http://www.collembola.org/taxa/pteninae.htm - Last updated on 2024.05.29 by Frans Janssens
Checklist of the Collembola: Ptenothricinae

Subfamilia Ptenothricinae Richards WR, 1968:25 sensu Bretfeld, G, 1999:128

Genus Number
of species
Bothriovulsus4
Papirioides12
Ptenothrix102
Ptenothricinae118
Genera with number of species
  Syn.: Dicyrtomidae Pterothricinae(sic) Deharveng, L, 2004:427 ISS
Ptenothrix sp. from Canada
Anterior dorsal spinelike setae
2008.12.11 © McCann, S.
Ptenothrix sp. from Canada
Small abdomen with 3M+2N setae
M-setae in transverse row
2023.02.18 © Barbin, T.
Ptenothrix sp. from Canada
Two unpaired midfacial setae
2023.12.17 © Chans, J.




Genus Ptenothrix Börner, C, 1906:184 key  key

F
  Syn.: ... Papirius Lubbock, 1862 partim
  Syn.: Dicyrtoma fusca Yosii, 1940
  Syn.: Dicyrtoma fusca Yosii, R, 1977:166
  Syn.: Smynthurelles(sic) Dycirtoma(sic) Lucas, H, 1849:Pl.2 ISS
  Syn.: Dicyrtomidae Dicyrtoma Greenslade, P, Boyer, S, Shields, MW et Wratten, SD, 2016:5 partim


  Syn.: Collembola Ptenothryx(sic) Uchida, 1953 ISS
  Syn.: Collembola Ptenothryx(sic) Ptenothryx(sic) {Collembola}3520:824 ISS
  Syn.: Collembola Ptenothryx(sic) Ptenothryx(sic) {Collembola}3520:825 ISS
  Syn.: Collembola Ptenothryx(sic) Yosii, R, 1955:380 ISS
  Syn.: Collembola Ptenothryx(sic) Yosii, R, 1955:397 ISS
  Syn.: Collembola Ptenothryx(sic) Yosii, R, 1955:399 ISS
  Syn.: Sminthuridae Ptenothryx(sic) Yosii, R, 1956:101 ISS
  Syn.: ... Ptenothryx(sic) Yosii, 1961 ISS
  Syn.: Dicyrtominae Ptehothrix(sic) Yosii, R, 1977:166 ISS
Ptenothrix sp. from the USA
Long tube-like everted vesicles of collophore
2009.03.30 © Bradford, A.
Ptenothrix castanea from Canada
Ant.3 apically with pseudo-annulations
2011.07.26 © Hochhalter, R.
Ptenothrix sp. from Canada
Ocelli with transparent crystal lenses
2011.10.26 © Hochhalter, R.
Ptenothrix sp. from Canada
"Eyebrow" and composed eye
2012.03.24 © Hochhalter, R.
Ptenothrix sp. from Canada
Footcomplex with long empodium
2012.05.07 © Hochhalter, R.
Ptenothrix sp. from Canada
Two thick unpaired midfacial macrosetae (*)
2015.10.30 © Barbin, T.
Ptenothrix sp. from Canada
Labrum with 3 ridges
2016.02.06 © Barbin, T.
Ptenothrix maculosa from the USA
Abd.6 with spinelike macrosetae : 3M+2N
2016.03.08 © Dobbs, Br.

  Syn.: Collembola Ptenotrix(sic) Duarte Núñez, DA, Pinzón Florián, OP et Palacios-Vargas, JG, 2020:1207 ISS
  Syn.: Dicyrtomidae Ptenotrix(sic) Nardi, F, Cucini, C, Leo, C, Frati, F, Fanciulli, PP et Carapelli, A, 2020:3122 ISS
  Syn.: Dicyrtomidae Ptenotrix(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:8 ISS
  Syn.: Dicyrtomidae Ptenotrix(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:9 ISS
  Syn.: Dicyrtomidae Ptenotrix(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:12 ISS
  Syn.: Dicyrtomidae Ptenotrix(sic) Ramos-Díaz, A, Palacios-Vargas, JG et Pinzón-Florián, OP, 2020:14 ISS
Ptenothrix sp. from Canada
Mid trochanter with suddenly bent macroseta
2021.11.14 © Barbin, T.
Ptenothrix sp. from Japan
2023.09.08 © Featherstone, A.W.
Ptenothrix sp. from the USA
With 2 unpaired midfacial setae
2024.02.03 © G, S.
Ptenothrix sp. juv. from Canada
2024.02.08 © Chan, J.



Species sp.1 Janssens F sensu Yosii R, 1969

            Syn.: Ptenothrix sp.independent near gracillicornis(sic) Yosii, R, 1969:224


Species sp.3 Janssens F & Moorehead C, 2008

            Syn.: Ptenothrix sp.3 f.1 Janssens F & Moorehead C, 2015
Remark by Janssens, F. 2015.01.dd: Middorsal posterior patch split in 2 patches. Middorsal anterior stripe pale.
Ptenothrix sp. from the USA
2007.12.25 © Moorehead, C.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.14 © Kyron, T.F.
Ptenothrix sp. from the USA
Split butt patch
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
Split butt patch
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.12.03 © Johnson, S.T.

            Syn.: Ptenothrix sp.3 f.1a Janssens F & Dobbs B, 2019
Remark by Janssens, F. 2019.02.07: Middorsal posterior patch split in 2 patches. Middorsal anterior stripe dark.
Ptenothrix sp. from the USA
2009.01.12 © Davis, J.
Ptenothrix sp. from the USA
2009.01.12 © Davis, J.
Ptenothrix sp. from the USA
2010.02.05 © Schimming, L.
Ptenothrix sp. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.10 © Dobbs, Br.
Ptenothrix sp. from the USA
Tubelike bipartite collophore vesicle:
warted apical part + smooth basal part
2018.01.15 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.12.28 © Duncan, W.L.fig

            Syn.: Ptenothrix sp.3 f.1aa Janssens F & Pearson RR, 2019
Remark by Janssens, F. 2023.02.25: Middorsal posterior patch split in 2 patches. Great abdomen bicoloured : anterior dark, posterior pale.
Ptenothrix sp. from the USA
2019.01.31 © Pearson, R.R.
Ptenothrix sp. from the USA
2019.01.31 © Pearson, R.R.
Ptenothrix sp. from the USA
2019.01.31 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
2020.12.02 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
2020.12.02 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
3+1+3 macrosetae on posterior margin of dorsal anal valve
2020.12.02 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
High contrast form
2021.01.06 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
High contrast form
2021.01.06 © Pearson, R.R.
Ptenothrix sp. ♀ from the USA
3+1+3 macrosetae on posterior margin of dorsal anal valve
2021.01.06 © Pearson, R.R.
Ptenothrix sp. from Canada
Dorsal posterior patch
2023.02.18 © Barbin, T.

            Syn.: Ptenothrix sp.3 f.2 Janssens F & Leppin C, 2015
Remark by Janssens, F. 2015.01.dd: Middorsal posterior patch reduced to middorsal stripe.
Ptenothrix sp. from the USA
2009.01.01 © Leppin, M.
Ptenothrix sp. from the USA
2009.01.01 © Leppin, M.

            Syn.: Ptenothrix sp.3 f.3 Janssens F & Kyron TF, 2016
Remark by Janssens, F. 2016.03.14: Middorsal posterior patch fused with lateral pigmentation.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix sp. ♀ from Canada
2017.01.21 © Barbin, T.
Ptenothrix sp. from the USA
2017.12.02 © Dobbs, Br.
Ptenothrix sp. from the USA
2019.01.25 © Wiser N.
Ptenothrix sp. ♀ juv. from the USA
2022.11.30 © Charny, V.
Ptenothrix sp. ♀ juv. from the USA
2022.11.30 © Charny, V.
Ptenothrix sp. ♀ juv. from the USA
2022.11.30 © Charny, V.
Ptenothrix sp. ♀ juv. from the USA
2022.11.30 © Charny, V.
Ptenothrix sp. from the USA
Grazing from jelly mushroom
2022.11.30 © Charny, V.
Ptenothrix sp. juv. from the USA
2022.12.13 © Charny, V.
Ptenothrix sp. from the USA
2022.12.13 © Charny, V.
Ptenothrix sp. from the USA
2022.12.14 © Charny, V.
Ptenothrix sp. from Canada
2023.01.28 © Barbin, T.
Ptenothrix sp. from Canada
2023.01.28 © Barbin, T.
Ptenothrix sp. from Canada
Frontal aspect
2023.01.28 © Barbin, T.
Ptenothrix sp. from Canada
Caudal aspect
2023.01.28 © Barbin, T.

            Syn.: Ptenothrix sp.3 f.4 Janssens F & Dobbs Br, 2016
Remark by Janssens, F. 2016.12.27: Middorsal posterior patch with many cross bars.
Ptenothrix sp. from the USA
2016.11.25 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.25 © Dobbs, Br.
Ptenothrix sp. from the USA
2019.01.21 © Duncan, W.L.

            Syn.: Ptenothrix sp.3 f.5 Janssens F & Dobbs Br, 2018
Remark by Janssens, F. 2018.01.11: With broad pale longitudinal middorsal band.
Ptenothrix sp. from Canada
2010.01.04 © McCann, S.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.10 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.11 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2017.12.11 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2018.01.08 © Dobbs, Br.
Ptenothrix sp. from the USA
Form with pale dorsum
2019.01.31 © Wiser N.

            Syn.: Ptenothrix sp.3 f.6 Janssens F & Wiser N, 2019
Remark by Janssens, F. 2019.01.29: Pale. Middorsal pattern reduced to a few dots.
Ptenothrix sp. from the USA
Pale form with a few dorsal dots
2019.01.25 © Wiser N.
Ptenothrix sp. from the USA
Pale form with a few dorsal dots
2019.01.25 © Wiser N.




Remark by Janssens, F. 2018.01.11: Principal form with pale middorsal anterior stripe and dark middorsal posterior patch.
Ptenothrix sp. from the USA
2005 © Leonard, W.
Ptenothrix sp. from the USA
2007.03.08 © Moorehead, C.
Ptenothrix sp. from the USA
2007.03.08 © Moorehead, C.
Ptenothrix sp. from the USA
2007.12.25 © Moorehead, C.
Ptenothrix sp. from the USA
2008.02.03 © Moorehead, C.
Ptenothrix sp. from the USA
2009.01.04 © Leppin, M.
Ptenothrix sp. from the USA
2009.01.04 © Leppin, M.
Ptenothrix sp. from the USA
2009.02.06 © Hall, K.
Ptenothrix sp. from the USA
2009.02.20 © Hall, K.
Ptenothrix sp. from the USA
2009.02.20 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.27 © Hall, K.
Ptenothrix sp. from the USA
2009.03.31 © LeMons, J.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. juv. from Canada
2013.01.17 © Kyron, T.F.
Ptenothrix sp. from the USA
2015.01.20 © Faberha, J.
Ptenothrix sp. from the USA
2015.02.15 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.01.21 © Dobbs, Br.
Ptenothrix sp. ♀ from the USA
2016.01.21 © Dobbs, Br.
Ptenothrix sp. from the USA
Two unpaired facial setae
2016.01.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Juvenile and adult
2016.02.22 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.02.23 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.02.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.02 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.06 © Dobbs, Br.
Ptenothrix sp. ♂ from the USA
Raised male genital papilla (*)
2016.11.06 © Johnson, S.T.
Ptenothrix sp. ♂ from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix sp. from Canada
2018.01.12 © Barbin, T.
Ptenothrix sp. from Canada
2021.01.14 © Rennert, M.
Ptenothrix sp. from Canada
2021.01.14 © Rennert, M.
Ptenothrix sp. from the USA
2023.01.28 © Barbin, T.
Ptenothrix sp. from Canada
Two unpaired midfacial setae
2023.12.17 © Chans, J.
Ptenothrix sp. from Canada
2023.12.17 © Chans, J.
Ptenothrix sp. from Canada
Hindfoot
2023.12.17 © Chans, J.
Ptenothrix sp. from Canada
2023.12.17 © Chans, J.
Ptenothrix sp. from Canada
2024.02.03 © Lypkie, P.
Ptenothrix sp. from Canada
Unguiculus of foot3 with sharp basal spine
and long apical filament
2024.02.03 © Lypkie, P.
Ptenothrix sp. from Canada
Unguis of foot3 with lateral teeth
2024.02.03 © Lypkie, P.
Ptenothrix sp. from Canada
Mucro with serrate edge
2024.02.03 © Lypkie, P.
Ptenothrix sp. from Canada
Distal external dental setae
E1 smooth; E2 serrate at base
2024.02.03 © Lypkie, P.
Ptenothrix sp. from Canada
Retinaculum lateral aspect
with anterior (al) and posterior lobe (pl)
2024.02.03 © Lypkie, P.

Species sp.4 Janssens F & Moorehead C, 2008

Remark by Janssens, F. 2021.01.29: With dark middorsal anterior stripe and dark middorsal posterior patch.
Ptenothrix sp. from the USA
2007.12.24 © Moorehead, C.
Ptenothrix sp. from Canada
2008.11.27 © McCann, S.
Ptenothrix sp. from Canada
2008.12.11 © McCann, S.
Ptenothrix sp. from Canada
Two unpaired facial setae
2008.12.11 © McCann, S.
Ptenothrix sp. from Canada
2008.12.01 © McCann, S.
Ptenothrix sp. from Canada
2008.12.02 © McCann, S.
Ptenothrix sp. from the USA
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.19 © Haley, R.
Ptenothrix sp. from the USA
2016.03.12 © Necturus.
Ptenothrix sp. from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Dark form
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Cephalic dorsal pattern
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Collophore vesicles extended
2017.10.26 © Dobbs, Br.
Ptenothrix sp. from the USA
Caudal aspect
2017.10.26 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.26 © Dobbs, Br.
Ptenothrix sp. from Canada
2020.06.27 © Barbin, T.
Ptenothrix sp. from the USA
2021.01.09 © Parker, H.S.

Species sp.5 Janssens F & LeMons J, 2008

Ptenothrix sp. from the USA
2007.03.05 © LeMons, J.
Ptenothrix sp. from the USA
2007.03.05 © LeMons, J.
Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from Canada
2016.02.09 © Barbin, T.
Ptenothrix sp. from the USA
With pale femur and dark tibiotarsus
2016.12.03 © Johnson, S.T.
Ptenothrix sp. from the USA
2016.12.03 © Johnson, S.T.

Species sp.6 Janssens F & Ten Eyck C, 2010

Ptenothrix sp. from the USA
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from the USA
Two thick unpaired facial setae
2010.01.17 © Ten Eyck, C.
Ptenothrix sp. from Canada
2011.03.21 © Hochhalter, R.
Ptenothrix sp. from Canada
2011.03.26 © Hochhalter, R.
Ptenothrix sp. from Canada
2011.03.26 © Hochhalter, R.
Ptenothrix sp. from Canada
2016.02.08 © Barbin, T.
Ptenothrix sp. from Canada
2016.02.08 © Barbin, T.
Ptenothrix sp. from the USA
2016.11.18 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.18 © Dobbs, Br.
Ptenothrix sp. juv. from the USA
2016.11.30 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.11.29 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.01.02 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.12.28 © Pearson, R.R.
Ptenothrix sp. from the USA
2019.01.12 © Dobbs, Br.
Ptenothrix sp. from the USA
2019.01.05 © Pearson, R.R.
Ptenothrix sp. from the USA
2019.01.05 © Pearson, R.R.
Ptenothrix sp. from Canada
2021.01.14 © Rennert, M.
Ptenothrix sp. from Canada
2021.01.14 © Rennert, M.

Species sp.7 Janssens F & Alexander C, 2012

Ptenothrix sp. from the USA
2012.12.27 © Alexander, C.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. from the USA
2013.01.09 © Alexander, C.
Ptenothrix sp. from the USA
2017.11.27 © Dobbs, Br.

Species sp.8 Janssens F & Ten Eyck, 2008

Ptenothrix sp. from the USA
2008.01.13 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
2010.01.10 © Ten Eyck, C.
Ptenothrix sp. from the USA
Unguis with outer tooth and 2 lateral teeth
2010.01.10 © Ten Eyck, C.

Species sp.9 Janssens F & Kyron TF, 2013

Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.
Ptenothrix sp. from Canada
2013.01.18 © Kyron, T.F.

Species sp.10 Janssens F & Dobbs B, 2016

Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.24 © Dobbs, Br.
Ptenothrix sp. from the USA
Unguiculus of 3rd leg with long knobbed filament
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Abd.6 dorsally with 3M+2N spinelike setae
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Eyepatch with reddish background
2016.11.27 © Dobbs, Br.
Ptenothrix sp. from the USA
Two unpair midfacial setae
2016.11.27 © Dobbs, Br.

Species sp.11 Janssens F & Dobbs B, 2017

Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.11.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
Antennae
2016.12.07 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.22 © Dobbs, Br.
Ptenothrix sp. from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Antenna pigmentation
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
Abd.6 with dark dorsal patch
2017.10.23 © Dobbs, Br.
Ptenothrix sp. from the USA
2018.02.22 © Dobbs, Br.
Ptenothrix sp. from Canada
2020.05.11 © Rennert, M.
Ptenothrix sp. from Canada
Botriotricha ABC
2020.05.11 © Rennert, M.
Ptenothrix sp. from Canada
Anterior dorsal spiky macrosetae
2020.05.11 © Rennert, M.

Species sp.12 Janssens F & Dobbs B, 2017

Ptenothrix sp. from the USA
Dark uniform body colour
2017.11.06 © Dobbs, B.
Ptenothrix sp. from the USA
Pale ant.1+2; pale tibiotarsi
2017.11.06 © Dobbs, B.
Ptenothrix sp. from the USA
Pale mouth region
2017.11.06 © Dobbs, B.
Ptenothrix sp. from the USA
2017.11.07 © Dobbs, B.
Ptenothrix sp. from the USA
2024.05.11 © Coogler, J.
Ptenothrix sp. from the USA
Antenna
2024.05.11 © Coogler, J.

Species sp.12A Janssens F & Barbin T, 2017

Ptenothrix sp.12A from Canada
2017.05.17 © Barbin, T.
Ptenothrix sp.12A from Canada
2017.05.17 © Barbin, T.
Ptenothrix sp.12A from Canada
2017.05.17 © Barbin, T.
Ptenothrix sp.12A from the USA
2018.02.06 © Dobbs, B.
Ptenothrix sp.12A from the USA
2018.02.06 © Dobbs, B.
Ptenothrix sp.12A from Canada
2020.12.20 © Reader-Lee, R.
Ptenothrix sp.12A from Canada
2020.12.20 © Reader-Lee, R.
Ptenothrix sp.12A from Canada
Dentes with dark pigmentation
2020.12.20 © Reader-Lee, R.
Ptenothrix sp.12A ♀ from the USA
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A♀ from the USA
Ant.3 with apical pseudo-annulations
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
Cephalic dorsal pattern
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
Small abdomen with anterior white spot
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
2 unpaired midfacial macrosetae
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
3 spiky M setae; 2 spiky N setae
Sub-anal appendages (SAA) of female
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A ♀ from the USA
Ventral aspect
2021.10.27 © O'Shea, B.
Ptenothrix sp.12A from Canada
2024.05.11 © Barbin, T.
Ptenothrix sp.12A from Canada
2024.05.11 © Barbin, T.

Species sp.13 Janssens F & Charny V, 2021

Ptenothrix sp. from the USA
2021.11.09 © Charny, V.
Ptenothrix sp. from the USA
2021.11.09 © Charny, V.
Ptenothrix sp. from the USA
2021.11.09 © Charny, V.
Ptenothrix sp. from the USA
2021.11.09 © Charny, V.

Species sp.14 Janssens F & Cheng H-J, 2021

Ptenothrix nr marmorata ♀ from Taiwan
2021.03.19 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2021.11.30 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2021.11.30 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2022.01.16 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2022.01.16 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2022.02.05 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2023.10.08 © Cheng, H.-J.
Ptenothrix nr marmorata from Taiwan
2023.10.08 © Cheng, H.-J.

Species sp.15 Janssens F & Cheng H-J, 2022

Ptenothrix sp. from Taiwan
Reddish colour form
2022.09.18 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Reddish colour form
2022.09.18 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Yellow colour form
2022.09.18 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Yellow colour form
2022.09.18 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Midfacial macrosetae 1+1+2
2022.09.20 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Forefoot
Unguiculus apically clavate
2022.09.22 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Bothriotricha A+B+C+D
2022.09.26 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Bothriotrichum D
2022.09.26 © Cheng, H.-J.
Ptenothrix sp. ♂ from Taiwan
2022.09.27 © Cheng, H.-J.
Ptenothrix sp. ♂ from Taiwan
Genital papilla (*)
2022.09.27 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
2022.09.28 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Bothriotricha A+B+C+D
2022.09.28 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
2022.09.28 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Setae M'-M-M' in transverse row
2022.09.28 © Cheng, H.-J.
Ptenothrix sp. from Taiwan
Setae M'-M-M' in transverse row
2022.09.30 © Cheng, H.-J.

Species sp.16 Janssens F & Cheng H-J, 2022

Ptenothrix sp. nr higumai? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix sp. nr higumai? juv. from Taiwan
2022.07.20 © Cheng, H.-J.
Ptenothrix sp. nr higumai? juv. from Taiwan
2022.07.20 © Cheng, H.-J.
Ptenothrix sp. nr higumai? juv. from Taiwan
2022.08.10 © Cheng, H.-J.
Ptenothrix sp. nr higumai? juv. from Taiwan
2022.08.10 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
M'-M-M' in transverse row
2022.10.12 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
2022.10.14 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
2022.10.14 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
2023.10.08 © Cheng, H.-J.
Ptenothrix sp. nr higumai? from Taiwan
2023.10.08 © Cheng, H.-J.

Species sp.16A Janssens F & Cheng H-J, 2022

Ptenothrix nr saxatilis from Taiwan
2011.12.30 © Abi-Aad, A.
Ptenothrix nr saxatilis from Taiwan
2011.12.30 © Abi-Aad, A.
Ptenothrix nr saxatilis from Taiwan
2013.11.17 © Abi-Aad, A.
Ptenothrix nr saxatilis from Taiwan
2014.01.25 © Abi-Aad, A.
Ptenothrix nr saxatilis from Taiwan
2014.03.01 © Abi-Aad, A.
Ptenothrix nr saxatilis from Taiwan
2021.11.21 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
2022.04.19 © Cheng, H.-J.
Leg.: Lin, T.-P.
Ptenothrix nr saxatilis from Taiwan
2022.04.19 © Cheng, H.-J.
Leg.: Lin, T.-P.
Ptenothrix nr saxatilis from Taiwan
2022.10.03 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
2022.10.03 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Midfacial setae 1+1+2+2+1
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Papilla of bothriotrichum A not included in circumlateral band
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Dorsal papilla with bothriotrichum A directed forwards
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Mucro with distinct apical notch
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Ant.1 (short and dark) + ant.2 (long and light)
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Mechanism to restrict downwards rotation of antenna
Ant.1 spur (*) + ant.2 spur (**)
2022.10.18 © Cheng, H.-J.
Ptenothrix nr saxatilis from Taiwan
Ant.3 (long) + ant.4 (short)
Ant.3 with apical pseudo-annulations
2022.10.18 © Cheng, H.-J.

Species sp.17 Janssens F & Cheng H-J, 2024

Ptenothrix nr denticulata var. catenata juv. from Taiwan
2021.03.03 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.03.15 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.03.15 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.03.19 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Grazing
2021.07.31 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.10.26 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.11.01 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
With twinkling abdomen
2021.11.01 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2021.11.23 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Ant.1 with subapical spur (*)
2021.11.23 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Facial pattern
2022.07.29 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Head with longitudinal stripe
2022.07.29 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Ant.3 without apical pseudosubdivisions
2022.07.29 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Ant.4
2022.07.29 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Dental apical outer setae serrated
E1 very short; E2 distinctly longer
2022.08.01 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata juv. from Taiwan
2022.10.02 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata juv. from Taiwan
2022.10.02 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata ♀ from Taiwan
Spotted
2022.11.11 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata ♀ from Taiwan
Sub-anal appendage (saa) on basal papilla
2022.11.11 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2022.11.11 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Patched
2022.11.11 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Forward directed bothriotrichum A on dorsal papilla
2022.11.11 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Modified macroseta of hind tibiotarsus
2022.11.14 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
6 prelabral setae
2022.11.14 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Inter ocular setae spiky and pointed
2022.11.14 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Midfacial macrosetae 1+1+2
2022.11.14 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Hindfoot
2022.11.17 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Hindfoot
Unguicular tubercle
2022.11.17 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Ant.3 organ
2022.11.17 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Mid trochanter with bent macroseta
Oval organ (Ov)
2022.11.21 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Mid trochanter with oval organ
2022.11.21 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Labium with labial palps
Ventral aspect
2022.11.23 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata ♂ from Taiwan
Male genital papilla
2022.11.23 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata ♂ from Taiwan
Ant.2 with cup-sensillum
2022.11.23 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
Dorsal cephalic spines short and not acuminate
2022.11.23 © Cheng, H.-J.

            Syn.: Ptenothrix denticulata Cheng, H-J, Janssens, F et Chang, C-H, 2022:137
            Syn.: Ptenothrix sp.1 Cheng H-J, 2024:il
Ptenothrix sp.1 from Taiwan
2024.01.02 © Cheng, H.-J.




Species sp.18 Janssens F & Cheng H-J, 2024

Ptenothrix nr denticulata var. catenata from Taiwan
2022.05.05 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2022.05.05 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.05.29 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.05.31 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.05.31 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.12.04 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.12.04 © Cheng, H.-J.
Ptenothrix nr denticulata var. catenata from Taiwan
2023.12.04 © Cheng, H.-J.

            Syn.: Ptenothrix sp.2 Cheng H-J, 2024:il
Ptenothrix sp.2 from Taiwan Matsu Island
2024.01.02 © Cheng, H.-J.




Species sp.19 Janssens F & Cheng H-J, 2024

Ptenothrix nr denticulata from Taiwan
2023.10.15 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2023.10.15 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.

            Syn.: Ptenothrix sp.3 Cheng H-J, 2024:il
Ptenothrix sp.3 from Taiwan
2024.01.02 © Cheng, H.-J.




Species sp.20 Janssens F & Cheng H-J, 2024

Ptenothrix nr denticulata from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
2024.01.01 © Cheng, H.-J.

            Syn.: Ptenothrix sp.4 Cheng H-J, 2024:il
Ptenothrix sp.4 from Taiwan
2024.01.02 © Cheng, H.-J.




Ptenothrix denticulata-group from Taiwan
2024.03.12 © Cheng, H.-J.

Species sp.21 Janssens F & Cheng H-J, 2024

Ptenothrix nr denticulata from Taiwan
Unguiculus with 2 basal teeth
2023.01.11 © Cheng, H.-J.
Ptenothrix nr denticulata from Taiwan
Mucro with apical notch
2024.01.01 © Cheng, H.-J.

            Syn.: Ptenothrix sp.7 Cheng H-J, 2024:il
Ptenothrix sp.7 from Taiwan
2024.01.02 © Cheng, H.-J.




Species albosignata (Parona, 1888) Stach, J, 1957:90

Species alveola (Lucas, H, 1846:255) Janssens F, 2020:mihi

            Syn.: Dicyrtoma alveolus(sic) Lucas, H, 1846:255 IOS
            Syn.: Ptenothrix italica Dallai, R, 1973:570
Ptenothrix italica from Italy
After Dallai R, 1973 Fig.25a
Ptenothrix italica from Italy
Ant.2 after Dallai R, 1973 Fig.25d
Compared with specimen from France, 2015.11.01 © Gros, P.
Ptenothrix italica from Italy
After Dallai R & al, 2000 Fig.3
Ptenothrix italica from France
2010.12.24 © Nicolosi., B
Ptenothrix italica from Spain
2013.11.04 © Encarna, Q.V.
Ptenothrix italica from France
Marbled body
2015.11.01 © Gros, P.
Ptenothrix italica from France
Thick mid-facial setae: 1,1,3,1,1,1
2015.11.01 © Gros, P.
Ptenothrix italica from France
Antennae with 3 broad black and white rings
2015.11.01 © Gros, P.
Ptenothrix italica from France
2015.11.01 © Gros, P.
Ptenothrix italica from France
Bicoloured eyepatch
2015.11.01 © Gros, P.
Ptenothrix italica from France
Abd.6 with backwards directed dorsal macrosetae (*)
2015.11.01 © Gros, P.
Ptenothrix italica from France
Lateral aspect
2015.11.14 © Gros, P.
Ptenothrix italica from France
Caudal aspect
2015.11.14 © Gros, P.
Ptenothrix italica from France
Ant.3 and 4
2015.11.14 © Gros, P.
Ptenothrix italica from France
Short dorsal spines
2015.11.14 © Gros, P.
Ptenothrix italica ♀ from France
Female gonopore (*)
2015.11.17 © Gros, P.
Remark by Janssens, F. 2015.11.17: Absent strong macrosetae on ant.2, absent dorsal spiky macrosetae in the anterior part of the great abdomen, and presence of backwards directed macrosetae on abd.6 are atypical for Ptenothrix and more characteristic to Dicyrtoma.

Ptenothrix italica from Italy
2017.02.05 © Rinaldo, R.
Ptenothrix italica from Italy
Midfacial setae: 1+1+1+1+1+1
2017.02.05 © Rinaldo, R.
Ptenothrix italica from Italy
Antenna
2017.02.05 © Rinaldo, R.
Ptenothrix italica from France
2017.10.02 © Tiky, S.
Ptenothrix italica juv. from Algeria
After Bendjaballah M & al, 2018 Fig.22
Ptenothrix italica from France
Spider mimicry
2018.06.10 © Tiky, S.
Ptenothrix italica from France
Dorsal setae of abd.6 pointing backwards suggests Dicyrtoma...
2018.06.10 © Tiky, S.
Ptenothrix italica from France
2018.10.25 © Tiky, S.
Ptenothrix italica from France
More than 2 unpaired midfacial macrosetae suggest Dicyrtoma...
2018.10.25 © Tiky, S.
Ptenothrix italica from France
2018.11.01 © Gros, P.
Ptenothrix italica from France
2018.11.01 © Gros, P.
Ptenothrix italica from France
Caudal aspect
2018.11.01 © Gros, P.

            Syn.: Dicyrtomina ornata Seminario-Córdova, RA, Baquero, E, Jordana, R et Vadell, M, 2018:55
Dicyrtomina ornata from Mallorca
After Seminario-Córdova RA et al, 2018 Fig.5B
= Ptenothrix alveola; Det.: Janssens F, 2023.06.08





            Syn.: Dicyrtoma alveolus(sic) Nicolet, H, 1847:362 IOS
            Syn.: Dicyrtoma alveolus(sic) Lucas, H, 1849:379 IOS
            Syn.: Dycirtoma(sic) alveolus(sic) Lucas, H, 1849:Pl.2 IOS
Dycirtoma(sic) alveolus(sic) from Algeria
After Lucas H, 1849 Pl.2 Fig.5

             Syn.: Dicyrtoma Alveolus(sic) Kolenati, FA, 1858:134 ISS
            Syn.: Dicyrtoma alveolus(sic) Lubbock, J, 1873:119 IOS
            Syn.: Dicyrtoma alveolus(sic) Stach, J, 1957:65,si IOS
            Syn.: Dicyrtoma alveolus(sic) Salmon, JT, 1964:634 IOS
             Syn.: Dicyrtoma alveolus(sic) Thibaud, J-M, 2013:234 ISS
Ptenothrix alveola from Portugal
2020.02.18 © Delgado, F.
Ptenothrix alveola from France
2020.11.25 © Tiky, S.
Ptenothrix alveola from France
Antenna
2020.11.25 © Tiky, S.
Ptenothrix alveola from France
Dorsal pattern
2020.11.25 © Tiky, S.
Ptenothrix alveola from France
Midfacial setae 1?+2+1+1+1
2020.11.25 © Tiky, S.
Ptenothrix alveola from Portugal
2020.12.12 © Delgado, F.





Species annulata Lin & Xia, 1985

             Syn.: Ptenothrix annulatus(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:130 ISS

Species argentina Delamare Deboutteville & Massoud, 1963:281

Ptenothrix argentina from Peru
2009.01-03.dd © Contreras Valdez, A.

Species atra (Linnæus, C, 1758:608) Börner, 1906, t.t., B

            Syn.: Podura antennis longioribus Hill, 1752 i
            Syn.: Podura polypoda Linnæus, 1761
             Syn.: Podura Polypoda(sic) Walckenær, CA, 1802:4 ISS
             Syn.: Podura Polypus(sic) Bourlet, C, 1845:152 ISS
             Syn.: Podura Polypoda(sic) Bourlet, C, 1845:154 ISS
             Syn.: Podura Polypoda(sic) Bourlet, C, 1845:155 ISS
Papirius polypodus from the UK
After Lubbock, J., 1873 Pl.11.

            Syn.: Podura globosa fusca de Geer, C, 1778:35
            Syn.: Papirius chloropus Stach, J, 1957:84 =?,si
            Syn.: Papirius annulatus Becker, 1902
            Syn.: Dicyrtoma setosa-ceratoleucaspis Börner, 1903
            Syn.: Ptenothrix setosa Denis, 1931
            Syn.: Papirius pini Christiansen, K et Bellinger, P, 1981:1231
            Syn.: Papirius pini Christiansen, K et Bellinger, P, 1998:1405



            Syn.: Podura atra Linnæus, 1746 i
             Syn.: Smynthurus(sic) atra(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:97 ISS
             Syn.: Podura Atra(sic) Bourlet, C, 1845:152 ISS
             Syn.: Podura Atra(sic) Bourlet, C, 1845:154 ISS
             Syn.: Sminthurus Ater(sic) Bourlet, C, 1845:160 ISS
             Syn.: Ptenothrix ater(sic) Denis, J-R, 1933:314 ISS
             Syn.: Ptenothrix ater(sic) Kloet, GS et Hincks, WD, 1945:9 ISS
             Syn.: Ptenothrix ater(sic) Denis, J-R, 1947:12 ISS
Ptenothrix atra from the UK
2004 © Hopkin, S.P.
Ptenothrix atra from the UK
2006.06.22 © Botting, J.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra from the UK
2008.08.16 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.10.10 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from the UK
2008.12.03 © Ardron, P.A.
Ptenothrix atra juv. from France
2009.09.dd © Lebeaux, P.
Ptenothrix atra juv. from the UK
2010.01.16 © Henton-Jones, D.
Ptenothrix atra juv. from Holland
2011.12.02 © van Duinen, J.
Ptenothrix atra from the UK
Trunk dorsal anterior macrosetae strongly spinelike
2013.02.10 © Prince, M.
Ptenothrix atra from the UK
2015.02.17 © Murray, A.
Ptenothrix atra from the UK
Cheek reticulate
2015.02.17 © Murray, A.
Ptenothrix atra from the UK
Ant.4 short and white
2015.02.17 © Murray, A.
Ptenothrix atra juv. from the UK
2015.02.17 © Murray, A.
Ptenothrix atra from the UK
2015.02.18 © Murray, A.
Ptenothrix atra from the UK
2015.02.18 © Murray, A.
Ptenothrix atra from the UK
Posterior dorsal great abdomen
2015.02.18 © Murray, A.
Ptenothrix atra from Sweden
2016.07.26 © Al Abbasi, S.
Ptenothrix atra from the UK
First instar 0.4mm
2016.12.03 © Holla, D.
Ptenothrix atra from the USA
2018.06.04 © Caterino, M.
Ptenothrix atra from the USA
Cheek reticulate
2018.06.04 © Caterino, M.
Ptenothrix atra from Poland
2020.10.01 © Antosik-Zoladek, J.
Ptenothrix atra from Poland
Spiky thoracic macrosetae AA-CC
2020.10.01 © Antosik-Zoladek, J.
Ptenothrix atra from Poland
2020.10.01 © Antosik-Zoladek, J.
Ptenothrix atra from Poland
Ant.3 apically white, ant.4 white
2020.10.01 © Antosik-Zoladek, J.
Ptenothrix atra from the USA
2023.03.31 © Wang, G.
Ptenothrix atra from the USA
2023.03.31 © Wang, G.
Ptenothrix atra juv. from the USA
Considering an escape jump
2024.05.04 © Coogler, J.
Ptenothrix atra juv. from the USA
Cheek and body reticulate
2024.05.04 © Coogler, J.



Species australis Snider, RJ, 1990:247

Ptenothrix (Ptenothrix) australis from the USA
After Snider R, 1990 Fig.1
Ptenothrix (Ptenothrix) australis from the USA
Dorsal pattern
After Snider R, 1990 Fig.1
Ptenothrix (Ptenothrix) australis from the USA
After Snider R, 1990 Fig.2
Ptenothrix (Ptenothrix) australis from the USA
Antenna pattern
After Snider R, 1990 Fig.2
Ptenothrix (Ptenothrix) australis from the USA
Tibiotarsus ringed
After Snider R, 1990 Fig.2

             Syn.: Ptenothrix (Ptenothrix) austalis(sic) Snider, RJ, 1990:247 ISS


Species beta (Christiansen, K & Bellinger, P, 1981:1233) Christiansen, K & Bellinger, P, 1998:1407

Ptenothrix (Ptenothrix) beta from the USA
After Christiansen K & Bellinger P, 1998 Fig.1035A
Ptenothrix beta from the USA
2004 © Maddison, D.R.
Ptenothrix beta from the USA
2004 © Maddison, D.R.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.03.30 © Gross, J.
Ptenothrix beta from the USA
2007.10.28 © Gross, J.
Ptenothrix beta from the USA
2007.10.28 © Gross, J.
Ptenothrix beta from the USA
2007.11.07 © Ten Eyck, C.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.10.25 © Hall, K.
Ptenothrix beta from the USA
2008.12.09 © Bradford, A.
Ptenothrix beta from the USA
2009.03.30 © Bradford, A.
Ptenothrix beta from the USA
2009.04.01 © Bradford, A.
Ptenothrix beta from the USA
2009.04.08 © Zungri, D.
Ptenothrix beta from the USA
2009.10.22 © Ten Eyck, C.
Ptenothrix beta from the USA
2009.10.22 © Ten Eyck, C.
Ptenothrix beta from the USA
2010.01.02 © LeMons, J.
Ptenothrix beta from the USA
2010.01.03 © Zungri, D.
Ptenothrix beta from the USA
2010.01.07 © LeMons, J.
Ptenothrix beta from the USA
2010.01.21 © Bradford, A.
Ptenothrix beta from the USA
2010.06.11 © Matthews, E.
Ptenothrix beta from the USA
2012.02.26 © Elliot.
Ptenothrix beta from the USA
Pale colour form
2012.07.11 © Goff, M.
Ptenothrix beta from the USA
2012.10.06 © Baker, A.
Ptenothrix beta from the USA
2016.02.16 © Lee, C.
Ptenothrix beta juv. from the USA
Preparing to moult
2017.01.02 © Abela, A.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from the USA
2017.01.12 © Gross, J.
Ptenothrix beta from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix beta from the USA
2018.03.18 © Schneider, K.
Ptenothrix beta from the USA
2018.03.18 © Schneider, K.
Ptenothrix beta from the USA
Colour form with pale dorsum
2018.07.01 © Parker, H.S.
Ptenothrix beta from the USA
Colour form with pale dorsum
2018.07.01 © Parker, H.S.
Ptenothrix beta from the USA
Colour form with pale dorsum
2018.07.06 © Parker, H.S.
Ptenothrix beta from the USA
2019.01.23 © Bairstow, A.
Ptenothrix beta from the USA
2020.01.24 © Lowe, E.
Ptenothrix beta from the USA
2020.01.30 © Kummel, M.
Ptenothrix beta juv. from the USA
2022.11.19 © Wang, G.
Ptenothrix beta juv. from the USA
2022.11.19 © Wang, G.
Ptenothrix beta juv. from the USA
2022.11.19 © Wang, G.
Ptenothrix beta from the USA
2023.01.08 © Wang, G.
Ptenothrix beta from the USA
2023.01.08 © Wang, G.
Ptenothrix beta from Canada
2023.12.11 © Chan, J.
Ptenothrix beta from Canada
With 2 unpaired midfacial setae
2023.12.11 © Chan, J.

Species blidana Bretfeld, G, 2001:119

Ptenothrix blidana from Algeria
After Bretfeld G, 2001 Fig.43
Ptenothrix blidana from Algeria
Facial chaetotaxy
After Bretfeld G, 2001 Fig.44
Ptenothrix blidana from Algeria
Anal valves chaetotaxy
After Bretfeld G, 2001 Fig.49
Ptenothrix ?blidana? from Portugal
2006 © Almeida, J.
Ptenothrix ?blidana? juv. from Portugal
2011.02.25 © Almeida, J.
Ptenothrix blidana from La Palma
Ant.2 with 1 distal white ring
2012.12.01 © Barton, T.
Ptenothrix blidana from La Palma
2012.12.01 © Barton, T.
Ptenothrix blidana from La Palma
2012.12.04 © Barton, T.
Ptenothrix blidana from La Palma
More than 2 unpaired facial setae
2012.12.04 © Barton, T.
Ptenothrix ?blidana? from Portugal
2018.02.18 © Almeida, J.
Ptenothrix ?blidana? from Portugal
2018.11.05 © Almeida, J.

Species borincana Soto-Adames, FN, 1988:65

Ptenothrix borincana from Puerto Rico
After Soto-Adames FN, 1988 Fig.34

Species brasiliensis Delamare Deboutteville & Massoud, 1963:285

             Syn.: Ptenothrix braziliensis(sic) Yosii, R, 1969:220 ISS
             Syn.: Ptenothrix braziliensis(sic) Yosii, R, 1969:222 ISS
Ptenothrix ca brasiliensis from Brazil
After Cipola NG & al, 2019 Fig.15



Species brouquissei Nayrolles, 1989

Species californica (Christiansen, K & Bellinger, P, 1981:1234) Christiansen, K & Bellinger, P, 1998:1408

            Syn.: ... marmorata Scott & Yosii, 1972


Ptenothrix (Ptenothrix) californica from the USA
After Christiansen K & Bellinger P, 1998 Fig.1036A
Ptenothrix cf. californica from the USA
2005 © Leonard, W.
Ptenothrix californica from the USA
2009.12.31 © Zungri, D.

Species castanea Snider, RJ, 1985:582

Dicyrtoma (Ptenothrix) castanea from the USA
After Snider RJ, 1985 Fig.1
Dicyrtoma (Ptenothrix) castanea from the USA
After Snider RJ, 1985 Fig.2
Ptenothrix castanea from Canada
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
Lateral bothriotricha form distinct angle
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
Bothriotricha A-D
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
Dorsal papilla with bothriotrichum A directed forwards
2011.07.12 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.14 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.14 © Hochhalter, R.
Ptenothrix castanea from Canada
2011.07.24 © Hochhalter, R.
Ptenothrix ?castanea? from the USA
2014.04.28 © Peterson, M.A.
Ptenothrix castanea from Canada
2022.05.21 © Rennert, M.
Ptenothrix castanea from Canada
2022.10.02 © Rennert, M.
Ptenothrix castanea from Canada
Dorsal spike-like macrosetae EE+DD
Positions of bothriotricha A-D
203.04.20 © Hochhalter, R.
Ptenothrix castanea from Canada
Dorsal spike-like macrosetae EE+DD
Dorsal papilla and bothriotrichum A
203.04.20 © Hochhalter, R.
Ptenothrix castanea from Canada
Small abdomen with 3 dorsal M setae in tranverse row
203.04.20 © Hochhalter, R.

Species catenata (Schött, 1921) Richards, 1968

            Syn.: Ptenothrix catenata (Schött, 1921) nec Börner, 1909 sensu Bellinger, 1996

Species cavicola Cassagnau, P et Delamare Deboutteville, C, 1955:392

Species ciliata Stach, J, 1957:102

Ptenothrix ciliata from Poland
After Stach J, 1957 Pl.VIII fig.1

            Syn.: Ptenothrix atra Berg, M, 2008:50
Ptenothrix atra from Holland
After Berg, M, 2008 Fig.1b
Ptenothrix atra from Holland
After Berg, M, 2008 Fig.1a




Ptenothrix ciliata from Holland
After Berg, M, 2009 Fig.1

Species ciliophora Yosii, R et Lee, C-E, 1963:17

Ptenothrix ciliophora from Korea
After Yosii R & Lee C-E, 1963 Fig.9A
Ptenothrix ?ciliophora? from Taiwan
2022.07.13 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
2022.07.13 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
2022.07.15 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
2022.09.16 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
Mid trochanter with suddenly bent macroseta
2022.09.16 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
2022.09.16 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
Ant.3 organ
2022.09.19 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
Bothriotrichum A on dorsal papilla
2022.09.19 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
Hind tibiotarsus with sparsely serrate modified macrosetae
2022.09.19 © Cheng, H.-J.
Ptenothrix ?ciliophora? from Taiwan
Femur bluish
2022.09.19 © Cheng, H.-J.

Species corynophora Börner, C, 1909:130

            Syn.: Ptenothrix corynophora var. cincta Börner, C, 1909:130
            Syn.: Ptenothrix corynephora(sic) var. cincta Yosii, R, 1977:166
            Syn.: Ptenothrix corynophora var. sellata Börner, C, 1909:130
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4C

            Syn.: Ptenothrix corynephora(sic) var. sellata Yosii, R, 1977:166
Ptenothrix corynophora f. sellata from Taiwan
With constricted dorsal spot
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata from Taiwan
With constricted dorsal spot
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2022.07.11 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? juv. from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? juv. from Taiwan
2022.12.23 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? juv. from Taiwan
2022.12.23 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2023.10.22 © Cheng, H.-J.
Ptenothrix corynophora f. sellata? from Taiwan
2023.10.22 © Cheng, H.-J.

            Syn.: Ptenothryx(sic) corynephora(sic) f. shibanaii Yosii, R, 1955:400
Ptenothryx(sic) corynephora(sic) f. shibanaii from Tokara Island
After Yosii R, 1955 Abb.10A

            Syn.: Ptenothrix corynephora(sic) f. shibanaii Yosii, R, 1965:64
            Syn.: Ptenothrix corynephora(sic) f. shibanaii Yosii, R, 1977:166
            Syn.: Ptenothrix tokarensis Yosii, R, 1965:64
             Syn.: Ptenothrix takarensis(sic) Yosii, R, 1965:64 ISS
Ptenothrix takarensis(sic) from Japan
After Yosii R, 1965 Fig.40A
Ptenothrix corynophora f. tokarensis from Taiwan
2021.11.22 © Tsai, Y.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2021.11.22 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2021.11.23 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
Antenna
2021.11.23 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2021.12.20 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.04.19 © Cheng, H.-J.
Leg.: Lin, T.-P.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.04.19 © Cheng, H.-J.
Leg.: Lin, T.-P.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.05.13 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.05.13 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
Mucro with apical notch
2022.10.18 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2022.10.18 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2023.03.01 © Cheng, H.-J.
Ptenothrix corynophora f. tokarensis from Taiwan
2023.03.01 © Cheng, H.-J.

            Syn.: Ptenothrix corynophora var. withpaleface Janssens & Cheng, 2022
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4A
Ptenothrix corynophora from Japan
After Uchida H, 1957 Fig.4B
Ptenothrix corynophora from Taiwan
With pale face
With pale diffuse antennae
2022.04.09 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
With pale face
With pale diffuse antennae
2022.05.11 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
With pale face
With pale diffuse antennae
2022.05.17 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
With pale face
With pale diffuse antennae
2022.07.12 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
With pale face
With pale diffuse antennae
2022.07.12 © Cheng, H.-J.




             Syn.: Ptenothrix corynephora(sic) Yosii, 1954 ISS
             Syn.: Ptenothryx(sic) corynephora(sic) Yosii, R, 1955:380 ISS
             Syn.: Ptenothrix corynephora(sic) Yosii, R, 1955:399 ISS
             Syn.: Ptenothrix corynephora(sic) Stach, J, 1957:78 ISS
             Syn.: Ptenothrix corynephora(sic) Yosii, R et Lee, C-E, 1963:26 ISS
Ptenothrix corynephora(sic) from Japan
After Yosii R & Lee E-C, 1963 Fig.14A

             Syn.: Ptenothrix corynephora(sic) Yosii, R, 1965:64 ISS
             Syn.: Ptenothrix corynephopa(sic) Yosii, R, 1965:65 ISS
             Syn.: Ptenothrix corynephora(sic) Yosii, R, 1977:166 ISS
Ptenothrix corynophora from Japan
After Uchida, 1957 in Itoh R & al, 2012 Fig.53a
Ptenothrix corynophora from Japan
2018.07.30 © Yoshida, Y.
Ptenothrix corynophora from Taiwan
2022.05.06 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
2022.05.06 © Cheng, H.-J.
Ptenothrix corynophora from Taiwan
Mucro with distinct apical notch (*)
2022.06.15 © Cheng, H.-J.



Species curvilineata (Wray, 1949) Snider, RJ, 1990:256

Ptenothrix (Ptenothrix) curvilineata from the USA
After Snider, R., 1990 Fig.76.
Ptenothrix (Ptenothrix) curvilineata from the USA
After Snider, R., 1990 Fig.77.
Ptenothrix curvilineata from the USA
2012.01.30 © DeSantis, T.
Ptenothrix curvilineata from the USA
2018.01.20 © Abbott, J.C.
Ptenothrix curvilineata from the USA
2024.02.07 Van Epps, R.
Ptenothrix curvilineata from the USA
2024.02.07 Van Epps, R.

Species dalii Zeppelini D, Ferreira AS et Oliveira JVLC in Oliveira, JVLC, Ferreira, AS et Zeppelini, D, 2020:2

Ptenothrix dalii from Brazil
After Oliveira JVLC et al, 2020 Fig.2 left
Ptenothrix dalii from Brazil
After Oliveira JVLC et al, 2020 Fig.2 right

Species deharvengi Nayrolles, 1989

Species delongi (Christiansen & Bellinger, 1981) Christiansen, K & Bellinger, P, 1998:1409

Ptenothrix (Ptenothrix) delongi from the USA
After Christiansen K & Bellinger P, 1998 Fig.1037A

Species denticulata (Folsom, JW, 1899:268) Börner, C, 1909:131

Papirius denticulatus from Japan
After Folsom JW, 1899 Pl.3 Fig.33
Papirius denticulatus from Japan
Unguiculus with 2 basal teeth (*)
After Folsom JW, 1899 Pl.3 Fig.34
Papirius denticulatus syntype from Japan
2012.09.21 © Farnum W @MCZ
Museum of Comparative Zoology Harvard
Papirius denticulatus syntype from Japan
2012.09.21 © Farnum W @MCZ
Museum of Comparative Zoology Harvard
Cephalic pattern

            Syn.: Ptenothrix denticulata var. catenata Börner, C, 1909:131
Ptenothrix denticulata from Japan
After Kinoshita S, 1932 X17
Ptenothrix denticulata var. catenata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata var. catenata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata var. catenata from Japan
2008.09.15 © Tsukiji, T.
Ptenothrix denticulata var. catenata from Japan
2019.06.28 © Anth, F.
Ptenothrix denticulata var. catenata from Japan
2019.06.28 © Anth, F.
Ptenothrix nr denticulata ♀ var. catenata from the USA
With female sub-anal appendage (SAA)
2021.09.01 © Karighattam, A.
Ptenothrix denticulata ♀ var. catenata from the USA
2021.09.01 © Karighattam, A.
Ptenothrix denticulata var. catenata from the USA
2021.09.01 © Karighattam, A.
Ptenothrix denticulata var. catenata from the USA
Grazing
2021.10.21 © Karighattam, A.
Ptenothrix denticulata var. catenata from the USA
Caudal aspect
2021.10.21 © Karighattam, A.
Ptenothrix denticulata ♀ var. catenata from the USA
2021.10.21 © Karighattam, A.
Ptenothrix denticulata ♀ var. catenata from the USA
Caudal aspect
2021.10.21 © Karighattam, A.
Ptenothrix denticulata var. catenata from the USA
2021.10.22 © Karighattam, A.
Ptenothrix denticulata ♀ var. catenata from the USA
2021.10.22 © Karighattam, A.
Ptenothrix denticulata ♀ var. catenata from the USA
7 macrosetae on posteriorly on dorsal anal valve
2021.10.22 © Karighattam, A.
Ptenothrix denticulata juv. var. catenata from the USA
2021.10.22 © Karighattam, A.
Ptenothrix denticulata var. catenata from Japan
Cephalic dorsal pattern
Modified after Folsom, 1899
2022.08.12 Janssens, F.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
Cephalic dorsal pattern
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata juv. from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
2023.01.18 © Cheng, H.-J.
Leg.: Nakamori, T.
Ptenothrix denticulata var. catenata from Japan
Hind unguicular basal tooth with denticle
2023.02.10 © Cheng, H.-J.
Leg.: Nakamori, T.




Papirius denticulatus from Japan
After Matsumura S, 1907 Plate 1 Fig.9
Ptenothrix denticulata from Japan
After Yosii R & Lee E-C, 1963 Fig.13A
Ptenothrix denticulata from Japan
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
Modified rough setae on posterior side of 3rd tibiotarsus
2015.11.03 © Murray, A.
Ptenothrix denticulata from Japan
2015.11.07 © Murray, A.
Ptenothrix denticulata from China
2023.02.09 © Yeung, D.

Species denticulata (Yosii R, 1954:824 nec Folsom) Yosii, R, 1965:65

            Syn.: Ptenothryx(sic) denticulata Yosii R, 1954:824 nec Folsom
Ptenothryx(sic) denticulata from Japan
Cephalic setae sharply pointed
After Yosii R, 1954 Abb.36
Ptenothrix denticulata from Japan
2014.08.29 © Hara, Y.
Ptenothrix denticulata from Japan
2014.08.29 © Hara, Y.
Ptenothrix denticulata from Japan
2016.07.01 © Hara, Y.
Ptenothrix denticulata from Japan
Cleaning dorsum with drop of saliva
2016.07.04 © Hara, Y.
Ptenothrix denticulata from Japan
Just moulted
2018.10.31 © Hara, Y.
Ptenothrix denticulata from Japan
Ant.3 with apical pseudosubdivisions (*)
2018.10.31 © Hara, Y.



Species dinghuensis Lin & Xia, 1985

             Syn.: Ptenothrix diinghuensis(sic) Chen, J-X et Christiansen, K, 1996:587 ISS

Species dominicana Mari Mutt, JA, 1977:285

Species dorsalis (Reuter, OM, 1876:85) Linnaniemi, 1912

            Syn.: Papirius ater var. albosignatus Parona, 1888
            Syn.: Dicyrtoma setosa var. ceratoleucaspis Börner, 1903 =?


Remark by Bretfeld, G. 1999:158: The colour variety dorsalis Reuter, 1876 of atra is incompletely known and may represent a separate species.

Species fiscellata Handschin, 1925

            Syn.: Ptenothrix fiscellata var. obscura Handschin, E, 1926:448


Species flavescens (Axelson, WM, 1905:794) Axelson, 1907

Remark by Bretfeld, G. 1999:158: The colour variety flavescens Axelson, 1905 of atra is incompletely known and may represent a separate species.

Species fuscamaculata Bretfeld & Trinklein, 2000

Species gaoligongshanensis Itoh, R et Zhao, L-J, 2000:156

Ptenothrix gaoligongshanensis from China
After Itoh R & Zhao L-J, 2000 Fig.7A
Ptenothrix gaoligongshanensis from China
After Itoh R & Zhao L-J, 2000 Fig.7B

Species gibbosa Börner, C, 1906:185

            Syn.: Ptenothrix pulchella Handschin, 1928
             Syn.: Ptenothrix pulchellus(sic) Stach, J, 1957:77 ISS
             Syn.: Ptenothrix pulchellus(sic) Uchida, H, 1957:29 ISS
             Syn.: Ptenothrix pulchellus(sic) Yayuk, RS(sic), 1989:20 ISS


Species gracilicornis (Schäffer, C, 1898:423) Börner, 1903

             Syn.: Sminthurus gracillicornis(sic) Yosii, R, 1969:217 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:217 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:218 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:219 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:220 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:221 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:222 ISS
             Syn.: Ptenothrix gracillicornis(sic) Yosii, R, 1969:224 ISS
Ptenothrix gracillicornis(sic) from the Solomon Islands
After Yosii R, 1969 Fig.1A

             Syn.: Ptenothrix gracilis(sic) Oliveira, JVLC, Ferreira, AS et Zeppelini, D, 2020:1 lapsus,gr ISS


Species hawaiiensis Snider, RJ, 1990:74

Ptenothrix (Ptenothrix) hawaiiensis from Hawaii
After Snider RJ, 1990 Fig.1

Species higashihirajii Yosii, R, 1965:67

Ptenothrix higashihirajii from Japan
After Yosii R, 1965 Fig.42A

            Syn.: Ptenothrix higashihirajii f. bilineata Yosii, R, 1965:68
Ptenothrix higashihirajii f. bilineata from Japan
After Yosii R, 1965 Fig.42B




Species higumai Yosii, R, 1965:68

Ptenothrix higumai from Japan
After Yosii R, 1965 Fig.43A

Species himalayensis Yosii, 1966

Species huangshanensis Chen, J-X et Christiansen, K, 1996:587

Ptenothrix huangshanensis from China
After Chen J-X & Christiansen K, 1996 Fig.1A
             Syn.: Ptenothrix huanshanensis(sic) Chen, J-X et Christiansen, K, 1996:589 ISS


Species iriomotensis Uchida, H, 1965:92

Ptenothrix iriomotensis from Japan
After Uchida, 1965 in Itoh R & al, 2012 Fig.62a
Ptenothrix iriomotensis from Japan
After Uchida, 1965 in Itoh R & al, 2012 Fig.62b

Species janthina Börner, C, 1909:132

            Syn.: Papirius unicolor Börner, C, 1909:132 =?
            Syn.: Ptenothrix setosa var. picta Börner, C, 1909:131
             Syn.: Ptenothrix setosa var. picata(sic) Yosii, R, 1977:166 ISS
            Syn.: Ptenothrix atra var. setosa Uchida, 1952
            Syn.: Ptenothrix atra var. setosa Uchida, 1959
            Syn.: Ptenothrix atra ssp. setosa Salmon, JT, 1964:641
            Syn.: Ptenothrix setosa f. principalis Yosii, R, 1977:166


Ptenothrix janthina from Japan
2014.01.13 © Komatsu, T.
Ptenothrix janthina juv. from Japan
2015.01.16 © Jiku.
Ptenothrix janthina juv. from Japan
2015.01.16 © Jiku.
Ptenothrix janthina juv. from Japan
Pre-jump posture
2015.01.16 © Jiku.

Species keralae Prabhoo, 1971

Species koreanensis Jo, S-H et Park, K-H, 2023:126

Ptenothrix koreanensis from South Korea
After Jo S-H & Park K-H, 2023 Fig.1A
Ptenothrix koreanensis from South Korea
After Jo S-H & Park K-H, 2023 Fig.1C

Species kuraschvilii Djanaschvili, 1970

Species lactea Uchida, 1953

Ptenothrix lactea from Japan
After Uchida, 1953 in Itoh R & al, 2012 Fig.50a

Species leucostrigata Stach, J, 1957:98

Ptenothrix leucostrigata from Poland
After Stach J, 1957 Pl.VIII fig.6
            Syn.: Ptenothrix leucostrigata var. simplificata Stach, J, 1957:101



Ptenothrix leucostrigata from Estonia
2006.09.17 © Tartes, U.
Ptenothrix leucostrigata from Estonia
2006.10.01 © Tartes, U.
Ptenothrix leucostrigata from Denmark
2015.08.05 © Jäschke, J.
Ptenothrix leucostrigata from Germany
2016.07.02 © Jäschke, J.
Ptenothrix leucostrigata from Germany
Antenna colouration pattern
2016.07.02 © Jäschke, J.
Ptenothrix leucostrigata from Poland
2017.09.29 © Ryszard.
Ptenothrix leucostrigata from Poland
Dark form
2017.10.18 © Ryszard.
Ptenothrix leucostrigata from Poland
Dark form with asymmetric dorsal pattern
2017.10.18 © Ryszard.
Ptenothrix leucostrigata from Poland
2020.10.01 © Antosik-Zoladek, J.

Species macomba (Wray, 1967) Snider, RJ, 1990:255

Ptenothrix (Ptenothrix) macomba from the USA
After Snider R, 1990 Fig.74
Ptenothrix (Ptenothrix) macomba from the USA
After Snider R, 1990 Fig.75
Ptenothrix macomba from the USA, juvenile
2007.06.22 © Montgomery, G.
Det. Snider, R.
Ptenothrix macomba from the USA
2007.07.07 © Montgomery, G.
Det. Snider, R.
Ptenothrix cf. macomba from New Zealand
2007.12.17 © van de Vyver, L.
Ptenothrix macomba from the USA
2020.11.15 © Ekker, S.
Ptenothrix macomba from the USA
2020.11.15 © Ekker, S.
Ptenothrix macomba from the USA
2020.11.15 © Ekker, S.
Ptenothrix macomba from the USA
Caudal aspect
2020.11.15 © Ekker, S.
Ptenothrix macomba from the USA
Frontal aspect
2020.11.15 © Ekker, S.

Species maculosa (Schött, H, 1891:14) Matsumura & Ishida, 1931

Papirius maculosus from the USA
After Schött H, 1891 Taf.III Fig.1
Papirius maculosus from the USA
Mucro
After Schött H, 1891 Taf.III Fig.2
Papirius maculosus from the USA
Foot of first leg
After Schött H, 1891 Taf.III Fig.3

            Syn.: Ptenothrix marmorata Uchida, H, 1954:64 =?
Ptenothrix marmorata from Japan
After Uchida H, 1954 Fig.3A
Ptenothrix marmorata from Japan
Facial pattern
After Uchida H, 1954 Fig.3A
Ptenothrix marmorata from Japan
Dorsal pattern
After Uchida H, 1954 Fig.3A




             Syn.: Ptenothrix maculosus(sic) Richards, WR, 1968:26 ISS
Dicyrtoma (Ptenothrix) maculosa from the USA
After Christiansen K & Bellinger P, 1981 Fig.1038B
Dicyrtoma (Ptenothrix) maculosa from the USA
After Christiansen K & Bellinger P, 1981 Fig.1038C

             Syn.: Dicyrtoma maculosus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix maculosa from the USA
2017.01.19 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.01.20 © Dobbs, Br.
Ptenothrix maculosa from the USA
Dark colour form
2017.01.27 © Dobbs, Br.
Ptenothrix maculosa from the USA
Very dark colour form
2017.01.29 © Dobbs, Br.
Ptenothrix maculosa from the USA
2017.02.12 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern weak
2018.01.13 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern intermediate
2018.01.13 © Dobbs, Br.
Ptenothrix maculosa from the USA
Cephalic pattern strong
2018.01.13 © Dobbs, Br.

  Subspecies olympia MacGillivray, AD, 1894:110

            Syn.: Papirius olympius MacGillivray, AD, 1894:110
            Syn.: Papirius maculosus Scott & Yosii, 1972
            Syn.: Dicyrtoma (Ptenothrix) maculosa Christiansen, K et Bellinger, P, 1981:1237 partim
Dicyrtoma (Ptenothrix) maculosa from Japan
After Scott & Yosii, 1972 in Christiansen K & Bellinger P, 1981 Fig.1038A

            Syn.: Ptenothrix maculosa Itoh, R, Hasegawa, M, Ichisawa, K, Furuno, K, Suma, Y, Tanakas, S, Hasegawa, M et Niijim, K, 2012:152
Ptenothrix maculosa from Japan
After Scott & Yosii, 1972 in Itoh R & al, 2012 Fig.65a

            Syn.: Ptenothrix maculosa ssp. olympia f.1 Janssens F & Moorehead C, 2008 sensu Janssens F, 2017
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.01.14 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.02.15 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2008.02.16 © Moorehead, C.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2010.01.02 © LeMons, J.
Ptenothrix sp. f.1 ♂ from the USA
Dorsum pale
2010.01.02 © LeMons, J.
Ptenothrix sp. f.1 from the USA
Dorsum pale
2010.01.03 © LeMons, J.
Ptenothrix sp. f.1 from Canada
Dorsum pale
2015.02.14 © Gilmore, S.
Ptenothrix sp. f.1 from Canada
Dorsum pale
2015.02.14 © Gilmore, S.
Ptenothrix maculosa f.1 from the USA
Dorsum pale
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa f.1 from the USA
2016.03.14 © Dobbs, Br.
Ptenothrix maculosa f.1 from the USA
Facial pigmentation
2016.03.14 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.02.27 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2018.12.15 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1 from Canada
2019.01.31 © Gilmore, S.
Ptenothrix maculosa ssp. olympia f.1 from the USA
2020.12.10 © Duncan, W.L.
Ptenothrix maculosa ssp. olympia f.1♀ the USA
2020.12.26 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1♀ the USA
Dorso-caudal aspect
2020.12.26 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1 ♀ the USA
Caudal aspect
2021.01.16 © Pearson, R.R.
Ptenothrix maculosa ssp. olympia f.1 ♂ from Canada
Male genital papilla
2024.02.03 © Rennert, M.

            Syn.: Ptenothrix maculosa ssp. olympia f.2 Janssens F & Moorehead C, 2008 sensu Janssens F, 2017
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.09 © Moorehead, C.
Ptenothrix sp. f.2 from the USA
Anterior dark + posterior pale
2008.02.11 © Moorehead, C.
Ptenothrix maculosa f.2 from the USA
Anterior dark + posterior pale
2016.02.04 © Dobbs, Br.
Ptenothrix maculosa f.2 from the USA
Anterior dark + posterior pale
2016.02.04 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.2 from the USA
2018.02.07 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.2 from the USA
2018.02.07 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f.2 from Canada
2019.01.31 © Gilmore, S.

            Syn.: Ptenothrix maculosa ssp. olympia f.3 Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix sp. from the USA
Only pigmented middorsally
2008.01.12 © Moorehead, C.
Ptenothrix sp. from the USA
Only pigmented middorsally
2008.01.12 © Moorehead, C.
Ptenothrix maculosa from the USA
2009.01.31 © Leppin, M.
Ptenothrix maculosa f.3 from the USA
Midfacial setae : 1+1+2+2
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from the USA
Pigmented middorsally, unpigmented laterally
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from the USA
Caudal aspect
2016.03.05 © Dobbs, Br.
Ptenothrix maculosa f.3 from Canada
2016.11.11 © Barbin, T.

            Syn.: Ptenothrix maculosa ssp. olympia f.4 Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from the USA
Completely dark
2016.03.06 © Dobbs, Br.
Ptenothrix maculosa f.4 from Canada
2016.11.11 © Barbin, T.
Ptenothrix maculosa f.4 from Canada
1+1+2+2+1+1 midfacial setae
2017.01.19 © Barbin, T.

            Syn.: Ptenothrix maculosa ssp. olympia f. subadult Janssens F & Dobbs B, 2016 sensu Janssens F, 2017
Ptenothrix maculosa subadult from the USA
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa subadult from the USA
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.11 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.11 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.15 © Dobbs, Br.
Ptenothrix maculosa subadult ♀ from the USA
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
Male genital papilla (*)
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa subadult ♂ from the USA
Roundish male genital papilla (*)
2016.03.20 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f. subadult from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f. subadult from the USA
With pale dorsum
2018.12.03 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f. subadult from the USA
With pale dorsum
2018.12.03 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia f. subadult from Canada
With pale dorsum
2020.06.23 © Barbin, T.
Ptenothrix maculosa ssp. olympia f. subadult from Canada
2020.06.23 © Barbin, T.




             Syn.: Ptenothrix olympius(sic) Mills & Rolfs, 1933 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1981:1218 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1981:1237 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1981:1238 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1998:1411 ISS
             Syn.: Papirius olympia(sic) Christiansen, K et Bellinger, P, 1998:1412 ISS
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2006 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.02.23 © Peden, S.
Ptenothrix maculosa from the USA
2007.03.19 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.12.06 © Ten Eyck, C.
Ptenothrix maculosa from the USA
2007.12.29 © Moorehead, C.
Ptenothrix maculosa from the USA
2007.12.29 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.29 © McDonald, G.
Ptenothrix maculosa from the USA
2008.01.30 © McDonald, G.
Ptenothrix maculosa from the USA
2008.02.09 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.02.10 © Moorehead, C.
Ptenothrix maculosa from the USA
2008.02.16 © Moorehead, C.
Ptenothrix maculosa from Canada
2008.12.08 © Turner, E.
Ptenothrix maculosa from Canada
2008.12.08 © Turner, E.
Ptenothrix maculosa from the USA
2009.01.01 © Schneider, K.
Ptenothrix maculosa from the USA
2009.01.01 © Schneider, K.
Ptenothrix maculosa from the USA
2009.02.22 © Schimming, L.
Ptenothrix maculosa from the USA
2010.01.07 © LeMons, J.
Ptenothrix maculosa from the USA
2010.01.22 © LeMons, J.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2011.11.20 © Peden, S.
Ptenothrix maculosa from the USA
2012.02.07 © Hochhalter, R.
Ptenothrix maculosa from Canada
Grooming with mouth secreted drop
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from Canada
2013.01.22 © Kyron, T.F.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.01.22 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.02.12 © Dobbs, Br.
Ptenothrix maculosa from the USA
Mid-facial setae : 1+1+2+2+1+1
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.03.04 © Dobbs, Br.
Ptenothrix maculosa from the USA
Unguis 1st leg slender + unguiculus with long filament
2016.03.07 © Dobbs, Br.
Ptenothrix maculosa ♂ from the USA
Raised genital papilla (*)
2016.03.08 © Dobbs, Br.
Ptenothrix maculosa ♂ from the USA
Dark lateral body
2016.03.08 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
Transvers split genital papilla (*)
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa ♀ from the USA
2016.03.13 © Dobbs, Br.
Ptenothrix maculosa from Canada
2016.03.20 © Gatten, J.
Ptenothrix maculosa from Canada
2016.04.06 © Barbin, T.
Ptenothrix maculosa from Canada
2016.04.06 © Barbin, T.
Ptenothrix maculosa from Canada
Dorsal macrosetae AA+BB indistinctly spinelike
2016.09.26 © Barbin, T.
Ptenothrix maculosa from Canada
Caudal aspect pigmentation
2016.09.26 © Barbin, T.
Ptenothrix maculosa juv. from the USA
With extended tubelike collophore vesicles
2016.10.17 © Dobbs, Br.
Ptenothrix maculosa from the USA
2016.11.24 © Gross, J.
Ptenothrix maculosa from the USA
2016.11.24 © Gross, J.
Ptenothrix maculosa from the USA
Greenish colour variant
2016.11.29 © Dobbs, Br.
Ptenothrix maculosa from the USA
Midfacial setae: 1+1+2+2+1+1
2016.11.29 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
Caudal aspect
2017.09.08 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.09.09 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.10.07 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia ♂ from the USA
2017.10.16 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia ♂ from the USA
2017.10.16 © Johnson, S.T.
Ptenothrix maculosa ssp. olympia from the USA
2017.10.23 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.21 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.21 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2017.12.22 © Dobbs, Br.
Ptenothrix maculosa ssp. olympia from the USA
2018.12.29 © Schneider, K.
Ptenothrix maculosa ssp. olympia from the USA
2019.02.21 © Parker, H.S.
Ptenothrix maculosa ssp. olympia from the USA
2019.02.21 © Parker, H.S.
Ptenothrix maculosa ssp. olympia from Canada
2020.07.01 © Barbin, T.
Ptenothrix maculosa ssp. olympia from Canada
2022.05.22 © Rennert, M.
Ptenothrix maculosa ssp. olympia from Canada
2022.05.22 © Rennert, M.
Ptenothrix maculosa ssp. olympia from the USA
2022.11.27 © G, S.



Species marmorata (Packard, AS, 1873:42) Folsom, 1928

            Syn.: Papirius testudineatus Folsom, JW, 1896:345
             Syn.: Papirius Testudineatus(sic) Jackson, AD, 1907:173 ISS
             Syn.: Ptenothrix testudineatus(sic) Handschin, 1928 ISS
             Syn.: Ptenothrix testudineatus(sic) Bird, 1930 ISS
             Syn.: Dicyrtoma testudineatus(sic) James, 1933 ISS
            Syn.: Papirius testudineatus Folsom, 1934
             Syn.: Ptenothrix testudineatus(sic) Maynard, EA, 1951:286 ISS
             Syn.: Dicyrtoma testudineatus(sic) Maynard, EA, 1951:286 ISS
             Syn.: Ptenothrix testudinesatus(sic) Uchida, H, 1954:64 ISS
             Syn.: Ptenothrix testudineatus(sic) Judd, 1965 ISS
             Syn.: Ptenothrix testudinatus(sic) Richards, WR, 1968:26 ISS
             Syn.: Ptenothrix testudineatus(sic) Skidmore, 1995 ISS
Ptenothrix marmorata f. testudineata from the USA
2006 © Murray, T.
Ptenothrix marmorata f. testudineata from the USA
2006 © Murray, T.
Ptenothrix marmorata ?f. testudineata? from the USA
2006 © Cresswell, S.
Ptenothrix marmorata f. testudineata from the USA
2007.07.04 © McClarin, J.
Ptenothrix marmorata f. testudineata from the USA
2007.08.02 © Murray, T.
Ptenothrix marmorata f. testudineata from Canada
2007.09.20 © Pilon, C.
Ptenothrix marmorata f. testudineata from the USA
2008.02.07 © Lowery, J.M.
Ptenothrix marmorata f. testudineata from the USA
2008.10.04 © Murray, T.
Ptenothrix marmorata f. testudineata from the USA
2008.10.25 © Murray, T.
Ptenothrix marmorata f. testudineata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata f. testudineata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata f. testudineata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata f. testudineata from the USA
2011.08.26 © Eiseman, C.
Ptenothrix marmorata f. testudineata from the USA
2016.08.13 © Schulz, K.
Ptenothrix marmorata f. testudineata from the USA
2016.08.13 © Schulz, K.
Ptenothrix marmorata f. testudineata from the USA
Dorsal aspect
Cephalic middorsal asymmetric pattern
2019.09.19 © Wechsler, D.
Ptenothrix marmorata f. testudineata from the USA
2021.10.30 © Shahan, T.
Ptenothrix marmorata f. testudineata from the USA
2021.10.30 © Shahan, T.
Ptenothrix marmorata f. testudineata from the USA
2021.10.30 © Shahan, T.
Ptenothrix marmorata f. testudineata ♀ from the USA
2023.04.27 © Gruver, Be.
Ptenothrix marmorata f. testudineata ♀ from the USA
2023.04.27 © Gruver, Be.
Ptenothrix marmorata f. testudineata ♀ from the USA
Ocelli C+D distinctly smaller
Ocellus C indented
2023.04.27 © Gruver, Be.
Ptenothrix marmorata f. testudineata from the USA
2024.01.21 © Gruver, Be.

            Syn.: Papirius maculosus Guthrie, JE, 1903:58 =?
Papirius maculosus from the USA
Ant.3 apically with 8 subdivisions
After Guthrie JE, 1903 Plate I Fig.1
Papirius maculosus from the USA
Foot of 3rd leg
After Guthrie JE, 1903 Plate I Fig.2
Papirius maculosus from the USA
Foot of 1st leg
After Guthrie JE, 1903 Plate I Fig.3
Papirius maculosus from the USA
Right eyepatch
Modified after Guthrie JE, 1903 Plate I Fig.4
Papirius maculosus from the USA
Mucro: dorsal (a), lateral (c), notched seta (b)
After Guthrie JE, 1903 Plate I Fig.5

            Syn.: Papirius maculosus James, 1933 =?
            Syn.: Ptenothrix oswegatchiensis Maynard, EA, 1951:283
Ptenothrix oswegatchiensis from the USA
After Maynard EA, 1951 Plate XXIV Fig.435

            Syn.: Papirius oswegatchiensis Christiansen, K et Bellinger, P, 1981:1239
Ptenothrix marmorata f. oswegatchiensis from the USA
Vinaceous colour
2007.06.07 © Murray, T.
Ptenothrix marmorata f. oswegatchiensis from the USA
Vinaceous colour
2007.06.07 © Murray, T.
Ptenothrix marmorata f. oswegatchiensis from the USA
Vinaceous colour
2007.06.07 © Murray, T.
Ptenothrix marmorata f. oswegatchiensis from the USA
2021.12.10 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2021.12.10 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2021.12.13 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Distinct thoracic dorsal pattern
2021.12.13 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Middorsal macrosetae CC-EE distinctly spinelike
2021.12.13 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Facial aspect
2021.12.13 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2021.12.29 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Caudal aspect
2021.12.31 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Caudal pattern
with transverse bands
2021.12.31 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Caudal pattern
with 3 transverse bands
2021.12.31 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Dorsal aspect
Cephalic middorsal asymmetric pattern
2021.12.31 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Facial aspect
2022.01.12 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2022.12.14 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2022.12.14 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2022.12.14 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2022.12.18 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Asymmetric pattern on vertex
2022.12.18 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
Abdominal pattern
2022.12.18 © Charny, V.
Ptenothrix marmorata f. oswegatchiensis from the USA
2024.05.11 © Coogler, J.
Ptenothrix marmorata f. oswegatchiensis from the USA
2024.05.25 © Coogler, J.
Ptenothrix marmorata f. oswegatchiensis from the USA
2024.05.25 © Coogler, J.




             Syn.: Ptenothrix marmoratus(sic) Folsom, 1928 ISS
             Syn.: Ptenothrix marmoratus(sic) Denis, J-R, 1933:310 ISS
             Syn.: Ptenothrix marmoratus(sic) Denis, J-R, 1933:312 ISS
             Syn.: Ptenothrix marmoratus(sic) Denis, J-R, 1933:314 ISS
             Syn.: Ptenothrix marmoratus(sic) Brimley, 1938 ISS
Ptenothrix marmorata from the USA
After Maynard EA, 1951 Plate XXIV Fig.434

             Syn.: Ptenothrix marmolatus(sic) Terrón-Sierra & Palacios-Vargas, 1991 ISS
             Syn.: Ptenothrix marmoratus(sic) Palacios-Vargas & Gómez-Anaya, 1993 ISS
             Syn.: Dicyrtoma marmoratus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix (Ptenothrix) marmorata from the USA
After Christiansen K & Bellinger P, 1998 Fig.1039B
Ptenothrix (Ptenothrix) marmorata from the USA
After Christiansen K & Bellinger P, 1998 Fig.1039C

             Syn.: Ptenothrix marmoratus(sic) Palacios-Vargas, JG et Castaño-Meneses, G, 2002:399 ISS
Ptenothrix marmorata from the USA
Climbing on smooth glass wall
Everting collophore vesicles at high speed
2007 © Bernard, E.C.
Ptenothrix marmorata juv. from the USA
2009.10.06 © Houston, S.
Ptenothrix marmorata from the USA
3rd & 4th antennal segment
2012.04.18 © Kouri, J.
Ptenothrix marmorata from the USA
2012.04.18 © Kouri, J.
Ptenothrix marmorata from the USA
2012.04.18 © Kouri, J.
Ptenothrix marmorata from the USA
2012.04.18 © Kouri, J.
Ptenothrix marmorata from the USA
2012.04.18 © Kouri, J.
Ptenothrix marmorata from the USA
2012.05.16 © Kouri, J.
Ptenothrix marmorata from the USA
2013.01.28 © Fleming, K.
Ptenothrix marmorata from the USA
2013.12.13 © Fleming, K.
Ptenothrix marmorata juv. from the USA
2015.05.04 © Anderson, J.
Ptenothrix marmorata from the USA
2015.09.07 © Brown, M.H.
Ptenothrix marmorata from the USA
2016.09.13 © Fleming, K.
Ptenothrix marmorata from the USA
2018.02.03 © Fleming, K.
Ptenothrix marmorata from the USA
2019.06.20 © Skitterbug.
Ptenothrix marmorata from the USA
2020.04.21 © Skitterbug.
Ptenothrix marmorata from the USA
2021.12.09 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.09 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.10 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.10 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.10 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.15 © Charny, V.
Ptenothrix marmorata from the USA
Head with transverse posterior band
2021.12.15 © Charny, V.
Ptenothrix marmorata from the USA
Legs ringed
2021.12.15 © Charny, V.
Ptenothrix marmorata from the USA
Facial aspect
2021.12.15 © Charny, V.
Ptenothrix marmorata from the USA
Facial aspect
2021.12.17 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.29 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.29 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.29 © Charny, V.
Ptenothrix marmorata from the USA
Caudal aspect
Small adomen
2021.12.31 © Charny, V.
Ptenothrix marmorata from the USA
2021.12.31 © Charny, V.
Ptenothrix marmorata from the USA
Head
Lateral pattern
2021.12.31 © Charny, V.
Ptenothrix marmorata from the USA
2022.01.02 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
2022.01.06 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
2022.01.06 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
2022.01.06 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
Ocelli C+D smaller
2022.01.06 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
Dorsal anterior pattern
2022.01.06 © Charny, V.
Ptenothrix marmorata from the USA
Preparing to moult
2 unpaired midfacial macrosetae
2022.01.06 © Charny, V.
Ptenothrix marmorata juv. from the USA
Ant.2 ringed
2022.01.18 © Charny, V.
Ptenothrix marmorata juv. from the USA
Tibiotarsus ringed
2022.01.18 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Reproductive instar
With distinctly raised genital papilla (*)
2022.01.20 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Reproductive instar
With distinctly raised genital papilla (*)
2022.01.20 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Feeding instar
Without genital papilla (*)
2022.01.20 © Charny, V.
Ptenothrix marmorata ♀ from the USA
Abd.6 with spiky macrosetae
M'+M+M' in transverse row
2022.01.20 © Charny, V.
Ptenothrix marmorata ♀ from the USA
Abd.6 with 3+1+3 macrosetae
2022.01.20 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Reproductive instar
With distinctly raised genital papilla (*)
2022.01.21 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Reproductive instar
With distinctly raised genital papilla (*)
2022.01.21 © Charny, V.
Ptenothrix marmorata ♂ from the USA
Reproductive instar
With distinctly raised genital papilla (*)
2022.03.26 © Charny, V.
Ptenothrix marmorata ♂ & ♀ from the USA
2022.04.05 © Charny, V.
Ptenothrix marmorata ♀ from the USA
Egg deposition
2022.11.21 © Gruver, Be.
Ptenothrix marmorata ♀ from the USA
Freshly deposited eggs
2022.11.21 © Gruver, Be.
Ptenothrix marmorata ♀ from the USA
Facial aspect
2022.11.21 © Gruver, Be.
Ptenothrix marmorata from the USA
Collophore vesicle pressed in wet surfacefilm to adsorb water
2022.12.14 © Gruver, Be.
Ptenothrix marmorata ♀ from the USA
Depositing egg
Shaking vigorously
2022.12.15 © Gruver, Be.
Ptenothrix marmorata from the USA
Collophore vesicle pressed in wet surfacefilm to adsorb water
2022.12.16 © Gruver, Be.
Ptenothrix marmorata from the USA
Antennae held back
2022.12.18 © Charny, V.
Ptenothrix marmorata from the USA
Facial pattern
2022.12.18 © Charny, V.
Ptenothrix marmorata from the USA
Abdominal pattern
2022.12.18 © Charny, V.
Ptenothrix marmorata from the USA
2023.01.03 © Shahan, T.
Ptenothrix marmorata from the USA
2023.01.03 © Shahan, T.
Ptenothrix marmorata from the USA
2023.01.03 © Shahan, T.
Ptenothrix marmorata from the USA
Antennae ringed
2024.03.10 © Charny, V.
Ptenothrix marmorata from the USA
Ant.2 ringed
Ant.1 light
2024.03.10 © Charny, V.



Species mongolica Yosii, 1954

            Syn.: Ptenothrix atra Yosii, 1949


Ptenothrix mongolica from Mongolia
Antenna
After Yosii R & Lee C-E, 1963 Fig.17A

Species monochroma Yosii, R et Lee, C-E, 1963:20

Ptenothrix monochroma from Japan
2013.05.30 © Hara, Y.
Ptenothrix monochroma from Japan
1,1,2,2,1,1 thick midfacial setae
2013.05.30 © Hara, Y.
Ptenothrix monochroma from Japan
Grooming
2013.05.30 © Hara, Y.
Ptenothrix monochroma from China
2014.08.05 © Yang, N.
Ptenothrix monochroma from China
2014.08.05 © Yang, N.
Ptenothrix monochroma from Taiwan
2014.10.04 © Huang, D.
Ptenothrix monochroma from Taiwan
Collophore vesicle pressed in watersurfacefilm
2021.11.29 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2021.11.29 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2021.11.29 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2021.11.29 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2022.02.05 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2022.06.12 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
Mucro with apical incision
2022.06.29 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2022.09.16 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
2022.09.16 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
M'-M-M' setae in transverse row
2022.09.16 © Cheng, H.-J.
Ptenothrix monochroma from Taiwan
M'-M-M' setae in transverse row
2022.10.03 © Cheng, H.-J.

Species narumii Uchida, 1940

Ptenothrix narumii from China
After Uchida H, 1940 in Bretfeld G, 1999 Fig.387
Ptenothrix nr narumii from Japan
2019.02.05 © Hara, Y.
Ptenothrix nr narumii from Japan
2019.02.05 © Hara, Y.
Ptenothrix nr narumii from Japan
Antenna
2019.02.05 © Hara, Y.
Ptenothrix nr narumii from Japan
Midfacial setae : 1+1+2+2+1+1
2019.02.05 © Hara, Y.

Species nigricornis (Ionesco, 1916) Stach, J, 1957:91

Remark by Bretfeld, G. 1999:158: The colour variety nigricornis Ionesco, 1916 of atra is incompletely known and may represent a separate species.

Species ocellata (Becker, 1902)

Species palawanensis Yosii, R, 1969:223

Species palmata (Folsom, JW, 1902:102) Stach, J, 1957:77

             Syn.: Ptenothrix palmatus(sic) Richards, WR, 1968:26 ISS
             Syn.: Dicyrtoma palmatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS


Ptenothrix palmata from the USA
2008.11.08 © Fleming, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from the USA
2008.11.22 © Hall, K.
Ptenothrix palmata from Canada
2011.07.24 © Hochhalter, R.
Ptenothrix palmata from Canada
2011.10.24 © Hochhalter, R.
Ptenothrix palmata from the USA
2016.06.22 © Dobbs, Br.
Ptenothrix palmata from the USA
2016.06.22 © Dobbs, Br.
Ptenothrix palmata from the USA
2017.07.15 © Dobbs, Br.
Ptenothrix palmata from the USA
2017.07.15 © Dobbs, Br.
Ptenothrix palmata from the USA
2018.01.15 © Dobbs, Br.
Ptenothrix palmata from the USA
2018.01.15 © Dobbs, Br.
Ptenothrix palmata from the USA
2018.01.15 © Dobbs, Br.
Ptenothrix palmata from Canada
2018.04.20 © Hochhalter, R.
Ptenothrix palmata from Canada
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
Protergum distinctly white
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
Ant.3 with apical pseudo-annulations (*)
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
2020.06.23 © Barbin, T.
Ptenothrix palmata from Canada
Small abdomen caudal aspect
Three spiky M macrosetae directed upwards
2020.06.24 © Barbin, T.
Ptenothrix palmata from Canada
Two unpaired midfacial macrosetae
2020.06.24 © Barbin, T.
Ptenothrix palmata from Canada
Dorsal macrosetae AA-CC spinelike
2020.06.24 © Barbin, T.
Ptenothrix palmata from Canada
Abdomen covered in spores
2020.07.21 © Spencer, Br.
Ptenothrix palmata from Canada
2020.07.21 © Spencer, Br.
Ptenothrix palmata from Canada
Ant.3 with apical pseudo-annulations (*)
2020.07.21 © Spencer, Br.
Ptenothrix palmata from the USA
2020.12.21 © Parker, H.S.
Ptenothrix palmata from the USA
2020.12.21 © Parker, H.S.
Ptenothrix palmata from Canada
2021.01.14 © Rennert, M.
Ptenothrix palmata from Canada
2021.01.14 © Rennert, M.
Ptenothrix palmata from Canada
2021.01.14 © Rennert, M.
Ptenothrix palmata ♀ from Canada
2022.05.21 © Rennert, M.
Ptenothrix palmata ♀ from Canada
Sub-anal appendages
2022.05.21 © Rennert, M.
Ptenothrix palmata from Canada
Dorsal macrosetae AA on light spots
2024.04.01 © Barbin, T.

Species palmisetacea Lin & Feng, 1985

            Syn.: Ptenothrix palmisetaceus(sic) Lin & Feng, 1985 IOS
             Syn.: Ptenothrix palmisetaceus(sic) Chen, J-X et Christiansen, K, 1996:587 ISS
             Syn.: Ptenothrix palmsetaceus(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:130 ISS



Species plumiseta Kang J-S et Lee B-H in Kang, J-S, Lee, B-H et Park, K-H, 2005:235

Ptenothrix plumiseta from Korea
After Kang J-S, Lee B-H & Park K-H, 2005 Fig.1A
             Syn.: Ptenothrix pluniseta(sic) Park, K-H et Kang, J-S, 2007:A106 ISS


Species quadrangularis (Mills, HB, 1934:106)

Species renateae (Snider, RJ, 1985:561) Snider, 1990

Dicyrtoma (Ptenothrix) renateae from the USA
After Snider RJ, 1985 Fig.1
Dicyrtoma (Ptenothrix) renateae from the USA
After Snider RJ, 1985 Fig.2

             Syn.: Ptenothrix (Ptenothrix) renatae(sic) Christiansen, K et Bellinger, P, 1998:1404 ISS
Ptenothrix renateae from the USA
2006 © Bernard, E.C.
Ptenothrix renateae from the USA
2008.02.28 © Bradford, A.
Ptenothrix renateae from the USA
2008.12.05 © Bradford, A.
Ptenothrix renateae from the USA
2009.02.24 © Bradford, A.
Ptenothrix renateae from the USA
2009.03.26 © Burishkin, J.
Ptenothrix renateae from the USA
Drinking from the watersurfacefilm with everted vesicles of collophore
2009.03.29 © Bradford, A.
Ptenothrix renateae from the USA
2009.03.30 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.07 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.07 © Bradford, A.
Ptenothrix renateae from the USA
2009.12.13 © Hart, J.
Ptenothrix renateae ♂ from the USA
2010.01.21 © Bradford, A.
Ptenothrix renateae from the USA
Bulby distinctly pale upper clypeus
2010.01.21 © Bradford, A.
Ptenothrix renateae from the USA
2010.03.09 © Zimlich, R.L.
Ptenothrix renateae from the USA
2013.02.11 © Anderson, J.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2013.04.17 © Schnepp, K.
Ptenothrix renateae from the USA
2016.01.01 © Gallagher, J.
Ptenothrix renateae from the USA
2016.01.01 © Gallagher, J.
Ptenothrix renateae from the USA
2016.01.07 © Fleming, K.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
Antenna
2016.02.01 © Anderson, J.
Ptenothrix renateae from the USA
2016.03.08 © Childs, K.
Ptenothrix renateae juv. from the USA
2016.12.06 © Fleming, K.
Ptenothrix renateae from the USA
Bulby distinctly pale upper clypeus
4 unpaired midfacial setae
2017.01.03 © Fleming, K.
Ptenothrix renateae from the USA
2018.01.12 © Fleming, K.
Ptenothrix renateae from the USA
2020.12.12 © Fleming, K.
Ptenothrix renateae from the USA
2021.01.05 © Wechsler, D.
Ptenothrix renateae from the USA
2021.12.12 © Charny, V.
Ptenothrix renateae from the USA
2021.12.12 © Charny, V.
Ptenothrix renateae from the USA
2021.12.12 © Charny, V.
Ptenothrix renateae from the USA
Wax crystals on dorsal wax tubes
2021.12.12 © Charny, V.
Ptenothrix renateae juv. from the USA
2021.12.12 © Charny, V.
Ptenothrix renateae from the USA
2021.12.29 © Charny, V.
Ptenothrix renateae from the USA
2021.12.29 © Charny, V.
Ptenothrix renateae from the USA
Antenna
2021.12.29 © Charny, V.
Ptenothrix renateae from the USA
2022.01.02 © Charny, V.
Ptenothrix renateae from the USA
2022.01.12 © Charny, V.
Ptenothrix renateae from the USA
More then 2 unpaired midfacial macrosetae
2022.01.12 © Charny, V.
Ptenothrix renateae from the USA
2022.01.23 © Charny, V.
Ptenothrix renateae from the USA
Cephalic + thoracic pattern
2022.01.23 © Charny, V.
Ptenothrix renateae from the USA
Caudal aspect
2022.01.23 © Charny, V.
Ptenothrix renateae from the USA
Small abdomen
2022.01.23 © Charny, V.
Ptenothrix renateae from the USA
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
Anterior dorsal pattern
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
More then 2 midfacial unpaired setae
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
More then 2 midfacial unpaired setae
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
Antenna
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
Ant.3 long and apically with pseudodivisions
Ant.4 distinctly short and annulated
2024.01.13 © Coogler, J.
Ptenothrix renateae from the USA
2024.02.08 © AVNJ.
Ptenothrix renateae from the USA
2024.03.15 © Wechsler, D.
Ptenothrix renateae from the USA
2024.03.15 © Wechsler, D.



Species reticulata Stach, J, 1957:105

Ptenothrix reticulata from Poland
After Stach J., 1957 Pl.IX fig.1.

Species rubra Bretfeld & Trinklein, 2000

Species ryoheii Uchida, 1953

Ptenothrix ryoheii from Japan
After Uchida, 1953 in Itoh R & al, 2012 Fig.49a
Ptenothrix ryoheii from Japan
After Uchida, 1953 in Itoh R & al, 2012 Fig.49b

Species saxatilis Yosii, R et Lee, C-E, 1963:19

Ptenothrix saxatilis from Korea
After Yosii R & Lee C-E, 1963 Fig.10A

Species setosa (Krausbauer, 1898:24) Handschin, 1929:125, B

            Syn.: Papirius palmatus Folsom, 1902


             Syn.: Ptenothrix setosus(sic) Denis, J-R, 1933:312 ISS
             Syn.: Ptenothrix setosus(sic) Denis, J-R, 1933:314 ISS
             Syn.: Ptenothryx(sic) setoas(sic) Yosii, R, 1954:824 ISS

Species simplificata Stach, 1957

Species sinensis Lin & Feng, 1985

Species tateyamana Uchida, H, 1958:76

Ptenothrix tateyamana from Japan
After Uchida, 1958 in Itoh R & al, 2012 Fig.55a

Species texensis (Packard, AS, 1873:42) Stach, J, 1957:80

            Syn.: Papirius Texensis(sic) Packard, AS, 1873:42 IOS

Species tranvansoii Yoshii, 1994

Species tricycla Uchida, 1953

Ptenothrix tricycla from Japan
After Uchida, 1953 in Bretfeld G, 1999 Fig.394
Ptenothrix tricycla from Japan
After Uchida, 1953 in Itoh R & al, 2012 Fig.57a

Species tsutsuii Yosii, R, 1955:398


Ptenothryx(sic) tsutsuii from Tokara Island
After Yosii R, 1955 Abb.9D
Ptenothryx(sic) tsutsuii from Tokara Island
Facial pattern
After Yosii R, 1955 Abb.9E
Ptenothryx(sic) tsutsuii from Tokara Island
Facial pattern
After Yosii R, 1955 Abb.9F
Ptenothrix tutsuii from Japan
After Yosii R, 1965 Fig.39A

Species unicolor (Harvey, 1893) Folsom, 1928

            Syn.: Papirius pini Folsom, JW, 1896:344
            Syn.: Papirius pini Folsom, 1934
            Syn.: Ptenothrix? setosa Denis, 1931
            Syn.: Ptenothrix setosa Denis, J-R, 1933:310
            Syn.: Dicyrtoma purpurata Maynard, EA, 1951:280
Dicyrtoma purpurata from the USA
After Maynard EA PlateXXII Fig.377b

            Syn.: Dicyrtoma purpurata Christiansen, K et Bellinger, P, 1981:1231
            Syn.: Dicyrtoma purpurata Christiansen, K et Bellinger, P, 1998:1405
            Syn.: Ptenothrix atra ssp. apigmentata Stach, 1966
            Syn.: Ptenothrix atra ssp. apigmentata Christiansen, K et Bellinger, P, 1998:1405
            Syn.: Ptenothrix atra ssp. apigmentata Bretfeld, G, 1999:158



Papirius unicolor from the USA
After Jackson, A.D., 1907 Pl.XI.
Ptenothrix unicolor from the USA
After Maynard EA, 1951 Plate XXIV Fig.430
Ptenothrix unicolor from the USA
After Maynard EA, 1951 Plate XXXVII Fig.663
Ptenothrix unicolor from the USA
2007.05.17 © Murray, T.
Ptenothrix unicolor from the USA
2007.05.17 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.09 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.09 © Murray, T.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.10 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.23 © McClarin, J.
Ptenothrix unicolor from the USA
2007.10.23 © McClarin, J.
Ptenothrix unicolor from the USA
2007.11.02 © Murray, T.
Ptenothrix unicolor from the USA
2007.11.02 © Murray, T.
Ptenothrix unicolor from the USA
2008.08.20 © Nanz, S.
Ptenothrix unicolor from the USA
2008.08.20 © Nanz, S.
Ptenothrix unicolor from the USA
2008.10.07 © Murray, T.
Ptenothrix unicolor from the USA
2008.10.15 © Murray, T.
Ptenothrix unicolor from the USA
2008.11.09 © Justis, S.
Ptenothrix unicolor from the USA
2008.11.09 © Justis, S.
Ptenothrix unicolor from the USA
2008.12.20 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.03.21 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.03.21 © Justis, S.
Ptenothrix unicolor juv. from the USA
2009.05.29 © Fleming, K.R.
Ptenothrix unicolor from the USA
2011.07.04 © Nanz, S.
Ptenothrix unicolor from the USA
2011.07.04 © Nanz, S.
Ptenothrix unicolor from the USA
2011.11.13 © Justis, S.
Ptenothrix unicolor from the USA
2011.12.24 © Justis, S.
Ptenothrix unicolor from the USA
Very short 4th antennal segment
2011.12.24 © Justis, S.
Ptenothrix unicolor from the USA
2012.06.03 © Murray, T.
Ptenothrix unicolor from the USA
2012.10.25 © Pugh, D.E.
Ptenothrix unicolor from the USA
2014.07.03 © Dankowicz, E.
Ptenothrix unicolor from the USA
2018.09.29 © Schulz, K.
Ptenothrix unicolor from the USA
Seta sa spinelike (*)
2018.09.29 © Schulz, K.
Ptenothrix unicolor from the USA
2018.11.01 © Webb, D.
Ptenothrix unicolor from the USA
2022.02.03 © Charny, V.
Ptenothrix unicolor from the USA
Two unpaired midfacial macrosetae
2022.02.03 © Charny, V.
Ptenothrix unicolor from the USA
2022.02.03 © Charny, V.
Ptenothrix unicolor from the USA
2022.02.03 © Charny, V.
Ptenothrix unicolor from the USA
2022.06.22 © Caterino, M.

Species unicolor Bueker, 1939 nec Harvey, 1893

Species utingae Arlé & Guimaraes, 1976

Species vinnula Uchida, H, 1957:26

Ptenothrix vinnula from Japan
After Uchida H, 1957 Fig.3A
Ptenothrix vinnula from Japan
2008.04.05 © Tsukiji, T.
Ptenothrix vinnula from Japan
2008.04.05 © Tsukiji, T.
Ptenothrix vinnula from Japan
2014.01.13 © Komatsu, T.
Ptenothrix vinnula from Japan
2015.01.12 © Jiku.
Ptenothrix vinnula from Japan
2015.01.12 © Jiku.
Ptenothrix vinnula from Japan
Frontal spines
2015.01.12 © Jiku.
Ptenothrix nr vinnula from Taiwan
2021.12.31 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
2021.12.31 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
2021.12.31 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
Antenna
Ant.1:2:3:4 = 1:4:5:1
2021.12.31 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
2022.01.02 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
2022.01.02 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
Dorsal cephalic + thoracic pattern
2022.01.02 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
Mouthparts
Labrum with 3 superfacial ridges
2022.01.02 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
Antenna
2022.03.26 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
Foot with unguis and unguiculus
Apex of unguiculus not swollen
2022.03.26 © Cheng, H.-J.
Ptenothrix nr vinnula from Taiwan
2022.09.19 © Cheng, H.-J.

Species violacea Handschin, 1929

Species violaceopa Hüther, 1967

            Syn.: Ptenothrix violaceopus(sic) Hüther, 1967 IOS
            Syn.: Ptenothrix violaceopus(sic) Yosii, R, 1969:222 IOS
            Syn.: Ptenothrix violaceopus(sic) Thibaud, J-M, 2013:235 IOS

Species vittata (Folsom, 1896) Folsom, 1934

            Syn.: Ptenothrix fasciatus(sic) Womersley, H, 1932:40 IOS
            Syn.: Ptenothrix fasciatus(sic) Womersley, H, 1939:261 IOS
Ptenothrix fasciata from Australia
After Womersley H, 1939 Fig.75N

            Syn.: Ptenothrix fasciatus(sic) Yosii, R et Lee, C-E, 1963:30 IOS
            Syn.: Ptenothrix fasciatus(sic) Salmon, JT, 1964:642 IOS
            Syn.: Ptenothrix fasciatus(sic) Greenslade, P in Houston, WWK, 1994:138 IOS
            Syn.: Ptenothrix fasciata Bretfeld, G, 1999:170



Ptenothrix vittata from Japan
After Yosii R & Lee C-E, 1963 Fig.16A
Ptenothrix vittata from Japan
Modified after Yosii R & Lee C-E, 1963 Fig.16B
Dicyrtoma (Ptenothrix) vittata from the USA
After Folsom in Christiansen K & Bellinger P, 1981 Fig.1043C
Dicyrtoma (Ptenothrix) vittata from the USA
After Folsom in Christiansen K & Bellinger P, 1981 Fig.1043D

             Syn.: Dicyrtoma vittatus(sic) Poole, RW in Poole, RW et Gentili, P, 1997:55 ISS
Ptenothrix nr vittata from Japan
2015.11.02 © Murray, A.
Ptenothrix nr vittata from Japan
2015.11.03 © Murray, A.
Ptenothrix ?vittata? from Australia
2023.07.16 © Watts, Br.





Species yakushimana Yosii, R, 1965:65

Ptenothrix yakushimana from Japan
After Yosii R, 1965 Fig.41A
Ptenothrix yakushimana from Japan
After Yosii R, 1965 Fig.41B

Species yunnana Itoh, R et Zhao, L-J, 2000:159

            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:159 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:159 nec Chen J-X & Christiansen K, 1996:587 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:159 nec Liu Y-Q, Hou D-B & Li Z-C, 1998:130 IOS
Ptenothrix yunnanus(sic) from China
With low dorsal hump
suggesting Papirioides...
After Itoh R & Zhao L-J, 2000 Fig.9A
Ptenothrix yunnanus(sic) ♀ from China
Anal valves
After Itoh R & Zhao L-J, 2000 Fig.10G


            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:147 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:160 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:161 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:162 IOS
            Syn.: Ptenothrix yunnanus(sic) Itoh, R et Zhao, L-J, 2000:176 IOS






Genus Papirioides Folsom, 1924

  Syn.: Ptenothricini Papiriodes(sic) Richards, WR, 1968:25 ISS
  Syn.: Sminthuridae Papiriodes(sic) Zeppelini Filho, D, 1996:17 ISS
  Syn.: Dicyrtominae Papiriodes(sic) Hopkin, SP, 1997:219 ISS

Species aequituberculatus Stach, J, 1965:369

            Syn.: Papirioides aequituberculata(sic) Stach, J, 1965:369 IOS
             Syn.: Ptenothrix aequituberculatus(sic) Chen, J-X et Christiansen, K, 1996:587 ISS
             Syn.: Ptenothrix aequituberculatus(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:130 ISS



Species caishijiensis (Wu, M & Chen, J-X, 1996:138) Guo & Chen, 1996

            Syn.: Ptenothrix (Papirioides) guanjiensis(sic) Wu, M et Chen, J-X, 1996:140 lapsus nn
            Syn.: Ptenothrix (Papirioides) guanjiensis(sic) Wu, M et Chen, J-X, 1996:141 lapsus nn
            Syn.: Ptenothrix (Papirioides) guanjiensis(sic) Wu, M et Chen, J-X, 1996:142 lapsus nn
            Syn.: Ptenothrix (Papirioides) guanjiensis(sic) Wu, M et Chen, J-X, 1996:143 lapsus nn
Ptenothrix (Papirioides) guanjiensis(sic) from China
After Wu M & Chen J-X, 1996 Fig.1




Papirioides caishijiensis from Hong Kong
2020.04.24 © Portioid.
Papirioides caishijiensis from Hong Kong
2020.04.24 © Portioid.
Papirioides caishijiensis from Hong Kong
2020.04.24 © Portioid.
Papirioides caishijiensis from Taiwan
2021.11.12 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
2021.11.12 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
2021.11.12 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
2022.05.05 © Cheng, H.-J.
Papirioides caishijiensis ♂ from Taiwan
Genital papilla
2022.06.28 © Cheng, H.-J.
Papirioides caishijiensis ♂ from Taiwan
Genital papilla with apical ring of circumgenital setae
2022.06.28 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
2022.10.18 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
Large abd. with posterior dorsal hump
2022.10.18 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
Posterior dorsal hump with dagger-like setae
2022.10.18 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
M'-M-M' in transverse row
2022.10.18 © Cheng, H.-J.
Papirioides caishijiensis from Taiwan
Retinaculum with anterior lobe (al), posterior lobe (pl) and ramus (r)
2022.10.18 © Cheng, H.-J.

Species dubius (Folsom, JW, 1932:74) Stach, 1965

Ptenothrix dubia from Hawaii
After Folsom JW, 1932 Plate12 Fig.133
Ptenothrix dubia from Hawaii
After Folsom JW, 1932 Plate12 Fig.134

Species jacobsoni Folsom, 1924, t.t.

Papiriodes jacobsoni from Japan
After Yosii R & Lee C-E, 1963 Fig.18A
Papiriodes jacobsoni from Japan
After Yosii R & Lee C-E, 1963 Fig.18B
Papirioides jacobsoni from Indonesia
2013.07.13 © Lam, P.
Papirioides jacobsoni from Taiwan
2017.10.16 © Anonymous.
Papirioides jacobsoni from Taiwan
2017.10.25 © Anonymous.
Papirioides jacobsoni from Taiwan
2018.12.15 © Anonymous.
Papirioides jacobsoni from Taiwan
2019.04.16 © Aerobird.
Papirioides jacobsoni from Taiwan
2019.04.16 © Aerobird.
Papirioides jacobsoni from Taiwan
2021.10.30 © Anonymous.
Papirioides jacobsoni from Taiwan
2021.10.30 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Grooming with drop of saliva
2021.10.30 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
2021.11.01 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Mouth ventral aspect
Labrum with 3 ridges
2021.11.22 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
2021.12.03 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Dorsal pattern
2021.12.03 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Antenna
2021.12.03 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Head
2021.12.03 © Cheng, H.-J.
Papirioides jacobsoni from Taiwan
Dorsal posterior protuberance
2021.12.03 © Cheng, H.-J.
Papirioides jacobsoni juv. from Taiwan
2022.09.28 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
2022.09.28 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
Sub-anal appendage (SAA)
2022.09.28 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
Genital orifice with anterior genital valve (agv) and posterior genital valve (pgv)
2022.10.11 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
Genital orifice in ventral aspect
2022.10.11 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
Sub-anal appendage (SAA) on basal papilla
2022.10.11 © Cheng, H.-J.
Papirioides jacobsoni ♀ from Taiwan
Anterior genital valve with transverse row of genital setae
2022.10.11 © Cheng, H.-J.

Species kauaiensis (Snider, RJ, 1990:75)

Species mirabilis (Denis, 1929)

Species serratus (Snider, RJ, 1990:79)

Ptenothrix (Papirioides) serrata from Hawaii
After Snider RJ, 1990 Fig.66

Species suzhouensis (Guo & Chen, 1996)

Species tonsori (Greenslade, P, 1994:48)

Species uenoi Uchida, 1957

Papirioides uenoi from Japan
After Itoh R & al, 2012 Fig.48a
Papirioides uenoi from Japan
After Itoh R & al, 2012 Fig.48b
Papirioides uenoi from Japan
After Itoh R & al, 2012 Fig.48c
Papirioides uenoi from Japan
2020.10.19 © Yoshida, Y.

Species yunnanus Itoh, R et Zhao, L-J, 1993:847

Papirioides yunnanus ♀ from China
After Itoh R & Zhao L-J, 1993 Fig.1
Papirioides yunnanus ♂ from China
After Itoh R & Zhao L-J, 1993 Fig.2
Papirioides yunnanus from China
After Itoh R & Zhao L-J, 1993 Fig.3
Papirioides yunnanus from China
Dorsal protuberance
After Itoh R & Zhao L-J, 1993 Fig.12
Papirioides yunnanus from China
Anal valves
After Itoh R & Zhao L-J, 1993 Fig.17

             Syn.: Ptenothrix yunnanus(sic) Chen, J-X et Christiansen, K, 1996:587 ISS
             Syn.: Ptenothrix yunnanus(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:130 ISS


Species zhejiangensis Li, Z-H, Chen, J-X et Li, J, 2007:?


Genus Bothriovulsus Richards, WR, 1968:26

Species sp. Christiansen, K et Bellinger, P, 1981:1219

Species kymgangensis Weiner & Betsch, 1992

Species pineolae (Wray, 1946 tt) Richards, WR, 1968:26

             Syn.: Ptenothrix pineola(sic) Richards, WR, 1968:26 ISS
             Syn.: Bothriovulsus pineolus(sic) Richards, WR, 1968:26 ISS
Dicyrtoma (Bothriovulsus) pineolae from the USA
After Christiansen K & Bellinger P, 1981 Fig.1025A
Dicyrtoma (Bothriovulsus) pineolae from the USA
After Christiansen K & Bellinger P, 1981 Fig.1025A
Bothriovulsus pineolae from the USA
2020.12.03 © Richman, M.
Bothriovulsus pineolae from the USA
2020.12.03 © Richman, M.



Species sohungensis Weiner & Betsch, 1992


References