesachorutes) quadriocellata Marlier, G, 1947:162

Genus Ceratophysella Börner in Brohmer, 1932

Hypogastruridae Ceratophysella Dethier, M et Hubart, J-M, 2000:27 ^sp
Hypogastruridae Ceratophysella Janssens, F et De Bruyn, L, 2005:ip ^sp
Hypogastruridae Ceratophysella Janssens, F et Dethier, M, 2005:151 ^sp

Species 12. armata (Nicolet, H, 1842:57) Börner, 1932

Achorutes armatus Lameere, A, 1895:594
Achorutes armatus Willem, V, 1900:6
Achorutes (Euachorutes) armatus f. principalis Börner, C, 1901:131
Achorutes armatus Willem, V, 1902:278
Hypogastrura (Hypogastrura) armata Bonet, F, 1931:373
Hypogastrura (Hypogastrura) armata Wolf, B, 1938:107
Hypogastrura (Hypogastrura) armata Leruth, R, 1939:196
Hypogastrura armata Marlier, GJ, 1942:8
Achorutes armatus Geyskens, J, 1945:16
Ceratophysella armata Verlinden, A, 1977:114
Ceratophysella armata Janssens, F et De Bruyn, L, 2005:ip

Species 13. bengtssoni (Ågren, H, 1904:2) Stach, J, 1949:142

Hypogastrura (Hypogastrura) stygia Leruth, R, 1939:196
Hypogastrura stygia Marlier, GJ, 1942:8
Hypogastrura bengtssoni Dethier, M, 1998:55
Hypogastrura bengtssoni Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Ceratophysella bengtssoni Dethier, M et Hubart, J-M, 2000:27
Ceratophysella bengtssoni Janssens, F et Dethier, M, 2005:151

Species 14. cavicola (Börner, 1901) Thibaud, J-M, 1970:88

Hypogastrura cavicola Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Hypogastrura cavicola Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Hypogastrura cavicola Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Hypogastrura cavicola Pollet, M et Desender, K, 1985:583
Hypogastrura cavicola Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Hypogastrura cavicola Berbiers, P et Mertens J, 1989:236
Schaefferia cavicola Dethier, M, 1998:55

Species 15. denticulata (Bagnall, 1941) Stach, 1949

Hypogastrura denticulata Heungens, A et Van Daele, E, 1973:136
Ceratophysella denticulata Jacobs, W, 1997:71
Ceratophysella denticulata Jacobs, W, De Bruyn, L et Lust, N, 1999:57
Ceratophysella denticulata Ponge, J-F, 2000:510
Ceratophysella denticulata Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Ceratophysella denticulata Lock, K, Janssens, F et Janssen, CR, 2003:27
Ceratophysella denticulata Gillet, S et Ponge, J-F, 2004:223
Ceratophysella denticulata Janssens, F et De Bruyn, L, 2005:ip
Ceratophysella denticulata Janssens, F et Witters, J, 2007:ip

Genus Schaefferia Absolon, K, 1900:265

Species 16. emucronata Absolon, K, 1900:266

Schaefferia emucronata Gillet, S et Ponge, J-F, 2004:223

Species 17. willemi (Bonet, F, 1930:125) Denis, J-R, 1936:82

Hypogastrura (Schaefferia) willemi Bonet, F, 1930:125
Octomma longispina Willem, V, 1902:279 nec Tullberg, 1876 ^h
Hypogastrura (Schötella(sic)) willemi Bonet, F, 1931:272
Hypogastrura (Schaefferia) willemi Bonet, F, 1931:374
Hypogastrura (Schäfferia(sic)) Willemi(sic) Denis, J-R, 1937:2 ^iss
Hypogastrura (Schaefferia) willemi Wolf, B, 1938:110
Hypogastrura (Schaefferia) Willemi(sic) Leruth, R, 1939:197 ^iss
Schäfferia(sic) willemi Marlier, GJ, 1942:8
Schaefferia willemi Gisin, H, 1960:55
Schaefferia willemi Thibaud, J-M et Massoud, Z, 1977:6
Schaefferia willemi Hubart, J-M, 1982:343
Schaefferia willemi Jordana, R et Arbea, JI in Ramos, MA et al, 1997:206
Hypogastrura willemi Dethier, M, 1998:38
Schaefferia willemi Dethier, M, 1998:55
Schaefferia willemi Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Schaefferia willemi Hubart, J-M et Dethier, M, 1999:167
Schaefferia willemi Dethier, M et Hubart, J-M, 2000:28
Schaefferia willemi Ponge, J-F, 2000:510
Schaefferia willemi Dethier, M et Hubart, J-M, 2003:52
Schaefferia willemi Dethier, M et Hubart, J-M, 2005:33
Schaefferia willemi Janssens, F et Dethier, M, 2005:151
Schaefferia willemi Stomp, N et Weiner, WM, 2005:231
Schaefferia willemi Dethier, M et Hubart, J-M, 2010:25
Schaefferia willemi Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:46

Genus Schoettella Schäffer, 1896

Species 18. ununguiculata (Tullberg, 1869) Schäffer, 1896

Schoetella(sic) ununguiculata Lock, K, Janssens, F et Janssen, CR, 2003:27

Genus Choreutinula Paclt, 1944

Species 19. inermis (Tullberg, 1871) Stach, 1955

Achorutes (Euachorutes) inermis Börner, C, 1901:131

Genus Mesogastrura Bonet, F, 1930:132

Species 20. sp. Leruth, R, 1939:197

Hypogastrura (Mesogastrura) sp. nov. Leruth, R, 1939:197

Genus Xenylla Tullberg, 1869

Species 21. brevicauda Tullberg, 1869

Xenylla brevicauda Jacquemart, S, 1967:5

Species 22. grisea Axelson, 1900

Xenylla grisea Ponge, J-F, 2000:510

Species 23. humicola (Fabricius O, 1780) Reuter, 1876

Xenylla humicola Janssens, F, 1993:68
Xenylla humicola Janssens, F et Bonte, D, 2004:ip

Species 24. tullbergi Börner, 1903

Xenylla tullbergi André, HM, 1983:274
Xenylla tullbergi Ponge, J-F, 2000:510

Species 25. welchi Folsom, JW, 1916:497

Xenylla welchi Delamare Deboutteville, C, 1948:133
Xenylla Welchi(sic) Marlier, G in Pasteels, M, 1948:59 ^iss

Species 26. brevisimilis Stach, J, 1949:232

Xenylla brevisimilis André, H, 1977:215
Xenylla brevisimilis ssp. brevisimilis André, H, 1977:222
Xenylla brevisimilis André, HM, 1983:274

Species 27. maritima Tullberg, 1869

Xenylla maritima Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Xenylla maritima Jacobs, W, Janssens, F et De Bruyn, L in Peeters, M, Franklin, A et Van Goethem, JL, 2003:127
Xenylla maritima Lock, K, Janssens, F et Janssen, CR, 2003:27

Familia Neanuridae Börner, 1901 sensu Yosii, 1956

Genus Friesea von Dalla Torre, 1895

Species 28. mirabilis (Tullberg, 1871) Stach, 1949

... mirabilis Lameere
Frisea(sic) mirabilis Marlier, GJ, 1942:8
Friesea mirabilis Heungens, A et Van Daele, E, 1973:138
Friesea mirabilis Jacobs, W, 1997:71
Friesea mirabilis Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Species 29. truncata Cassagnau, P, 1958:20

Friesea truncata Ponge, J-F, 2000:510
Friesea truncata Gillet, S et Ponge, J-F, 2004:223

Genus Gisinea Massoud, Z, 1965:431

Species 30. delhezi Massoud, Z, 1965:432

Gisinea delhezi Massoud, Z, 1965:432
Gisinea delhezi Delhez, F, 1965:11
Gisinea delhezi Massoud, Z, 1967:141
Gisinea delhezi Delhez, F, Gilson, R et Hubart, J-M, 1973:411
Gizinea(sic) delhezi Delhez, F, Gilson, R et Hubart, J-M, 1973:411
Gisinea delhezi Hubart, J-M, 1976:282
Gisinea delhezi Thibaud, J-M et Massoud, Z, 1977:6
Gisinea delhesi(sic) Hubart, J-M, 1982:343 ^iss
Gisinea delhezi Dethier, M, 1998:33
Gisinea delhezi Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Gisinea delhezi Hubart, J-M et Dethier, M, 1999:167
Gisinea delhezi Delhez, Dethier & Hubart, 2000 in prep
Gisinea delhezi Dethier, M et Hubart, J-M, 2010:25

Genus Neanura MacGillivray, AD, 1893:314

Species 31. muscorum (Templeton, R in Templeton, R & Westwood, JO, 1836:97) MacGillivray, AD, 1893:314

Anura muscorum Lameere, A, 1895:595
Anura muscorum Willem, V, 1900:34
Neanura muscorum Börner, C, 1901:133
Neanura muscorum Willem, V, 1902:280
Achorutes muscorum Bonet, F, 1931:376
Achorutes muscorum Wolf, B, 1938:112
Achorutes muscorum Leruth, R, 1939:197
Anura muscorum Lameere, A, 1940:8
Achorutes muscorum Marlier, GJ, 1942:8
Achorutes muscorum Leleup, N, 1948:10
Neanura muscorum Heungens, A et Van Daele, E, 1973:136
Neanura muscorum Verlinden, A, 1977:113
Neanura muscorum Berbiers, P et Mertens J, 1989:236
Neanura muscorum Berbiers, P et Segers, H, 1991:83
Neanura muscorum Jacobs, W, 1997:72
Neanura muscorum Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Neanura muscorum Dethier, M et Hubart, J-M, 2000:28 ^cf
Neanura muscorum Dethier, M et Hubart, J-M, 2000:28
Neanura muscorum Ponge, J-F, 2000:510
Neanura muscorum Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Neanura muscorum Lock, K, Janssens, F et Janssen, CR, 2003:27
Neanura muscorum Dethier, M et Hubart, J-M, 2003:52
Neanura muscorum Gillet, S et Ponge, J-F, 2004:223
Neanura muscorum Janssens, F et De Bruyn, L, 2005:ip
Neanura muscorum Janssens, F et Dethier, M, 2005:150

Genus Pseudachorutes Tullberg, 1871

Neanuridae Pseudachorutes Janssens, F et De Bruyn, L, 2005:ip ^sp

Species 32. boerneri Schött, 1902

Pratanurida boerneri Janssens, F et De Bruyn, L, 2005:ip

Species 33. parvulus Börner, C, 1901:701

Pseudachotures(sic) parvulus Verlinden, A, 1977:114

Species 34. subcrassus Tullberg, 1871

Pseudachorutes subcrassus Jacquemart, S, 1967:5

Genus Micranurida Börner, C, 1901:702

Species 35. forsslundi Gisin, 1949

Micranurida forsslundi Ponge, J-F, 2000:510

Species 36. papillosa Axelson, 1902

Micranurida papillosa Leleup, N, 1948:10

Species 37. pygmaea Börner, C, 1901:702

Anurida pygmaea Heungens, A et Van Daele, E, 1973:136
Micranurida pygmaea Ponge, J-F, 2000:510
Micranurida pygmaea Gillet, S et Ponge, J-F, 2004:223

Genus Anurida Laboulbène, 1865

Species 38. granaria (Nicolet, H, 1847:387) Tullberg, 1869

Anurida granaria Lameere, A, 1895:595
Aphoromma granaria Willem, V, 1902:279
Anurida granaria Bonet, F, 1931:376
Anurida granaria Wolf, B, 1938:112
Anurida granaria Leruth, R, 1939:197
Anurida granaria Marlier, GJ, 1942:8
Anurida granaria Delhez, F et Kersmaekers, M, 1973:8
Anurida granaria Dethier, M, 1998:55
Anurida granaria Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Anurida granaria Janssens, F et Dethier, M, 2005:150

Species 39. granulata Agrell, 1943

Anurida granulata Ponge, J-F, 2000:510
Micranurida granulata Gillet, S et Ponge, J-F, 2004:223

Species 40. maritima (Guérin-Méneville, 1836 tt) Laboulbène, 1865

Anurida maritima Willem, V, 1900:31
Anurida maritima Börner, C, 1901:133
Anurida maritima Imms, AD in Herdman, WA, 1906:11
Anurida maritima Willem, 1925
Anurida maritima Marlier, GJ, 1942:8
Anurida maritima Stach, J, 1949:21
Anurida maritima Hansson, HG, 1998:11
Anurida maritima Janssens, F et Bonte, D, 2004:ip

Familia Brachystomellidae Stach, 1949

Genus Brachystomella Ågren, 1903

Species 41. parvula (Schäffer, 1896:176) Stach, 1926

Brachystomella parvula Marlier, GJ, 1942:8
Brachystomella parvula Heungens, A et Van Daele, E, 1973:140
Brachystomella parvula Grégoire-Wibo, C, 1981:632

Familia Onychiuridae Lubbock J in Börner C, 1913:319

Genus Kalaphorura Absolon, K, 1901:388

Species 42. burmeisteri (Lubbock, J, 1873:190) Handschin, 1920

Kalaphorura Burmeisteri(sic) Leruth, R, 1939:198
Kalaphorura burmeisteri Marlier, GJ, 1942:8
Kalaphorura burmeisteri Dethier, M et Hubart, J-M, 2000:28
Kalaphorura burmeisteri Janssens, F et Dethier, M, 2005:154

Species 43. paradoxa (Schäffer, 1900) Ellis, WN & Bellinger, PF, 1973:26

Onychiurus paradoxus Godissart, J et Delhez, F, 1969:17
Onychiurus paradoxus Dethier, M, 1998:56
Onychiurus paradoxus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus paradoxus Hubart, J-M et Dethier, M, 1999:168

Genus Hymenaphorura Bagnall, 1948

Species 44. sibirica (Tullberg, T, 1877:40) Bagnall, 1948:637

Hymenaphorura sibirica Ponge, J-F, 2000:510

Genus Oligaphorura Bagnall, 1949

Species 45. absoloni (Börner, 1901) Dallai & Malatesta in Calderini, 1995:7

Archaphorura absoloni Ponge, J-F, 2000:510

Genus Supraphorura Stach, J, 1954:26

Species 46. furcifera (Börner, C, 1901:3) Pomorski, 1996:27

Onychiurus subfurcifer Jacquemart, S, 1967:1
Onychiurus subfurcifer Fjellberg, A, 1998:106
Protaphorura furcifer(sic) Thibaud, J-M et Massoud, Z, 1977:6 ^iss
Onychiurus furcifer(sic) Dethier, M, 1998:56 ^iss
Onychiurus furcifer(sic) Delhez, F(), Dethier, M et Hubart, J-M, 1999:34 ^iss
Kalaphorura furcifera Ponge, J-F, 2000:510
Supraphorura furcifera Janssens, F et Dethier, M, 2005:154

Genus Protaphorura Absolon, K, 1901:387

Species 47. armata (Tullberg, 1869) Börner, 1909

Lipura armata Lameere, A, 1895:595
Lipura armata Willem, V, 1900:24
Aphorura armata f. principalis Börner, C, 1901:132
Onychiurus armatus Marlier, GJ, 1942:8
Onychiurus armatus Leleup, N, 1948:10
Onychiurus armatus Heungens, A et Van Daele, E, 1973:140
Onychiurus armatus Verlinden, A, 1977:113
Protaphorura armata Jacobs, W, 1997:73
Protaphorura armata Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Protaphorura armata Janssens, F et Dethier, M, 2005:154

Species 48. campata (Gisin, 1952) Salmon, JT, 1964:170

Onychiurus campatus Heungens, A et Van Daele, E, 1973:136

Species 49. eichhorni (Gisin, 1954) Salmon, JT, 1964:171

Onychiurus eichhorni Heungens, A et Van Daele, E, 1973:136
Protaphorura eichhorni Ponge, J-F, 2000:510
Protaphorura eichhorni Gillet, S et Ponge, J-F, 2004:223

Species 50. octopunctata (Tullberg, T, 1877:40) Kloet, GS & Hincks, WD, 1945:5

Onychiurus octopunctatus Marlier, GJ, 1942:8
Onychiurus octopunctatus Jacquemart, S, 1967:2
Onychiurus octopuntata(sic) Jacquemart, S, 1967:4 ^iss

Species 51. orthacantha (Handschin, 1920,sd) Salmon, JT, 1964:173

Onychiurus orthacanthus Leleup, N, 1948:10

Species 52. quadriocellata (Gisin, 1947) Salmon, JT, 1964:170

Onychiurus quadriocellatus Heungens, A et Van Daele, E, 1973:136

Species 53. subuliginata (Gisin, H, 1956:339,sd) Salmon, JT, 1964:175

Protaphorura subuliginata Jacobs, W, 1997:73
Protaphorura subuliginata Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Genus Thalassaphorura Bagnall, 1949

Species 54. encarpata (Denis, 1931) Fjellberg, A, 1998:111

Onychiurus hortensis Heungens, A, 1970:721
Onychiurus hortensis Heungens, A et Van Daele, E, 1973:136

Genus Onychiurus Gervais, 1841

Onychiuridae Onychiurus Berbiers, P et Segers, H, 1991:83 ^sp
Onychiuridae Onychiurus Jacobs, W, 1997:72 ^sp
Onychiuridae Onychiurus Janssens, F et Dethier, M, 2005:153 ^sp

Species 55. sp.2 Leruth, R, 1939:198

Onychiurus sp.2 Leruth, R, 1939:198

Species 56. ambulans (Linnæus, C, 1758:609) Gervais, 1841

Lipura ambulans Lameere, A, 1895:595
Aphorura ambulans Willem, V, 1902:277
Onychiurus ambulans Bonet, F, 1931:378
Onychiurus fimetarius Bonet, F, 1931:272 ^e
Onychiurus ambulans Wolf, B, 1938:115
Onychiurus ambulans Wolf, B, 1938:773
Onychiurus ambulans Leruth, R, 1939:198
Onychiurus ambulans Marlier, GJ, 1942:8
Aphorura ambulans Geyskens, J, 1945:15
Onychiurus ambulans Thibaud, J-M et Massoud, Z, 1977:6 ^cf
Onychiurus ambulans Gilson, R et Hubart, J-M, 1980:61 ^gr
Onychiurus ambulans Hubart, J-M, 1982:343 ^gr
Onychiurus ambulans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus ambulans Janssens, F et Dethier, M, 2005:153
Onychiurus ambulans Dethier, M et Hubart, J-M, 2010:27

Species 57. arans Gisin, 1952

Onychiurus arans Dethier, M et Hubart, J-M, 2000:28 ^cf
Onychiurus arans Dethier, M et Hubart, J-M, 2003:52 ^cf
Onychiurus arans Janssens, F et Dethier, M, 2005:153

Species 58. circulans Gisin, 1952

Onychiurus circulans Dethier, M, 1998:56
Onychiurus circulans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Species 59. dissimulans Gisin, 1952

Onychiurus dissimulans Dethier, M et Hubart, J-M, 2000:28
Onychiurus dissimulans Janssens, F et Dethier, M, 2005:153

Species 60. sublegans Gisin, 1960

Onychiurus sublegans Delhez, F et Houssa, M, 1969:206
Onychiurus sublegans Delhez, Gilson & Hubart, 1973
Onychiurus sublegans Dethier, M, 1998:56
Onychiurus sublegans Delhez, F(), Dethier, M et Hubart, J-M, 1999:34

Genus Deuteraphorura Absolon, K, 1901:387

Species 61. imperfecta (Denis, 1938) Dányi, L & Traser, G, 2008:21

Onychiurus imperfectus Gilson, R et Hubart, J-M, 1980:61

Species 62. inermis (Tullberg, 1869) Absolon, 1901

Lipura fimetaria Lameere, A, 1895:595
Onychiurus fimetarius Leruth, R, 1939:198
Onychiurus fimetarius Marlier, GJ, 1942:8
Onychiurus fimetarius Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Aphorura spelea(sic) Willem, V, 1902:278 ^iss
Onychiurus spelaeus Wolf, B, 1938:118
Lipura inermis Willem, V, 1900:6
Aphorura inermis Börner, C, 1901:132
Deuteraphorura inermis Janssens, F et Dethier, M, 2005:153

Genus Bionychiurus Pomorski, RJ, 1996:92

Species 63. normalis (Gisin, 1949) Pomorski, RJ, 1996:92

Onychiurus normalis Heungens, A et Van Daele, E, 1973:140

Genus Deharvengiurus Weiner, WM, 1996:183

Species 64. severini (Willem, V, 1902:277) Pomorski, RJ in Christiansen, K, 2001:il

Aphorura severini Willem, V, 1902:277
Onychiurus argus Marlier, GJ, 1942:8 ^un
Onychiurus (Onychiurus) argus Gisin, H, 1960:136 ^un
Onychiurus (Paronychiurus) argus Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:120 ^un
Paronychiurus argus Salmon, JT, 1964:185 ^un
Onychiurus argus Thibaud, J-M et Massoud, Z, 1977:6 ^un
Onychiurus argus Hubart, J-M, 1982:343 ^un
Deharvengiurus argus Dethier, M et Hubart, J-M, 2000:28 ^un
Onychiurus severini Bonet, F, 1931:378
Onychiurus Severini(sic) Denis, J-R, 1932:365 ^iss
Onychiurus severini Wolf, B, 1938:116
Onychiurus Severini(sic) Leruth, R, 1939:198 ^iss
Onychiurus severini Marlier, GJ, 1942:8
Onychiurus severini Stach, J, 1954:61
Onychiurus severini Stach, J, 1954:76
Onychiurus severini Stach, J, 1954:153
Onychiurus severini Stach, J, 1954:163
Onychiurus (Onychiurus) severini Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:110
Protaphorura severini Salmon, JT, 1964:175
Onychiurus severini Godissart, J et Delhez, F, 1969:38
Onychiurus severini Delhez, Gilson & Hubart, 1973
Onychiurus severini Deharveng, L et Gouze, A, 1984:390
Onychiurus severini Dethier, M, 1998:56
Onychiurus severini Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Onychiurus severini Hubart, J-M et Dethier, M, 1999:167
Protaphorura severini Dethier, M et Hubart, J-M, 2000:28 ^cf
Deharvengiurus severini Janssens, F et Dethier, M, 2005:153
Deharvengiurus severini Dethier, M et Hubart, J-M, 2010:25

Genus Onychiuroides Bagnall, 1948

Species 65. pseudogranulosus (Delamare Deboutteville & Gisin, 1951) Weiner, WM, 1996:186

Onychiurus pseudogranulosus Thibaud, J-M et Massoud, Z, 1977:6

Familia Tullbergiidae Bagnall RS, 1935:238

Genus Stenaphorura Absolon, K, 1900:411

Species 66. denisi Bagnall, 1935

Tullbergia denisi Heungens, A et Van Daele, E, 1973:140

Species 67. quadrispina Börner, C, 1901:699

Stenaphorura quadrispina Jacobs, W, 1997:73
Stenaphorura quadrispina Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Genus Mesaphorura Börner, C, 1901:1

Species 68. betschi Rusek, 1979

Mesaphorura betschi Ponge, J-F, 2000:510

Species 69. hylophila Rusek, J, 1982:14

Mesaphorura hylophila Ponge, J-F, 2000:510

Species 70. isochaeta Arbea, JI et Jordana, R, 1989:8

Mesaphorura leitzaensis Ponge, J-F, 2000:510
Mesaphorura leitzaensis Gillet, S et Ponge, J-F, 2004:223

Species 71. italica (Rusek, J, 1971:203) Ellis, 1974:109

Mesaphorura italica Ponge, J-F, 2000:510

Species 72. jevanica Rusek, J, 1996:208

Mesaphorura jevanica Ponge, J-F, 2000:510
Mesaphorura jevanica Gillet, S et Ponge, J-F, 2004:223

Species 73. krausbaueri Börner, C, 1901:2

Tullbergia krausbaueri Jacquemart, S, 1967:5
Tullbergia krausbaueri Heungens, A et Van Daele, E, 1969:1106
Tullbergia krausbaueri Heungens, A, 1970:721
Tullbergia krausbaueri Heungens, A et Van Daele, E, 1973:136
Mesaphorura krausbaueri Janssens, F et Dethier, M, 2005:154

Species 74. macrochaeta Rusek, 1976

Mesaphorura macrochaeta Jacobs, W, 1997:72
Mesaphorura macrochaeta Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Mesaphorura macrochaeta Ponge, J-F, 2000:510
Mesaphorura macrochaeta Gillet, S et Ponge, J-F, 2004:223

Species 75. pongei Rusek, 1982

Mesaphorura pongei Ponge, J-F, 2000:510

Species 76. tenuisensillata Rusek, 1974

Mesaphorura tenuisensillata Ponge, J-F, 2000:510
Mesaphorura tenuisensillata Gillet, S et Ponge, J-F, 2004:223

Species 77. yosiii (Rusek, J, 1967:191) Rusek, 1973

Mesaphorura yosiii Dethier, M et Hubart, J-M, 2000:28 ^cf
Mesaphorura yosii(sic) Ponge, J-F, 2000:510 ^iss
Mesaphorura yosii(sic) Gillet, S et Ponge, J-F, 2004:223 ^iss
Mesaphorura yosii(sic) Janssens, F et Dethier, M, 2005:154 ^iss

Genus Neotullbergia Bagnall, RS, 1935:241

Species 78. crassicuspis (Gisin, 1943) Salmon, JT, 1964:148

Neotullbergia crassicuspis Jacobs, W, 1997:72
Neotullbergia crassicuspis Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Genus Paratullbergia Womersley, 1930

Species 79. callipygos (Börner, 1902) Salmon, JT, 1964:147

Tullbergia callipygos Heungens, A, 1970:721
Tullbergia callipygas(sic) Heungens, A et Van Daele, E, 1973:136 ^iss
Paratullbergia callipygos Jacobs, W, 1997:73
Paratullbergia callipygos Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Paratullbergia callipygos Ponge, J-F, 2000:510
Paratullbergia callipygos Gillet, S et Ponge, J-F, 2004:223
Paratullbergia callipygos Janssens, F et Dethier, M, 2005:154

Familia Odontellidae Massoud Z, 1967:61

Genus Superodontella Stach, 1949

Species 80. lamellifera (Axelson, WM, 1903:3) Arbea, JI & Jordana, R in Ramos, MA & al, 1997:244

Odontella lamellifera Lock, K, Janssens, F et Janssen, CR, 2003:27
Superodontella lamellifera Janssens, F et De Bruyn, L, 2005:ip

Genus Xenyllodes Axelson, WM, 1903:3

Species 81. armatus Axelson, WM, 1903:4

Xenyllodes armata(sic) Berbiers, P et Segers, H, 1991:81 ^iss
Xennylodes(sic) armata(sic) Jacobs, W, 1997:73 ^iss
Xenyllodes armata(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:59 ^iss
Xenyllodes armatus Ponge, J-F, 2000:510

Ordo Neelipleona Massoud Z, 1971:198

Familia Neelidae Folsom JW, 1896:391

Neelipleona Neelidae Janssens, F et Dethier, M, 2005:ip

Genus Neelus Folsom, JW, 1896:391

Species 82. murinus Folsom, JW, 1896:391

Neelus murinus Willem, V, 1902:282
Neelus murinus Bonet, F, 1931:390
Neelus murinus Wolf, B, 1938:135
Neelus murinus Leruth, R, 1939:201
Neelus murinus Marlier, GJ, 1942:10
Neelus murinus Delamare Deboutteville, C et Bassot, J-M, 1957:77
Neelus sp. Dethier, M et Hubart, J-M, 2000:29 ^sp
Neelus murinus Janssens, F et Dethier, M, 2005:159
Neelus murinus Dethier, M et Hubart, J-M, 2010:25

Genus Megalothorax Willem, V, 1900:7

Neelidae Megalothorax Schneider, C, Cruaud, C et D'Haese, C, 2011:388 ^sp

Species 83. minimus Willem, V, 1900:9

Megalothorax minimus Willem, V, 1900:9
Megalothorax minimus Willem, V, 1900:65
Megalothorax minimus Börner, C, 1901:137
Megalothorax minimus Marlier, GJ, 1942:10
Megalothorax minimus Barlet, J, 1951:142
Neelus minimus Heungens, A et Van Daele, E, 1973:136
Megalothorax minimus Blancquaert, J-P et Mertens, J, 1979:125
Megalothorax minimus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Neelus minimus Berbiers, P et Mertens J, 1989:237
Megalothorax minimus Jacobs, W, 1997:72
Neelus minimus Dethier, M, 1998:57
Megalothorax minimus Bretfeld, G, 1999:21
Neelus minimus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Megalothorax minimus Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Megalothorax sp. Dethier, M et Hubart, J-M, 2000:29
Megalothorax minimus Ponge, J-F, 2000:510
Megalothorax minimus Gillet, S et Ponge, J-F, 2004:223
Megalothorax minimus Janssens, F et Dethier, M, 2005:160
Megalothorax minimus Schneider, C et D'Haese, CA, 2013:319

Species 84. willemi Schneider, C et D'Haese, CA, 2013:347

Megalothorax willemi Schneider, C et D'Haese, CA, 2013:320

Ordo Entomobryomorpha Börner, 1913:319, sensu Soto-Adames FN et al., 2008:501

Familia Oncopoduridae Carl J & Lebedinsky J, 1905:565

Genus Oncopodura Carl, J et Lebedinsky, J, 1905:564

Oncopoduridae Oncopodura Janssens, F et Dethier, M, 2005:154 ^sp

Species 85. crassicornis Shoebotham, 1911

Oncopodura crassicornis Heungens, A et Van Daele, E, 1973:140

Species 86. dethieri Janssens, F et De Bruyn, L, 2010:50

Oncopodura sp. nov. Leruth, R, 1939:201
Oncopodura sp. Marlier, GJ, 1942:10
Entomobryidae Oncopodura Delhez, F(), Dethier, M et Hubart, J-M, 1999:44
Oncopodura sp. Hubart, J-M et Dethier, M, 1999:168
Oncopoduridae Oncopodura Dethier, M et Hubart, J-M, 2000:29 ^sp
Oncopoduridae Oncopodura Dethier, M et Hubart, J-M, 2003:52 ^sp
Oncopodura sp. Janssens, F et Dethier, M, 2005:154 ^sp
Oncopodura reyersdorfensis Delhez, F(), Dethier, M et Hubart, J-M, 1999:35 ^e
Oncopodura reyersdorfensis Hubart, J-M et Dethier, M, 1999:168 ^e
Oncopodura reyersdorfensis Delhez, Dethier & Hubart, 2000 in prep ^e
Oncopodura dethieri Janssens, F et De Bruyn, L, 2010:50

Familia Tomoceridae Schäffer, 1896

Collembola Tomoceridae Janssens, F et Dethier, M, 2005:154 ^sp

Genus Tomocerus Nicolet, H, 1842:67

Tomoceridae Tomocerus Dethier, M et Hubart, J-M, 2000:29 ^sp
Tomoceridae Tomocerus Janssens, F et De Bruyn, L, 2005:ip ^sp
Tomocerus Tomocerus Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:3,cf ^cf

Species 87. minor (Lubbock, J, 1862:598) Schäffer, 1900

Tomocerus plumbeus Lameere, A, 1940:8
Tomocerus plumbeus Geyskens, J, 1945:16
Tomocerus tridentiferus Willem, V, 1900:6
Tomocerus tridentiferus Willem, V, 1902:281
Tomocerus minor Börner, C, 1901:134
Tomocerus minor Bonet, F, 1931:388
Tomocerus minor Wolf, B, 1938:131
Tomocerus (Tomocerus) minor Leruth, R, 1939:200
Tomocerus minor Marlier, GJ, 1942:10
Tomocerus minor Delhez, F et Kersmaekers, M, 1973:8
Tomocerus minor Delhez, Gilson & Hubart, 1973
Tomocerus minor Heungens, A et Van Daele, E, 1973:138
Tomocerus minor Thibaud, J-M et Massoud, Z, 1977:6
Tomocerus minor Verlinden, A, 1977:114
Tomocerus minor Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Tomocerus minor Berbiers, P et Mertens J, 1989:236
Tomocerus minor Janssens, F, 1996:53
Tomocerus minor Jacobs, W, 1997:73
Tomocerus minor Dethier, M, 1998:57
Tomocerus minor Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Tomocerus minor Hubart, J-M et Dethier, M, 1999:168
Tomocerus minor Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Tomocerus minor Dethier, M et Hubart, J-M, 2000:19
Tomocerus minor Dethier, M et Hubart, J-M, 2000:29
Tomocerus minor Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Tomocerus minor De Boey, B, 2002:31
Tomocerus minor Lock, K, Janssens, F et Janssen, CR, 2003:27
Tomocerus minor Janssens, F et De Bruyn, L, 2005:ip
Tomocerus minor Janssens, F et Dethier, M, 2005:154
Tomocerus minor Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 88. vulgaris (Tullberg, 1871) Brook, 1883

Tomocerus plumbeus Lameere, 1895:591, nec Lubbock, 1873 ^h
Tomocerus vulgaris Lameere, A, 1895:591
Tomocerus vulgaris Willem, V, 1900:6
Tomocerus vulgaris Börner, C, 1901:134
Tomocerus vulgaris Marlier, GJ, 1942:10
Tomocerus (Tomocerus) vulgaris Yosii, R, 1967:29
Tomocerus vulgaris Berbiers, P et Segers, H, 1991:83
Tomocerus vulgaris De Boey, B, 2002:31
Tomocerus vulgaris Lock, K, Janssens, F et Janssen, CR, 2003:27
Tomocerus vulgaris Janssens, F et Bonte, D, 2004:ip
Tomocerus vulgaris Janssens, F et De Bruyn, L, 2005:ip
Tomocerus vulgaris Janssens, F et Dethier, M, 2005:155
Tomocerus vulgaris Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Tomocerus vulgaris Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Genus Pogonognathellus Paclt, 1944

Tomocerinae Pogonognathellus Janssens, F et Dethier, M, 2005:155 ^sp

Species 89. flavescens (Tullberg, 1871) Stach, 1929

Tomocerus plumbeus Willem, V, 1900:44
Tomocerus niger Lameere, A, 1895:591
Tomocerus flavescens Lameere, A, 1895:591
Tomocerus (Pogonognathus) flavescens Leruth, R, 1939:200
Tomocerus flavescens Marlier, GJ, 1942:10
Pogonognathellus flavescens Yosii, R, 1967:10 p
Tomocerus flavescens Delhez, F et Kersmaekers, M, 1973:8
Tomocerus flavescens Berbiers, P et Segers, H, 1991:83
Pogonognathellus flavescens Janssens, F, 1996:53
Tomocerus flavescens Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Pogonognathellus flavescens Janssens, F in Bruers, J et Viskens, G, 2000:15
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29 ^cf
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29
Pogonognathellus flavescens Ponge, J-F, 2000:510
Pogonognathellus flavescens De Boey, B, 2002:32
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2003:52
Pogonognathellus flavescens Gillet, S et Ponge, J-F, 2004:223
Pogonognathellus flavescens Janssens, F et Bonte, D, 2004:ip
Pogonognathellus flavescens Janssens, F et De Bruyn, L, 2005:ip
Pogonognathellus flavescens Janssens, F et Dethier, M, 2005:155
Pogonognathellus flavescens Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Pogonognathellus flavescens Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Species 90. longicornis (Müller OF, 1776) Paclt, 1944

Tomocerus plumbeus Börner, C, 1901:134
Tomocerus longicornis Lameere, A, 1895:591
... longicornis Willem
Tomocerus longicornis Marlier, GJ, 1942:10
Tomocerus longicornis Berbiers, P et Mertens, J, 1989:280
Tomocerus longicornis Berbiers, P et Mertens J, 1989:236
Tomocerus longicornis Janssens, F, 1993:22
Pogonognathellus longicornis Janssens, F in Bruers, J et Viskens, G, 2000:15
Tomocerus longicornis Lock, K, Janssens, F et Janssen, CR, 2003:27
Pogonognathellus longicornis Janssens, F et Bonte, D, 2004:ip
Pogonognathellus longicornis Janssens, F et Dethier, M, 2005:155
Pogonognathellus longicornis Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Plutomurus Yosii, R, 1956:78

Species 91. unidentatus (Börner, 1901) Dobat, 1979

Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29 ^cf
Tomocerus (Tomocerus) unidentatus Leruth, R, 1939:200
Tomocerus unidentatus Marlier, GJ, 1942:10
Tomocerus unidentatus Dethier, M, 1998:57
Tomocerus unidentatus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Tomocerus unidentatus Hubart, J-M et Dethier, M, 1999:167
Plutomurus unidentatus Janssens, F et Dethier, M, 2005:156
Plutomurus unidentatus Stomp, N et Weiner, WM, 2005:230
Plutomurus unidentatus Janssens, F et Dethier, M, 2006:ip
Plutomurus unidentatus Dethier, M et Hubart, J-M, 2010:27

Familia Isotomidae Schäffer, 1896

Entomobryomorpha Isotomidae Janssens, F et Dethier, M, 2005:ip ^sp

Genus Proisotoma Börner, 1901

Species 92. minima (Absolon, K, 1901:33) Linnaniemi, 1907

Proisotoma minima Heungens, A et Van Daele, E, 1973:136
Proisotoma minima Ponge, J-F, 2000:510
Proisotoma minima Gillet, S et Ponge, J-F, 2004:223

Species 93. minuta (Tullberg, 1871) Linnaniemi, 1907

Isotoma stagnalis Willem, V, 1900:28
Isotoma stagnalis Willem, V, 1900:131
Isotoma stagnalis Willem, V, 1901:193
Isotoma stagnalis var? tenebricola Willem, V, 1900:29
Isotoma tenebricola Willem, V, 1900:29
Isotoma (Proisotoma) stagnalis var. tenebricola Börner, C, 1901:133
Isotoma stagnalis var. tenebricola Willem, V, 1901:195
Isotoma (Proisotoma) stagnalis f. principalis Börner, C, 1901:133
Proisotoma minuta Marlier, GJ, 1942:9
Proistoma(sic) minuta Leleup, N, 1948:10
Proisotoma minuta Heungens, A et Van Daele, E, 1969:1106
Proisotoma minuta Heungens, A, 1970:721
Proisotoma minuta Heungens, A et Van Daele, E, 1973:136
Proisotoma minuta Verlinden, A, 1977:113
Proisotoma minuta Janssens, F et Witters, J, 2007:ip

Genus Folsomides Stach, 1922

Species 94. angularis (Axelson, WM, 1905:791) Gisin, 1942

Folsomides angularis Jacquemart, S, 1967:5

Genus Folsomina Denis, 1931

Species 95. onychiurina Denis, 1931

Folsomia(sic) onychiurina Heungens, A et Van Daele, E, 1973:140

Genus Hemisotoma Bagnall, RS, 1949:94

Species 96. thermophila (Axelson, 1900:9) Bagnall, RS, 1949:95

Cryptopygus thermophilus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Genus Proisotomodes Bagnall, 1949

Species 97. bipunctatus (Axelson, WM, 1903:9) Rusek, J, 2002:307

Isotomina bipunctata Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotomina bipunctata Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotomina bipunctata Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotomina bipunctata Berbiers, P et Mertens J, 1989:236
Isotomina bipunctata Berbiers, P et Segers, H, 1991:83

Genus Isotomodes Axelson, 1907

Species 98. productus (Axelson, WM, 1906:11) Linnaniemi, 1907

Isotomodes productus Heungens, A et Van Daele, E, 1973:140

Genus Pseudanurophorus Stach, 1922

Species 99. binoculatus (Kseneman, 1934) Christiansen, 1951:134

Pseudanurophorus binoculatus Ponge, J-F, 2000:510
Pseudanurophorus binoculatus Gillet, S et Ponge, J-F, 2004:223

Genus Anurophorus Nicolet, 1842

Species 100. laricis Nicolet, H, 1842:53

Anurophorus corticinus Lameere, A, 1895:595
Anurophorus Laricis(sic) Lameere, A, 1895:595 ^iss
Anurophorus laricis Willem, V, 1900:28
Anurophorus laricis f. principalis Börner, C, 1901:133
Anurophorus laricis Lameere, A, 1940:8
Anurophorus laricis Marlier, GJ, 1942:9
Anurophorus laricis André, HM, 1983:274

Genus Mucrosomia Bagnall, 1949

Species 101. garretti (Bagnall, 1939 tt) Bagnall, 1949

Mucrosomia garretti Janssens, F et Dethier, M, 2005:156

Genus Archisotoma Linnaniemi, WM, 1912:118

Species 102. pulchella (Moniez, 1890) Denis, 1923

Archisotoma pulchella Mertens, J, Beladjal, L, Janssens, F et Matthys, P, 2007:178

Genus Folsomia Willem, 1902

Isotomidae Folsomia Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20 ^sp
Isotomidae Folsomia Janssens, F et Dethier, M, 2005:156 ^sp
Isotomidae Folsomia Janssens, F et Dethier, M, 2006:ip ^sp
Pseudosinella decipiens Dethier, M et Hubart, J-M, 2000:29 ^cf

Species 103. candida Willem, V, 1902:280

Folsomia candida Willem, V, 1902:280
Folsomia fimetaria Bonet, F, 1931:380 ^e
Folsomia fimetaria Wolf, B, 1938:120 ^e
Folsomia fimetaria Leruth, R, 1939:199 ^e
Folsomia candida Delhez, F et Houssa, M, 1969:207
Folsomia candida Delhez, Gilson Hubart, 1973
Folsomia candida Thibaud, J-M et Massoud, Z, 1977:6
Folsomia candida Hubart, J-M, 1982:343
Folsomia candida Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Folsomia candida Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Folsomia candida Berbiers, P et Mertens J, 1989:236
Folsomia candida Janssens, F, 1994:6
Folsomia candida Dethier, M, 1998:56
Folsomia candida Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Folsomia candida Dethier, M et Hubart, J-M, 2000:19
Desoria(sic) candida Dethier, M et Hubart, J-M, 2000:28
Folsomia candida Dethier, M et Hubart, J-M, 2003:52
Folsomia candida Janssens, F et Dethier, M, 2005:156

Species 104. diplophthalma (Axelson, 1902) Axelson, 1907

Folsomia diplophtalma(sic) Jacobs, W, 1997:71 ^iss

Species 105. fimetaria (Linnæus, C, 1758:609) Willem, V, 1925:626

Isotoma (Proisotoma) fimetaria Börner, C, 1901:133
Folsomia fimetaria Marlier, GJ, 1942:9
Folsomia fimetaria Leleup, N, 1948:10
Folsomia fimetaria Heungens, A, 1970:721
Folsomia fimetaria Heungens, A et Van Daele, E, 1973:136

Species 106. fimetarioides (Axelson, WM, 1903:8) Handschin, 1929:59

Folsomia fimetarioides Heungens, A et Van Daele, E, 1973:136
Folsomia fimetarioides Jacobs, W, 1997:71
Folsomia fimetarioides Jacobs, W, De Bruyn, L et Lust, N, 1999:57
Folsomia fimetarioides Janssens, F et Dethier, M, 2005:156

Species 107. manolachei Bagnall, 1939

Folsomia nana Heungens, A et Van Daele, E, 1973:140
Folsomia manolachei Dethier, M et Hubart, J-M, 2000:28
Folsomia manolachei Ponge, J-F, 2000:510
Folsomia manolachei Janssens, F et Dethier, M, 2005:156

Species 108. penicula Bagnall, 1939

Folsomia multiseta Heungens, A, 1970:721
Folsomia multiseta Heungens, A et Van Daele, E, 1973:136

Species 109. quadrioculata (Tullberg, 1871) Linnaniemi, 1907

... quadrioculata Willem
Folsomia quadrioculata Leruth, R, 1939:199
Folsomia quadrioculata Marlier, GJ, 1942:9
Folsomia quadrioculata Jacquemart, S, 1967:4
Folsomia quadrioculata Heungens, A et Van Daele, E, 1973:136
Folsomia quadrioculata Grégoire-Wibo, C, 1974:207
Folsomia quadrioculata Grégoire-Wibo, C, 1980:43
Folsomia quadrioculata Berbiers, P et Segers, H, 1991:83
Folsomia quadrioculata Jacobs, W, 1997:71
Folsomia quadrioculata Jacobs, W, De Bruyn, L et Lust, N, 1999:58
Folsomia quadrioculata Dethier, M et Hubart, J-M, 2000:28
Folsomia quadrioculata Ponge, J-F, 2000:510
Folsomia quadrioculata Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Folsomia quadrioculata De Boey, B, 2002:28
Folsomia quadrioculata Gillet, S et Ponge, J-F, 2004:223
Folsomia quadrioculata Janssens, F et Dethier, M, 2005:157
Folsomia quadrioculata Dethier, M et Hubart, J-M, 2010:27

Species 110. sexoculata (Tullberg, 1871) Handschin, 1929:57

Isotoma sexoculata Willem, 1900
Folsomia sexoculata Marlier, GJ, 1942:9

Species 111. similis Bagnall, 1939:57

Folsomia similis Heungens, A et Van Daele, E, 1973:140

Species 112. spinosa Kseneman, 1936

Folsomia spinosa Jacobs, W, 1997:71
Folsomia spinosa Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Genus Isotomiella Bagnall, 1939

Isotomidae Isotoma Leleup, N, 1948:10 ^sp

Species 113. minor (Schäffer, 1896) Yosii, 1939

Isotoma (Isotoma) minor Leruth, R, 1939:199
Isotomiella minor Heungens, A, 1970:721
Isotomiella minor Heungens, A et Van Daele, E, 1973:136
Isotomiella minor Jacobs, W, 1997:72
Isotomiella minor Dethier, M, 1998:56
Isotomiella minor Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Isotomiella minor Ponge, J-F, 2000:510
Isotomiella minor Gillet, S et Ponge, J-F, 2004:223

Genus Isotomurus Börner, 1903

Isotomidae Isotomurus De Boey, B, 2002:56 ^sp
Isotominae Isotomurus Janssens, F et De Bruyn, L, 2005:ip ^sp
Isotominae Isotomurus Janssens, F et Dethier, M, 2005:157 ^sp

Species 114. antennalis (Bagnall, 1940) Gisin, 1944

Desoria antennalis Janssens, F et De Bruyn, L, 2005:ip

Species 115. graminis Fjellberg, A, 2007:96

Isotomurus prasinus Janssens, F et De Bruyn, L, 2005:ip
Isotomurus prasinus Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 116. maculatus (Schäffer, 1896) Ågren, 1903

Isotomurus palustris f. maculata(sic) Verlinden, A, 1977:114 ^iss

Species 117. palustris (Müller OF, 1776) Börner, 1903

Isotomurus sp. Dethier, M et Hubart, J-M, 2000:29
Isotoma palustris Lameere, A, 1895:594
Isotoma palustris Willem, V, 1901:195
Isotomurus palustris Marlier, GJ, 1942:9
Isotoma palustris Geyskens, J, 1945:16
Isotomurus palustris Heungens, A et Van Daele, E, 1973:136
Isotomurus palustis(sic) Heungens, A et Van Daele, E, 1973:141 ^iss
Isotomurus palustris Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotomurus palustris Grégoire-Wibo, C, 1983:41
Isotomurus palustris Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma(sic) palustris Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotomurus palustris Pollet, M et Desender, K, 1986:962
Isotomurus palustris Pollet, M et Desender, K, 1988:1120
Isotomurus palustris Berbiers, P et Mertens, J, 1989:280
Isotomurus palustris Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotomurus palustris Berbiers, P et Mertens J, 1989:236
Isotomurus palustris Berbiers, P et Segers, H, 1991:83
Isotomurus palustris Jacobs, W, 1997:72
Isotomurus palustris Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Isotomurus palustris De Boey, B, 2002:29
Isotomurus palustris Janssens, F et Bonte, D, 2004:ip
Isotomurus palustris Janssens, F et De Bruyn, L, 2005:ip
Isotomurus palustris Janssens, F et Dethier, M, 2005:157

Species 118. plumosus Bagnall, 1940

Isotomurus plumosus De Boey, B, 2002:29
Isotomurus plumosus Janssens, F et De Bruyn, L, 2005:ip
Isotomurus plumosus Janssens, F et Dethier, M, 2005:157

Species 119. pseudopalustris Carapelli, A, Frati, F, Fanciulli, PP et Dallai, R, 2001:127

Isotoma palustris var. aquatilis Börner, C, 1901:134

Species 120. unifasciatus (Börner, C, 1901:50) Stach, J, 1947:459

Isotomurus unifasciatus Janssens, F et De Bruyn, L, 2005:ip
Isotomurus unifasciatus Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Agrenia Börner, C, 1906:171

Species 121. bidenticulata (Tullberg, T, 1877:35) Denis, J-R, 1924:231

Agrenia bidenticulata Marlier, GJ, 1942:9
Agrenia bidenticulata Collart, 1943
Agrenia bidenticulata Leleup, N in Pasteels, M, 1948:16
Agrenia bidenticulata Collart, A in Cooreman, J, 1958:33
Agrenia bidenticulata Jacquemart, S, 1975:83

Genus Isotoma Bourlet, C, 1839:399

Isotomidae Isotoma Janssens, F et De Bruyn, L, 2005:ip ^sp

Species 122. caerulea Bourlet, C, 1839:401

Isotoma caerulea Fjellberg, A, 2003:94 2003
Isotoma caerulea Janssens, F et Bonte, D, 2004:ip
Isotoma caerulea Janssens, F et Witters, J, 2007:ip

Species 123. riparia (Nicolet, H, 1842:62) Nicolet, H, 1847:373

Isotoma viridis var. riparia Marlier, GJ, 1942:3
Isotoma riparia Janssens, F et Bonte, D, 2004:ip

Species 124. viridis Bourlet, C, 1839:401

Isotoma viridis f. genuina Börner, C, 1901:134
Isotoma viridis Lameere, A, 1895:594
Isotoma viridis Willem, V, 1900:39
Isotoma (Isotoma) viridis Leruth, R, 1939:199
Isotoma viridis Marlier, GJ, 1942:9
Isotoma viridis Geyskens, J, 1945:16
Isotoma viridis Jacquemart, S, 1967:5
Isotoma viridis Heungens, A et Van Daele, E, 1973:138
Isotoma viridis Verlinden, 1977:114
Isotoma viridis Grégoire-Wibo, C, 1981:632
Isotoma viridis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotoma viridis Grégoire-Wibo, C, 1983:41
Isotoma viridis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma viridis Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotoma viridis Pollet, M et Desender, K, 1986:962
Isotoma viridis Pollet, M et Desender, K, 1988:1120
Isotoma viridis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotoma viridis Berbiers, P et Mertens J, 1989:236
Isotoma viridis Berbiers, P et Segers, H, 1991:83
Isotoma viridis Janssens, F, 1991:66
Isotoma viridis Viskens, G et Bruers, J, 1994:29
Isotoma viridis Janssens, F, 1995:19
Isotoma viridis Jacobs, W, 1997:72
Isotoma viridis Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Isotoma viridis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Isotoma viridis De Boey, B, 2002:28
Isotoma viridis Lock, K, Janssens, F et Janssen, CR, 2003:27
Isotoma viridis Janssens, F et Bonte, D, 2004:ip
Isotoma viridis Janssens, F et De Bruyn, L, 2005:ip
Isotoma viridis Janssens, F et Dethier, M, 2005:157
Isotoma viridis Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Pseudisotoma Handschin, 1924

Species 125. monochaeta (Kos, 1942) Cassagnau, P, 1954:623

Pseudisotoma monochaeta Jacquemart, S, 1967:4

Species 126. sensibilis (Tullberg, T, 1877:36) Agrell, I, 1939:8

Pseudisotoma sensibilis Marlier, GJ, 1942:9
Pseudisotoma sensibilis Jacquemart, S, 1967:4
Isotoma (Pseudisotoma) sensibilis Heungens, A et Van Daele, E, 1973:136
Pseudisotoma sensibilis Janssens, F in Bruers, J et Viskens, G, 2000:15
Pseudisotoma sensibilis Ponge, J-F, 2000:510
Pseudisotoma sensibilis Gillet, S et Ponge, J-F, 2004:223
Pseudisotoma sensibilis Janssens, F et Dethier, M, 2005:157

Genus Desoria Agassiz M & Nicolet H in Desor E, 1841:384

Species 127. intermedia (Schött, 1902) Yosii, 1963

Isotoma intermedia Marlier, GJ, 1942:5

Species 128. olivacea (Tullberg, 1871) Yosii, R, 1977:156

Isotoma olivacea Marlier, GJ, 1942:9
Isotoma olivacea Heungens, A et Van Daele, E, 1973:138
Isotoma olivacea Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Isotoma olivacea Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Isotoma olivacea Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Isotoma olivacea Berbiers, P et Mertens, J, 1989:280
Isotoma olivacea Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Isotoma olivacea Berbiers, P et Mertens J, 1989:236
Isotoma olivacea Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 129. tigrina (Tullberg, 1871) Parona, 1875

Desoria tigrina Dethier, M et Hubart, J-M, 2000:28
Isotoma tigrina De Boey, B, 2002:28
Desoria tigrina Janssens, F et De Bruyn, L, 2005:ip
Isotoma tigrina Janssens, F et Dethier, M, 2005:157

Species 130. violacea (Tullberg, T, 1877:36)

Isotoma violacea Marlier, GJ, 1942:9
Isotoma violacea Heungens, A et Van Daele, E, 1973:138
Isotoma violacea Dethier, M et Hubart, J-M, 2000:29
Isotoma violacea Gillet, S et Ponge, J-F, 2004:223
Isotoma violacea Janssens, F et Dethier, M, 2005:157

Genus Parisotoma Bagnall, RS, 1940:171

Species 131. agrelli (Delamare Deboutteville, 1950)

Isotoma agrelli Jacobs, W, 1997:71
Isotoma agrelli Jacobs, W, De Bruyn, L et Lust, N, 1999:58

Species 132. notabilis (Schäffer, 1896) Bagnall, RS, 1940:171

Isotoma notabilis Marlier, GJ, 1942:9
Isotoma notabilis Jacquemart, S, 1967:5
Isotoma notabilis Heungens, A et Van Daele, E, 1969:1106
Isotoma notabilis Heungens, A, 1970:721
Isotoma notabilis Heungens, A et Van Daele, E, 1973:136
Isotoma notabilis Grégoire-Wibo, C, 1983:41
Parisotoma notabilis Dethier, M et Hubart, J-M, 2000:29
Parisotoma notabilis Ponge, J-F, 2000:510
Parisotoma notabilis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Parisotoma notabilis Gillet, S et Ponge, J-F, 2004:223
Parisotoma notabilis Janssens, F et De Bruyn, L, 2005:ip
Parisotoma notabilis Janssens, F et Dethier, M, 2005:157

Genus Vertagopus Börner, C, 1906:171

Species 133. arboreus (Linnæus, C, 1758:609) Ågren, 1903

Vertagopus arborea(sic) André, HM, 1983:274 ^iss
Vertagopus arboreus Potapov, M, 2001:528
Vertagopus arboreus Lock, K, Janssens, F et Janssen, CR, 2003:27

Species 134. cinereus (Nicolet, H, 1842:60) Marlier, GJ, 1942:9

Isotoma cinerea Lameere, A, 1895:594
Isotoma cinerea Willem, 1900
Vertagopus cinerea(sic) Marlier, GJ, 1942:9 ^iss

Genus Marisotoma Fjellberg, A, 1997:26

Species 135. tenuicornis (Axelson, WM, 1903:10) Bulimar & Hutu, 1984

Isotoma tenuicornis Marlier, GJ, 1942:4

Genus Paranurophorus Denis, J-R, 1929:167

Species 136. simplex Denis, J-R, 1929:167

Coloburella simplex Heungens, A et Van Daele, E, 1973:140

Familia Actaletidae Börner, 1902, sensu Soto-Adames FN et al., 2008:506

Genus Actaletes Giard, 1889

Species 137. neptuni Giard, 1889

Actaletes Neptuni(sic) Willem, V, 1900:6 ^iss
Actaletes neptuni Willem, V, 1900:50
Actaletes Neptuni(sic) Willem, V, 1901:474 ^iss
Actaletes neptuni Marlier, GJ, 1942:10
Actaletes neptuni Bellinger, PF, 1962:91

Familia Orchesellidae Börner C, 1906:162 sensu Zhang F et al, 2019:273

Genus Orchesella Templeton, R in Templeton, R et Westwood, JO, 1836:92

Entomobryidae Orchesella Janssens, F et De Bruyn, L, 2005:ip ^sp
Entomobryidae Orchesella Janssens, F et Dethier, M, 2005:159 ^sp

Species 138. flavescens (Bourlet, C, 1839:395) Nicolet, H, 1847:365

Orchesella rufescens Lameere, A, 1895:593
Orchesella rufescens Willem, V, 1900:49
Orchesella flavescens var. pallida Marlier, GJ, 1942:5
Orchesella rufescens f. principalis Börner, C, 1901:135
Orchesella flavescens Marlier, GJ, 1942:10
Orchesella flavescens Berbiers, P et Mertens J, 1989:236
Orchesella flavescens De Boey, B, 2002:30
Orchesella flavescens Janssens, F et Bonte, D, 2004:ip
Orchesella flavescens Janssens, F et De Bruyn, L, 2005:ip
Orchesella flavescens Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 139. spectabilis Tullberg, 1871:60

Orchesella spectabilis Janssens, F, 1993:21
Orchesella spectabilis Janssens, F et Bonte, D, 2004:ip

Species 140. bifasciata Nicolet, H, 1842:80

Orchesella bifasciata Berbiers, P et Mertens J, 1989:236
Orchesella bifasciata Berg, M, 2007:83

Species 141. cincta (Linnæus, C, 1758:609) Burmeister, 1835

Orchesella cincta f. principalis Börner, C, 1901:135
Orchesella cincta Lameere, A, 1895:593
Orchesella cincta Willem, V, 1900:49
Orchesella cincta Willem, V, 1901:654
Orchesella cincta Marlier, GJ, 1942:10
Orchesella cincta Geyskens, J, 1945:16
Orchesella cincta Thibaud, J-M et Massoud, Z, 1977:6
Orchesella cincta Mertens, J et Blancquaert, J-P, 1980:301
Orchesella cincta Mertens, Coessens & Blancquaert, 1982:9
Orchesella cincta André, HM, 1983:274
Orchesella cincta Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Orchesella cincta van Straalen, NM, 1986:71
Orchesella cincta Berbiers, P et Mertens, J, 1989:280
Orchesella cincta Berbiers, P et Mertens J, 1989:236
Orchesella cincta Berbiers, P et Segers, H, 1991:82
Orchesella cincta Janssens, F, 1993:21
Orchesella cincta Posthuma & van Straalen, 1993
Orchesella cincta Janssens, F, 1995:19
Orchesella cincta Janssens, F, 1995:140
Orchesella cincta Janssens, F, 1996:53
Orchesella cincta Janssens, F in Bruers, J et Viskens, G, 2000:15
Orchesella cincta Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Orchesella cincta De Boey, B, 2002:30
Orchesella cincta Lock, K, Janssens, F et Janssen, CR, 2003:27
Orchesella cincta Janssens, F et Bonte, D, 2004:ip
Orchesella cincta Janssens, F et De Bruyn, L, 2005:ip
Orchesella cincta Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Orchesella cincta Janssens, TKS, Mariën, J, Cenijn, P, Legler, P, van Straalen, NM et Roelofs, D, 2007:1

Species 142. quinquefasciata (Bourlet, 1842) Bourlet, 1843

Orchesella quinquefasciata Marlier, GJ, 1942:10
Orchesella quinquefasciata Berbiers, P et Mertens J, 1989:236

Species 143. villosa (von Linné, C, 1767:1014) Gervais in Walckenaër, 1844

Orchesella villosa Lameere, A, 1895:593
Orchesella villosa Willem, V, 1900:49
Orchesella villosa Börner, C, 1901:135
Orchesella villosa Willem, V, 1901:654
Orchesella villosa Lameere, A, 1940:8
Orchesella villosa Marlier, GJ, 1942:10
Orchesella villosa Verlinden, A, 1977:114
Orchesella villosa Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Orchesella villosa Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Orchesella villosa Berbiers, P et Mertens J, 1989:236
Orchesella villosa Berbiers, P et Segers, H, 1991:83
Orchesella villosa Janssens, F, 1993:21
Orchesella villosa Viskens, G et Bruers, J, 1994:29
Orchesella villosa Janssens, F, 1996:53
Orchesella villosa Dethier, M et Hubart, J-M, 2000:29
Orchesella villosa Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Orchesella villosa De Boey, B, 2002:30
Orchesella villosa Lock, K, Janssens, F et Janssen, CR, 2003:27
Orchesella villosa Janssens, F et Bonte, D, 2004:ip
Orchesella villosa Janssens, F et De Bruyn, L, 2005:ip
Orchesella villosa Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Orchesella villosa Janssens, TKS, Mariën, J, Cenijn, P, Legler, P, van Straalen, NM et Roelofs, D, 2007:2

Genus Heteromurus Wankel, 1860 nc , 1976

Species 144. major (Moniez, 1889) Absolon, 1901

Heteromurus major Janssens, F et Witters, J, 2007:ip
Heteromurus major Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Species 145. nitidus (Templeton, R in Templeton, R & Westwood, JO, 1836:94) Megnin, 1888

Templetonia crystallina Lameere, A, 1895:593
Heteromurus margaritarius? Willem, V, 1902:281
Heteromurus margaritarius Wolf, B, 1938:180
Heteromurus (Heteromurus) nitidus Bonet, F, 1931:387
Heteromurus (Heteromurus) nitidus Leruth, R, 1939:200
Heteromurus nitidus Marlier, GJ, 1942:10
Heteromurus nitidus Leleup, N, 1948:10
Heteromurus nitidus Delhez, F et Houssa, M, 1969:207
Heteromurus nididus(sic) Godissart, J et Delhez, F, 1969:38 ^iss
Heteromurus nitidus Delhez, F et Kersmaekers, M, 1973:8
Heteromurus nitidus Delhez, Gilson & Hubart, 1973
Heteromurus nitidus Heungens, A et Van Daele, E, 1973:140
Heteromurus nitidus Thibaud, J-M et Massoud, Z, 1977:6
Heteromerus(sic) nitidus Hubart, J-M, 1982:343
Heteromurus nitidus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Heteromurus nitidus Berbiers, P et Mertens J, 1989:236
Heteromurus nitidus Jacobs, W, 1997:71
Heteromurus nitidus Dethier, M, 1998:56
Heteromurus nitidus Delhez, F(), Dethier, M et Hubart, J-M, 1999:34
Heteromurus nitidus Jacobs, W, De Bruyn, L et Lust, N, 1999:58
Heteromurus nitidus Dethier, M et Hubart, J-M, 2000:29
Heteromurus nitidus De Boey, B, 2002:31
Heteromurus nitidus Janssens, F et De Bruyn, L, 2005:ip
Heteromurus nitidus Janssens, F et Dethier, M, 2005:159
Heteromurus nitidus Janssens, F et Dethier, M, 2006:ip
Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29 ^cf
Heteromurus nitidus Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Familia Paronellidae Börner, 1906:162 sensu Zhang F & al, 2019:275

Genus Cyphoderus Nicolet H, [1842:63]

Species 146. albinus Nicolet, H, 1842:67

Cyphoderus albinos(sic) Lameere, A, 1895:591 ^ios
Cyphoderus albinus Willem, V, 1900:49
Cyphoderus albinos(sic) Börner, C, 1901:136 ^ios
Cyphoderus albinus Lameere, A, 1940:8
Cyphoderus albinus Marlier, GJ, 1942:10
Cyphoderus albinos(sic) Geyskens, J, 1945:16 ^ios
Cyphoderus albinus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Cyphoderus albinus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Cyphoderus albinus Berbiers, P et Mertens J, 1989:236
Cyphoderus albinus Janssens, F, 1995:80
Cyphoderus albinus Janssens, F et Dethier, M, 2005:158
Cyphoderus albinus Dekoninck, W, Lock, K et Janssens, F, 2007:159
Cyphoderus albinus Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Familia Entomobryidae Schäffer, 1896 sensu Zhang F et al, 2019:252

Collembola Entomobryidae Hubart, J-M, Xhaard, P, Dumoulin, P, Monsieur, R, Roosens, J-P et Dethier, M, 2003:105 ^sp

Genus Willowsia Shoebotham, 1917

Species 147. buski (Lubbock, J, 1870:280) Schoebotham, 1917

Seira Buski(sic) Lameere, A, 1895:593 ^iss
Sira(sic) buski Marlier, GJ, 1942:9
Willowsia buski Heungens, A, 1970:721
Willowsia buski Heungens, A et Van Daele, E, 1973:136
Willowsia buski Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20

Species 148. nigromaculata (Lubbock, J, 1873:146) Shoebotham, 1917

Sira(sic) nigromaculata Börner, C, 1901:136

Species 149. platani (Nicolet, H, 1842:72) Palissa, 1959

Willowsia platani André, HM, 1983:274
Willowsia platani Berbiers, P et Mertens J, 1989:236
Willowsia platani Janssens, F et De Bruyn, L, 2005:ip

Genus Entomobrya Rondani, 1861

Entomobryidae Entomobrya Janssens, F et De Bruyn, L, 2005:ip ^sp
Entomobryidae Entomobrya Janssens, F et Dethier, M, 2005:158 ^sp

Species 150. albocincta (Templeton, R in Templeton, R & Westwood, JO, 1836:95) Rondani, 1861

Entomobrya albocincta Lameere, A, 1895:592
Entomobrya albocincta Marlier, GJ, 1942:9
Entomobrya albocincta Berbiers, P et Mertens, J, 1989:280
Entomobrya albocincta Berbiers, P et Mertens J, 1989:236
Entomobrya albocincta Berbiers, P et Segers, H, 1991:82
Entomobrya albocincta Lock, K, Janssens, F et Janssen, CR, 2003:27
Entomobrya albocincta Mertens, J, 2004:il

Species 151. arborea (Tullberg, 1871) Brook, G, 1884:271

Entomobrya arborea Lameere, A, 1895:592
Entomobrya arborea Marlier, GJ, 1942:9

Species 152. corticalis (Nicolet, H, 1842:72) Rondani, 1861

Entomobrya corticalis f. principalis Börner, C, 1901:135
Entomobrya corticalis Lameere, A, 1895:592
Entomobrya corticalis Willem, V, 1900:50
Entomobrya corticalis Marlier, GJ, 1942:9
Entomobrya corticalis Berbiers, P et Mertens, J, 1989:280
Entomobrya corticalis Berbiers, P et Mertens J, 1989:236
Entomobrya corticalis Berbiers, P et Segers, H, 1991:82
Entomobrya corticalis Janssens, F et De Bruyn, L, 2005:ip

Species 153. lanuginosa (Nicolet, H, 1842:74) Rondani, 1861

Entomobrya maritima Börner, C, 1901:135
Entomobrya lanuginosa Lameere, A, 1895:592
Entomobrya lanuginosa Börner, C, 1901:135
Entomobrya lanuginosa Heungens, A et Van Daele, E, 1973:136
Entomobrya lanuginosa Dethier, M et Hubart, J-M, 2000:29
Entomobrya lanuginosa Janssens, F et Dethier, M, 2005:158

Species 154. marginata (Tullberg, 1871) Brook, G, 1884:271

Entomobrya marginata Marlier, GJ, 1942:9
Entomobrya marginata Heungens, A et Van Daele, E, 1973:140

Species 155. multifasciata (Tullberg, 1871) Brook, G, 1884:275

Entomobrya decemfasciata Marlier, GJ, 1942:9
Entomobrya multifasciata Lameere, A, 1895:592
Entomobrya multifasciata Willem, V, 1900:50
Entomobrya nicoleti var. multifasciata Börner, C, 1901:135
Entomobrya multifasciata Marlier, GJ, 1942:9
Entomobrya multifasciata Heungens, A et Van Daele, E, 1973:138
Entomobrya multifasciata André, H, 1977:217
Entomobrya multifasciata Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Entomobrya multifasciata Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Entomobrya multifasciata Berbiers, P et Mertens, J, 1989:280
Entomobrya multifasciata Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Entomobrya multifasciata Berbiers, P et Mertens J, 1989:236
Entomobrya multifacciata Janssens, F, 1991:101
Entomobrya multifasciata Janssens, F, 1995:19
Entomobrya multifasciata Lock, K, Janssens, F et Janssen, CR, 2003:27
Entomobrya multifasciata Janssens, F et De Bruyn, L, 2005:ip
Entomobrya multifasciata Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 156. muscorum (Nicolet, H, 1842:75) Rondani, 1861

Entomobrya muscorum Lameere, A, 1895:592
Entomobrya muscorum Börner, C, 1901:135
Entomobrya muscorum Marlier, GJ, 1942:9
Entomobrya muscorum Berbiers, P et Mertens J, 1989:236
Entomobrya muscorum De Boey, B, 2002:29
Entomobrya muscorum Janssens, F et De Bruyn, L, 2005:ip
Entomobrya muscorum Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 157. nicoleti (Lubbock, J, 1870:299) Brook, G, 1884:271

Entomobrya nicoleti Berbiers, P et Mertens J, 1989:236
Entomobrya nicoleti Janssens, F, 1995:19

Species 158. nivalis (Linnæus, C, 1758:609) Rondani, 1861

Entomobrya nivalis Lameere, A, 1895:592
Entomobrya nivalis Marlier, GJ, 1942:9
Entomobrya nivalis Geyskens, J, 1945:16
Entomobrya nivalis Heungens, A et Van Daele, E, 1973:140
Entomobrya nivalis Verlinden, A, 1977:114
Entomobrya nivalis André, HM, 1983:274
Entomobrya nivalis Berbiers, P et Mertens J, 1989:234
Entomobrya nivalis Berbiers, P et Mertens, J, 1989:280
Entomobrya nivalis Berbiers, P et Mertens J, 1989:236
Entomobrya nivalis Janssens, F, 1991:101
Entomobrya nivalis Viskens, G et Bruers, J, 1994:29
Entomobrya nivalis Janssens, F, 1996:53
Entomobrya nivalis Ponge, J-F, 2000:510
Entomobrya nivalis Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Entomobrya nivalis De Boey, B, 2002:30
Entomobrya nivalis Lock, K, Janssens, F et Janssen, CR, 2003:27
Entomobrya nivalis Janssens, F et De Bruyn, L, 2005:ip

Species 159. quinquelineata Börner, C, 1901:69

Entomobrya quinquelineata Marlier, GJ, 1942:9

Species 160. schoetti Stach, 1922

Entomobryoides purpurascens Janssens, F et Dethier, M, 2005:158 ^e

Species 161. spectabilis Reuter, OM, 1892:25

Entomobrya spectabilis Berbiers, P et Mertens J, 1989:236

Genus Coecobrya Yosii, 1956

Species 162. tenebricosa (Folsom, JW, 1902:365) Gruia, 1998

Sinella caeca Marlier, GJ, 1942:9
Sinella tenebricosa Bellinger, PF, 2000:il
Sinella tenebricosa Janssens, F et Dethier, M, 2005:158

Genus Lepidocyrtus Bourlet, C, 1839:391

Entomobryidae Lepidocyrtus Grégoire-Wibo, C, 1981:631 ^sp
Entomobryidae Lepidocyrtus Dethier, M et Hubart, J-M, 2000:29 ^sp
Entomobryidae Lepidocyrtus Janssens, F et De Bruyn, L, 2005:ip ^sp

Species 163. lignorum (Fabricius O, 1775) Gervais, 1844

Lepidocyrtus lignorum Willem, V, 1900:48
Lepidocyrtus lignorum Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Lepidocyrtis(sic) lignorum Janssens, F in Bruers, J et Viskens, G, 2000:15
Lepidocyrtus lignorum Ponge, J-F, 2000:510
Lepidocyrtus lignorum Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20
Lepidocyrtus lignorum De Boey, B, 2002:31
Lepidocyrtus lignorum Lock, K, Janssens, F et Janssen, CR, 2003:27
Lepidocyrtus lignorum Gillet, S et Ponge, J-F, 2004:223
Lepidocyrtus lignorum Janssens, F et De Bruyn, L, 2005:ip
Lepidocyrtus lignorum Janssens, F et Dethier, M, 2005:158
Lepidocyrtus lignorum Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Lepidocyrtus lignorum Janssens, F et Dethier, M, 2006:ip
Entomobryidae Lepidocyrtus Dethier, M et Hubart, J-M, 2000:29 ^sp
Lepidocyrtus lignorum Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Species 164. violaceus (Geoffroy in Fourcroy, 1785) Lubbock, J, 1873:154

Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:190 ^iss
Lepidocyrtus cyanus(sic) van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:321 ^iss
Lepidocyrtus cyaneus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:22 ^e
Lepidocyrtus violaceus Lameere, A, 1895:591
Lepidocyrtus violaceus van den Bruel, WE in de Jonghe d'Ardoye, E, 1937:190
Lepidocyrtus violaceus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:20 ^cf
Lepidocyrtus violaceus Janssens, F et De Bruyn, L, 2005:ip

Species 165. curvicollis Bourlet, C, 1839:392

Lepidocyrtus Börneri(sic) Willem, V, 1902:281 ^ios
Lepidocyrtus sp. var. Börneri(sic) Willem, V, 1902:282 ^?
Lepidocyrtus börneri(sic) Bonet, F, 1931:383 ^iss
Lepidocyrtus börneri(sic) Wolf, B, 1938:123
Lepidocyrtus curvicollis Lameere, A, 1895:591
Lepidocyrtus curvicollis Leruth, R, 1939:199
Lepidocyrtus curvicollis Marlier, GJ, 1942:9
Lepidocyrtus (Lepidocyrtus) curvicollis Yosii, 1959
Lepidocyrtus curvicollis Delhez, F et Kersmaekers, M, 1973:8
Lepidocyrtus curvicolis(sic) Thibaud, J-M et Massoud, Z, 1977:6 ^iss
Lepidocyrtus curvicollis Berbiers, P et Segers, H, 1991:83
Lepidocyrtus curvicollis Jacobs, W, 1997:72
Lepidocyrtus curvicollis Dethier, M, 1998:56
Lepidocyrtus curvicollis Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Lepidocyrtus curvicollis Dethier, M et Hubart, J-M, 2000:29
Lepidocyrtus curvicollis Dethier, M et Hubart, J-M, 2003:52
Lepidocyrtus curvicollis Janssens, F et Dethier, M, 2005:158
Lepidocyrtus (curvicollis-group) curvicollis Mateos, E et Petersen, H, 2012:61

Species 166. paradoxus Uzel, 1890

Lepidocyrtus paradoxus Heungens, A et Van Daele, E, 1973:140
Lepidocyrtus paradoxus Lock, K, Janssens, F et Janssen, CR, 2003:27
Lepidocyrtus (curvicollis-group) paradoxus Mateos, E et Petersen, H, 2012:61

Species 167. cyaneus Tullberg, 1871

Lepidocyrtus cyaneus Leruth, R, 1939:199
Lepidocyrtus cyaneus Marlier, GJ, 1942:9
Lepidocyrtus cyaneus Jacquemart, S, 1967:5
Lepidocyrtus cyaneus Heungens, A et Van Daele, E, 1973:136
Lepidocyrtus cyaneus Verlinden, 1977:114
Lepidocyrtus cyaneus Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Lepidocyrtus cyaneus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Lepidocyrtus cyaneus Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Lepidocyrtus cyaneus Berbiers, P et Mertens, J, 1989:280
Lepidocyrtus cyaneus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Lepidocyrtus cyaneus Berbiers, P et Mertens J, 1989:236
Lepidocyrtus cyaneus Dethier, M et Hubart, J-M, 2003:52
Lepidocyrtus cyaneus Lock, K, Janssens, F et Janssen, CR, 2003:27
Lepidocyrtus (Lanocyrtus) cyaneus Janssens, F et De Bruyn, L, 2005:ip
Lepidocyrtus cyaneus Janssens, F et Dethier, M, 2005:158
Lepidocyrtus cyaneus Vanthournout, B, Rousaki, A, Parmentier, T, Janssens, F, Mertens, J, Vandenabeele, P, D'Alba, L et Shawkey, M, 2021:4

Species 168. lanuginosus (Linnaeus in Gmelin, 1788) Tullberg, 1871

Lepidocyrtus lanuginosus Lameere, A, 1895:591
Lepidocyrtus lanuginosus Börner, C, 1901:136
Lepidocyrtus lanuginosus Leruth, R, 1939:200
Lepidocyrtus lanuginosus Marlier, GJ, 1942:9
Lepidocyrtus lanuginosus Jacquemart, S, 1967:5
Lepidocyrtus lanuginosus Heungens, A, 1970:721
Lepidocyrtus lanuginosus Heungens, A et Van Daele, E, 1973:136
Lepidocyrtus lanuginosus Verlinden, 1977:114
Lepidocyrtus lanuginosus Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Lepidocyrtus lanuginosus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Lepidocyrtus lanuginosus Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Lepidocyrtus lanuginosus Berbiers, P et Mertens J, 1989:236
Lepidocyrtus lanuginosus Viskens, G et Bruers, J, 1994:29
Lepidocyrtus lanuginosus Jacobs, W, 1997:72
Lepidocyrtus lanuginosus Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Lepidocyrtus lanuginosus Ponge, J-F, 2000:510
Lepidocyrtus lanuginosus Gillet, S et Ponge, J-F, 2004:223
Lepidocyrtus lanuginosus Janssens, F et De Bruyn, L, 2005:ip

Species 169. pallidus Reuter, OM, 1892:24

Lepidocyrtus cyaneus Heungens, A, 1967:243 ^e
Lepidocyrtus pallidus Heungens, A, 1967:243
Lepidocyrtus pallidus Heungens, A et Van Daele, E, 1973:140

Genus Pseudosinella Schäffer, 1897

Entomobryidae Pseudopsinella(sic) Leleup, N, 1948:10 ^sp
Entomobryidae Pseudosinella Dethier, M et Hubart, J-M, 2000:29 ^sp

Species 170. alba (Packard, AS, 1873:37) Absolon, K, 1901:86

Pseudosinella alba Marlier, GJ, 1942:10
Pseudopsinella(sic) alba Leleup, N, 1948:10
Pseudosinella alba Jacobs, W, 1997:73
Pseudosinella alba Jacobs, W, De Bruyn, L et Lust, N, 1999:59
Pseudosinella alba Ponge, J-F, 2000:510
Pseudosinella alba Gillet, S et Ponge, J-F, 2004:223
Pseudosinella alba Janssens, F et De Bruyn, L, 2005:ip
Pseudosinella alba Janssens, F et Dethier, M, 2005:158

Species 171. decipiens Denis, J-R, 1924:198

Pseudosinella decipiens Janssens, F et Dethier, M, 2005:158

Species 172. duodecimocellata Handschin, 1928

Pseudosinella duodecimoculata(sic) Marlier, GJ, 1942:9 ^iss

Species 173. insubrica Gisin, H et da Gama, MM, 1969:292

Pseudosinella monoculata Thibaud, J-M et Massoud, Z, 1977:7

Species 174. ksenemani Gisin, 1944

Pseudosinella ksenemani Jacobs, W, 1997:73

Species 175. mauli Stomp, N, 1972:279

Pseudosinella mauli Ponge, J-F, 2000:510
Pseudosinella mauli Gillet, S et Ponge, J-F, 2004:223

Species 176. octopunctata Börner, C, 1901:705

Pseudosinella octopunctata Marlier, GJ, 1942:10
Pseudosinella octopunctata Jacobs, W, 1997:73

Species 177. petterseni Börner, C, 1901:707

Pseudosinella petterseni Heungens, A et Van Daele, E, 1973:140

Species 178. vandeli Denis, 1923

Pseudosinella vandeli Thibaud, J-M et Massoud, Z, 1977:6
Pseudosinella vandeli Hubart, J-M, 1982:343
Pseudosinella vandeli Dethier, M, 1998:38
Pseudosinella vandeli Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Pseudosinella vandeli Hubart, J-M et Dethier, M, 1999:167
Pseudosinella vandeli Dethier, M et Hubart, J-M, 2010:25

Genus Seira Lubbock, J, 1870:279

Species 179. domestica (Nicolet, H, 1842:76) Lubbock, J, 1870:279

Seira elongata Lameere, A, 1895:593
Sira(sic) elongata Willem, V, 1900:47
Sira(sic) domestica Börner, C, 1901:136
... domesticus Willem
Lepidocyrtinus domesticus Marlier, GJ, 1942:9
Lepidocyrtinus domesticus van den Bruel, WE in Collart, A, 1944:210
Sira(sic) domestica Geyskens, J, 1945:16
Lepidocyrtinus domesticus van den Bruel, WE, 1945:36
Seira domestica Heungens, A et Van Daele, E, 1969:1106
Seira domestica Heungens, A, 1970:721
Seira domestica Heungens, A et Van Daele, E, 1973:136
Seira domestica Berbiers, P et Mertens J, 1989:236
Seira domestica Dethier, M, 1998:57
Seira domestica Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Lepidocyrtinus domesticus Dethier, M et Hubart, J-M, 2003:52
Seira domestica Mertens, J, 2004:il
Seira domestica Janssens, F et Dethier, M, 2005:159

Ordo Symphypleona Börner C, 1901:5 sensu Massoud, 1971

Familia Sminthurididae Börner, 1906:163 sensu Betsch J-M & Massoud Z, 1970:199

Genus Sminthurides Börner, 1900

Species 180. aquaticus (Bourlet, 1842) Börner, C, 1901:96

Sminthurides aquaticus f. principalis Börner, C, 1901:137
Sminthurus aquaticus Lameere, A, 1895:589
Prosminthurus aquaticus Willem, V, 1900:6
Sminthurus aquaticus Willem, V, 1900:55
Prosminthurus aquaticus Lameere, A, 1940:9
Sminthurides aquaticus Marlier, GJ, 1942:10
Sminthurus aquaticus Geyskens, 1945:16
Sminthurides aquaticus Heungens, A et Van Daele, E, 1973:140
Sminthurus aquaticus Verlinden, A, 1977:114
Sminthurides aquaticus Blancquaert, J-P, 1981:2
Sminthurides aquaticus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sminthurides aquaticus Berbiers, P et Mertens J, 1989:237
Sminthurides aquaticus Viskens, G et Bruers, J, 1994:29
Sminthurides aquaticus Janssens, F, 1995:19

Species 181. penicillifer (Schäffer, 1896) Börner, C, 1901:94

Sminthurides malmgreni Janssens, F, 1992:99 ^e
Sminthurides penicillifer Janssens, F, 1995:20

Species 182. malmgreni (Tullberg, T, 1877:30) Axelson, 1905

Sminthurides malmgreni Marlier, GJ, 1942:10
Sminthurides malmgreni Heungens, A et Van Daele, E, 1973:140
Sminthurides malmgreni Blancquaert, J-P, 1981:1
Sminthurides malmgreni Berbiers, P et Mertens J, 1989:237
Sminthurides malmgreni Janssens, F et De Bruyn, L, 2005:ip

Species 183. parvulus (Krausbauer, 1898) Heymons, R & Heymons, H in Brauer, A, 1909:13

Sminthurides parvulus Blancquaert, J-P, 1981:1
Sminthurides parvulus Berbiers, P et Mertens J, 1989:237

Species 184. schoetti Axelson, WM, 1903:12

Smithurides(sic) schoetti Heungens, A et Van Daele, E, 1973:136
Sminthurides schoetti Blancquaert, J-P, 1981:1
Sminthurides schoetti Berbiers, P et Mertens J, 1989:237

Genus Sphaeridia Linnaniemi, WM, 1912:248

Species 185. pumilis (Krausbauer, 1898) Agrell, 1934

Sminthurides pumilis Heungens, A et Van Daele, E, 1973:138
Sphaeridia pumilis Blancquaert, J-P et Mertens, J, 1977:343
Sphaeridia pumilis Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sphaeridia pumilis Blancquaert, J-P, Mertens, J et Coessens, R, 1982:605
Sphaeridia pumilis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sphaeridia pumilis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sphaeridia pumilis Pollet, M et Desender, K, 1985:583
Sphaeridia pumilis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurides pumilis Berbiers, P et Mertens J, 1989:237
Sphaeridia pumilis Ponge, J-F, 2000:510
Sphaeridia pumilis Gillet, S et Ponge, J-F, 2004:223
Sphaeridia pumilis Janssens, F et De Bruyn, L, 2005:ip

Familia Katiannidae Börner, 1913:321 sensu Bretfeld G, 1999:13

Genus Sminthurinus Börner, C, 1901:99

Species 186. aureus (Lubbock, J, 1862:589) Börner, C, 1901:103

Sminthurinus aureus var. ochropus Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:355
Sminthurinus aureus f. alba(sic) Marlier, GJ, 1942:10 ^iss
Sminthurinus aureus Grégoire-Wibo, C, 1981:632
Sminthurinus aureus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Sminthurinus aureus Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurinus aureus Berbiers, P et Mertens J, 1989:237
Sminthurinus aureus Berbiers, P et Segers, H, 1991:83
Sminthurinus aureus Jacobs, W, 1997:73
Sminthurinus aureus Ponge, J-F, 2000:510
Sminthurinus aureus Janssens, F et De Bruyn, L, 2005:ip
Sminthurinus aureus Janssens, F et Dethier, M, 2005:160

Species 187. elegans (Fitch, A, 1863:674) Folsom, 1928

Sminthurinus elegans Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sminthurinus elegans Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sminthurinus elegans Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurinus elegans Berbiers, P et Mertens J, 1989:237
Sminthurinus elegans Janssens, F, 1996:54

Species 188. lawrencei Gisin, H, 1963:87

Sminthurinus lawrencei Gillet, S et Ponge, J-F, 2004:223
Sminthurinus lawrencei Gillet, S et Ponge, J-F, 2004:227

Species 189. niger (Lubbock, 1862) Börner, C, 1901:101

Sminthurus niger Lameere, A, 1895:589
Sminthurinus niger Marlier, GJ, 1942:10
Sminthurinus niger Heungens, A et Van Daele, E, 1973:140
Sminthurinus niger Janssens, F, 1993:22
Sminthurinus niger Ponge, J-F, 2000:510

Species 190. trinotatus Axelson, WM, 1905:794

Sminthurinus trinotatus Heungens, A et Van Daele, E, 1973:137

Familia Arrhopalitidae Stach, 1956, sensu Bretfeld G, 1999:13

Genus Arrhopalites Börner, C, 1906:182

Arrhopalitidae Arrhopalites Dethier, M et Hubart, J-M, 2000:30 ^sp
Arrhopalitidae Arrhopalites Janssens, F et Dethier, M, 2005:160 ^sp

Species 191. caecus (Tullberg, 1871:146 tt) Börner, C, 1906:182

Arrhopalites caecus Selga, D, 1963:457
Arrhopalites caecus Heungens, A et Van Daele, E, 1973:140
Arrhopalites caecus Janssens, F et Dethier, M, 2005:160

Species 192. microphthalmus Cassagnau, P et Delamare Deboutteville, C, 1953:133

Arrhopalites microphtalmus(sic) Jacobs, W, 1997:71 ^iss
Arrhopalites microphthalmus Jacobs, W, De Bruyn, L et Lust, N, 1999:57
Arrhopalites microphtalmus(sic) Jacobs, W, De Bruyn, L et Lust, N, 1999:58 ^iss

Genus Pygmarrhopalites Vargovitsh, RS, 2009:20

Species 193. bifidus (Stach, J, 1945:34) Vargovitsh, RS, 2009:20

Arrhopalites bifidus Selga, D, 1963:454
Arrhopalites bifidus Selga, D, 1963:456
Arrhopalites bifidus Selga, D, 1963:457
Arrhopalites bifidus Dethier, M, 1998:57
Arrhopalites bifidus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Arrhopalites bifidus Hubart, J-M et Dethier, M, 1999:168
Arrhopalites bifidus Delhez, Dethier & Hubart, 2000 in prep
Arrhopalites bifidus Janssens, F et Dethier, M, 2005:160

Species 194. principalis (Stach, J, 1945:37) Vargovitsh, RS, 2009:9

Arrhopalites principalis Berbiers, P et Mertens J, 1989:237

Species 195. pygmaeus (Wankel, 1860:256 tt) Vargovitsh, RS, 2009:8

Sminthurinus affinis Willem, V, 1902:283
Sminthurinus binoculatus Willem, V, 1902:283
Arrhopalites binoculatus Denis, J-R, 1937:3
Arrhopalites pygmaeus Bonet, F, 1931:391
Arrhopalites pygmaeus Wolf, B, 1938:136
Arrhopalites pygmaeus Leruth, R, 1939:201
Arrhopalites pygmaeus Marlier, GJ, 1942:10
Arrhopalites pygmaeus Selga, D, 1963:453
Arrhopalites pygmaeus Selga, D, 1963:454
Arrhopalites pygmaeeus(sic) Selga, D, 1963:454 ^iss
Arrhopalites pygmaeus Selga, D, 1963:455
Arrhopalites pygmaeu(sic) Selga, D, 1963:455 ^iss
Arrhopalites pygmaeus Selga, D, 1963:456
Arrhopalites pygmaeus Selga, D, 1963:457
Arrhopalites pygmaeus Godissart, J et Delhez, F, 1969:17
Arrhopalites pygmaeus Godissart, J et Delhez, F, 1969:38
Arrhopalites pygmaeus Delhez, F et Kersmaekers, M, 1973:8
Arrhopalites pygmaeus Delhez, Gilson & Hubart, 1973
Arrhopalites pygmaeus Thibaud, J-M et Massoud, Z, 1977:6
Arrhopalites pygmaeus Hubart, J-M, 1982:343
Arrhopalites pygmaeus Dethier, M, 1998:57
Arrhopalites pygmaeus Delhez, F(), Dethier, M et Hubart, J-M, 1999:35
Arrhopalites pygmaeus Hubart, J-M et Dethier, M, 1999:168
Arrhopalites pygmaeus Delhez, Dethier & Hubart, 2000 in prep
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:19
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:30 ^cf
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2000:30
Arrhopalites pygmaeus Janssens, F et Dethier, M, 2005:160
Arrhopalites pygmaeus Stomp, N et Weiner, WM, 2005:229
Arrhopalites pygmaeus Dethier, M et Hubart, J-M, 2010:25

Species 196. sericus (Gisin, 1947) Vargovitsh, RS, 2009:e18

Arrhopalites sericus Blancquaert, J-P, Coessens, R et Mertens, J, 1981:116
Arrhopalites sericus Berbiers, P et Mertens J, 1989:237
Arrhopalites sericus Janssens, F et Dethier, M, 2005:161

Species 197. spinosus (Rusek, 1967:23) Vargovitsh, RS, 2009:20

Arrhopalites spinosus Gillet, S et Ponge, J-F, 2004:223

Familia Sminthuridae Lubbock, 1862, sensu Deharveng, L, 2004:427

Genus Sphyrotheca Börner, C, 1906:183

Sminthuridae Sphyroteca(sic) Jacobs, W, 1997:73 ^sp

Genus Lipothrix Börner, C, 1906:183

Species 198. lubbocki (Tullberg, T, 1872:33) Cassagnau, 1961

Sminthurus lubbocki Jacquemart, S, 1967:4
Sphyrotheca lubbocki Berbiers, P et Mertens J, 1989:237
Lipothrix lubbocki Ponge, J-F, 2000:510
Lipothrix lubbocki Gillet, S et Ponge, J-F, 2004:223

Genus Allacma Börner, C, 1906:183

Species 199. fusca (Linnæus, C, 1758:608) Börner, C, 1906:183

Sminthurus fuscus Lameere, A, 1895:589
Sminthurus fuscus Willem, V, 1900:56
Sminthurus (Eusminthurus) fuscus f. principalis Börner, C, 1901:138
... fuscus Willem, 1901
Allacma fusca Marlier, GJ, 1942:11
Sminthurus fuscus Geyskens, J, 1945:16
Allacma fusca Berbiers, P et Mertens, J, 1989:280
Allacma fusca Berbiers, P et Mertens J, 1989:237
Allacma fusca Dall'Asta, 1990:135
Allacma fusca Lock, K, Janssens, F et Janssen, CR, 2003:27
Allacma fusca Janssens, F et Dethier, M, 2005:ip
Sminthurus multipunctatus Dethier, M et Hubart, J-M, 2000:30 ^cf
Allacma fusca Janssens, F et De Bruyn, L, 2005:ip
Allacma fusca Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Sminthurus Latreille in Sonnini, [1802]

Species 200. multipunctatus Schäffer, 1896

Sminthurus multipunctatus Marlier, GJ, 1942:11

Species 201. viridis (Linnæus, C, 1758:608) Latreille in Sonnini, [1802-03]

Sminthurus viridis Lameere, A, 1895:589
... viridis Willem
Sminthurus (Eusminthurus) viridis f. principalis Börner, C, 1901:138
Sminthurus viridis Lameere, A, 1940:9
Sminthurus viridis Marlier, GJ, 1942:11
Sminthurus viridis Heungens, A et Van Daele, E, 1973:136
Sminthurus viridis Verlinden, A, 1977:114
Sminthurus viridis Grégoire-Wibo, C, 1981:632
Sminthurus viridis Mertens, J, Desender, K, Coessens, R et Deleersnijder, R in Hulselmans, J, 1982:115
Sminthurus viridis Grégoire-Wibo, C, 1983:41
Sminthurus viridis Desender, K, Mertens, J, d'Hulster, M et Berbiers, P, 1984:354
Sminthurus viridis Berbiers, P, Maelfait, J-P et Mertens, J, 1989:309
Sminthurus viridis Berbiers, P et Mertens J, 1989:237
Sminthurus viridis Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Caprainea Dallai, 1970

Species 202. marginata (Schött, H, 1893:25) Dallai, 1972

Caprainea marginata Jacobs, W, 1997:71
Caprainea marginata Jacobs, W, De Bruyn, L et Lust, N, 1999:59

Familia Bourletiellidae Börner, 1913:322, sensu Bretfeld, 1994

Genus Deuterosminthurus Börner, 1901 nc ICZN , 1976

Species 203. bicinctus (Koch in Herrich-Schäffer, 1840) Linnaniemi, 1912

Deuterosminthurus flavus Janssens, F et De Bruyn, L, 2005:ip
Deuterosminthurus flavus Janssens, F, De Tré, E et Tolleneer, P, 2005:ip
Deuterosminthurus bicinctus Marlier, GJ, 1942:10
Bourletiella bicincta Heungens, A et Van Daele, E, 1973:138
Deuterosminthurus bicinctus Verlinden, 1977:114
Deuterosminthurus bicinctus Berbiers, P et Mertens J, 1989:237
Deuterosminthurus bicinctus Janssens, F et De Bruyn, L, 2005:ip

Species 204. pallipes (Bourlet, 1843) Handschin, 1924

Deuterosminthurus repandus Berbiers, P et Mertens J, 1989:237
Sminthurus pallipes Lameere, A, 1895:589
Bourletiella pallipes Heungens, A et Van Daele, E, 1973:138
Deuterosminthurus pallipes Berbiers, P et Mertens J, 1989:237

Species 205. sulphureus (Koch, 1840) Stach, 1956:159

Sminthurus luteus Lameere, A, 1895:589
Bourletiella lutea Marlier, GJ, 1942:10
Bourletiella luteus(sic) Janssens, F, 1995:19 ^iss
Bourletiella lutea Janssens, F, 1995:22
Bourletiella sulphurea Berbiers, P et Mertens J, 1989:237

Genus Bourletiella Banks, 1899 ICZN

Sminthuridae Bourletiella Marlier, GJ, 1942:5 ^sp

Species 206. hortensis (Fitch, A, 1863:668) Brown, 1918

Bourletiella hortensis Grégoire-Wibo, C, 1983:41
Bourletiella hortensis Crappe, D, Cannoodt, M et De Grisse, A, 1985:123
Bourletiella hortensis Berbiers, P et Mertens J, 1989:237
Bourletiella hortensis Viskens, G et Bruers, J, 1994:29

Genus Heterosminthurus Stach, 1955

Species 207. bilineatus (Bourlet, 1842) Stach, J, 1956:182

Sminthurus bilineatus Lameere, A, 1895:589
Heterosminthurus bilineatus Berbiers, P et Mertens J, 1989:237
Heterosminthurus bilineatus Janssens, F et Dethier, M, 2005:161

Species 208. claviger (Gisin, 1958) Bretfeld, G, 1971:150

Heterosminthurus claviger Berbiers, P et Mertens J, 1989:237

Species 209. insignis (Reuter, OM, 1876:83) Bretfeld, G, 1988:1

Deuterosminthurus insignis Marlier, GJ, 1942:10

Genus Fasciosminthurus Gisin, H, 1960:281

Species 210. quinquefasciatus (Krausbauer, 1898) Bretfeld, 1970

Bourtellia(sic) quinquefasciata Jacquemart, S, 1967:5
Bourletellia(sic) quinquefasciata Jacquemart, S, 1967:7

Familia Dicyrtomidae Börner C, 1906:163, sensu Deharveng, L, 2004:427

Genus Ptenothrix Börner, C, 1906:184

Species 211. atra (Linnæus, C, 1758:608) Börner, 1906

Ptenothrix atra Ball, A, 1926:261

Species 212. setosa (Krausbauer, 1898:24) Handschin, 1929:125

Ptenothrix setosa Leruth, R, 1939:201
Ptenothrix setosa Marlier, GJ, 1942:11

Genus Dicyrtoma Bourlet, 1842

Species 213. fusca (Lubbock, J, 1873:120) Axelson, 1903

Papirius fuscus Lameere, A, 1895:590
Papirius fuscus Willem, V, 1900:6
Papirius fuscus Marlier, GJ, 1942:11
Dicyrtoma fusca Berbiers, P et Mertens J, 1989:237
Dicyrtoma fusca Janssens, F et De Bruyn, L, 2005:ip
Dicyrtoma fusca Janssens, F et Dethier, M, 2005:161
Dicyrtoma fusca Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Genus Dicyrtomina Börner, 1903

Species 214. flavosignata (Tullberg, 1871) Linnaniemi, 1912

Papirius flavo-signatus(sic) Willem, V, 1900:63 ^iss
Papirius flavosignatus f. principalis Börner, C, 1901:139

Species 215. minuta (Fabricius O, 1783) Börner, 1906

Papirius minutus Lameere, A, 1895:590
Papirius minutus Willem, V, 1900:63
Papirius minutus f. principalis Börner, C, 1901:139
Dicyrtoma minuta Marlier, GJ, 1942:11
Dicyrtoma minuta Heungens, A et Van Daele, E, 1973:140
Dicyrtomina minuta Thibaud, J-M et Massoud, Z, 1977:6
Dicyrtoma minuta Berbiers, P et Mertens J, 1989:237
Dicyrtomina minuta Janssens, F, 1996:54
Dicyrtomina minuta Ponge, J-F, 2000:510
Dicyrtomina minuta Gillet, S et Ponge, J-F, 2004:223
Dicyrtomina minuta Janssens, F et De Bruyn, L, 2005:ip
Dicyrtomina minuta Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 216. ornata (Nicolet, H, 1842:83) Collinge & Shoebotham, 1910

Dicyrtomina ornata Thibaud, J-M et Massoud, Z, 1977:6
Dicyrtoma ornata Berbiers, P et Mertens J, 1989:237
Dicyrtoma ornata Berbiers, P et Segers, H, 1991:83
Dicyrtomina ornata Lock, K, Janssens, F et Janssen, CR, 2003:27
Dicyrtomina ornata Janssens, F et De Bruyn, L, 2005:ip
Dicyrtomina ornata Janssens, F et Dethier, M, 2005:161
Dicyrtomina ornata Janssens, F, De Tré, E et Tolleneer, P, 2005:ip

Species 217. saundersi (Lubbock, J, 1862:438) Stach, 1922

Dicyrtoma saundersi Berbiers, P et Mertens J, 1989:237

Discussion

Chen & Christiansen (1997) consider European records of Sinella caeca as records of the widespread Sinella tenebricosa.

Acknowledgements

We thank Wim Jacobs for his contribution based on his unpublished thesis. We thank Peter Bellinger for reviewing the species list. Especially for pointing out the problematic issue of the European records of Sinella caeca versus Sinella tenebricosa. We thank Jean-François Ponge for providing a species list from beech forests of the Belgian Ardennes. We thank Michel Dethier for the provisional catalogue (including addenda & corrigenda) of the Delhez Collection, for providing additional locality data and for providing additional papers on Belgian cave Collembola. We thank Koen Lock and Jos Bruers for their contribution in the form of an exhaustive bibliographic research.

Appendix

Janssens, F. 1996-2022. Collembola Belgica. http://www.collembola.be
Janssens, F. 2006-2022. Loci Collembola Belgica.
Janssens, F. 2008-2017. Atlas Collembola Belgica.