http://www.collembola.org/taxa/dicyinae.htm - Last updated on 2024.08.12 by Frans Janssens
Checklist of the Collembola: Dicyrtominae

Subfamilia Dicyrtominae Richards WR, 1968:26 sensu Bretfeld, G, 1999:128 key

Genus Number
of species
Calvatomina48
Dicyrtoma31
Dicyrtomina26
Gibberathrix1
Jordanathrix3
Pseudodicyrtomina3
Dicyrtominae112
 Genera with number of species
Dicyrtomina sp. from New Zealand
2014.02.25 © Murray, A.
Dicyrtoma fusca from Belgium
Abd.6 dorsally curved (*)
2016.02.19 © Huskens, M.-L.
Dicyrtominae from Taiwan
Abd.6 with 3 M-setae + 4 N setae
M-setae form a distinct angle
2023.02.07 © Cheng, H.-J.
Dicyrtomina ornata from France
More than 2 thick unpaired midfacial setae
2023.03.12 © Clerc, O.


Genus Dicyrtoma Bourlet, 1842

  Syn.: Sminthuridæ(sic) Papirius Lubbock, J, 1862:431
  Syn.: Poduridæ(sic) Symnthurus(sic) Fitch, A, 1863:674 ISS
  Syn.: ... Symnthurus(sic) Fitch in Van Benthuysen, 1867 ISS
  Syn.: Dicyrtomidae Ptenothrix Palacios-Vargas, JG et Cruz-Leal, JI, 2020:89
Ptenothrix sp. ♂ from Mexico
After Murray A in Palacios-Vargas JG & Cruz-Leal JI, 2020 Fig.6




  Syn.: ... Dicyrtomina(sic) Maes, J-M et Palacios-Vargas, JG, 1988:6 ISS
Remark by ICZN. 2007:246: Placed on the Official List of Names in Zoology. Opinion 1092.


  Syn.: Dicyrtomidae Dicyrtona(sic) Dutra, CC, Meotti, C et Fernandes, MG, 2009:550 ISS


Dicyrtoma sp. from the USA
2009.08.06 © Trager, J.C.
Dicyrtoma sp. from the Azores
2009.09.15 © E.D.E.N.
Dicyrtoma sp. ♀ from the Philippines
2014.06.09 © Cosalan, E.
Dicyrtoma fusca from the UK
Very short ant.4
Ant.3 apically annulated
2015.12.13 © Horton, M.
Dicyrtoma sp. from the Philippines
2020.05.28 © Engler, P.
Dicyrtoma fusca from France
Unguis (U) long narrow
Unguiculus (u) with inner tooth (*) + filament (**)
2020.11.22 © Reinach, Br.
Dicyrtoma sp. from Taiwan
2022.04.09 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.04.09 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Cephalic + thoracic spines absent
Eyepatch with yellow background
2022.04.09 © Cheng, H.-J.
Dicyrtoma sp. ♂ from China
2023.06.18 © Wang, G.
Dicyrtoma sp. from China
2023.06.18 © Wang, G.
Dicyrtoma sp. juv. from China
2023.06.18 © Wang, G.
Dicyrtoma sp. from China
Third foot
Tunica absent
2023.06.24 © Wang, G.
Dicyrtoma sp. from China
2023.06.29 © Wang, G.

Species sp.0 Delamare Deboutteville, C et Massoud, Z, 1963:281

Species sp.1 Greenslade, P in Houston, WWK, 1994:137

Dicyrtoma sp. from Australia
After Thurmer J in Greenslade P, 1994 p.137
Dicyrtoma sp. from New Zealand
Abd.6 pointing down (*)
2016.02.04 © Murray, A.

Species sp.2 Bellini, BC et Zeppelini, D, 2004:588

Species sp.3 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.4 Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1166

Species sp.5 Janssens F & Barbin T, 2016

Dicyrtoma sp. from Canada
2016.02.09 © Barbin, T.
Dicyrtoma sp. from Canada
2016.02.09 © Barbin, T.
Dicyrtoma sp. from Canada
Ant.1 with ventral spur (*)
2016.02.09 © Barbin, T.
Dicyrtoma sp. from Canada
Long slender unguis and unguiculus
2016.02.09 © Barbin, T.

Species sp.6 Janssens F & Duran DP, 2018

Dicyrtoma sp. from the USA
2018.12.19 © Duran, D.P.
Dicyrtoma sp. from the USA
2018.12.19 © Duran, D.P.
Dicyrtoma sp. from the USA
2018.12.19 © Duran, D.P.
Dicyrtoma sp. from the USA
2018.12.19 © Duran, D.P.
Dicyrtoma sp. juv.? from the USA
2018.12.19 © Duran, D.P.

Species sp.7 Janssens F & Jacobs I, 2019

Dicyrtoma sp. from Thailand
2019.06.25 © Jacobs, I.
Dicyrtoma sp. from Thailand
2019.06.26 © Jacobs, I.
Dicyrtoma sp. from Thailand
2019.07.08 © Jacobs, I.
Dicyrtoma sp. from Thailand
2019.12.01 © Jacobs, I.

Species sp.8 Janssens F & Cheng H-J, 2021

Dicyrtoma sp. from Taiwan
2021.11.08 © Cheng, H.-J.
Leg. : Tsai, Y.-J.
Dicyrtoma sp. from Taiwan
2021.11.08 © Cheng, H.-J.
Leg. : Tsai, Y.-J.
Dicyrtoma sp. from Taiwan
2021.11.09 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2021.11.09 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Ventral aspect
2021.11.09 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
Early instar
2021.11.12 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
2021.11.13 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2021.11.15 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
At least 4 unpaired midfacial setae
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
Abd.6 with N+N' setae spine-like
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
Diagnostic thoracic pattern
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
Abd.6 with N+N' setae spine-like
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. juv. from Taiwan
Unguis slender without tunica
2021.11.18 © Cheng, H.-J.
Dicyrtoma sp. ♀ from Taiwan
2021.11.19 © Cheng, H.-J.
Leg. : Tsai, Y.-J.
Dicyrtoma sp. ♀ from Taiwan
7 macrosetae at posterior margin of abd.6
a3-1 a0 a1-3
2021.11.19 © Cheng, H.-J. Leg. : Tsai, Y.-J.
Dicyrtoma sp. ♀ from Taiwan
2021.11.19 © Cheng, H.-J.
Leg. : Tsai, Y.-J.

Species sp.9 Janssens F & Cheng H-J, 2022

Dicyrtoma sp. from Taiwan
2022.08.10 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.08.10 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.08.10 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.08.10 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.08.31 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
2022.08.31 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Midfacial setae subequal in length
2022.08.31 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Abd.6 dorsal spines pointing backwards
2022.08.31 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Dorsal anterior macrosetae not spine-like
2022.08.31 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Midfacial setae pattern: 1+1+2+2
2022.09.01 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Midfacial setae subequal in length
Pattern: 1+1+2+2?+1?
2022.09.14 © Cheng, H.-J.
Dicyrtoma sp. from Taiwan
Midfacial setae subequal in length
Pattern: 1+1+2+2+1
2022.09.14 © Cheng, H.-J.

Species aurata (Mills, HB, 1934:107) Christiansen, K & Bellinger, P, 1981:1220

             Syn.: Dicyrtoma atra(sic) Vázquez, M et Palacios-Vargas, JG, 2004:100,ca ISS
Dicyrtoma aurata from the USA
2007.10.19 © McClarin, J.
Dicyrtoma aurata from the USA
2008.10.15 © Murray, T.
Dicyrtoma aurata from the USA
2010.12.23 © Eiseman, C.
Dicyrtoma aurata from the USA
2010.12.23 © Eiseman, C.
Dicyrtoma aurata from the USA
2014.12.07 © Montgomery, G.
Dicyrtoma aurata from the USA
Producing drop of saliva
2015.12.10 © Schrayer, G.
Dicyrtoma aurata from the USA
2016.11.23 © Schulz, K.
Dicyrtoma aurata from the USA
2016.11.23 © Schulz, K.
Dicyrtoma aurata from the USA
2023.01.04 © Hendrix, S.
Dicyrtoma aurata from the USA
2023.01.04 © Hendrix, S.
Dicyrtoma aurata from the USA
Great abdomen reticulate
2023.01.04 © Hendrix, S.
Dicyrtoma aurata from the USA
Cheek posteriorly spotted
2023.01.04 © Hendrix, S.
Dicyrtoma aurata from the USA
2023.11.03 © Tell, O.
Dicyrtoma aurata from the USA
Antenna
2023.11.03 © Tell, O.



Species balicrura Lin & Feng, 1985

Species chloropa (Tullberg, T, 1877:31) Börner, 1906

            Syn.: Dicyrtoma chloropus(sic) var. pallens Börner, C, 1909:129
            Syn.: Dicyrtoma chlorops(sic) var. pallens Yosii, R, 1977:166,si
Dicyrtoma pallens from Japan
After Itoh R & al, 2012 Fig.43a




             Syn.: Dicyrtoma chloropus(sic) Börner, C, 1909:129 ISS
             Syn.: Dicyrtoma chloropus(sic) Stach, J, 1957:65,si ISS
             Syn.: Dicyrtoma chloropus(sic) Salmon, JT, 1964:634 ISS
             Syn.: Dicyrtoma chlorops(sic) Yosii, R, 1977:166,si ISS

Species christinae Szeptycki, 1981

Species cirtana Lucas, H, 1846:255

            Syn.: Dicyrtoma Cirtanus(sic) Lucas, H, 1846:255 IOS
             Syn.: Dicyrtoma cirtanus(sic) Nicolet, H, 1847:362 ISS
Dycirtoma(sic) cirtana from Algeria
After Lucas H, 1849 Pl.2 Fig.6

             Syn.: Dicyrtoma Cirtanus(sic) Kolenati, FA, 1858:134 ISS
             Syn.: Dicyrtoma citrana(sic) Handschin, 1925 ISS




Species dorsosignata Stach, J, 1924:125

            Syn.: Dicyrtoma fusca var. dorsosignata Stach, J, 1924:125
Dicyrtoma fusca var. dorsosignata from Malta
Dark colour form
After Stach J, 1924 Taf.II Fig.1
Dicyrtoma fusca var. dorsosignata from Malta
Typical colour form
After Stach J, 1924 Taf.II Fig.2
Dicyrtoma fusca var. dorsosignata from Malta
Pale colour form
After Stach J, 1924 Taf.II Fig.3

            Syn.: Dicyrtoma fusca Stach, J, 1924:123 partim
            Syn.: Dicyrtoma fusca var. pallida Stach, J, 1924:125
Dicyrtoma fusca var. pallida from Malta
After Stach J, 1924 Taf.II Fig.4

            Syn.: Dicyrtoma fusca f. strigatus(sic) Stach, J, 1924:124 IOS
            Syn.: Dicyrtoma melitensis Stach, J, 1957:73
Dicyrtoma melitensis from Malta
After Stach J, 1957 Pl.V Fig.1
Dicyrtoma melitensis from Malta
After Stach J, 1967 Pl.XXXVI Fig.1
Dicyrtoma melitensis from Malta
After Stach J, 1967 Pl.XXXVI Fig.2
Dicyrtoma melitensis from Malta
Weaker colouration
After Stach J, 1967 Pl.XXXVI Fig.3




             Syn.: Papirius fuscus var. dorsosignata(sic) Stach, J, 1930:83 ISS
Dicyrtoma dorsosignata from France
2016.11.23 © Combes, R.
Dicyrtoma dorsosignata from France
2017.03.14 © Tiky, S.
Dicyrtoma dorsosignata from France
2017.03.14 © Tiky, S.
Dicyrtoma dorsosignata from France
2017.03.14 © Tiky, S.
Dicyrtoma dorsosignata juv. from France
2017.03.14 © Tiky, S.
Dicyrtoma dorsosignata from France
Eyepatch with reddish background
2017.10.29 © Tiky, S.
Dicyrtoma dorsosignata from Portugal
2018.02.25 © Almeida, J.
Dicyrtoma dorsosignata from Portugal
Head in profile
Bulby clypeus with clypeal suture (cs)
2018.02.25 © Almeida, J.
Dicyrtoma dorsosignata from Portugal
Distinct laterodorsal pigmentation
2018.02.25 © Almeida, J.
Dicyrtoma dorsosignata from Portugal
Unguis without tunica
2018.02.25 © Almeida, J.
Dicyrtoma dorsosignata from Portugal
Eversed collophore vesicles
2018.02.25 © Almeida, J.
Dicyrtoma dorsosignata from Portugal
2018.11.01 © Almeida, J.
Dicyrtoma dorsosignata from Malta
Midfacial unpaired macrosetae
2018.11.01 © Ujvári, Z.
Dicyrtoma dorsosignata from Malta
Clypeolabral suture (cls)
2018.11.01 © Ujvári, Z.
Dicyrtoma dorsosignata from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata ♂ from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata juv. from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata from Malta
2018.11.02 © Ujvári, Z.
Dicyrtoma dorsosignata from Spain
2019.12.19 © Zeeders, M.
Dicyrtoma dorsosignata from Spain
2019.12.19 © Zeeders, M.
Dicyrtoma dorsosignata from Spain
2019.12.19 © Zeeders, M.
Dicyrtoma dorsosignata from Spain
Dark specimen
2020.11.22 © Miralles, A.
Dicyrtoma dorsosignata from Holland
2020.11.27 © Christant, F.
Dicyrtoma dorsosignata from Holland
2020.11.29 © Christant, F.
Dicyrtoma dorsosignata from Holland
Dorsal pigmentation pattern
2020.12.06 © Berg, M.
Dicyrtoma dorsosignata from Holland
2020.12.06 © Berg, M.
Dicyrtoma dorsosignata from Holland
Anterior outer dental setae smooth
E4:E3=3
2020.12.06 © Berg, M.
Dicyrtoma dorsosignata from Holland
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
Tibiotarsus dark, "knee" pale
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
Unguis (U) long narrow
Unguiculus (u) with long apical filament (f)
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Holland
Posterior patch with cross stripes
2020.12.08 © van Duinen, J.
Dicyrtoma dorsosignata from Spain
2021.12.21 © Zeeders, M.
Dicyrtoma dorsosignata ♂ from Spain
2021.12.21 © Zeeders, M.
Dicyrtoma dorsosignata from Sardinia
2024.02.10 © Krásenský, P.
Dicyrtoma dorsosignata from Sardinia
2024.02.11 © Krásenský, P.





Species flammea Maynard, EA, 1951:281

Dicyrtoma flammea from the USA
After Maynard EA, 1951 PlateXXIII Fig.399

            Syn.: Calvatomina rossi Dukes, CD, Janssens, F, Recuero, E et Caterino, MS, 2022:11
Calvatomina rossi from the USA
After Dukes CD & al, 2020 Fig.4F
2023.12.22 Janssens F =? Dicyrtoma flammea juv.

            Syn.: Dicyrtoma flammea Janssens, 2023 = juv.?



Dicyrtoma flammea from Canada
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
More than 2 unpaired midfacial setae
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Apically weakly annulated ant.3
2015.08.08 © Murray, A.
Dicyrtoma flammea juv. from Canada
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Unpaired median facial setae
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Six medial 'unpaired' midfacial setae
Marked setal socket positions
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Dorsal abdominal pattern
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Darker colouration
2015.08.08 © Murray, A.
Dicyrtoma flammea from Canada
Distinctly shorter medial outer setae (*) on tibiotarsus
2015.08.13 © Murray, A.
Dicyrtoma flammea juv. from Alaska
2016.06.26 © Palhof, M.
Dicyrtoma flammea from Alaska
2016.07.23 © Palhof, M.
Dicyrtoma flammea from Alaska
2016.07.23 © Palhof, M.
Dicyrtoma flammea from the USA
2020.07.02 © Caterino, M.
Dicyrtoma flammea from Canada
2020.08.16 © Barbin, T.
Dicyrtoma flammea from Canada
2020.08.16 © Barbin, T.
Dicyrtoma flammea juv. from the USA
2022.01.01 © Charny, V.
Dicyrtoma flammea juv. from the USA
More then 2 midfacial unpaired macrosetae
2022.01.01 © Charny, V.
Dicyrtoma flammea juv. from the USA
2022.01.05 © Charny, V.
Dicyrtoma flammea juv. from the USA
Unguis long, narrow, slender
2022.01.05 © Charny, V.
Dicyrtoma flammea from the USA
2023.01.14 © Shahan, T.
Dicyrtoma flammea from the USA
2023.01.14 © Shahan, T.
Dicyrtoma flammea from the USA
More then 2 unpaired midfacial macrosetae
2023.01.14 © Shahan, T.
Dicyrtoma flammea from the USA
Eye with 8 ocelli
Ocelli C+D distinctly smaller
2023.01.14 © Shahan, T.

Species fusca (Lubbock, J, 1873:120) Axelson, 1903, t.t., B

Papirius fuscus from the UK
After Lubbock J, 1873 Pl.8 Fig.1
Papirius fuscus from the UK
After Lubbock J, 1873 Pl.8 Fig.2
Papirius fuscus from the UK
After Lubbock J, 1873 Pl.8 Fig.3
            Syn.: Podura atra Linnæus, C, 1746:342
             Syn.: Smynthurus(sic) Atra(sic) Fabricius O, 1783 ISS
            Syn.: Dicyrtoma atro-purpurea(sic) Bourlet, 1843
             Syn.: Dicyrtoma atro-purpurea(sic) Gervais in Walckenaër, 1844 ISS
             Syn.: Dicyrtoma atro-purpurea(sic) Nicolet, H, 1847:361 ISS
             Syn.: Dicyrtoma Atropurpurea(sic) Kolenati, FA, 1858:134 ISS
            Syn.: Papirius cursor Lubbock, J, 1862:436
             Syn.: Papirius cursore(sic) Reuter, OM, 1892:24 ISS
             Syn.: Papirius coursor(sic) Rusek, 1977 ISS
            Syn.: Papirius silvaticus Tullberg, 1871
             Syn.: Podura fusco nigra(sic) Lubbock, J, 1873:107 ISS
            Syn.: Papirius longicornis von Dalla Torre, 1895
            Syn.: Dicyrtoma fusca f. typica Ågren, H, 1904:26
            Syn.: Dicyrtoma albella Wahlgren, 1906
Dicyrtoma fusca f. albella? from Holland
2015.12.06 © van Duinen, J.

            Syn.: Dicyrtoma fusca f. fenestratus(sic) Stach, J, 1924:124
            Syn.: Papirius fuscus f. principalis Lubbock, 1873
Dicyrtoma fusca f. principalis from Tunis
After Stach J, 1924 Taf.II Fig.5

            Syn.: Dicyrtoma fusca var. maculifera Stach, J, 1957:70
Dicyrtoma fusca var. maculifera from Poland
After Stach, J, 1957 Pl.IV Fig.1

            Syn.: Dicyrtoma fusca f. signata Fabricius JC, 1793 sensu Janssens F, 2019
            Syn.: Podura (Globulosæ(sic)) fusco-nigra(sic) Geoffroy, M, 1762:607 r IOS
             Syn.: Podura (Globulosæ(sic)) fusco-nigra(sic) Geoffroy, M, 1764:607 r ISS
             Syn.: Podura (Globulosæ(sic)) fusco-nigra(sic) Geoffroy, M, 1799:607 ISS
             Syn.: Podura fusco-nigra(sic) Walckenær, CA, 1802:3 ISS
             Syn.: Podura fusco-nigra(sic) de Lamarck, M, 1818:20 ISS
             Syn.: Podura fusco nigra(sic) Lubbock, J, 1873:107 ISS
            Syn.: Podura signata Fabricius, JC, 1793:65
            Syn.: Podura signata Fabricius, JC, 1796:138
            Syn.: Podura signata Walckenær, CA, 1802:3
Dicyrtoma fusca var. from France
2006 © Ollivier, E.

Dicyrtoma fusca var.1 from Holland
2007.11.08 © Baas, A.H.
Dicyrtoma fusca var.1 from Holland
2007.11.24 © Baas, A.H.
Remark by Janssens, F. 2007.12.10: Colour variant with distinct middorsal dark line, and yellowish lateral pigmentation.

Dicyrtoma fusca var.1 from the UK
2007.12.10 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.19 © Campbell, A.
Dicyrtoma fusca var.1 from the UK
2007.12.22 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.24 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.24 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.24 © Barton, T.
Dicyrtoma fusca var.1 from the USA
2007.12.25 © Moorehead, C.
Dicyrtoma fusca var.1 from the UK
2007.12.26 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.26 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.26 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.29 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2007.12.29 © Kilford., B
Dicyrtoma fusca var.1 from the UK
2008.01.01 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2008.01.01 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2008.01.12 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2008.01.12 © Barton, T.
Dicyrtoma fusca var.1 from Holland
2008.01.20 © Baas, A.H.
Dicyrtoma fusca var.1 from Holland
2008.01.22 © Baas, A.H.
Dicyrtoma fusca var.1 from the UK
2008.02.07 © Campbell, A.
Dicyrtoma fusca var.1 from the USA
2008.02.11 © Moorehead, C.
Dicyrtoma fusca var.1 from the USA
2008.02.11 © Moorehead, C.
Dicyrtoma fusca var.1 from the UK
2008.02.12 © Campbell, A.
Dicyrtoma fusca var.1 from Belgium
2008.02.16 © Doornaert. A.
Dicyrtoma fusca var.1 from Belgium
2008.02.16 © Doornaert. A.
Dicyrtoma fusca var.1 from the UK
2008.02.17 © Campbell, A.
Dicyrtoma fusca var.1 from the UK
2008.02.17 © Campbell, A.
Dicyrtoma fusca var.1 from the UK
2008.03.03 © Robertson, A.
Dicyrtoma fusca var.1 from the UK
2009.01.18 © Farrell, S.
Dicyrtoma fusca var.1 from the UK
2009.01.18 © Farrell, S.
Dicyrtoma fusca var.1 from the UK
2009.02.20 © Michaelson, J.
Dicyrtoma fusca var.1 from France
2009.02.25 © Lefebvre, V.
Dicyrtoma fusca var.1 from the UK
2009.03.16 © Farrell, S.
Dicyrtoma fusca var.1 from the UK
2009.12.06 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2009.12.06 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2009.12.06 © Barton, T.
Dicyrtoma fusca var.1 from the UK
2010.01.24 © Barton, T.
Dicyrtoma fusca var.1 from France
2010.03.03 © Legrand, T.
Dicyrtoma fusca var.1 from France
Composed eye
2010.03.05 © Legrand, T.
Dicyrtoma fusca var.1 from France
2010.03.05 © Legrand, T.
Dicyrtoma fusca var.1 from the UK
2011.12.12 © Kent, J.J.
Dicyrtoma fusca var.1 from the UK
2012.02.03 © Nurcombe, E.
Dicyrtoma fusca var.1 from Holland
2012.02.17 © van Duinen, J.
Dicyrtoma fusca var.1 from the UK
2012.02.22 © Talbot, M.E.
Remark by Janssens, F. 2012.02.24: Possibly a case of ecomorphosis: a winter form.

Dicyrtoma fusca var.1 from the UK
2012.02.24 © Nurcombe, E.
Dicyrtoma fusca var.1 from France
2012.12.09 © Sandra.
Dicyrtoma fusca var.1 from the UK
2012.12.28 © Phillips, E.
Dicyrtoma fusca var.1 from the UK
2013.01.13 © Phillips, E.
Dicyrtoma fusca var.1 from the UK
2013.02.03 © Smith, S.U.
Dicyrtoma fusca var.1 from the UK
Dorsolateral pattern of dark spots
2013.03.19 © Ford, S.
Dicyrtoma fusca var.1 from the UK
With 1+1+2+2+1+1 midfacial setae
2014.11.29 © Henton-Jones, D.
Dicyrtoma fusca var.1 from Holland
2015.02.15 © van Duinen, J.
Dicyrtoma fusca var.1 from the UK
Lateral bothriotricha B and C
2016.03.08 © Phillips, E.
Dicyrtoma fusca var.1 from the UK
Midfacial setae : 1+1+2+2+1+1
Two duplicated unpaired setae (*)
2016.03.08 © Phillips, E.
Dicyrtoma fusca var.1 from Belgium
2016.12.11 © Huskens, M.-L.
Dicyrtoma fusca var.1 from France
2016.12.31 © Garcelon, P.
Dicyrtoma fusca var.1 from Belgium
Sub-anal appendages (*)
2017.01.10 © Huskens, M.-L.
Dicyrtoma fusca var.1 from France
Feeding instar
2017.11.11 © Picard, J.
Dicyrtoma fusca var.1 from Belgium
2017.12.30 © Louvigny, R.
Dicyrtoma fusca var.1 ♀ from Belgium
Caudal aspect
2018.01.05 © Huskens, M.-L.
Dicyrtoma fusca var.1 ♀ from Belgium
Sub-anal appendages on papillae (*)
2018.01.05 © Huskens, M.-L.
Dicyrtoma fusca var.1 from Belgium
Pre-jump posture
2018.01.06 © Huskens, M.-L.
Dicyrtoma fusca var.1 from Holland
Midfacial setae = 1+1+2+2+1+1
2018.12.23© Kamsteeg, G.
Dicyrtoma fusca var.1 from Germany
Midfacial setae = 1+2+1+2+1+1
2019.01.03 © Kluck, A.

Dicyrtoma fusca f. signata from Belgium
Midfacial setae : 1+1+2+1+2+1
2019.01.04 © Huskens, M.-L.
Dicyrtoma fusca f. signata from Holland
Bothriotricha A-B-C forming obtuse angle
2019.01.04 © Kamsteeg, G.
Dicyrtoma fusca f. signata from Belgium
Midfacial setae : 1+1+2+2+1+1
Labial palps (*)
2019.01.09 © Huskens, M.-L.
Dicyrtoma fusca f. signata ♀ from France
2019.01.19 © Garcelon, P.
Dicyrtoma fusca f. signata ♀ from France
Sub-anal appendages on papillae (*)
2019.01.19 © Garcelon, P.
Dicyrtoma fusca f. signata ♀ from France
Dorso-caudal dot pattern
2019.01.19 © Garcelon, P.
Dicyrtoma fusca f. signata from Holland
Cuticular wax glands enlarged = ready to moult
2019.02.09 © Kamsteeg, G.
Dicyrtoma fusca f. signata from Holland
Dorsal pattern of enlarged cuticular wax glands
2019.02.09 © Kamsteeg, G.
Dicyrtoma fusca f. signata from France
2020.02.18 © Dejoux, P.
Dicyrtoma fusca f. signata from Belgium
Tibiotarsus with 2 blunt setae (*)
2020.12.03 © Huskens, M.-L.
Dicyrtoma fusca f. signata from Belgium
Dorsal anal valve with 7 macrosetae on posterior edge (a3-1+a0+a1-3)
2020.12.03 © Huskens, M.-L.
Dicyrtoma fusca f. signata from Belgium
Midfacial macrosetae 1+3+2+1+1+1
2020.12.03 © Huskens, M.-L.
Dicyrtoma fusca f. signata from Belgium
Frontal area with frontal eye
2020.12.03 © Huskens, M.-L.
Dicyrtoma fusca f. signata from Belgium
Anterior tentorial pit
2020.12.03 © Huskens, M.-L.
Dicyrtoma fusca f. signata from France
Dark adult specimen
2021.01.18 © Garcelon, P.
Dicyrtoma fusca f. signata from France
Pale juvenile specimen
2021.01.18 © Garcelon, P.


            Syn.: Dicyrtoma fusca var.2 Janssens F, 2010
Remark by Janssens, F. 2010.08.14: Colour variant with middorsal pale stripe; ant.3 basally black, apically white; ant.4 white; feet white.

Dicyrtoma fusca var.2 from Czechia
2006.09.03 © Krásenský, P.
Dicyrtoma fusca var.2 from Czechia
2006.09.03 © Krásenský, P.
Dicyrtoma fusca var.2 from Estonia
2006.09.17 © Tartes, U.
Dicyrtoma fusca var.2 from Czechia
2010.09.00 © Deml, M.
Dicyrtoma fusca var.2 from Czechia
2010.09.00 © Deml, M.
Dicyrtoma fusca var.2 from Germany
Drinking from watersurfacefilm using everted collophore vesicles
2013.07.19 © Schmitt, R.
Dicyrtoma fusca var.2 from Czechia
2013.07.25 © Krásenský, P.
Dicyrtoma fusca var.2 from Czechia
2013.07.25 © Krásenský, P.
Dicyrtoma fusca var.2 from Czechia
Ant.3 basally black, apically white; ant.4 white
2013.07.25 © Krásenský, P.
Dicyrtoma fusca var.2 from Holland
2013.10.05 © van Duinen, J.
Dicyrtoma fusca var.2 from Poland
2014.08.23 © Kukla, M.
Dicyrtoma fusca var.2 from Czechia
2015.12.07 © Krásenský, P.
Dicyrtoma fusca var.2 from Bulgaria
2016.06.21 © van Duinen, J.
Dicyrtoma fusca var.2 from Holland
2016.07.09 © Wieringa, P.
Dicyrtoma fusca var.2 from Holland
2016.07.09 © Wieringa, P.
Dicyrtoma fusca var.2 from Portugal
Red-eyed albino form
2018.10.20 © Almeida, J.
Dicyrtoma fusca var.2 from Portugal
Red-eyed albino form
2018.10.21 © Almeida, J.
Dicyrtoma fusca var.2 from Holland
2020.06.20 © Heemskerk, R.
Dicyrtoma fusca var.2 from Holland
2020.06.20 © Heemskerk, R.
Dicyrtoma fusca var.2 from the USA
2021.03.08 © Pavy, R.
Dicyrtoma fusca var.2 from the USA
Ant.4 white
Apical part of ant.3 white
2021.03.08 © Pavy, R.
Dicyrtoma fusca var.2 from the USA
Grooming
Extended long collophore vesicle (*)
2021.03.08 © Pavy, R.


            Syn.: Dicyrtoma fusca var. rufescens Janssens F & Gutekunst V, 2016
Remark by Janssens, F. 2016.05.23: Colour variant with yellow eyepatch; ant.3+4 darker than ant.1+2; previously misidentified as Calvatomina rufescens.

"Calvatomina" rufescens from the UK
1975.05.25 © Storey, M.
"Calvatomina" rufescens from Sweden
2005 © Hall, K.
"Calvatomina" rufescens from Russia
2005 © Macroclub.ru.
"Calvatomina" rufescens from Czechia
2006.12.28 © Krásenský, P.
"Calvatomina" rufescens from Czechia
2006.12.28 © Krásenský, P.
"Calvatomina" rufescens from France
2007.07.20 © Lebeaux, P.
"Calvatomina" rufescens from the UK
2008.11.09 © Rodrigues, T.
"Calvatomina" rufescens from Czechia
2010.05.29 © Krejcik, S.
"Calvatomina" rufescens from Czechia
2011.02.09 © Krejcik, S.
"Calvatomina" rufescens from Holland
2011.07.26 © van Duinen, J.
"Calvatomina" rufescens from the UK
2012.01.07 © Kent, J.J.
"Calvatomina" rufescens from the UK
2012.06.19 © Jones, A.
"Calvatomina" rufescens from the UK
2012.06.19 © Jones, A.
"Calvatomina" rufescens from Holland
2012.06.21 © van Duinen, J.
"Calvatomina" rufescens juv. from Holland
2012.08.31 © van Duinen, J.
"Calvatomina" rufescens from France
2012.10.28 © Quintin, C.
"Calvatomina" rufescens from the UK
2012.11.18 © Nurcombe, E.
"Calvatomina" rufescens from Holland
Anal segment
2013.06.02 © van Duinen, J.
"Calvatomina" rufescens from the UK
2013.08.22 © Murray, A.
"Calvatomina" rufescens from the UK
Eyepatch with yellowish background and black ocelli
2013.08.22 © Murray, A.
"Calvatomina" rufescens from the UK
Foot of 1st leg with long sticky unguiculus
2013.08.22 © Murray, A.
"Calvatomina" rufescens ♂ from the UK
Raised male genital papilla (*)
2013.08.28 © Nurcombe, E.
"Calvatomina" rufescens from the UK
Ready to moult
2013.10.19 © Nurcombe, E.
"Calvatomina" rufescens from Holland
2013.10.26 © van Duinen, J.
"Calvatomina" rufescens from Germany
Mucro
2014.08.16 © Kluck, A.
"Calvatomina" rufescens juv. from the UK
Just moulted
2014.08.18 © Nurcombe, E.
"Calvatomina" rufescens from Poland
Unguis
2014.08.30 © Beck, K.
Dicyrtoma fusca var. rufescens from Germany
Head back and anterior great abdomen dorsally with spines
2014.09.10 © Gutekunst, V.
"Calvatomina" rufescens from Germany
Adult and juvenile
2014.09.27 © Beck, K.
"Calvatomina" rufescens from Germany
Dorsal tubercle with forward directed bothriotrichum
2014.09.27 © Beck, K.
"Calvatomina" rufescens from Germany
"Eyebrow" with spiky setae
2014.09.27 © Beck, K.
"Calvatomina" rufescens from Germany
Three backward directed ocelli
2014.09.29 © Beck, K.
"Calvatomina" rufescens from Germany
Eyepatch with central macroseta
2014.10.12 © Kluck, A.
"Calvatomina" rufescens from Germany
Forward directed bothriotrichum on dorsolateral tubercle
2014.11.15 © Beck, K.
"Calvatomina" rufescens from Germany
Six median unpaired facial setae
2015.08.09 © Kluck, A.
"Calvatomina" rufescens from Germany
Strong outer spur on ant.1 (*)
Caudal aspect
2015.08.30 © Kluck, A.
"Calvatomina" rufescens from Germany
2015.08.30 © Kluck, U.
Dicyrtoma fusca var. "rufescens" from Germany
Filament (f) of unguiculus (ung) longer than unguis (UNG); ung. with inner tooth (it)
2016.05.10 © Janssens F.
Leg.: Gutekunst, V.
Dicyrtoma fusca var. "rufescens" from Germany
Head back dorsally with distinct spines (*)
2016.05.19 © Gutekunst, V.
Dicyrtoma fusca var. "rufescens" from Germany
Unguiculus of mesoleg longer than unguis (*)
2016.05.19 © Gutekunst, V.
Dicyrtoma fusca var. "rufescens" from Germany
Dental setae serrate (*)
2016.05.20 © Janssens F.
Leg.: Gutekunst, V.
Dicyrtoma fusca var. "rufescens" from Holland
Head back dorsally with distinct spines (*)
2016.10.10 © van Duinen, J.
Dicyrtoma fusca var. "rufescens" from Belgium
2016.12.14 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Belgium
2016.12.14 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Belgium
2016.12.14 © Huskens, M.-L.
Dicyrtoma fusca var.1 & var. "rufescens" from Belgium
2016.12.14 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Hungary
2017.10.01 © Ujvári, Z.
Dicyrtoma fusca var. "rufescens" from France
Unguis without tunica
2017.10.03 © Picard, J.
Dicyrtoma fusca var. "rufescens" from Hungary
2017.11.23 © Ujvári, Z.
Dicyrtoma fusca var. "rufescens" ♂ from Belgium
Male genital papilla with circumgenital setae (*)
2018.09.16 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Finland
2018.10.15 © Ujvári, Z.
Dicyrtoma fusca var. "rufescens" from Belgium
2020.08.31 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Belgium
Left eyepatch
Ocelli C+D distinctly smaller
2020.08.31 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Belgium
Bothriotrichum A on papilla
2020.08.31 © Huskens, M.-L.
Dicyrtoma fusca var. "rufescens" from Germany
Midfacial setae 1+1+1+2+1+1
2020.11.05 © Jäschke, J.


            Syn.: Dicyrtoma fusca ab.1 Janssens F & Nurcombe E, 2012
Remark by Janssens, F. 2012.01.29: Colour aberration lacking black pigmentation. Kind of albino form with reddish eyes.

Dicyrtoma fusca ab.1 from the UK
Reddish ocelli on reddish eyepatch
2012.01.27 © Nurcombe, E.
Dicyrtoma fusca ab.1 from the UK
Reddish ocelli on reddish eyepatch
2012.01.27 © Nurcombe, E.
Dicyrtoma fusca ab.1 from the UK
Reddish ocelli on reddish eyepatch
2012.01.27 © Nurcombe, E.
Dicyrtoma fusca ab.1 from Holland
Reddish ocelli on reddish eyepatch
2015.12.28 © van Duinen, J.
Dicyrtoma fusca ab.1 from Holland
Reddish ocelli on reddish eyepatch
2018.01.11 © van Duinen, J.
Dicyrtoma fusca ab.1 ♂ subadult from Holland
Reddish ocelli on reddish eyepatch
2022.10.11 © van Duinen, J.


            Syn.: Dicyrtoma fusca ab.2 Janssens F, 2012
Remark by Janssens, F. 2012.01.29: Colour aberration with white feet indicating the specimen is ready to moult.

Dicyrtoma fusca ab.2 from the UK
2005 © Hopkin, S.P.
Dicyrtoma fusca ab.2 from the USA
2007.02.18 © Moorehead, C.
Dicyrtoma fusca ab.2 from France
2007.04.10 © Lebeaux, P.
Dicyrtoma fusca ab.2 from France
with left vesicle eversed from collophore
2007.04.10 © Lebeaux, P.
Dicyrtoma fusca ab.2 from France
2007.04.10 © Lebeaux, P.
 
Dicyrtoma fusca ab.2 from France
2007.04.10 © Lebeaux, P.
 
Dicyrtoma fusca ab.2 from the UK
2007.12.29 © Kilford., B
Dicyrtoma fusca ab.2 from the UK
2009.03.22 © Robertson, A.
Dicyrtoma fusca ab.2 from the UK
Transparant foot reveals new foot has been redrawn
2009.03.22 © Robertson, A.
Dicyrtoma fusca ab.2 from the UK
2011.12.01 © Nurcombe, E.
Dicyrtoma fusca ab.2 from the UK
2012.01.10 © van Duinen, J.
Dicyrtoma fusca ab.2 from the UK
Speckled appearance reveals new cuticula in preparation
2012.12.11 © Phillips, E.
Dicyrtoma fusca ab.2 from the UK
Speckled appearance reveals new cuticula in preparation
2013.01.07 © Phillips, E.




             Syn.: Papirius fusco(sic) Reuter, OM, 1892:24 ISS
Dicyrtoma fusca from Siberia
After Stebaeva SK, 1966 Fig.3.9
Dicyrtoma (Dicyrtoma) fusca from the USA
After Snider, R., 1990 Fig.72.
Dicyrtoma (Dicyrtoma) fusca from the USA
After Snider, R., 1990 Fig.73.
Dicyrtoma fusca from the UK
Foot complex, claw without tunica
2004 © Hopkin, S.P.
Dicyrtoma fusca from Estonia
2006.09.17 © Tartes, U.
Dicyrtoma fusca from Estonia
2006.10.01 © Tartes, U.
Dicyrtoma fusca from the USA
2007.03.13 © Moorehead, C.
Dicyrtoma fusca from the USA
2007.05.05 © Moorehead, C.
Dicyrtoma fusca from the USA
2007.05.05 © Moorehead, C.
Dicyrtoma fusca from the USA
2007.05.06 © Moorehead, C.
Dicyrtoma fusca from the USA
2007.05.31 © Cowen, R.
Dicyrtoma fusca from Norway
2007.07.23 © Fauskanger, I.
Dicyrtoma fusca from Slovakia
2007.09.01 © Stefanovie, P.
Dicyrtoma fusca from the UK
2007.11.14 © Campbell, A.
Dicyrtoma fusca from the UK
2007.12.23 © Kilford., B
Dicyrtoma fusca from the UK
2007.12.24 © Barton, T.
Dicyrtoma fusca from the UK
2007.12.26 © Barton, T.
Dicyrtoma fusca from the UK
2007.12.29 © Barton, T.
Dicyrtoma fusca from the USA
2007.12.29 © Moorehead, C.
Dicyrtoma fusca from the UK
2007.12.29 © Barton, T.
Dicyrtoma fusca from the UK
2008.01.03 © Campbell, A.
Dicyrtoma fusca from the Netherlands
2008.03.02 © Waardenburg, J.
Dicyrtoma fusca from the UK
2008.03.04 © Farrell, S.
Dicyrtoma fusca from the UK
2008.03.04 © Farrell, S.
Dicyrtoma fusca from the UK
2008.03.28 © Farrell, S.
Dicyrtoma fusca from the UK
2008.10.06 © Robertson, A.
Dicyrtoma fusca from Holland
2008.11.13 © Baas, A.H.
Dicyrtoma fusca from the UK
2009.02.13 © Talbot, M.
Dicyrtoma fusca from the UK
2009.02.20 © Michaelson, J.
Dicyrtoma fusca from Belgium
2009.09.13 © Wallays, H.
Dicyrtoma fusca from the USA
2009.11.16 © Bergen, L.
Dicyrtoma fusca from the UK
2009.11.21 © Barton, T.
Dicyrtoma fusca from Belgium
2009.12.16 © De Bruyn, L.
Dicyrtoma fusca from Belgium
2010.01.13 © De Bruyn, L.
Dicyrtoma fusca from Belgium
2010.01.13 © De Bruyn, L.
Dicyrtoma fusca from France
Bothriotrichum A on tubercle
2010.03.05 © Legrand, T.
Dicyrtoma fusca from the USA
2011.02.19 © Coulter.
 , B
Dicyrtoma fusca (bar=2µm)
Hexagonal epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.6B
Dicyrtoma fusca juv. from the UK
Preparing to moult
2012.01.05 © Kent, J.J.
Dicyrtoma fusca juv. from the UK
2012.09.07 © Murray, A.
Dicyrtoma fusca juv. from the UK
2012.09.15 © Nurcombe, E.
Dicyrtoma fusca juv. from Holland
2012.10.02 © van Duinen, J.
Dicyrtoma fusca from Holland
Ready to moult
2012.10.28 © van Duinen, J.
Dicyrtoma fusca from the UK
More than 2 unpaired facial setae
2012.11.16 © Phillips, E.
Dicyrtoma fusca from the UK
Speckled appearance indicating preparing to moult
2012.12.11 © Phillips, E.
Dicyrtoma fusca from the UK
Annulated apical part of 3rd antennal segment
2012.12.12 © Phillips, E.
Dicyrtoma fusca ♂ from the UK
Raised male genital papilla
2012.12.12 © Phillips, E.
Dicyrtoma fusca from the UK
Facial tentorial pits
2012.12.12 © Phillips, E.
Dicyrtoma fusca from the UK
Atrium : opening of buccal cone
From which mouthparts protrude
2013.01.01 © Henton-Jones, D.
Dicyrtoma fusca from the UK
Predated by Acari
2013.01.06 © Holt, H. & A.
Dicyrtoma fusca from the UK
Grazing
2013.01.12 © Holt, H. & A.
Dicyrtoma fusca from the UK
Predated by Acari: Bdellidae
2013.01.12 © Holt, H. & A.
Dicyrtoma fusca from the UK
Labral chaetotaxy
Labial palps
2013.02.07 © Phillips, E.
Dicyrtoma fusca from Holland
Just moulted. Footcomplex with slender claw and needle-like empodium longer than claw
2013.10.25 © van Duinen, J.
Dicyrtoma fusca juv. from the UK
2013.10.25 © Phillips, E.
Dicyrtoma fusca ♂ from the UK
Raised genital papilla of reproductive male
2013.11.15 © Garlick, T.
Dicyrtoma fusca ♀ from Germany
2014.01.01 © Beck, K.
Dicyrtoma fusca from the UK
Labrum
2014.02.22 © Henton-Jones, D.
Dicyrtoma fusca from Germany
Unguiculus (ung) & unguis (Ung) of 1st leg
2014.11.09 © Kluck, A.
Dicyrtoma fusca from the UK
Mottled: preparing to moult
2015.01.17 © Phillips, E.
Dicyrtoma fusca from the UK
Hindfoot with unguiculus having long exad bent apex
2015.01.17 © Phillips, E.
Dicyrtoma fusca from Germany
Ready to moult
2015.10.31 © Kluck, A.
Dicyrtoma fusca from Holland
With 6 thick unpaired midfacial setae
2015.11.06 © Wieringa, P.
Dicyrtoma fusca from the UK
Midfacial thick setae: 1+1+2+2+1+1
2015.11.21 © Phillips, E.
Dicyrtoma fusca from the USA
2015.12.22 © Turley, N.
Dicyrtoma fusca from New Zealand
2014 © Murray, A.
After Greenslade P et al, 2016 Fig.4b
Dicyrtoma fusca from New Zealand
2014 © Murray, A.
After Greenslade P et al, 2016 Fig.4a
Dicyrtoma fusca from Belgium
Labrum
Labial palps
2016.02.10 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Proleg with slender unguiculus with inner tooth
2016.02.10 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Ant.1 with ventral spur (*)
2016.02.10 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Dorsal papilla with forward directed bothriotrichum A
2016.02.11 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Pre-jump posture
2016.11.27 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Speckled: preparing to moult
2016.12.02 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Bothriotricha B+C
2016.12.05 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Spermatophore
2016.12.11 © Huskens, M.-L.
Dicyrtoma fusca ♂ from Belgium
Raised genital papilla with apical ring of setae
2017.01.24 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Preparing to moult
2017.10.20 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Preparing to moult
2017.11.14 © Huskens, M.-L.
Dicyrtoma fusca ♂ from Belgium
2018.03.26 © Huskens, M.-L.
Dicyrtoma fusca from France
Antenna
2018.10.27 © Picard, J.
Dicyrtoma fusca from the UK
Midfacial setae : 1+1+2+2+1+1
2018.12.30 © Barton, T.
Dicyrtoma fusca from Belgium
Mouthparts ventral aspect
Labrum with 3 parallel sclerotic ridges
2019.01.05 © Huskens, M.-L.
Dicyrtoma fusca from Belgium
Strong ant.1 spur (**); weak ant.2 spur (*)
2019.01.28 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Sub-anal appendages
2019.12.25 © Huskens, M.-L.
Dicyrtoma fusca juv. from France
Bulby clypeus
2019.12.30 © Garcelon, P.
Dicyrtoma fusca from France
Mouthparts ventral : labrum with 3 ridges; split labium with labial palps
2020.03.25 © Garcelon, P.
Dicyrtoma fusca from Belgium
Pale spots mark fat tissue
2020.10.11 © Huskens, M.-L.
Dicyrtoma fusca from the UK
Head outline dorso-frontal aspect
2020.10.20 © Cann, A.J.
Dicyrtoma fusca ♂ from Belgium
Midfacial setae : 1+1+2+2+1+1
2020.10.28 © Huskens, M.-L.
Dicyrtoma fusca ♂ from Belgium
Genital papilla
With apical crown of circumventing setae
2020.10.28 © Huskens, M.-L.
Dicyrtoma fusca juv. ♂? from Belgium
Head pale
Midfacial setae : 1+1+2+2?+1+1
2020.10.28 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Midfacial setae 1+1+1+1+1+1
2020.11.03 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Small abdomen
Macrosetae a0-a3 present
2020.11.03 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Midfacial setae 1+1+2+2+1+1
2020.11.06 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Sub-anal appendage (*)
2020.11.06 © Huskens, M.-L.
Dicyrtoma fusca ♀ from Belgium
Midfacial setae 1+1+2+1+1+1
2020.11.09 © Huskens, M.-L.
Dicyrtoma fusca from Holland
Prepairing to moult
2020.11.14 © Kamsteeg, G.
Dicyrtoma fusca ♂ from Belgium
Midfacial setae 1+1+2+2+1+1
2020.11.17 © Huskens, M.-L.

  Subspecies flavescens Axelson, 1902

            Syn.: Papirius fuscus var. cavernicola Schäffer, 1900 =?


  Subspecies medialis Stach, J, 1930:82

Papirius fuscus var. medialis from Spain
After Stach J, 1930 Abb.XV.19



Species ghilarovi Bretfeld, 1996

Species grinbergsi Stebaeva, SK, 1966:10

Dicyrtoma grinbergsi from Siberia
After Stebaeva SK, 1966 Fig.3.1
Dicyrtoma grinbergsi from Siberia
After Stebaeva SK, 1966 Fig.3.8

Species guttata (Say, 1821) Kolenati, FA, 1858:134

             Syn.: Sminthurus Guttatus Bourlet, C, 1845:158 ISS
             Syn.: Dicyrtoma Guttata(sic) Kolenati, FA, 1858:134 ISS

Species hageni (Folsom, JW, 1896:344) Stach, J, 1957:63

            Syn.: Papirius hagenii(sic) Folsom, JW, 1896:344 IOS
            Syn.: Dicyrtoma frontalis Banks, 1903
Ptenothrix frontalis from the USA
After Maynard EA, 1951 PlateXXXVII Fig.668
Ptenothrix frontalis from the USA
After Maynard EA, 1951 PlateXXXVII Fig.669
Dicyrtoma hageni f. frontalis from the USA
2014.03.02 © Bertone, M.
Dicyrtoma hageni f. frontalis from the USA
2018.06.04 © Caterino, M.
Dicyrtoma hageni f. frontalis from the USA
2020.03.12 © Caterino, M.

            Syn.: Dicyrtoma hageni f. alabamensis Janssens F & Charny V, 2022
Dicyrtoma hageni f. alabamensis from the USA
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
Facial aspect
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
Eye background reddish
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
Facial aspect
2022.01.12 © Charny, V.
Dicyrtoma hageni f. alabamensis from the USA
Reddish colour variant
2022.03.24 © Charny, V.

            Syn.: Dicyrtoma hageni f. canadensis Janssens F & Chan J, 2024
Dicyrtoma hageni f. canadensis ♀ from Canada
2024.01.07 © Chan, J.
Dicyrtoma hageni f. canadensis ♀ from Canada
2024.01.07 © Chan, J.
Dicyrtoma hageni f. canadensis ♀ from Canada
2024.01.10 © Chan, J.
Dicyrtoma hageni f. canadensis ♀ from Canada
2024.01.10 © Chan, J.
Dicyrtoma hageni f. canadensis ♀ from Canada
2024.01.10 © Chan, J.
Dicyrtoma hageni f. canadensis ♀ from Canada
Dorsal anal valve with row of 7 posterior setae
Ventral anal valve with subanal appendage (saa)
2024.01.10 © Chan, J.
Dicyrtoma hageni f. canadensis ♂ from Canada
With more then 2 midfacial unpaired setae
2024.01.26 © Chan, J.
Dicyrtoma hageni f. canadensis ♂ from Canada
Male genital papilla
2024.01.26 © Chan, J.
Dicyrtoma hageni f. canadensis ♂ from Canada
Upper unguis long and narrow
Lower unguiculus longer then unguis
2024.01.26 © Chan, J.





            Syn.: Papirius hagenii(sic) Jackson, AD, 1907:171 IOS
Dicyrtoma (Dicyrtoma) hageni from the USA
After Folsom JW in Christiansen KA & Bellinger PF, 1981 Fig.1029C
Dicyrtoma (Dicyrtoma) hageni from the USA
After Folsom JW in Christiansen KA & Bellinger PF, 1981 Fig.1029D

             Syn.: Dicyrtoma haegi(sic) Palacios-Vargas, JG et Castaño-Meneses, G, 2002:399 ISS
Dicyrtoma hageni from the USA
2009.12.29 © Zungri, D.
Dicyrtoma hageni from the USA
2010.01.03 © Zungri, D.
Dicyrtoma hageni from the USA
2010.01.09 © Zungri, D.
Dicyrtoma hageni from the USA
2010.01.09 © Zungri, D.
Dicyrtoma hageni from the USA
2010.01.09 © Zungri, D.





Species insularis Carpenter, 1935, n.n.

Species maculosa Ionesco in Denis, 1930, n.n.

Species mangle Soto-Adames, FN, 1988:60

Dicyrtoma mangle from Puerto Rico
After Soto-Adames FN, 1988 Fig.1

Species mithra Wray, 1949

            Syn.: Dicyrtoma (Dicyrtoma) flammea Christiansen, K et Bellinger, P, 1981:1226 =?
            Syn.: Dicyrtoma (Dicyrtoma) flammea Christiansen, K et Bellinger, P, 1998:1397 =?


Species novaeboracensis (Fitch, A, 1863:674) Barnes, JK, 1988:90

            Syn.: Symnthurus(sic) Novæboracensis(sic) Fitch, A, 1863:674 IOS
            Syn.: Dicyrtoma? sp. Barnes, JK, 1988:90



             Syn.: Papirius novoeboracensis(sic) Packard, 1873 ISS
             Syn.: Papirius novoeboreacensis(sic) Harvey, 1893 ISS
             Syn.: Papirius novoeboracensis(sic) Dalla Torre, 1895 ISS
             Syn.: Papirius novoeboracensis(sic) Jackson, AD, 1907:166 ISS



Species ochreoa Wray, 1949

Species oraniensis Lin. in Lucas, H, 1846:255

            Syn.: Dicyrtoma Oraniensis(sic) Lin. in Lucas, H, 1846:255 IOS
Dycirtoma(sic) oraniensis from Algeria
After Lucas H, 1849 Pl.2 Fig.7
             Syn.: Dicyrtoma Oraniensis(sic) Kolenati, FA, 1858:134 ISS




Species pakistanica Yosii & Ashraf, 1965

Species vinalis Wray, 1949


Genus Dicyrtomina Börner, 1903

  Syn.: ... Dicyrtoma Bourlet, 1842 partim
  Syn.: Dicyrtomidae Dicyrtoma(sic) Busmachiu, G, Bedos, A et Deharveng, L, 2015:103 lapsus
  Syn.: Sminthuridæ(sic) Papirius Lubbock, J, 1862:431
  Syn.: ... Papirirus(sic) Stach, 1930 lapsus ISS


  Syn.: Collembola Dicrytomina(sic) Paulus, HF in Horridge, GA, 1974:10 ISS
Remark by ICZN. 2007:246: Placed on the Official List of Names in Zoology. Opinion 1092.

Dicyrtomina sp. from the USA
2006 © Moorehead, C.
Dicyrtomina sp. from the USA
2008.01.06 © Moorehead, C.
Dicyrtomina sp. from the USA
2008.02.28 © Bradford, A.
Dicyrtomina sp. from the UK
2008.12.14 © Kilford., B
Dicyrtomina violacea from the UK
Unguis with outer tunica
2009.03.23 © Ardron, P.A.
Dicyrtomina violacea from the UK
Ant.1 organ : strong subapical ant.1 spur; weak basal ant.2 spur
2009.03.23 © Ardron, P.A.
Dicyrtomina saundersi from the UK
Foot: unguis with tunica
2010.01.23 © Robertson, A.
Dicyrtomina sp. from the USA
Foot: unguis with tunica
2010.01.24 © Krebs, C.
Dicyrtomina minuta f. ornata from Canada
Very short 4th antennal segment
2012.02.01 © Hochhalter, R.
Dicyrtomina saundersi & minuta from the UK
2012.12.01 © Phillips, E.
Dicyrtomina saundersi from the UK
Footcomplex with broad empodium with inner tooth (*)
2013.11.19 © Phillips, E.
Dicyrtomina sp. from the UK
Distinctly short 4th antennal segment
2014.01.15 © Phillips, E.
Dicyrtomina sp. from New Zealand
2014.02.25 © Murray, A.
Dicyrtomina sp. from New Zealand
2014.02.25 © Murray, A.
Dicyrtomina sp. from New Zealand
2014.05.08 © Murray, A.
Dicyrtomina sp. from Czechia
2015.03.05 © Krásenský, P.
Dicyrtomina sp. from Australia
2015.07.27 © Porch, N.
Dicyrtomina saundersi from Germany
Third foot: unguiculus with broad lamella
Thick unguis with tunica
2015.11.27 © Kluck, A.
Dicyrtomina minuta from Sweden
Dorsally upcurved gut
2016.09.19 © Neergaard, R.
Dicyrtomina saundersi from the UK
First foot: unguis with tunica (*);
unguiculus with long filament (**)
2017.01.24 © Horton, M.
Dicyrtomina sp. from the UK
Spermatophore
2017.01.24 © Horton, M.
?Dicyrtomina? sp. from Australia
2017.06.10 © Jeales, T.
?Dicyrtomina? sp. from Australia
2017.06.10 © Jeales, T.
Dicyrtomina saundersi from the UK
3rd foot: unguis with tunica
2017.11.12 © Horton, M.
Dicyrtomina ornata f. labellei from Belgium
Abd.6 dorsally with heavy spiky macrosetae M+N+E+H
2018.12.23 © Huskens, M.-L.
Dicyrtomina ornata f. couloni from Germany
Abd.6 dorsally with heavy spiky macrosetae M+N
2019.01.10 © Kluck, A.
Dicyrtomina ornata f. couloni from Belgium
Antennal organ with distinct ant.1 spur (**) + smaller ant.2 spur (*)
2019.01.28 © Huskens, M.-L.
?Dicyrtomina? sp. from Taiwan
2022.02.24 © Cheng, H.-J.
Dicyrtomina sp. from France
Infected by Neozygites sp.
2023.02.04 © Clerc, O.
?Dicyrtomina? sp. from China
2023.03.03 © El-Rady, J.
Dicyrtomina sp. from Taiwan
2023.05.31 © Cheng, H.-J.
Dicyrtomina sp. from Taiwan
2023.05.31 © Cheng, H.-J.
Dicyrtomina sp. from Taiwan
2023.05.31 © Cheng, H.-J.
?Dicyrtomina? sp. from China
2023.07.01 © Wang, G.
Dicyrtomina ornata from the UK
Abd.6 with 3 M setae + 4 N setae
2023.11.05 © Barton, T.
Dicyrtomina sp. from Belgium
Spermatophore
Stalk exits spermdrop apically
2023.12.22 © Huskens, M.-L.



Species sp.1 Yayuk, RS (sic), 1989:20

            Syn.: Dicyrtomina calva Yosii, 1959, nec Denis, 1948


Species sp.2 Janssens F & Coulter B, 2011

Dicyrtomina sp. from the USA
2011.02.27 © Coulter., B
Dicyrtomina sp. from the USA
2014.03.15 © Rosenfeld, J.
Dicyrtomina sp. from the USA
2014.03.15 © Rosenfeld, J.
Dicyrtomina sp. from the USA
2024.02.08 © AVNJ.
Dicyrtomina sp. juv.? from the USA
2024.02.08 © AVNJ.

Species sp.4 Janssens F & Jiku, 2015

Dicyrtomina sp. from Japan
2015.01.03 © Jiku.
Dicyrtomina sp. from Japan
2015.01.03 © Jiku.
Dicyrtomina sp. from Japan
2015.01.03 © Jiku.
Dicyrtomina sp. from Japan
Pale and dark colour form
2015.01.03 © Jiku.
Dicyrtomina sp. from Japan
2015.01.03 © Jiku.
Dicyrtomina sp. from Japan
Dark colour form
2015.01.03 © Jiku.

Species sp.5 Janssens F & Dobbs B, 2016

Dicyrtomina sp. from the USA
2016.12.01 © Dobbs, Br.
Dicyrtomina sp. from the USA
2016.12.31 © Dobbs, Br.
Dicyrtomina sp. from the USA
2016.12.31 © Dobbs, Br.
Dicyrtomina sp. from the USA
Six unpaired midfacial setae
2016.12.31 © Dobbs, Br.
Dicyrtomina sp. from the USA
Thick unguis with tunica (*)
2017.01.02 © Dobbs, Br.
Dicyrtomina sp. from the USA
2017.01.02 © Dobbs, Br.
Dicyrtomina sp. from the USA
Caudal aspect
2017.01.02 © Dobbs, Br.
Dicyrtomina sp. from the USA
2017.01.19 © Dobbs, Br.
Dicyrtomina sp. from the USA
2017.01.20 © Dobbs, Br.
Dicyrtomina sp. from the USA
2017.01.21 © Dobbs, Br.
Dicyrtomina sp. from the USA
Six unpaired midfacial setae
2017.11.29 © Dobbs, Br.
Dicyrtomina sp. juv. from the USA
2017.12.11 © Dobbs, Br.
Dicyrtomina sp. juv. from the USA
2017.12.11 © Dobbs, Br.
Dicyrtomina sp. from the USA
2018.12.30 © Duncan, W.L.
Dicyrtomina sp. from the USA
2018.12.30 © Duncan, W.L.
Dicyrtomina sp. from the USA
Bothriotrichum A on papilla and directed forwards
2019.01.10 © Dobbs, Br.
Dicyrtomina sp. from the USA
Abd.6 with dorsal setae 2M'+M+2N
2019.01.15 © Dobbs, Br.
Dicyrtomina sp. ♂ juv. from the USA
2019.01.15 © Dobbs, Br.
Dicyrtomina sp. ♂ juv. from the USA
2019.01.15 © Dobbs, Br.

Species sp.6 Janssens F & Goff M, 2022

Dicyrtomina sp.nov. from the USA
2022.03.29 © Goff, M.
Dicyrtomina sp.nov. from the USA
2022.03.29 © Goff, M.
Dicyrtomina sp.nov. from the USA
Dark colour form
2022.03.29 © Goff, M.
Dicyrtomina sp.nov. from the USA
Dark colour form
2022.03.29 © Goff, M.

Species africana Womersley, 1931

Species cyanea Merlo & Najt, 1974

Species dorsolineata Latzel, R, 1917:252

Dicyrtomina dorsolineata from France
2017.09.09 © Picard, J.
Dicyrtomina dorsolineata from France
2017.09.09 © Picard, J.
Dicyrtomina dorsolineata from France
2017.09.09 © Picard, J.
Dicyrtomina dorsolineata juv. from France
Dorsal pattern undeveloped
2017.09.09 © Picard, J.
Dicyrtomina dorsolineata from France
Six unpaired midfacial setae
2017.09.10 © Picard, J.
Dicyrtomina dorsolineata from France
Outer pair of longitudinal bands joined posteriorly
2017.09.10 © Picard, J.
Dicyrtomina dorsolineata from France
Wiggling gut signals
2017.09.17 © Picard, J.
Dicyrtomina dorsolineata from France
Laterally white background
2017.09.23 © Picard, J.
Dicyrtomina dorsolineata from France
Pale subadult
2017.09.24 © Picard, J.
Dicyrtomina dorsolineata ♂ from France
Pale subadult
2017.09.24 © Picard, J.
Dicyrtomina dorsolineata juv. from France
2017.09.29 © Picard, J.
Dicyrtomina dorsolineata ♀ from France
Sub-anal appendages (s.a.a.)
2017.09.30 © Picard, J.
Dicyrtomina dorsolineata ♀ from France
Mucro edges smooth
2017.09.30 © Picard, J.
Dicyrtomina dorsolineata from France
Ant.3 not subdivided
2017.09.30 © Picard, J.
Dicyrtomina dorsolineata ♂ juv. from France
Pale subadult
2017.09.30 © Picard, J.
Dicyrtomina dorsolineata ♀ juv. from France
Caudal aspect
2017.09.30 © Picard, J.
Dicyrtomina dorsolineata from France
Caudal aspect
2017.10.01 © Tiky, S.
Dicyrtomina dorsolineata from France
2017.10.01 © Tiky, S.
Dicyrtomina dorsolineata from France
2017.10.01 © Tiky, S.
Dicyrtomina dorsolineata from France
2018.04.04 © Garcelon, P.
Dicyrtomina dorsolineata from France
2018.04.04 © Garcelon, P.
Dicyrtomina dorsolineata from Hungary
2018.09.28 © Ujvári, Z.
Dicyrtomina dorsolineata from Hungary
2018.09.28 © Ujvári, Z.
Dicyrtomina dorsolineata ♂ from France
Very dark
Lateral longitudinal bands joined
2019.05.28 © Picard, J.
Dicyrtomina dorsolineata ♂ from France
Caudal aspect
2019.05.28 © Picard, J.
Dicyrtomina dorsolineata ♂ from France
Male genital papilla
2019.05.28 © Picard, J.
Dicyrtomina dorsolineata from France
2019.09.24 © Dejoux, P.
Dicyrtomina dorsolineata from Spain
2019.12.17 © Zeeders, M.
Dicyrtomina dorsolineata from Spain
2020.01.25 © Zeeders, M.
Dicyrtomina dorsolineata from Spain
2021.12.21 © Zeeders, M.
Dicyrtomina dorsolineata from Spain
2021.12.23 © Zeeders, M.
Dicyrtomina dorsolineata from Spain
Feeding instar
2021.12.23 © Zeeders, M.
Dicyrtomina dorsolineata from Spain
Feeding instar
Caudal aspect
2021.12.23 © Zeeders, M.

Species ealana Marlier, 1945

Species flavosignata (Tullberg, 1871) Linnaniemi, 1912, B

            Syn.: Papirius flavo-signatus(sic) Tullberg, 1871 IOS
             Syn.: Dicyrtoma Lucasii(sic) Nicolet, H, 1847:361 ISS
Dicyrtoma Lucasii(sic) from Switzerland
After Nicolet,, 1847 Pl.6 fig.6.

             Syn.: Dicyrtoma lucasii(sic) Stach, 1957,si ISS
            Syn.: Papirius flavosignatus var. cinerascens Reuter, OM, 1892:231 nn
            Syn.: Papirius flavosignatus var. orcina Absolon, K, 1901:213
            Syn.: Dicyrtoma minuta ab. pallida Ågren, 1903
            Syn.: Dicyrtoma minuta f. flavescens Wahlgren, 1906
            Syn.: Dicyrtomina variabilis Maynard, EA, 1951:278
Dicyrtomina variabilis from the USA
After Maynard, E.A., 1951 Pl.XXXVI Fig.647





            Syn.: Papirius flavo-signatus(sic) Tullberg, 1872 IOS
            Syn.: Papirius flavo-signatus(sic) Reuter, OM, 1876:80 IOS
             Syn.: Papirius flavo-signato(sic) Reuter, OM, 1876:85 ISS
             Syn.: Papirius flavosignato(sic) Reuter, OM, 1895:14 ISS
             Syn.: Papirius flavo-signatus(sic) Willem, V, 1900:63,B ISS
Dicyrtomina minuta var. flavosignata from Poland
After Stach J., 1957 Pl.III fig.1.
Dicyrtomina flavosignata from the UK
2008.08.18 © Robertson, A.
Dicyrtomina flavosignata from Russia
2008.12.22 © Macroclub.ru
Dicyrtomina flavosignata from Hungary
2009.05.17 © Keresztes, G.
Dicyrtomina flavosignata from Germany
2010.09.18 © Marquard, F.
Dicyrtomina flavosignata from Russia
2010.09.24 © Pavlov, D.
Dicyrtomina flavosignata from Germany
2015.06.28 © Wolf, M.
Dicyrtomina flavosignata from the UK
2016.01.23 © Barton, T.
Dicyrtomina flavosignata from France
2016.02.08 © Garcelon, P.
Dicyrtomina flavosignata from France
2016.02.25 © Garcelon, P.
Dicyrtomina flavosignata from France
2016.11.27 © Garcelon, P.
Dicyrtomina flavosignata from France
2016.11.27 © Garcelon, P.
Dicyrtomina flavosignata from France
2016.11.29 © Garcelon, P.
Dicyrtomina flavosignata ♂ from France
2016.12.17 © Garcelon, P.
Dicyrtomina ?flavosignata? from Sweden
2017.09.18 © Neergaard, R.
Dicyrtomina flavosignata from France
2018.03.11 © Garcelon, P.
Dicyrtomina flavosignata from France
2018.03.11 © Garcelon, P.





Species gigantisetae Lin & Xia, 1985

Species insularis Carpenter, 1934

Species minuta (Fabricius O, 1783) Börner, 1906, B

            Syn.: Podure(sic) noirâtre à taches fauves sous le ventre Geoffroy, 1762
            Syn.: Podure(sic) noirâtre à taches jaunes sous le ventre Nicolet, H, 1847:358
            Syn.: Smynthurus(sic) signata(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:97 IOS
Smynthurus(sic) signata(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.8

Smynthurus(sic) signatus from France
After Lucas H, 1840 Pl.1 Fig.5
Smynthurus(sic) signatus from France
After Lucas H, 1840 Pl.1 Fig.5A


             Syn.: Smynthurus(sic) Signata(sic) Lucas, H, 1840:567 ISS
             Syn.: Podura Signata(sic) Bourlet, C, 1845:153 ISS
             Syn.: Podura Signata(sic) Bourlet, C, 1845:154 ISS
             Syn.: Sminthurus Signata(sic) Bourlet, C, 1845:160 =? ISS
            Syn.: Dicyrtoma atropurpurea Bourlet, 1842
             Syn.: Dicyrtoma dorsi-maculata(sic) Bourlet, 1843 ISS
             Syn.: Dicyrtoma Dorsimalulata(sic) Bourlet, C, 1845:150 ISS
             Syn.: Dicyrtoma Dorsimalulata(sic) Bourlet, C, 1845:160 ISS
             Syn.: Dicyrtoma Dorsimalulata(sic) Bourlet, C, 1845:168 ISS
Dicyrtoma Dorsimaculata(sic) from France
With dorsal tubercles
After Bourlet C, 1845 Fig.15
Dicyrtoma Dorsimaculata(sic) from France
Ventral aspect
Collophore vesicles everted
After Bourlet C, 1845 Fig.16

             Syn.: Sminthurus Dorsimaculata (sic) var. Viridis(sic) Bourlet, C, 1845:160 =? ISS
            Syn.: Smynthure(sic) croisé(sic) Nicolet, H, 1847:358
            Syn.: Papirius nigromaculatus Lubbock, J, 1862:589
Papirius nigromaculatus from the UK
After Lubbock J, 1862 Pl.LIX Fig.4

            Syn.: Papirius nigromaculatus Lubbock, J, 1873:127
Papirius nigromaculatus from the UK
After Lubbock J, 1873 Pl.10
Dicyrtomina minuta f. nigromaculata from Czechia
2015.12.07 © Krásenský, P.
Dicyrtomina minuta f. nigromaculata from Belgium
2016.12.10 © Huskens, M.-L.
Dicyrtomina minuta f. nigromaculata from Belgium
2017.08.29 © Huskens, M.-L.
Dicyrtomina minuta f. nigromaculata from France
3 spots mark anterior tentorial pits
2020.11.18 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from France
Ocellus E with transparent dome
2020.11.18 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from France
Ant.1 with subapical ventral spur (sp1)
Ant.2 with basal ventral spur (sp2)
2020.11.18 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from France
Very dark form
With pale dorsum
2020.11.18 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from France
2020.11.21 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from France
2020.11.21 © Fiacre, R.
Dicyrtomina minuta f. nigromaculata from Belgium
2024.06.26 © Huskens, M.-L.

             Syn.: Smynthurus(sic) Coulonii(sic) Lubbock, J, 1870:281 ISS
             Syn.: Smynthurus(sic) culoni(sic) Tullberg, 1872 ISS
             Syn.: Smynthurus(sic) Coulonii(sic) Parona, 1876 ISS
             Syn.: Papirius Coulonii(sic) Uzel, 1891 ISS
             Syn.: Papirius Coulonii(sic) von Dalla Torre, 1895 ISS
             Syn.: Smynthurus(sic) Coulonii(sic) Stach ISS
            Syn.: Smynthurus(sic) couloni Palissa, 1964 partim
            Syn.: Papirius minutus var. fusca Krausbauer, 1902
            Syn.: Papirius minutus var. pulchella Krausbauer, 1902
            Syn.: Papirius minutus var. quadrimaculata Krausbauer, 1902
            Syn.: Dicyrtoma minuta ab. pallida Ågren, 1903
            Syn.: Dicyrtomina labellei Christiansen, K et Bellinger, P, 1981:1218
            Syn.: Dicyrtomina labellei Poole, RW in Poole, RW et Gentili, P, 1997:55
            Syn.: Dicyrtomina labellei Christiansen, K et Bellinger, P, 1998:1398
            Syn.: Dicyrtomina variabilis Maynard, EA, 1951:278
            Syn.: Dicyrtoma minuta f. typica Paclt, J, 1961:238



             Syn.: Papirius minuto(sic) Reuter, OM, 1895:14 ISS
Dicyrtomina minuta from Poland
After Stach, J, 1957 Pl.II Fig.1.
Dicyrtomina minuta from Europe
Abdomen
After Yosii R, 1965 Fig.36A
Dicyrtomina minuta from the UK
2005 © CEH.
Dicyrtomina minuta from Sweden
2005 © Hall, K.
Dicyrtomina minuta from Sweden
2006 © Hall, K.
Dicyrtomina minuta
2006.06.27 © Fauskanger, I.
Dicyrtomina minuta from the UK
2007 © Cornwall, N.J.
Dicyrtomina minuta from the UK
2007 © Cornwall, N.J.
Dicyrtomina minuta from the UK
2007.02.14 © Botting, J.
Dicyrtomina minuta from the UK
2008.10.14 © Talbot, M.
Dicyrtomina minuta from the UK
2009.02.21 © Ransom, T.
Dicyrtomina minuta from the UK
2010.01.10 © Henton-Jones, D.
Dicyrtomina minuta from the UK
2011.11.12 © Henton-Jones, D.
Dicyrtomina minuta from the UK
2011.11.27 © Jones, A.
Dicyrtomina minuta ♂ from the UK
2012.01.07 © Henton-Jones, D.
Dicyrtomina minuta from Norway
2012.05.23 © Fjellheim, K.
Dicyrtomina minuta from the UK
2012.11.02 © Jones, A.
Dicyrtomina minuta from the UK
Antenna
2012.12.01 © Phillips, E.
Dicyrtomina minuta from the UK
2013.01.19 © Holt, H. & A.
Dicyrtomina minuta ♂ from the UK
2012.12.23 © Henton-Jones, D.
Dicyrtomina minuta from Holland
2013.10.26 © van Duinen, J.
Dicyrtomina minuta ♂ from Holland
2013.12.18 © van Duinen, J.
Dicyrtomina minuta ♂ from Holland
2013.12.18 © van Duinen, J.
Dicyrtomina minuta from Holland
Posterior dorsal pigmentation patch
2014.09.20 © van Duinen, J.
Dicyrtomina minuta from the UK
2014.11.29 © Henton-Jones, D.
Dicyrtomina minuta from Germany
2 dark round touching spots at base of antennae
2015.08.08 © Kluck, A.
Dicyrtomina minuta from the UK
Without median interocular/interantennal stripe
2016.02.11 © Horton, M.
Dicyrtomina minuta ♀ from Belgium
2017.08.27 © Huskens, M.-L.
Dicyrtomina minuta from Belgium
2017.08.28 © Huskens, M.-L.
Dicyrtomina minuta from Belgium
Interocular stripe pale
2017.08.28 © Huskens, M.-L.
Dicyrtomina minuta from Belgium
2 dark round touching spots at base of antennae
2017.08.28 © Huskens, M.-L.
Dicyrtomina minuta from the UK
Interantennal spot
Interocular spot
2017.12.16 © O'Neill, N.
Dicyrtomina minuta from the UK
Interantennal spot
Interocular spot
2019.01.07 © Cann, A.J.
Dicyrtomina minuta from France
2024.01.28 © Quintin, C.
Dicyrtomina minuta from France
2024.01.28 © Quintin, C.





Species murphyi (Delamare Deboutteville, C et Massoud, Z, 1965:99)

Dicyrtoma murphyi from Angola
After Delamare Deboutteville C & Massoud Z, 1965 Planche I Fig.F
            Syn.: Dicyrtoma murphyi Delamare Deboutteville, C et Massoud, Z, 1964:99 u


Species nigra Bretfeld in Bretfeld, Poliakov & Broza, 2000

Species novazealandica Salmon, JT, 1941:419

            Syn.: Dicyrtomina nova-zealandica(sic) Salmon, JT, 1941:419 IOS
Dicyrtomina novazealandica from New Zealand
2009 © Te Papa Museum

            Syn.: Dicyrtomina spiculata Salmon, JT, 1943:11
Dicyrtomina spiculata from New Zealand
After Salmon JT, 1943 Pl.1 Fig.10
Dicyrtomina spicutalata ssp. spiculata from New Zealand
2009 © Te Papa Museum

            Syn.: Dicyrtomina spiculata ssp. spadica Salmon, JT, 1944:182
Dicyrtomina spicutalata ssp. spadica from New Zealand
2009 © Te Papa Museum




             Syn.: Dicyrtomina nova-zealandia(sic) Salmon, JT, 1943:10 ISS
             Syn.: Dicyrtomina nova-zealandica(sic) Salmon, JT, 1946:59 ISS
             Syn.: Dicyrtomina nova-zealandica(sic) Stach, J, 1957:37 ISS


Dicyrtomina novazealandica from New Zealand
2014.02.23 © Murray, A.
Dicyrtomina novazealandica from New Zealand
2014.02.23 © Murray, A.
Dicyrtomina novazealandica from New Zealand
2014.02.23 © Murray, A.



Species ochrea Womersley, H, 1939:261

Species ornata (Nicolet, H, 1842:83) Collinge & Shoebotham, 1910, B

            Syn.: Podura (Smynthurus(sic)) signata Latreille, X in Cuvier, G, 1817:162
             Syn.: Dicyrtoma Signatus(sic) Bourlet, C, 1845:149 =? ISS
            Syn.: Smynthurus(sic) Coulonii(sic) Nicolet, H, 1842:84 IOS
Smynthurus(sic) Coulonii(sic) from Switzerland
After Nicolet, H, 1842 Pl.9 fig.12

             Syn.: Smynthurus(sic) ornatus var. Coulonii(sic) Nicolet, H, 1842:84 ? ISS
            Syn.: Smynthurus(sic) Coulonii(sic) Oken, 1842 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Gervais in Walckenaër, 1844 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Lucas, 1844 IOS
            Syn.: Sminthurus Coulonii(sic) Bourlet, C, 1845:68 IOS
             Syn.: Dicyrtoma dorsimaculata var. Coulonii(sic) Bourlet, C, 1845:68 ISS
            Syn.: Smynthurus(sic) Coulonii(sic) Nicolet, H, 1847:359 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Elditt, HL, 1854:40 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Kolenati, FA, 1858:134 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Olfers, 1862 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Lubbock, 1873 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Parona, 1876 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) von Dalla Torre, 1888 IOS
            Syn.: Smynthurus(sic) Coulonii(sic) Schött, 1893 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Reuter, OM, 1895:5 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Reuter, OM, 1895:15 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Schäffer, 1896 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Carl, 1899 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Carl, 1899 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Carpenter & Evans, 1899 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Kieffer, 1900 IOS
            Syn.: Papirius minutus var. Coulonii(sic) Carl, 1901 IOS
             Syn.: Dicyrtomina minuta-couloni(sic) Handschin, 1925 ISS
             Syn.: Dicyrtomina minuta var. coulonii(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
            Syn.: Smynthurus(sic) couloni Palissa, 1964 partim
Dicyrtomina ornata from the UK
Three different instars
2006 © Valentine, Br.
Dicyrtomina ornata from the UK
2006 © Valentine, Br.
Dicyrtomina ornata from the UK
2006 © Valentine, Br.
Dicyrtomina ornata from the UK
2006 © Valentine, Br.
Dicyrtomina ornata from the UK
2006 © Valentine, Br.
Dicyrtomina ornata from France
2006 © Locasso, S.
Dicyrtomina ornata from France
2006 © Locasso, S.
Dicyrtomina ornata from the UK
2006 © Valentine., B
Dicyrtomina ornata from Germany
2007 © Stanjek, G.H.
Dicyrtomina ornata from the UK
2007 © Valentine, Br.
Dicyrtomina ornata from the UK
2007 © Valentine, Br.
Dicyrtomina ornata from the UK
2007 © Valentine, Br.
Dicyrtomina ornata from France
2007.04.14 © Lebeaux, P.
Dicyrtomina ornata from the UK
2007.11.21 © Valentine, Br.
Dicyrtomina ornata from Holland
2007.11.22 © Baas, A.H.
Dicyrtomina ornata from the UK
2007.12.05 © Barton, T.
Dicyrtomina ornata from the UK
2007.12.15 © Valentine, Br.
Dicyrtomina ornata from the UK
2007.12.16 © Barton, T.
Dicyrtomina ornata from the UK
2007.12.16 © Barton, T.
Dicyrtomina ornata from the UK
2008.01.12 © Barton, T.
Dicyrtomina ornata from the UK
2008.01.12 © Barton, T.
Dicyrtomina ornata from the UK
2008.01.13 © Valentine, Br.
Dicyrtomina ornata from the UK
2008.01.18 © Valentine, Br.
Dicyrtomina ornata from France
Drinking, with eversed collophore vesicles
2008.04.10 © Lebeaux, P.
Dicyrtomina ornata from the UK
2009.02.19 © Barton, T.
Dicyrtomina ornata from Belgium
2012.12.29 © Louvigny, R.
Dicyrtomina ornata from Germany
Cephalic areae verticalis and parietalis
2014.02.23 © Beck, K.
"Discotomina" ornata from the UK
Shake that ass ;-)
2014.11.29 © Horton, M.
Dicyrtomina ornata from Germany
Abdominal posterior patch
2015.01.14 © Landgraf, A.
Dicyrtomina ornata from Germany
Ant.3 apically annulated
2015.01.14 © Landgraf, A.
Dicyrtomina ornata from the UK
2015.11.08 © Valentine, Br.
Dicyrtomina ornata from France
2017.02.11 © Garcelon, P.
Dicyrtomina ornata from Belgium
2017.12.06 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
2018.12.19 © Huskens, M.-L.
Dicyrtomina ornata from France
2018.12.19 © Picard, J.

            Syn.: Dicyrtomina ornata f. couloni Janssens F, 2019
Dicyrtomina ornata f. couloni from France
Grazing
2020.11.15 © Faucheron, A.
Dicyrtomina minuta f. couloni from the USA
Extended furca
Everted collophore vesicles
2021.01.18 © Hendrix, S.

            Syn.: Dicyrtomina ornata f. couloni sf. stachi Janssens F, 2019
            Syn.: Dicyrtomina ornata Stach, J, 1957:51
Dicyrtomina ornata from Poland
After Stach J, 1957 Pl.II Fig.2

            Syn.: Dicyrtomina ornata f. couloni sf.4 Janssens F, 2019
Dicyrtomina ornata from the UK
2009.03.10 © Cole, M.
Dicyrtomina ornata from the UK
2009.03.10 © Cole, M.
Dicyrtomina ornata from Belgium
2016.12.22 © Janssens, F.
Dicyrtomina ornata f. couloni from Germany
2019.01.10 © Kluck, A.
Dicyrtomina minuta f. couloni ♀ from the USA
2020.01.14 © Scholnick, S.
Dicyrtomina minuta f. couloni ♀ from the USA
Ventral aspect
2020.01.14 © Scholnick, S.
Dicyrtomina minuta f. couloni ♀ from the USA
Collophore
2020.01.14 © Scholnick, S.

            Syn.: Dicyrtomina ornata f. couloni sf.4a Janssens F, 2021
Dicyrtomina ornata f. couloni ♀ from Belgium
2021.01.02 © Willems, J.

            Syn.: Dicyrtomina ornata f. couloni sf.5 Janssens F, 2019
Dicyrtomina ornata f. couloni from France
2019.01.19 © Garcelon, P.

            Syn.: Dicyrtomina ornata f. couloni sf.6 Janssens F, 2021
Dicyrtomina ornata f. couloni ♂ from Belgium
2021.01.02 © Willems, J.

             Syn.: Dicyrtoma dorsi-maculata(sic) Gervais in Walckenaër, 1844 ISS
             Syn.: Dicyrtoma dorsi-maculata(sic) Lucas, 1844 ISS
             Syn.: Dicyrtoma dorsi-maculata(sic) Nicolet, H, 1847:361 ISS
            Syn.: Dicyrtoma dorsimaculata var. maculata Bourlet, C, 1845:149
            Syn.: Sminthurus viridis Templeton, 1834
            Syn.: Papirius Saundersii(sic) Lubbock, 1862 IOS
            Syn.: Dicyrtomina variabilis Maynard, EA, 1951:278
Dicyrtomina variabilis from the USA
After Maynard, E.A., 1951 Pl.XXIV Fig.432
Dicyrtomina variabilis from the USA
After Maynard, E.A., 1951 Pl.XXIV Fig.433

            Syn.: Papirius carpenteri Collinge, WE et Shoebotham, JW, 1909:11
Papirius carpenteri from the UK
Ant. lateral; ant.4 ventral
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.8,9
Papirius carpenteri from the UK
Metafoot complex
Note: unguis with tunica
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.10
Papirius carpenteri from the UK
Left eyepatch: 6 large + 2 small ocelli
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.11
Papirius carpenteri from the UK
Mucro dentated with blunt teeth
After Collinge WE & Shoebotham JW, 1909 Pl.III Fig.12

            Syn.: Dicyrtomina labellei Maynard, EA, 1951:279
Dicyrtomina labellei from the USA
After Maynard EA, 1951 Pl.XXIV Fig.420
Dicyrtomina ornata f. labellei from Sweden
2005 © Hall, K.
Dicyrtomina ornata f. labellei from the UK
2006 © Cornwall, N.J.
Dicyrtomina ornata f. labellei from the UK
2006 © Cornwall, N.J.
Dicyrtomina ornata f. labellei from the UK
2006 © Cornwall, N.J.
Dicyrtomina minuta f. labellei from the USA
2007.06.12 © Murray, T.
Dicyrtomina ornata f. labellei from the UK
2007.12.23 © Campbell, A.
Dicyrtomina minuta f. labellei from the USA
2008.11.08 © Thombs, D.
Dicyrtomina minuta f. labellei from the USA
2008.11.15 © Justis, S.
Dicyrtomina minuta f. labellei from the USA
2008.11.22 © Justis, S.
Dicyrtomina minuta f. labellei from the USA
Pre-jump position: waving frontlegs
2011.12.25 © Justis, S.
Dicyrtomina ornata f. labellei from Germany
With eversed tube-like vesicle of collophore
2014.10.11 © Kluck, A.
Dicyrtomina ornata f. labellei from Belgium
2018.12.23 © Huskens, M.-L.
Dicyrtomina ornata f. labellei from Belgium
2019.10.20 © Huskens, M.-L.
Dicyrtomina minuta f. labellei from the USA
2019.12.14 © Carr, J.F.
Dicyrtomina minuta f. labellei from the USA
2019.12.14 © Carr, J.F.
Dicyrtomina minuta f. labellei from Canada
2023.12.06 © Chans, J.
Dicyrtomina minuta f. labellei from Canada
2023.12.06 © Chans, J.
Dicyrtomina minuta f. labellei from Canada
2023.12.06 © Chans, J.


            Syn.: Dicyrtomina minuta f. ornata sensu Janssens, 2010
Remark by Janssens, F. 2010.01.24: Awaiting a formal revision ornata lookalike specimens from Northern America are recorded as minuta f. ornata in line with Christiansen & Bellinger (1980, 1998) but tentatively considered as synonymous with ornata.

Dicyrtomina minuta f. ornata from the USA
Three different colour forms
2006 © McClarin, J.
Dicyrtomina minuta f. ornata from the USA
2006 © Murray, T.
Dicyrtomina minuta f. ornata from the USA
2007 © Moorehead, C.
Dicyrtomina minuta f. ornata from the USA
2007.03.13 © Moorehead, C.
Dicyrtomina minuta f. ornata from Canada
2007.11.18 © McCann, S.
Dicyrtomina minuta f. ornata from Canada
2007.12.09 © Marsman, I.
Dicyrtomina minuta f. ornata from Canada
2007.12.17 © McCann, S.
Dicyrtomina minuta f. ornata from the USA
2008.02.23 © Bradford, A.
Dicyrtomina minuta f. ornata from the USA
2008.02.23 © Bradford, A.
Dicyrtomina minuta f. ornata from Canada
2008.11.27 © McCann, S.
Dicyrtomina minuta f. ornata from the USA
2008.12.07 © Justis, S.
Dicyrtomina minuta f. ornata from the USA
2008.12.13 © Bradford, A.
Dicyrtomina minuta f. ornata from the USA
2009.02.20 © Bergen, L.
Dicyrtomina minuta f. ornata from the USA
2009.08.12 © Janssens, F.
Leg. Bradford, A., 2009.03.29
Dicyrtomina minuta f. ornata from the USA
2009.08.12 © Janssens, F.
Leg. Bradford, A., 2009.03.29
Dicyrtomina minuta f. ornata from the USA
2009.11.16 © Bergen, L.
Dicyrtomina minuta f. ornata from the USA
2009.11.20 © Thombs, D.
Dicyrtomina minuta f. ornata from Canada
2009.12.20 © McCann, S.
Dicyrtomina minuta f. ornata from the USA
White feet mark pre-moulting phase
2012.01.14 © Justis, S.
Dicyrtomina minuta f. ornata from the USA
White feet mark pre-moulting phase
2012.01.14 © Justis, S.
Dicyrtomina minuta f. ornata from the USA
Mottled pigmentation mark pre-moulting phase
2012.01.14 © Justis, S.
Dicyrtomina minuta f. ornata from Canada
2012.02.01 © Hochhalter, R.
Dicyrtomina minuta f. ornata from the USA
2022.01.18 © Charny, V.
Dicyrtomina minuta f. ornata from the USA
2022.01.18 © Charny, V.

            Syn.: Dicyrtoma minuta ssp. meridionalis Stach, J, 1930:76 partim
            Syn.: Dicyrtomina ornata f. meridionalis sensu Janssens, 2023
Dicyrtomina ornata f. meridionalis from Italy
2013.01.13 © Consolo, M.
Dicyrtomina ornata f. meridionalis from Italy
2013.01.13 © Consolo, M.
Dicyrtomina ornata f. meridionalis from France
2019.01.19 © Joachim, J.
Dicyrtomina ornata f. meridionalis from France
2019.11.30 © Joachim, J.
Dicyrtomina ornata f. meridionalis from France
2019.11.30 © Joachim, J.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.20 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.20 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
Abd.6 with 3M+4N setae
M-setae in an angle
2023.12.20 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.21 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.21 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.21 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2023.12.21 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
Preparing to moult
2023.12.22 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
Preparing to moult
2023.12.22 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
Interocular pattern
2023.12.22 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
2024.01.02 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis from Belgium
Distinct transverse pattern on vertex
2024.01.02 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis ♀ from Belgium
With dark face
2024.01.03 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis ♀ from Belgium
Typical pattern
With long transverse bars
2024.01.03 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis ♀ from Belgium
Dorsal anal valve
With 7 posterior macrosetae a3-a2-a1-a0-a1-a2-a3
2024.01.03 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis ♀ from Belgium
Dorsal anal valve
With 3 M-setae M'-M-M' in triangle
With 4 N setae N'-N-N-N'
2024.01.03 © Huskens, M.-L.
Dicyrtomina ornata f. meridionalis ♀ from Belgium
Hindfoot
Unguis thick with tunica
Unguiculus with distinct inner tooth
2024.01.03 © Huskens, M.-L.




Smynthurus(sic) ornatus from Switzerland
After Nicolet, 1842 Pl.9 Fig.11
             Syn.: Papirius ornato(sic) Reuter, OM, 1876:85 ISS
             Syn.: Dicyrtomina minuta var. ornatus(sic) Turk, 1933 ISS
             Syn.: Dicyrtomina minuta-ornata(sic) Handschin, E, 1942:445 ISS
Dicyrtomina ornata from the UK
Foot complex, claw with tunica
2004 © Hopkin, S.P.
Dicyrtomina ornata juv. from the UK
2007 © Cornwall, N.J.
Dicyrtomina ornata juv. from the UK
2007 © Cornwall, N.J.
Dicyrtomina ornata juv. from the UK
2007 © Cornwall, N.J.
Dicyrtomina ornata from the UK
2007 © Valentine., B
Dicyrtomina ornata from the UK
2007 © Valentine., B
Dicyrtomina ornata from Belgium
2007 © Vuijlsteke, M.
Dicyrtomina ornata from the UK
2007.03.23 © Valentine., B
Dicyrtomina ornata from the UK
2007.10.30 © Campbell, A.
Dicyrtomina ornata from the UK
Anal valves and female genital plate
2008.01.23 © Valentine., B
Dicyrtomina ornata from the UK
Anal valves and female genital plate
2008.01.25 © Valentine., B
Dicyrtomina ornata from the UK
2008.01.27 © Valentine., B
Dicyrtomina ornata from Belgium
2008.02.11 © Haentjens, R.
Dicyrtomina ornata from the UK
2008.02.29 © Valentine., B
Dicyrtomina ornata from Belgium
Bilaterally serrated mucro
2008.09.01 © Janssens, F.
Dicyrtomina ornata juv. from France
2008.11.13 © Lavoué, Br.
Dicyrtomina ornata from the UK
Male genital papilla
2008.11.22 © Valentine., B
Dicyrtomina ornata from Belgium
2008.12.15 © Haentjens, R.
Dicyrtomina ornata from the UK
Male genital papilla
2009.01.13 © Valentine., B
Dicyrtomina ornata ♂ from the UK
2009.11.18 © Valentine., B
Dicyrtomina ornata from the UK
2009.12.04 © Valentine., B
Dicyrtomina ornata from the UK
Antenna posterior aspect
2009.12.06 © Valentine., B
Dicyrtomina ornata from the UK
Spermatophore with consumed spermdrop
2009.12.10 © Valentine., B
Dicyrtomina ornata from the UK
Left composed eye
2009.12.18 © Valentine., B
Dicyrtomina ornata from the UK
With white feet
2009.12.24 © Cole, M.
Dicyrtomina ornata (bar=2µm)
Hexagonal epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.6A
Dicyrtomina ornata from France
With laterally deformed abdomen
2012.02.18 © Riffaud, E.
Dicyrtomina ornata ♀ from the UK
Female sub-anal appendage
2012.03.27 © Murray, A.
Dicyrtomina ornata ♂ from the UK
2012.07.30 © Murray, A.
 
Dicyrtomina ornata juv. from the UK
2012.08.19 © Murray, A.
Dicyrtomina ornata from Portugal
With dorsally deformed abdomen
2012.10.16 © Murray, A.
Dicyrtomina ornata from Portugal
Hind leg
2012.10.16 © Murray, A.
Dicyrtomina ornata from Spain
2012.11.19 © Herrera, J.M.
Dicyrtomina ornata from Spain
Dorso-lateral pigmentation
2012.11.19 © Herrera, J.M.
Dicyrtomina ornata juv. from the UK
2012.11.22 © Phillips, E.
Dicyrtomina ornata ♂ from France
Raised male genital papilla
2012.11.25 © Sandra.
Dicyrtomina ornata from Spain
Anogenital segments
2012.11.25 © Herrera, J.M.
Dicyrtomina ornata from Spain
2012.11.25 © Herrera, J.M.
Dicyrtomina ornata juv. from the UK
2012.12.04 © Phillips, E.


             Syn.: Dicyrtomina ornate(sic) Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:71 ISS
Dicyrtomina ornata from the UK
Cleaning antennae with a drop of mouth secretion
2013.11.03 © Garlick, T.
Dicyrtomina ornata ♀ from Holland
Ventral aspect: furca and collophore
2013.11.04 © van Duinen, J.
Dicyrtomina ornata ♀ from the UK
Ready to moult: white feet and mottled lateral pigmentation
2013.11.14 © Nurcombe, E.
Dicyrtomina ornata from Belgium
Facial spines
2013.12.17 © De Kesel, A.
Dicyrtomina ornata from Holland
Dark specimen
2014.02.06 © van Someren, M.
Dicyrtomina ornata from the UK
2014.02.23 © Garlick, T.
Dicyrtomina ornata from Germany
Two strong spines in frontal area
2014.03.02 © Beck, K.
Dicyrtomina ornata from Germany
Indistinct change of colour at joint of 2nd to 3rd antennal segment
2014.04.16 © Beck, K.
Dicyrtomina ornata from Germany
Eversed tube-like vesicle of collophore
2014.10.12 © Kluck, A.
Dicyrtomina ornata from Germany
Eversed tube-like vesicles of collophore
2014.11.01 © Kluck, A.
Dicyrtomina ornata from Germany
Annulated tube-like vesicle of collophore
2014.11.01 © Kluck, A.
Dicyrtomina ornata from Germany
Spiky setae on eyebrows and in eyepatch
2014.11.08 © Kluck, A.
Dicyrtomina ornata from Germany
Smiling ;-)
2014.11.08 © Kluck, A.
Dicyrtomina ornata from Germany
Unguiculus with broad lamella and distinct tooth
2014.11.22 © Kluck, A.
Dicyrtomina ornata from the UK
Ready to moult: white feet
2014.12.15 © Phillips, E.
Dicyrtomina ornata from the UK
With everted tubelike collophore vesicles
2014.12.19 © Horton, M.
Dicyrtomina ornata ♂ from Belgium
Pale cheek adult
2015.12.01 © Huskens, M.-L.
Dicyrtomina ornata ♀ from France
Form with long serrated butt patch
2016.12.06 © Garcelon, P.
Dicyrtomina ornata ♂ & ♀ from France
Female with long serrated butt patch
2016.12.06 © Garcelon, P.
Dicyrtomina ornata ♂ from Belgium
Pale cheek subadult
2017.01.08 © Huskens, M.-L.
Dicyrtomina ornata ♂ from Belgium
Non-reproductive genital papilla (*)
2017.01.08 © Huskens, M.-L.
Dicyrtomina ornata ♂ from Belgium
2017.01.30 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Eversed telescopic collophore vesicle
2017.02.03 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Collophore vesicle apically warted
2017.02.03 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Eversed collophore vesicle
2017.11.16 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
2017.12.09 © Huskens, M.-L.
Dicyrtomina ornata from France
2018.01.28 © Garcelon, P.
Dicyrtomina ornata from France
2018.01.28 © Garcelon, P.
Dicyrtomina ornata from France
2018.01.28 © Garcelon, P.
Dicyrtomina ornata from France
With saundersi-like dorsal patch
2018.03.13 © Garcelon, P.
Dicyrtomina ornata from Belgium
Wax producing tormogene cells enlarged
2018.09.09 © Huskens, M.-L.
Dicyrtomina ornata from Portugal
2018.12.28 © Gomes, M.
Dicyrtomina ornata from Belgium
2019.01.05 © Huskens, M.-L.
Dicyrtomina ornata from Germany
Spermatophore
2019.02.27 © Fabian, Be.
Dicyrtomina ornata from Germany
Stalk of spermatophore penetrates spermdrop
2019.02.27 © Fabian, Be.
Dicyrtomina ornata from Belgium
2019.12.05 © Huskens, M.-L.
Dicyrtomina ornata from Portugal
2020.01.25 © Delgado, F.
Dicyrtomina ornata ♂ from France
2020.11.21 © Fiacre, R.
Dicyrtomina ornata from the UK
3 spots mark anterior tentorial pit
2020.11.23 © Valentine, Br.
Dicyrtomina ornata from Belgium
2020.12.02 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Forward directed bothriotrichum A on papilla
2020.12.02 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Caudal aspect
2020.12.02 © Huskens, M.-L.
Dicyrtomina ornata ♂ from Belgium
2020.12.10 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Spermatophore
Stalk exits spermdrop apically
2020.12.10 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Spermdrop lower half not transparant
2020.12.10 © Huskens, M.-L.
Dicyrtomina ornata juv. from Belgium
2020.12.17 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
2020.12.17 © Huskens, M.-L.
Dicyrtomina ornata from Belgium
Dorsal macrosetae CC-EE spinelike
2020.12.17 © Huskens, M.-L.
Dicyrtomina ornata from the UK
Waving with frontlegs
Considering an escape jump
2021.01.01 © Barton, T.
Dicyrtomina ornata from France
Everted tube-like collophore vesicle
2023.11.19 © Clerc, O.





Species platensis (Merlo & Najt, 1974) Mari Mutt & Bellinger, 1990:145

Species saundersi (Lubbock, J, 1862:438) Stach, 1922, B

            Syn.: Papirius Saundersii(sic) Lubbock, J, 1862:438 IOS
Papirius Saundersii(sic) from the UK
After Lubbock J, 1862 Pl.XLV Fig.43


            Syn.: Smynthurus(sic) viridis Templeton, 1834
            Syn.: Smynthurus(sic) Coulonii(sic) Nicolet, H, 1842:84 IOS
            Syn.: Smynthurus(sic) smynthure croisé Gervais in Walckenaër, 1844
            Syn.: Papirius Saundersii(sic) var. a Lubbock, J, 1862:438
            Syn.: Papirius Saundersii(sic) var. b Lubbock, J, 1862:438
            Syn.: Smynthurus(sic) ornatus Lubbock, J, 1870:281
            Syn.: Papirius ornatus Lubbock, J, 1873:123
Papirius ornatus from the UK
After Lubbock J, 1873 Pl.9

            Syn.: Papirius ornatus Parona, 1885
            Syn.: Papirius ornatus Uzel, 1890
            Syn.: Papirius ornatus Schäffer, 1896
            Syn.: Papirius minutus var. ornata Börner, 1901
            Syn.: Papirius minutus var. ornata Krausbauer, 1902
            Syn.: Papirius minutus var. ornatus Schött, 1902
            Syn.: Papirius minutus var. ornata Ågren, 1903
            Syn.: Papirius minutus var. ornata Collinge & Shoebotham, 1910
            Syn.: Papirius minutus var. ornata Ionesco, 1915
            Syn.: Papirius minutus var. ornata Denis, 1921
            Syn.: Dicyrtomina minuta f. ornata Handschin, E in Dahl, F, 1929:124
            Syn.: Dicyrtoma minuta ab. ornata Stach, J, 1930:79
            Syn.: Dicyrtomina saundersi var. ornata Stach, J, 1957:56,is
            Syn.: Papirius ornatus var. cordatus Parona, 1888
            Syn.: Dicyrtoma minuta Stach, J, 1930:76
            Syn.: Dicyrtoma minuta f. principalis Stach, J, 1930:81
            Syn.: Dicyrtoma minuta ssp. meridionalis Stach, J, 1930:76 partim
            Syn.: Dicyrtoma minuta ssp. meridionalis f.1 Stach, J, 1930:76
Dicyrtoma minuta ssp. meridionalis from Spain
Incomplete dorsal superanal spot
After Stach J, 1930 Abb.XV.15
Dicyrtomina saundersi f. meridionalis from the UK
2009.03.05 © Barton, T.
Dicyrtomina saundersi f. meridionalis from the UK
2009.03.05 © Barton, T.
Dicyrtomina saundersi f. meridionalis from the UK
2009.03.05 © Barton, T.
Dicyrtomina saundersi f. meridionalis from Italy
2012.11.22 © Consolo, M.
Dicyrtomina saundersi f. meridionalis from France
2019.11.09 © Joachim, J.


            Syn.: Dicyrtoma minuta ssp. meridionalis f.2 Stach, J, 1930:76
Dicyrtoma minuta ssp. meridionalis from Spain
Broad dorsal superanal spot
After Stach J, 1930 Abb.XV.16
Dicyrtomina saundersi f. meridionalis from the UK
2007.12.23 © Kilford., B
Dicyrtomina saundersi f. meridionalis from the UK
2008.01.05 © Kilner, M.
Dicyrtomina saundersi f. meridionalis from the UK
2008.01.29 © Kilner, M.
Dicyrtomina saundersi f. meridionalis from the UK
2015.11.22 © Horton, M.
Dicyrtomina saundersi f. meridionalis from the UK
2015.12.11 © Valentine., B
Dicyrtomina saundersi f. meridionalis from the UK
2016.01.25 © Horton, M.
Dicyrtomina saundersi f. meridionalis from Germany
2016.02.28 © Kluck, A.
Dicyrtomina saundersi f. meridionalis from the UK
2017.11.30 © Horton, M.
Dicyrtomina saundersi f. meridionalis from France
2018.11.01 © Garcelon, P.
Dicyrtomina saundersi f. meridionalis from France
Dorsal posterior patch
2018.11.01 © Garcelon, P.
Dicyrtomina saundersi f. meridionalis from France
Preparing to moult
2018.12.10 © Garcelon, P.
Dicyrtomina saundersi f. meridionalis from France
Preparing to moult
2018.12.10 © Garcelon, P.
Dicyrtomina saundersi f. meridionalis from the UK
2019.02.11 © Barton, T.
Dicyrtomina saundersi f. meridionalis from the UK
2019.02.28 © Henton-Jones, D.
Dicyrtomina saundersi f. meridionalis from Belgium
2020.12.23 © Huskens, M.-L.


            Syn.: Dicyrtoma minuta ssp. meridionalis f.3 Stach, J, 1930:76
Dicyrtoma minuta ssp. meridionalis from Spain
Typical dorsal superanal spot
After Stach J, 1930 Abb.XV.17

            Syn.: Dicyrtoma minuta ssp. meridionalis f.4 Stach, J, 1930:76
Dicyrtoma minuta ssp. meridionalis from Spain
Confluent dorsal superanal spot
After Stach J, 1930 Abb.XV.18


            Syn.: Dicyrtoma minuta ab. pseudosaundersi Stach, J, 1930:81
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.01 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.01 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.01 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.03 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.04 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.04 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.06 © Barton, T.
Dicyrtomina minuta nr ab. pseudosaundersi from La Palma
2012.12.06 © Barton, T.


            Syn.: Dicyrtoma minuta ab. scalaris Stach, J, 1930:81
Dicyrtomina saundersi var. scalaris from Poland
After Stach, J, 1957 Pl.II Fig.4.

            Syn.: Dicyrtomina minuta Ponge, 1980
            Syn.: Dicyrtomina minuta f. saundersi sensu Janssens, 2010
Remark by Janssens, F. 2010.01.24: Awaiting a formal revision saundersi lookalike specimens from Northern America are recorded as minuta f. saundersi in line with Christiansen & Bellinger (1980,, 1998) but tentatively considered as synonymous with saundersi.

Dicyrtomina minuta f. saundersi from the USA
2007 © Moorehead, C.
Dicyrtomina minuta f. saundersi from Canada
2007.12.05 © Marsman, I.
Dicyrtomina minuta f. saundersi from the USA
2007.12.25 © Moorehead, C.
Dicyrtomina minuta f. saundersi from the USA
2008.12.16 © Paige.
Dicyrtomina minuta f. saundersi from Canada
2013.01.11 © Kyron, T.F.
Dicyrtomina minuta f. saundersi from the USA
2016.11.17 © Johnson, S.T.




             Syn.: Papirius Saundersii(sic) Lubbock, J, 1862:589 ISS
             Syn.: Papirius Saundersii(sic) Packard, AS, 1869:624 ISS
             Syn.: Papirius Saundersii(sic) Lubbock, J, 1870:281 ISS
             Syn.: Papirius Saundersii(sic) Packard, AS, 1872:624 ISS
             Syn.: Papirius Saundersii(sic) Packard, AS, 1889:624 ISS
             Syn.: Papirius saundersii(sic) Krausbauer, 1901 ISS
             Syn.: Papirius Saundersii(sic) Krausbauer, 1902 ISS
             Syn.: Dicyrtomina Saundersi(sic) Stach, 1922 ISS
             Syn.: Dicyrtoma Saundersi(sic) Stach, J, 1930:76 ISS
Dicyrtomina saundersi from the UK
2005 © Hopkin, S.P.
Dicyrtomina saundersi from Sweden
2005 © Hall, K.
Dicyrtomina saundersi from Sweden
2005 © Hall, K.
Dicyrtomina saundersi from the UK
2006 © Valentine., B
Dicyrtomina saundersi ♂ from the UK
2006 © Jones, D.
Dicyrtomina saundersi ♂ from the UK
2006 © Jones, D.
Dicyrtomina saundersi from Sweden
2006 © Hall, K.
Dicyrtomina saundersi from Sweden
2006 © Hall, K.
Dicyrtomina saundersi from the UK
2006 © Barton, T.
Dicyrtomina saundersi from France
2006 © Lavoué., B
Dicyrtomina saundersi from the UK
2006.12.16 © Barton, T.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Tonsbeek, M.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Cornwall, N.J.
Dicyrtomina saundersi from the UK
2007 © Barton, T.
Dicyrtomina saundersi from Belgium
2007 © Vuijlsteke, M.
Dicyrtomina saundersi from the UK
2007.02.13 © Jones, D.
Dicyrtomina saundersi from the UK
2007.02.18 © Barton, T.
Dicyrtomina saundersi from the UK
2007.03.18 © Cornwall, N.J.
Dicyrtomina saundersi from Belgium
2007.04.14 © Vuijlsteke, M.
Dicyrtomina saundersi from Norway
2007.10.26 © Fauskanger, I.
Dicyrtomina saundersi from the UK
2007.11.08 © Valentine., B
Dicyrtomina saundersi from the Netherlands
2007.11.10 © Baas, A.H.
Dicyrtomina saundersi from the UK
2007.11.11 © Kilner, M.
Dicyrtomina saundersi from the UK
2007.12.14 © Valentine., B
Dicyrtomina saundersi from the UK
2007.12.16 © Kilford., B
Dicyrtomina saundersi from the UK
2007.12.23 © Kilner, M.
Dicyrtomina saundersi from the UK
2007.12.24 © Barton, T.
Dicyrtomina saundersi from the UK
2007.12.29 © Campbell, A.
Dicyrtomina saundersi from the UK
2008.01.01 © Campbell, A.
Dicyrtomina saundersi from the UK
2008.01.08 © Edwards, A.
Dicyrtomina saundersi from the UK
2008.01.12 © Barton, T.
Dicyrtomina saundersi from the UK
2008.01.12 © Barton, T.
Dicyrtomina saundersi from the UK
With phoretic Nematoda
2008.01.14 © Farrell, S.
Dicyrtomina saundersi from the UK
2008.01.20 © Farrell, S.
 
Dicyrtomina saundersi from the UK
2008.01.21 © Rodrigues, T.
Dicyrtomina saundersi from the UK
Escaping
2008.01.23 © Farrell, S.
Dicyrtomina saundersi from the UK
Waving
2008.01.23 © Farrell, S.
Dicyrtomina saundersi from the UK
2008.01.30 © Bantock, T.
Dicyrtomina saundersi from the UK
2008.02.01 © Robertson, A.
Dicyrtomina saundersi from the UK
Front to back grazing technique
2008.02.02 © Farrell, S.
Dicyrtomina saundersi from the UK
Front to back grazing technique
2008.02.05 © Farrell, S.
Dicyrtomina saundersi from the UK
2008.02.07 © Campbell, A.
Dicyrtomina saundersi from the UK
2008.02.07 © Farrell, S.
Dicyrtomina saundersi from Belgium
2008.02.10 © Vuijlsteke, M.
Dicyrtomina saundersi from the UK
2008.02.26 © Oakshott, A.
Dicyrtomina saundersi from the UK
2008.04.30 © Farrell, S.
Dicyrtomina saundersi from the Netherlands
2008.10.31 © Baas, A.H.
Dicyrtomina saundersi from the UK
2008.11.09 © Barton, T.
Dicyrtomina saundersi from the UK
2008.11.10 © Dawes, D.
Dicyrtomina saundersi from France
2008.11.13 © Lavoué., B
Dicyrtomina saundersi from the UK
2008.11.15 © Oakshot, A.
Dicyrtomina saundersi from the UK
2008.11.26 © Ransom, T.
Dicyrtomina saundersi from Germany
2008.12.07 © Peters, J.
Dicyrtomina saundersi from the UK
2008.12.17 © Jeffery, J.
Dicyrtomina saundersi from the UK
2009.03.08 © Valentine., B
Dicyrtomina saundersi from the UK
2009.04.12 © Robertson, A.
Dicyrtomina saundersi from Belgium
2009.09.13 © Wallays, H.
Dicyrtomina saundersi from the UK
2009.11.25 © Ward, T.
Dicyrtomina saundersi from Belgium
2009.12.07 © D'Hert, D.
Dicyrtomina saundersi from the UK
2009.12.09 © Oakshott, A.
Dicyrtomina saundersi from the UK
2009.12.09 © Oakshott, A.
Dicyrtomina saundersi from the UK
Ready to moult: white feet
2009.12.13 © Barton, T.
Dicyrtomina saundersi from the UK
Dorso-lateral tubercle with bothriotrichum
2010.02.01 © Robertson, A.
Dicyrtomina saundersi from Belgium
2010.02.08 © Wallays, H.
Dicyrtomina saundersi from the UK
Raised genital papilla of reproductive male
2011.10.24 © Jones, A.
Dicyrtomina saundersi ♀ from the UK
2011.11.10 © Kilner, M.
Dicyrtomina saundersi ♂ from the UK
2011.11.28 © Jones, A.
Dicyrtomina saundersi from the UK
Prejump posture with uplifted forelegs
2012.01.07 © Barton, T.
Dicyrtomina saundersi ♂ from the UK
2012.01.25 © van Duinen, J.
Dicyrtomina saundersi ♂ from the UK
2012.01.30 © Dargie, S.
Dicyrtomina saundersi from Ireland
With everted collophore vesicles
2012.02.04 © Cluskey, D.
Dicyrtomina saundersi ♂ from the UK
2012.11.10 © Jones, A.
Dicyrtomina saundersi from the UK
Unguiculus with strong exad tooth
2012.11.16 © Phillips, E.
Dicyrtomina saundersi from the UK
Spermatophore with reduced spermdrop
2012.11.20 © Phillips, E.
Dicyrtomina saundersi from the UK
Prejump posture with uplifted front legs
2012.11.26 © Good, P.
Dicyrtomina saundersi from the UK
'Drinking' with exerted collophore vesicles
2012.11.28 © Phillips, E.
Dicyrtomina saundersi ♂ from the UK
2012.11.28 © Nurcombe, E.
Dicyrtomina saundersi ♂ from the UK
2012.11.30 © Phillips, E.
Dicyrtomina saundersi from the UK
2012.12.03 © Valentine., B
Dicyrtomina saundersi from the UK
Distinct change in colour at joint of 2nd to 3rd antennal segment
2012.12.04 © Nurcombe, E.
Dicyrtomina saundersi ♂ from the UK
Feeding instar with reduced genital papilla
2012.12.06 © Phillips, E.
Dicyrtomina saundersi from the UK
Mucro
2012.12.08 © Nurcombe, E.
Dicyrtomina saundersi ♀ from the UK
Composed eye with spiked eyebrow
2012.12.08 © Nurcombe, E.
Dicyrtomina saundersi ♀ from the UK
Female gonopore
2012.12.15 © Barton, T.
Dicyrtomina saundersi from the UK
Colour form with pale dorsum
2012.12.15 © Barton, T.
Dicyrtomina saundersi from the UK
Laterodorsal bothriotrichum A on papilla
2012.12.17 © Phillips, E.
Dicyrtomina saundersi from the UK
Labial palps
2012.12.18 © Phillips, E.
Dicyrtomina saundersi ♂ from the UK
2013.01.05 © Valentine., B
Dicyrtomina saundersi ♂ from the UK
Raised male genital papilla (*)
2013.01.16 © Phillips, E.
Dicyrtomina saundersi from the UK
Grooming using a mouth secreted drop
2013.01.12 © Holt, H. & A.
Dicyrtomina saundersi from the UK
Grooming using a mouth secreted drop
2013.01.12 \© Holt, H. & A.
Dicyrtomina saundersi from the UK
End of moulting
2013.01.27 © Holt, H. & A.
Dicyrtomina saundersi from the UK
Extended mouth parts
2013.02.22 © Phillips, E.
Dicyrtomina saundersi from the UK
Grazing
2013.02.19 © Cobby, P.
Dicyrtomina saundersi ♂ from the UK
2013.11.07 © Nurcombe, E.
Dicyrtomina saundersi from the UK
Moulting
2013.11.08 © Nurcombe, E.
Dicyrtomina saundersi from the UK
Ant.3 with apical annulation
2013.11.10 © Nurcombe, E.
Dicyrtomina saundersi from the UK
Distinct change of colour at the joint of Ant.2 (pale) and Ant.3 (dark)
2013.12.15 © Nurcombe, E.
Dicyrtomina saundersi ♂ from France
2014.01.25 © Sandra.
Dicyrtomina saundersi from Germany
Distinct change in colour at joint of 2nd to 3rd antennal segment
2014.11.01 © Kluck, A.
Sminthurides malmgreni ♀ & Dicyrtomina saundersi ♂ from Holland
Love at first sight ;-)
2014.11.11 © van Duinen, J.
Dicyrtomina saundersi ♂ from the UK
Dark cheek
2014.11.15 © Phillips, E.
Dicyrtomina saundersi from the UK
Dorsal posterior patch in shape of multibarred cross
2014.11.22 © Barton, T.
Dicyrtomina saundersi from the UK
Pre-jump posture with uplifted frontlegs
2014.12.17 © Horton, M.
Dicyrtomina saundersi from the UK
Pre-jump posture with uplifted frontlegs
2014.12.17 © Horton, M.
Dicyrtomina saundersi ♂ from the UK
Pre-jump posture with uplifted frontlegs
2015.01.06 © Horton, M.
Dicyrtomina saundersi from the UK
2015.01.17 © Phillips, E.
Dicyrtomina saundersi from the UK
Caudal aspect
2015.02.21 © Horton, M.
Dicyrtomina saundersi from the UK
Labial palps
2015.03.12 © Phillips, E.
Dicyrtomina saundersi from Germany
With black dew drops containing Hypoxylon spores
2015.11.01 © Kluck, A.
Dicyrtomina saundersi from the UK
Unguis with tunica
2015.11.29 © Henton-Jones, D.
Dicyrtomina saundersi from Germany
Cleaning antenna with drop of saliva
2015.11.15 © Kluck, A.
Dicyrtomina saundersi from the UK
Six unpaired midfacial setae
2015.11.22 © Horton, M.
Dicyrtomina saundersi ♂ from the UK
Distinct genital papilla (*)
2015.11.25 © Phillips, E.
Dicyrtomina saundersi from the UK
Everted tube-like collophore vesicles (^)
Drinking from watersurfacefilm
2015.12.02 © Branch, A.
Dicyrtomina saundersi ♂ from the UK
Checking spermatophore
2015.12.14 © Phillips, E.
Dicyrtomina saundersi from Belgium
2016.01.02 © Janssens, F.
Dicyrtomina saundersi from the UK
2016.01.06 © Gillean.
Dicyrtomina saundersi ♂ from the UK
Checking spermatophore
2016.10.31 © Phillips, E.
Dicyrtomina saundersi ♂ from the UK
Grooming
2016.12.08 © Cooper, P.
Dicyrtomina saundersi ♀ from the UK
Pale cheek
2017.02.01 © Horton, M.
Dicyrtomina saundersi ♂ from the UK
Dark cheek
2017.02.01 © Horton, M.
Dicyrtomina saundersi ♂ & ♀ from the UK
2017.02.06 © Henton-Jones, D.
Dicyrtomina saundersi from Belgium
2017.11.09 © Huskens, M.-L.
Dicyrtomina saundersi from the UK
With phoretic Nematoda
2017.12.07 © Horton, M.
Dicyrtomina saundersi from the UK
With phoretic Nematoda
2017.12.07 © Horton, M.
Dicyrtomina saundersi from the UK
Phoretic Nematoda in face
2017.12.07 © Horton, M.
Dicyrtomina saundersi ♂ from Belgium
Dark body colour
2018.12.19 © Huskens, M.-L.
Dicyrtomina saundersi ♂ from Belgium
Genital papilla with apical circumgenital setae
2018.12.19 © Huskens, M.-L.
Dicyrtomina saundersi ♂ from Belgium
Ventro-caudal aspect
Broad manubrium, genital papilla, anal valves
2019.10.31 © Huskens, M.-L.
Dicyrtomina saundersi ♀ from Belgium
2020.11.24 © Callens, M.
Dicyrtomina saundersi from the UK
Ant.1 spur (*)
2020.12.28 © Jones, S.
Dicyrtomina saundersi ♂ from Belgium
Dark body colour
2022.02.02 © Huskens, M.-L.
Dicyrtomina saundersi ♂ from Belgium
Male genital papilla
2022.02.02 © Huskens, M.-L.
Dicyrtomina saundersi ♀ from Belgium
Ready to moult
Old cuticula regionally released
2024.01.05 © Huskens, M.-L.
Dicyrtomina saundersi ♀ from Belgium
Moulting
2024.01.05 © Huskens, M.-L.
Dicyrtomina saundersi ♀ from Belgium
End of moult
2024.01.05 © Huskens, M.-L.
Dicyrtomina saundersi from the UK
More then 2 midfacial unpaired macrosetae
2024.01.05 © Barton, T.
Dicyrtomina saundersi ♀ from the UK
Dorsal anal valve with 7 macrosetae
a3-a2-a1-a0-a1-a2-a3
2024.02.24 © Barton, T.





Species signata Stach, 1919

Dicyrtomina ornata var. signata from Poland
After Stach J., 1957 Pl.II Fig.3
Dicyrtomina signata from France
2016.11.30 © Garcelon, P.
Dicyrtomina signata from France
2016.11.30 © Garcelon, P.
Dicyrtomina signata from France
2016.12.27 © Garcelon, P.
Dicyrtomina signata from France
2016.12.31 © Garcelon, P.
Dicyrtomina signata from France
2017.02.14 © Garcelon, P.
Dicyrtomina signata from France
2017.02.14 © Garcelon, P.
Dicyrtomina signata from France
2017.02.14 © Garcelon, P.
Dicyrtomina signata from France
Wiggling gut animation
2017.10.07 © Picard, J.
Dicyrtomina signata from France
2017.10.07 © Picard, J.
Dicyrtomina signata from France
Small dorsal posterior patch
2017.10.07 © Picard, J.
Dicyrtomina signata from France
2017.11.03 © Garcelon, P.
Dicyrtomina signata ♂ from France
2017.11.04 © Picard, J.
Dicyrtomina signata juv. from France
2017.11.12 © Tiky, S.
Dicyrtomina signata from France
2018.01.04 © Garcelon, P.
Dicyrtomina signata from France
2018.02.05 © Picard, J.
Dicyrtomina signata from France
2018.03.13 © Garcelon, P.
Dicyrtomina signata from France
Caudal aspect
2018.03.13 © Garcelon, P.
Dicyrtomina nr signata from Belgium
2018.05.19 © Huskens, M.-L.
Dicyrtomina nr signata from Belgium
2019.06.25 © Huskens, M.-L.
Dicyrtomina nr signata ♂ from Belgium
2023.12.29 © Huskens, M.-L.

Species torsolimata Latzel, 1917

Species tuberculata Womersley, 1933

Dicyrtomina tuberculata from Australia
After Womersley H, 1939 Fig.75H
Dicyrtomina tuberculata from Tasmania
2011.05.17 © Daley, A.
Dicyrtomina tuberculata from Tasmania
2011.05.17 © Daley, A.
Dicyrtomina tuberculata from Tasmania
2011.05.17 © Daley, A.
Dicyrtomina tuberculata from Tasmania
2011.07.dd © Zosterops, K.
Dicyrtomina tuberculata from Tasmania
2011.07.dd © Zosterops, K.
Dicyrtomina tuberculata from Australia
2013.08.14 © Flyman.
Dicyrtomina tuberculata from Tasmania
2016.07.16 © Zosterops, K.
Dicyrtomina tuberculata from Australia
2022.08.03 © Dashper, J.

Species turbotti Salmon, 1948

Dicyrtomina turbotti from New Zealand
2014.02.25 © Murray, A.

Species venusta Latzel, R, 1917:251

Species violacea (Krausbauer, 1898) Denis, J-R, 1924:197

            Syn.: Dicyrtomina minuta var. signata Denis, 1925 =?
            Syn.: Dicyrtomina minuta var. signata Handschin, E in Dahl, F, 1929:124
            Syn.: Dicyrtomina signata Salmon, JT, 1964:640


Dicyrtomina minuta f. violacea from the USA
2007.04.22 © Moorehead, C.
Dicyrtomina minuta f. violacea from the USA
2007.04.22 © Moorehead, C.
Dicyrtomina minuta f. violacea from the USA
2007.04.22 © Moorehead, C.
Dicyrtomina minuta f. violacea from the USA
2007.04.22 © Moorehead, C.
Dicyrtomina minuta f. violacea from the USA
2007.04.22 © Moorehead, C.
Dicyrtomina violacea from Holland
2007.11.09 © Baas, A.H.
Dicyrtomina cf. violacea from the UK
2008.07.22 © Ardron, A.P.
Dicyrtomina cf. violacea from the UK
2008.07.22 © Ardron, A.P.
Dicyrtomina violacea from Hungary
2014.05.01 © Rahmé, N.
Dicyrtomina violacea from the UK
2017.01.28 © Henton-Jones, D.
Dicyrtomina violacea from France
2017.10.22 © Picard, J.
Dicyrtomina violacea from France
2017.10.22 © Picard, J.
Dicyrtomina violacea from France
2017.11.06 © Tiky, S.
Dicyrtomina violacea from France
2017.11.06 © Tiky, S.
Dicyrtomina violacea from France
2017.11.06 © Tiky, S.
Dicyrtomina violacea from Hungary
2017.11.09 © Ujvári, Z.
Dicyrtomina violacea juv. from Hungary
2017.11.09 © Ujvári, Z.
Dicyrtomina violacea from Hungary
2017.11.14 © Ujvári, Z.
Dicyrtomina violacea from Hungary
2017.11.14 © Ujvári, Z.
Dicyrtomina violacea from Hungary
2018.01.02 © Ujvári, Z.
Dicyrtomina violacea from Hungary
2018.01.04 © Ujvári, Z.
Dicyrtomina violacea from Hungary
Pale colour form
2018.01.21 © Ujvári, Z.
Dicyrtomina violacea from Hungary
Interocular patch with 4 pale dots
2018.01.21 © Ujvári, Z.
Dicyrtomina violacea from the UK
2018.05.09 © Robson, S.
Dicyrtomina violacea from the UK
Broad interocular patch
2018.05.09 © Robson, S.
Dicyrtomina violacea from the UK
2019.01.25 © Cann, A.J.
Dicyrtomina violacea from the UK
Abd.6 with 2 pale spots
2019.01.25 © Cann, A.J.
Dicyrtomina violacea ♂ from France
2020.11.18 © Fiacre, R.
Dicyrtomina violacea juv. from Poland
2021.02.04 © Antosik-Zoladek, J.


Genus Calvatomina Yosii, 1966 key

  Syn.: Collembola Papirius Reuter, OM, 1892:18
  Syn.: ... Papiruis(sic) Jackson, AD, 1907:177 ISS


  Syn.: ... Calvastomina(sic) Mari Mutt, JA, 1987:364 ISS
Calvatomina sp. from Czechia
Red eyepatches with black ocelli
2013.06.26 © Krásenský, P.
Calvatomina sp. from Czechia
Ant.3 basally black, apically white; ant.4 white
2013.06.26 © Krásenský, P.
Calvatomina sp. from Greece
2014.10.05 © Antonopoulos, Z.
Calvatomina sp. juv. from Greece
2014.11.02 © Antonopoulos, Z.
Calvatomina sp. from Tasmania
2016.01.11 © Murray, A.
Calvatomina sp. from India
2019.08.12 © Rajtk, S.
Calvatomina sp. from New Caledonia
2020.10.02 © Brouste, D.
Calvatomina sp. from New Caledonia
Eyepatch with yellow background colour
2020.10.02 © Brouste, D.



Species sp.1 Janssens F sensu Bretfeld, 1999

            Syn.: Dicyrtomina rufescens Schött, 1927
            Syn.: Calvatomina sp.other Bretfeld, G, 1999:136

Species sp.2 Janssens F sensu Bretfeld, 1999

            Syn.: Calvatomina rufescens Mari Mutt, JA, 1987:378
Calvatomina rufescens from Colombia
After Mari Mutt JA, 1987 Fig.59

            Syn.: Calvatomina rufescens Heckman, CW, 2001:307
            Syn.: Calvatomina sp.other Bretfeld, G, 1999:136


Species sp.3 Janssens F sensu Bretfeld, 1999

            Syn.: Calvatomina rufescens Soto-Adames, FN, 1988:65
            Syn.: Calvatomina sp.other Bretfeld, G, 1999:136

Species sp.4 Janssens F sensu Samalot-Roque, 2006

            Syn.: Calvatomina sp.nov.nr. rossi Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:23

Species sp.5 Janssens F sensu Samalot-Roque, 2006

            Syn.: Calvatomina sp.nov.1 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:23

Species sp.6 Janssens F sensu Samalot-Roque, 2006

            Syn.: Calvatomina sp.nov.2 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:23

Species sp.7 Janssens F sensu Ospina-Sánchez, 2011

            Syn.: Calvatomina sp.nov.3 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:23

Species sp.8 Janssens F sensu Ospina-Sánchez, 2011

            Syn.: Calvatomina sp.nov.4 Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:23

Species sp.9 Janssens F & Snyder SL, 2014

Calvatomina sp. from the USA
2014.12.17 © Snyder, S.L.
Calvatomina sp. from the USA
2014.12.17 © Snyder, S.L.
Calvatomina sp. from the USA
Ventral aspect
2014.12.17 © Snyder, S.L.
Calvatomina sp. from the USA
2014.12.17 © Snyder, S.L.
Calvatomina sp. from the USA
2015.11.17 © Snyder, S.L.
Calvatomina sp. from the USA
2015.11.17 © Snyder, S.L.
Calvatomina sp. from the USA
2021.02.15 © Lampkin, J.
Calvatomina sp. from the USA
2021.02.15 © Lampkin, J.

Species sp.10 Janssens F & Fleming K, 2017

Calvatomina sp. from the USA
2014.mm.dd © Bernard, E.C.
Calvatomina sp. from the USA
2014.12.27 © Fleming, K.
Calvatomina sp. from the USA
2017.01.26 © Fleming, K.
Calvatomina sp. from the USA
2017.01.26 © Fleming, K.
Calvatomina sp. from the USA
2017.03.22 © Fleming, K.
Calvatomina sp. from the USA
2017.03.22 © Fleming, K.
Calvatomina sp. from the USA
Dark colour form
2018.01.22 © Fleming, K.
Calvatomina sp. from the USA
Pale colour form
2018.01.23 © Fleming, K.
Calvatomina sp. from the USA
2022.02.12 © Fleming, K.
Calvatomina sp. from the USA
2022.02.12 © Fleming, K.
Calvatomina sp. from the USA
Feeding instar
2022.02.22 © Charny, V.
Calvatomina sp. from the USA
Feeding instar
2022.02.22 © Charny, V.
Calvatomina sp. from the USA
Feeding instar
2022.02.22 © Charny, V.
Calvatomina sp. from the USA
2022.12.08 © Fleming, K.
Calvatomina sp. from the USA
2022.12.08 © Fleming, K.
Calvatomina sp. from the USA
Dark colour form
2022.12.09 © Fleming, K.
Calvatomina sp. from the USA
Dark colour form
2022.12.09 © Fleming, K.
Calvatomina sp. juv. from the USA
2023.01.03 © Fleming, K.
Calvatomina sp. juv. from the USA
2023.01.03 © Fleming, K.
Calvatomina sp. ♀ from the USA
2023.01.03 © Fleming, K.
Calvatomina sp. ♀ from the USA
2023.01.03 © Fleming, K.

Species sp.11 Janssens F & Daley T, 2017

Calvatomina sp. from Tasmania
Feeding from spermatophore; with two parasitic protonymph Acari
2014.09.29 © Murray, A.
Calvatomina sp. juv. from Tasmania
2016.08.01 © Daley, T.
Calvatomina sp. from Tasmania
2016.08.15 © Daley, T.
Calvatomina sp. from Tasmania
2016.08.15 © Daley, T.
Calvatomina sp. from Tasmania
2016.08.30 © Daley, T.
Calvatomina sp. from Tasmania
6 unpaired midfacial setae
2016.08.30 © Daley, T.
Calvatomina sp. from Tasmania
2017.05.13 © Daley, T.
Calvatomina sp. from Tasmania
2017.05.13 © Daley, T.
Calvatomina sp. from Tasmania
2017.05.13 © Daley, T.
Calvatomina sp. from Tasmania
2017.10.07 © Amanda.
Calvatomina sp. from Tasmania
2018.08.25 © Calmels, P.
Calvatomina sp. from Tasmania
2020.08.18 © Stylis, C.
Calvatomina sp. from Tasmania
2021.06.18 © Ellingsen, K.
Calvatomina sp. from Tasmania
2021.06.18 © Ellingsen, K.
Calvatomina sp. from Tasmania
2021.06.18 © Ellingsen, K.
Calvatomina sp. from Tasmania
2021.06.18 © Ellingsen, K.
Calvatomina sp. from Tasmania
2022.08.05 © Ellingsen, K.

Species alacris (Yosii, R, 1969:229)

Dicyrtomina (Calvatomina) alacris from the Solomon Islands
After Yosii R, 1969 Fig.7A

Species andrewarthai (Womersley, H, 1932:39) Betsch, J-M, 1980:147

             Syn.: Dicyrtomina andrewarthae(sic) Salmon, JT, 1964:629 ISS
             Syn.: Dicyrtomina andrewarthae(sic) Salmon, JT, 1964:630 ISS

Species antenna Nguyen, 2017

            Syn.: Dicyrtomina antenna Nguyen, 1994 nn
             Syn.: Calvatomina antena(sic) Nguyen, TT, Nguyen, TD, Nguyen, DA et Nguyen, VQ, 2008:609 nn ISS
             Syn.: Calvatomina antena(sic) Nguyen, TT, Nguyen, TD, Nguyen, DA et Nguyen, VQ, 2008:611 nn ISS
             Syn.: Calvatomina antena(sic) Nguyen, TT, Nguyen, TD, Nguyen, DA et Nguyen, VQ, 2008:612 nn ISS


             Syn.: Calvatomina antena(sic) Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:133 ISS
             Syn.: Calvatomina antena(sic) Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:152 ISS

Species bellingeri (Snider, RJ, 1990:90)

Dicyrtoma (Calvatomina) bellingeri from Hawaii
After Snider RJ, 1990 Fig.257
Dicyrtoma (Calvatomina) bellingeri from Hawaii
After Snider RJ, 1990 Fig.258

Species bougainvilleae (Yosii, 1960) Betsch, J-M, 1980:147

Dicyrtomina (Calvatomina) bougainvilleae from the Solomon Islands
After Yosii R, 1969 Fig.6A
             Syn.: Calvatomina bougainvillae(sic) Betsch, J-M, 1980:147 ISS


Species brevifibra (Snider, RJ, 1990:85)

Dicyrtoma (Calvatomina) brevifibra from Hawaii
After Snider RJ, 1990 Fig.168
Dicyrtoma (Calvatomina) brevifibra from Hawaii
After Snider RJ, 1990 Fig.169
Dicyrtoma (Calvatomina) brevifibra from Hawaii
After Snider RJ, 1990 Fig.170

Species calva (Denis, 1948) Betsch, J-M, 1980:147

Calvatomina cf. calva from Indonesia
2013.10.25 © Bertner, P.

Species christianseni (Delamare Deboutteville, C & Massoud, Z, 1964:74) Mari Mutt, JA, 1987:379

Calvatomina christianseni from Surinam
After Heckman CW, 2001 Fig.1.481 right
Calvatomina christianseni from Surinam
After Heckman CW, 2001 Fig.1.481 left

Species cruciata (Yosii, 1966) Betsch, J-M, 1980:147, t.t.

Species discolor (Schött, 1902) Betsch, J-M, 1980:147

Calvatomina discolor from Colombia
After Mari Mutt JA, 1987 Fig.66
Calvatomina discolor from Trinidad & Tobago
2016.01.05 © Fotografi, J.E.
Calvatomina discolor from Peru
2024.07.17 © R., S.

Species formosana (Yosii, R, 1965:61) Betsch, J-M, 1980:147

Dicyrtomina formosana from Taiwan
After Yosii R, 1965 Fig.38A
Calvatomina formosana from Taiwan
2022.04.28 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.04.28 © Cheng, H.-J.
Calvatomina formosana from Taiwan
Ant.4 inserted obliquely to ant.3
2022.04.28 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.07.15 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.07.15 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.10.06 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.10.06 © Cheng, H.-J.
Calvatomina formosana from Taiwan
M'-M' normal setae
M + N-N spiky macrosetae
2022.10.06 © Cheng, H.-J.
Calvatomina formosana from Taiwan
2022.10.12 © Cheng, H.-J.
Calvatomina formosana from Taiwan
Ant.4 inserted obliquely to ant.3
2022.10.12 © Cheng, H.-J.
Calvatomina formosana from Taiwan
Eyepatch with 8 ocelli (A-H)
2022.10.12 © Cheng, H.-J.

Species gladiata Medeiros GDS, Bellini BC et Cipola NG in Cipola, NG, Medeiros, GDS, Oliveira, VAMD, Barbosa, LGDM, Lycariao, TA et Bellini, BC, 2021:25

Calvatomina gladiata from Brazil
After Cipola NG & al, 2021 Fig.19A
Calvatomina gladiata from Brazil
After Cipola NG & al, 2021 Fig.19B

Species guyanensis Nayrolles & Betsch, 1996

Species lawrencei (Yosii, R, 1969:231)

Dicyrtomina (Calvatomina) lawrencei from the Solomon Islands
After Yosii R, 1969 Fig.8A

Species longidigita (Snider, RJ, 1990:90)

Dicyrtoma (Calvatomina) longidigita from Hawaii
After Snider RJ, 1990 Fig.238

Species madestris (Snider, RJ, 1990:80)

Dicyrtoma (Calvatomina) madestris from Hawaii
After Snider RJ, 1990 Fig.101
Dicyrtoma (Calvatomina) madestris from Hawaii
After Snider RJ, 1990 Fig.102

Species microdentata (Snider, RJ, 1990:89)

Dicyrtoma (Calvatomina) microdentata from Hawaii
After Snider RJ, 1990 Fig.226

Species modesta (Yosii, R, 1969:234) Soto-Adames, FN, 1988:65

Dicyrtomina (Calvatomina) modesta from the Solomon Islands
After Yosii R, 1969 Fig.10A

Species monodi (Delamare Deboutteville, C et Massoud, Z, 1965:102)

Dicyrtomina monodi from Angola
After Delamare Deboutteville C & Massoud Z, 1965 Planche I Fig.E
            Syn.: Dicyrtomina monodi Delamare Deboutteville, C et Massoud, Z, 1964:102 u


Species nymphascopula Soto-Adames, FN, 1988:61

Calvatomina nymphascopula from Puerto Rico
After Soto-Adames FN, 1988 Fig.22

Species opalina (Folsom, 1896) Snider, 1990

            Syn.: Papirius opalinus Folsom, 1896
Calvatomina opalina from the USA
2009.01.19 © Justis, S.
Calvatomina opalina from the USA
2009.01.29 © Justis, S.
Calvatomina opalina from the USA
2009.01.31 © Justis, S.
Calvatomina opalina from the USA
2010.03.07 © Bradford, A.
Calvatomina opalina from the USA
2010.03.07 © Bradford, A.
Calvatomina opalina from the USA
2015.04.26 © Dankowicz, E.



Species oryzae (Murphy, 1960)

Species pagoda Yosii, 1966

Species pallida Yosii, 1966

Species pseudorufescens (Hüther, 1965)

             Syn.: Dicyrtomina (Calvatomina) pseudoruiescens(sic) Thibaud, J-M, 2013:235 ISS

Species rossi (Wray, 1952) Soto-Adames, FN, 1988:65

Dicyrtomina (Dicyrtomina) rossi from the USA
After Snider, R., 1990 Fig.36
Dicyrtomina (Dicyrtomina) rossi from the USA
After Snider, R., 1990 Fig.37

Species rufescens (Reuter, OM, 1892:22) Betsch, J-M, 1980:147

Papirius rufescens from Finland
After Reuter OM, 1892 Tab.I Fig.3

            Syn.: Papirius rufescens var. discolor Linnaniemi, WM, 1912:323
            Syn.: Papirius rufescens ssp. discolor Salmon, JT, 1964:633
            Syn.: Calvatomina opalina Bretfeld, G, 1999:136



Papirius rufescens from Finland
After Schött H, 1893 Taf.III Fig.5
Calvatomina rufescens ♀ subadult from Tenerife
2022.12.03 © Huskens, M.-L.
Calvatomina rufescens ♀ subadult from Tenerife
2022.12.03 © Huskens, M.-L.
Calvatomina rufescens ♀ subadult from Tenerife
2022.12.03 © Huskens, M.-L.
Calvatomina rufescens ♀ subadult from Tenerife
2022.12.03 © Huskens, M.-L.
Calvatomina rufescens ♀ subadult from Tenerife
Circumanal chaetotaxy
Genital opening (*)
2022.12.03 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
Eye with smaller ocelli C + D
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
Facial pattern
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
Vertex and thorax without spiky macrosetae
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
Bothriotricha B + C
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
2022.12.05 © Huskens, M.-L.
Calvatomina rufescens ♀ from Tenerife
Bothriotrichum A with secondary seta
2022.12.05 © Huskens, M.-L.

Species scutellina Nguyen, 2001

Species solomonensis (Yosii, R, 1969:233) Soto-Adames, FN, 1988:65

Dicyrtomina (Calvatomina) solomonensis from the Solomon Islands
After Yosii R, 1969 Fig.9A
Dicyrtomina (Calvatomina) solomonensis from the Solomon Islands
After Yosii R, 1969 Fig.9B

            Syn.: Dicyrtomina (Calvatomina) solomonensis f. infuscata Yosii, R, 1969:234
Dicyrtomina (Calvatomina) solomonensis f. infuscata from the Solomon Islands
After Yosii R, 1969 Fig.9C
Dicyrtomina (Calvatomina) solomonensis f. infuscata from the Solomon Islands
After Yosii R, 1969 Fig.9D




Species strigata Bretfeld, G, 2005:10

Calvatomina strigata from Yemen
After Bretfeld G, 2005 Fig.15
Calvatomina strigata from Yemen
After Bretfeld G, 2005 Fig.16

Species superba (Salmon, JT, 1943:11) Betsch, J-M, 1980:147

Dicyrtomina superba from New Zealand
After Salmon J.T., 1943 Pl.1 Fig.7
Dicyrtomina superba from New Zealand
After Salmon J.T., 1943 Pl.2 Fig.35-38
Dicyrtomina superba from New Zealand
2014 © Te Papa Museum
Calvatomina superba from New Zealand
2014.02.25 © Murray, A.
Calvatomina superba from New Zealand
2014.02.25 © Murray, A.
Calvatomina superba from New Zealand
2014.02.25 © Murray, A.
Calvatomina superba from New Zealand
2014.02.25 © Murray, A.
Calvatomina superba from New Zealand
2014.09.16 © Murray, A.
Calvatomina superba from New Zealand
Metafoot with unguiculus (u) with lamella and unguis with tunica (t)
2014.09.16 © Murray, A.
Calvatomina superba from New Zealand
2023.07.17 © Ashwood, F.

Species sylvestratilis (Snider, RJ, 1990:84)

Dicyrtoma (Calvatomina) sylvestratilis from Hawaii
After Snider RJ, 1990 Fig.135
             Syn.: Dicyrtoma sylvestralis(sic) Christiansen, K et Bellinger, P, 1994:318 ISS


Species tesselata (Snider, RJ, 1990:87)

Dicyrtoma (Calvatomina) tesselata from Hawaii
After Snider RJ, 1990 Fig.193

Species trivandrana Prabhoo, 1971

Species tuberculata Nguyen, 2001

Species virgata (Schött, 1927) Betsch, J-M, 1980:147

Species yaeyamensis (Yosii, R, 1965:60)

Dicyrtomina yaeyamensis from Japan
After Yosii R, 1965 Fig.37A


Genus Pseudodicyrtomina Stach, J, 1957:30

  Syn.: Dicyrtomidae Calvatomina Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:132 sp


Pseudodicyrtomina sp. from Taiwan
2022.07.13 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.07.13 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.07.13 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.07.13 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.09.15 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.09.15 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.09.15 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
Unguis with tunica (tun)
2022.09.20 © Cheng, H.-J.
Pseudodicyrtomina sp. ♂ from Taiwan
Genital papilla (*)
2022.09.20 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.10.02 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
2022.10.03 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
Eyepatch background colour yellowish
2022.10.03 © Cheng, H.-J.
Pseudodicyrtomina sp. from Taiwan
M'-M-M' setae in triangular constellation
2022.10.04 © Cheng, H.-J.

Species bombayensis (Yosii, 1966) Hüther, 1967

Species trukana (Uchida, 1944) Stach, J, 1957:30, t.t.

Species verrucosa (Handschin, 1930) Stach, J, 1957:41


Genus Jordanathrix Bretfeld G et Arbea JI in Bretfeld, G, 1999:148

Jordanathrix from Spain
2014.11.16 © López, M.
Det.: Arbea, J.I.
Jordanathrix from Spain
2014.11.16 © López, M.
Det.: Arbea, J.I.

Species sp.4 (Janssens F & Murray A, 2014) Janssens F & Huskens ML, 2017

            Syn.: Jordanathrix articulata Ardron, PA, 2009:12 nr
            Syn.: Calvatomina superba Ardron, PA, 2009:12
Calvatomina superba from the UK
After Ardron PA, 2009 p.12




Calvatomina nr superba from the UK
2008.04.01 © Ardron, P.
Calvatomina nr superba from the UK
2008.04.01 © Ardron, P.
Calvatomina nr superba from the UK
2008.04.01 © Ardron, P.
Calvatomina nr superba from the UK
2008.04.01 © Ardron, P.
Calvatomina nr superba ♀ from the UK
Anogenital segment with sub-anal appendages
2008.11.17 © Ardron, P.
Calvatomina nr superba from the UK
2008.11.17 © Ardron, P.
 
Calvatomina nr superba from the UK
Mucro with both edges serrated
2008.11.17 © Ardron, P.
Calvatomina nr superba from the UK
2008.11.17 © Ardron, P.
Calvatomina nr superba from the UK
2008.11.24 © Ardron, P.
Calvatomina nr superba from the UK
2008.11.24 © Ardron, P.
Calvatomina nr superba from the UK
2008.12.08 © Ardron, P.
Calvatomina nr superba from the UK
2009.02.01 © Ardron, P.
Calvatomina nr superba from the UK
2009.02.01 © Ardron, P.
Calvatomina nr superba from the UK
2009.02.01 © Ardron, P.
Calvatomina nr superba from the UK
Leg 1: thick unguis with outer tunica
2009.04.01 © Ardron, P.
Calvatomina nr superba from the UK
2009.04.01 © Ardron, P.
Calvatomina nr superba from the UK
2009.11.15 © Robertson, A.
Calvatomina nr superba from the UK
2009.11.17 © Robertson, A.
Calvatomina nr superba from the UK
2009.11.17 © Robertson, A.
Calvatomina nr superba from the UK
2009.11.22 © Robertson, A.
Calvatomina nr superba from the UK
2010.01.23 © Robertson, A.
Calvatomina nr superba from the Azores
2010.09.22 © E.D.E.N.
Calvatomina nr superba from Holland
2011.10.30 © de Waard, H.
Calvatomina nr superba from the UK
2011.11.04 © Jones, A.
Calvatomina nr superba from the UK
2012.01.09 © Nurcombe, E.
Calvatomina nr superba from the UK
2012.01.09 © Nurcombe, E.
Calvatomina nr superba from the UK
2012.07.04 © Jones, A.
Calvatomina nr superba from the UK
2012.07.07 © Jones, A.
Calvatomina nr superba from the UK
2012.11.30 © Nurcombe, E.
Calvatomina nr superba from the UK
2012.12.10 © Nurcombe, E.
Calvatomina nr superba from France
2013.06.01 © Sandra.
Calvatomina nr superba from the UK
2013.12.09 © Nurcombe, E.
Calvatomina nr superba from Holland
Eyepatch
2014.11.09 © van Duinen, J.
Calvatomina nr superba from Holland
2014.11.09 © van Duinen, J.
Calvatomina nr superba from Holland
2014.11.09 © Wieringa, P.
Calvatomina nr superba from Holland
Cleaning antenna with drop of saliva on claw of proleg
2014.11.09 © Wieringa, P.
Calvatomina nr superba from Holland
2014.11.20 © Baas, A.H.
Calvatomina nr superba from Holland
Dark colour form
2015.11.20 © Wieringa, P.
Calvatomina nr superba from Belgium
2016.01.06 © Huskens, M.L.
Calvatomina nr superba from Belgium
Upside down triangular butt patch
2016.01.06 © Huskens, M.L.
Calvatomina nr superba from Belgium
Reddish eyepatch
2016.01.06 © Huskens, M.L.
Calvatomina nr superba from Holland
6 unpaired midfacial setae
2016.02.12 © van Duinen, J.
Calvatomina nr superba from Belgium
2016.02.16 © Janssens, F.
Calvatomina nr superba from the UK
2016.03.10 © Phillips, E.
Calvatomina nr superba from Holland
Leg 3: unguiculus (ung) with exad tooth; thick unguis (UNG)
2016.11.13 © van Duinen, J.
Calvatomina nr superba ♂ from the UK
Raised male genital papilla (*)
2016.11.15 © Phillips, E.
Calvatomina nr superba ♀ from the UK
2016.11.15 © Phillips, E.
Calvatomina nr superba ♂ from Belgium
2016.11.25 © Huskens, M.L.
Calvatomina nr superba ♂ from Belgium
Raised male genital papilla (*)
2016.11.25 © Huskens, M.L.
Calvatomina nr superba from Belgium
2016.12.01 © Huskens, M.L.
Calvatomina nr superba ♂ from Belgium
2016.12.17 © Huskens, M.L.
Calvatomina nr superba ♂ from Belgium
Raised genital papilla with apical ring (*)
2016.12.17 © Huskens, M.L.
Calvatomina nr superba ♂ from Belgium
Triangular dorsal posterior patch
2016.12.17 © Huskens, M.L.
Calvatomina nr superba ♀ from Belgium
Sub-anal appendages (*)
2017.01.20 © Huskens, M.L.
Calvatomina nr superba from the UK
Channel Islands L'Etac
2017.02.04 © Robson, S.
Calvatomina nr superba from France
2017.02.05 © Picard, J.
Calvatomina nr superba ♀ from Belgium
Sub-anal appendages
2017.02.21 © Huskens, M.L.
Jordanathrix nr superba from Belgium
Spines absent from head apex and dorsal anterior
2017.02.24 © Huskens, M.L.
Jordanathrix nr superba juv. from Belgium
Left-right wiggling midgut
2017.09.29 © Janssens, F.
Jordanathrix nr superba ♀ from Belgium
Backwards curved sub-anal appendages (SAA)
2017.11.23 © Huskens, M.L.
Jordanathrix nr superba from Belgium
2017.12.13 © Huskens, M.L.
Jordanathrix nr superba ♂ from Belgium
2018.01.25 © Huskens, M.L.
Jordanathrix nr superba ♂ from Belgium
2018.01.25 © Huskens, M.L.
Jordanathrix nr superba from Germany
Labrum with 3 superficial ridges
2018.03.06 © Jäschke, J.
Jordanathrix nr superba ♀ from Belgium
2020.09.11 © Huskens, M.L.
Jordanathrix nr superba ♀ from Belgium
Sub-anal appendages
2020.09.11 © Huskens, M.L.
Jordanathrix nr superba from Belgium
2020.10.28 © Huskens, M.L.
Jordanathrix nr superba from Belgium
2020.10.28 © Huskens, M.L.
Jordanathrix nr superba from Belgium
Ocelli A-H
2022.02.25 © Huskens, M.L.
Jordanathrix nr superba from Belgium
Facial aspect
2022.11.13 © Huskens, M.-L.
Jordanathrix nr superba from Belgium
Facial aspect
2022.12.29 © Huskens, M.-L.

Species articulata (Ellis, WN, 1974:148) Bretfeld G & Arbea JI in Bretfeld, G, 1999:148, t.t.

Dicyrtomina (Calvatomina) articulata from Rhodos
After Ellis WN, 1974 Fig.10c
Dicyrtomina (Calvatomina) articulata from Rhodos
After Ellis WN, 1974 Fig.9f
Jordanathrix articulata from Greece
2012.05.22 © László, D.
Jordanathrix nr articulata from Portugal
2012.10.11 © Murray, A.
 
Jordanathrix nr articulata from Portugal
2012.10.11 © Murray, A.
 
Jordanathrix nr articulata from Portugal
2012.10.11 © Murray, A.
Jordanathrix nr articulata from Portugal
2012.10.11 © Murray, A.
Jordanathrix nr articulata from Portugal
2012.10.11 © Murray, A.
Jordanathrix articulata from Spain
2012.11.19 © Herrera, J.M.
Jordanathrix articulata from Spain
2012.11.19 © Herrera, J.M.
Jordanathrix articulata from Spain
2012.11.25 © Herrera, J.M.
Jordanathrix articulata from Spain
2012.11.25 © Herrera, J.M.
Jordanathrix articulata from Spain
2016.12.13 © Herrera, J.M.
Det. Arbea J.I.
Jordanathrix articulata from Spain
2016.12.17 © Herrera, J.M.
Det. Arbea J.I.
Jordanathrix articulata from France
2017.10.02 © Tiky, S.
Jordanathrix articulata from France
2017.10.02 © Tiky, S.
Jordanathrix articulata from France
2017.10.02 © Tiky, S.
Jordanathrix articulata from France
Caudal aspect
2017.10.02 © Tiky, S.
Jordanathrix articulata from France
2017.11.02 © Tiky, S.
Jordanathrix articulata from France
2017.11.02 © Tiky, S.
Jordanathrix articulata from France
2017.11.02 © Tiky, S.
Jordanathrix articulata from France
2018.06.10 © Tiky, S.
Jordanathrix articulata from France
2018.10.19 © Tiky, S.
Jordanathrix articulata juv. from France
2019.11.10 © Garcelon, P.
Jordanathrix articulata from France
2020.10.01 © Tiky, S.

  Subspecies navarrae Bretfeld G et Arbea JI in Bretfeld, G, 1999:149

Species leptothrix (Börner, C, 1909:128) Bretfeld, G, 1999:151

            Syn.: Dicyrtomina rufescens f. nivalis Uchida, 1953
            Syn.: Dicyrtomina rufescens f. nivalis Yosii, R, 1965:56
            Syn.: Dicyrtomina rufescens f. nivalis Yosii, R, 1977:166
            Syn.: Dicyrtomina rufescens Uchida, 1953
            Syn.: Dicyrtomina rufescens Yosii, R, 1965:58
            Syn.: Dicyrtomina rufescens Uchida & Fujita, 1968


             Syn.: Dicyrtomina leptothryx(sic) Yosii, R, 1961:14 ISS
Dicyrtomina leptothrix from Japan
Pale form
After Yosii R, 1965 Fig.35A
Dicyrtomina leptothrix from Japan
After Itoh R & al, 2012 Fig.45b





Genus Gibberathrix Uchida, 1952

Species tsugarensis Uchida, 1952 tt

Dicyrtomina tsugarensis from Japan
After Yosii R, 1970 Fig.18A
Dicyrtomina tsugarensis from Japan
After Yosii R, 1970 Fig.19A
Gibberathrix tsugarensis from Japan
After Uchida, 1952 in Itoh R & al, 2012 Fig.46b

References