http://www.collembola.org/taxa/tomoceri.htm - Last updated on 2024.10.12 by Frans Janssens
Checklist of the Collembola: Tomoceridae

Familia Tomoceridae Schäffer, 1896 key  key

Genus Number
of species
Antennacyrtus1
Aphaenomurus1
Entomocerus1
Lasofinius2
Lepidophorella9
Lethemurus3
Monodontocerus7
Neophorella1
Novacerus3
Plutomurus39
Pogonognathellus21
Pseudolepidophorella1
Tomocerina14
Tomocerus (Ciliatomurus)2
Tomocerus (Coloratomurus)6
Tomocerus (Ocreatomurus)20
Tomocerus (Striatomurus)6
Tomocerus (Tomocerus)66
Tomocerus102
Tomolonus1
Tritomurus5
Yoshiicerus7
Tomoceridae218
Genera with number of species
  Syn.: Entomobryidae Tomocerinæ(sic) Schäffer, 1896 ISS


  Syn.: Entomobryidae Tomocerini(sic) Börner, C, 1901:15 ISS
  Syn.: Entomobryidae Tomocerini(sic) Absolon, K, 1903:108 ISS
  Syn.: ... Tomocerinæ(sic) Börner, 1906 ISS
  Syn.: Collembola Tomocerinæ(sic) Folsom, JW, 1913:451 ISS
  Syn.: Collembola Tomoceridæ(sic) Dethier, M, 1998:57 ISS


Pogonognathellus sp. from the UK
Body covered with iridescent scales
2007.12.19 © Campbell, A.
 
Tomocerus minor from the UK
3rd abdominal segment superequal to 4th
2008.09.24 © Robertson, A.

  Syn.: Collembola Tomocerida(sic) Yadav, RS, 2017:1339 ISS

Subfamilia Lepidophorellinae Absolon K, 1903

  Syn.: Collembola Arthropleona Glime, JM, 2015:3


Lepidophorellinae from Tasmania
2013.05.25 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.05.25 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.06.19 © Zosterops, K.
Lepidophorellinae from Tasmania
2013.06.19 © Zosterops, K.
Lepidophorellinae from Tasmania
2014.04.07 © Murray, A.
"Arthropleona oruarangi"(sic) from New Zealand
After Moore S in Glime JM, 2015 Fig3


Tribus Neophorellini Janssens F, 2017


Genus Neophorella Womersley, H, 1934:464

Species dubia Womersley, H, 1934:464, t.t.

Neophorella dubia from South Africa
After Womersley H, 1934 Fig.12a


Genus Lasofinius Ireson, JE et Greenslade, P, 1990:205

  Syn.: Lepidophorellinae Lasofinus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2004:41 ISS

Species gemini Ireson & Greenslade P, 1990:211

Species willi Ireson & Greenslade P, 1990:209, t.t.


Tribus Lepidophorellini Absolon K, 1903:108


Genus Lepidophorella Schäffer, 1897

  Syn.: ... Drepanura Moniez, 1894
  Syn.: Tomoceridae Pseudolepidophorella Salmon, 1941 =?
  Syn.: Isotomidae Millsia Womersley, H, 1942:26 =?
  Syn.: Tomoceridae Tomocerus(sic) Nunes, RC, Santos-Costa, RC et Bellini, BC, 2020:39 lapsus


Lepidophorella sp. from New Zealand
Descaled
2017.03.31 © Navest, M.P.
Lepidophorella sp. from New Zealand
Descaled
2019.10.09 © Navest, M.P.

Species australis Carpenter, GH, 1925:11

Lepidophorella australis from New Zealand
Mesothorax with small scales
After Hawes TC & Greenslade P, 2015 Fig.1d
Lepidophorella australis from New Zealand
Large body scales with longitudinal intermittant strong and weak ribs
After Hawes TC & Greenslade P, 2015 Fig.1a

  Subspecies fusca Salmon, JT, 1941:340

Lepidophorella australis ssp. fusca from New Zealand
2014 © Te Papa Museum

Species brachycephala (Moniez, 1894) Denis, 1923

             Syn.: Lepidophorella brachycephalla(sic) Womersley, H, 1936:321 ISS

Species communis Salmon, JT, 1937:353

Lepidophorella communis from New Zealand
After Salmon JT, 1937 Pl.49 Fig.d
Lepidophorella communis from New Zealand
After Salmon JT, 1941 Pl.54 Fig.246
Lepidophorella communis from New Zealand
2014 © Te Papa Museum

Species flavescens (Nicolet, H, 1847:383) Schäffer, 1897

            Syn.: Cyphoderus flaveus(sic) Nicolet, H, 1847:383 IOS
            Syn.: Cyphoderus giganteus Nicolet, H in Gay, C, 1849:92
Cyphoderus giganteus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.7.

            Syn.: Lepidophorella flava Schäffer, 1897,tt



             Syn.: Lepidocyrtus flaveus(sic) Lubbock, J, 1873:157 ISS
             Syn.: Cyphoderus flaveus(sic) Salmon, JT, 1964:536 ISS
Remark by Ellis, W.N. & Bellinger, P.F. 1973:27-28: Cyphoderus flaveus Nicolet, 1847 is an incorrect original spelling (art.32a(ii)). Nicolet, 1847:383 described the new species Cyphoderus giganteus and C. flaveus. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]:92-93. In the last paper however "flavescens" is used in stead of "flaveus". But in the first paper Nicolet cites as author of both species Nicol. in Gay, evidently the publications appeared in a reversed order. This, and the fact that "flaveus" is bad latin, brings us to the conclusion that "flaveus" is an incorrect original spelling.

Lepidophorella flavescens from Chile
2021.06.20 © Gargiulo, M.
Lepidophorella flavescens from Chile
2022.12.31 © Gargiulo, M.





Species nigra Salmon, JT, 1943:383

Lepidophorella nigra from New Zealand
After Salmon JT, 1943 Pl.42 Fig.48
Lepidophorella nigra from New Zealand
2014 © Te Papa Museum

Species rubicunda Salmon, JT, 1941:342

Lepidophorella rubicunda from New Zealand
2014 © Te Papa Museum

Species spadica Salmon, JT, 1944:152

Lepidophorella spadica from New Zealand
2014 © Te Papa Museum

Species tiegsi (Womersley, H, 1942:26)

Millsia tiegsi from Australia
After Womersley, H., 1942 Fig.3A
Remark by Greenslade, P. in Dallai, R. 1989:110: The type series consists of three specimens, all probably early instars. Millsia is similar to Lepidophorella and differs only in the lack of scales and the shorter, more cylindrical dens. It is possible that Millsia is a synonym of Lepidophorella.

Species unadentata Salmon, JT, 1941:341

Lepidophorella unadentata from New Zealand
2014 © Te Papa Museum


Genus Pseudolepidophorella Salmon, JT, 1941:342

Species longiterga (Salmon, JT, 1937:354) Salmon, JT, 1941:343, t.t.

Lepidophorella longiterga from New Zealand
After Salmon JT, 1937 Pl.49 Fig.h
Pseudolepidophorella longiterga from New Zealand
After Salmon JT, 1941 Pl.54 Fig.259
Pseudolepidophorella longiterga from New Zealand
2014 © Te Papa Museum
Pseudolepidophorella longiterga from New Zealand
2014.02.18 © Murray, A.
Pseudolepidophorella longiterga from New Zealand
2014.02.18 © Murray, A.


Genus Antennacyrtus Salmon, JT, 1941:343


Antennacyrtus insolitus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.261.
Species insolitus Salmon, JT, 1941:344, t.t.



Tribus Novacerini Salmon JT, 1941:345


Genus Novacerus Salmon, JT, 1942:259

Remark by Salmon, J.T. 1942:259: Novacerus Salmon JT, 1942 nomen novum to replace the name Neocerus Salmon, 1941, pre-occupied by Wasmann, 1893.

  Syn.: Neocerini Neocerus Salmon, JT, 1941:345 nec Wasmann, 1893


  Syn.: Lepidophorellinae Novocerus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2004:41 ISS

Species insolitatus (Salmon, JT, 1941:346) Salmon, JT, 1942:259


Neocerus spinosus from New Zealand
After Salmon, J.T. 1941 Pl.55 Fig.269.
Species spinosus (Salmon, JT, 1941:345) Salmon, JT, 1942:259, t.t.


Species tasmanicus (Womersley, H, 1937:154) Yosii, R, 1967:51

Tomocerus tasmanicus from Tasmania
After Womersley H, 1937 Fig.1g
Novacerus tasmanicus from Tasmania
Distinctly short ant.3
After Yosii R, 1967 Fig.29A
Novacerus tasmanicus from Tasmania
2014.10.24 © Murray, A.
Novacerus tasmanicus from Tasmania
2014.10.25 © Murray, A.


Subfamilia Tomocerinae Schäffer, 1896

  Syn.: ... Tomocerinæ(sic) Schäffer, 1896 ISS
  Syn.: ... Paratomocerinae Tarsia in Curia, 1938


Tomocerus sp. from France
Long 3rd antennal segment
2007.02.23 © Ollivier, E.
Tomocerinae juv. from Belgium
2008.06.09 © Vuijlsteke, M.
 
Tomocerinae juv. from the UK
Second instar with short 3rd antennomere
2008.09.28 © Barton, T.
Tomocerinae juv. from the USA
Long 3rd antennal segment
2009.03.14 © Justis, S.
Tomocerus sp. from Czechia
Descaled specimen revealing body colour
2010.11.09 © Fiala, M.
Tomocerinae juv. from the UK
First instar with short 3rd antennomere
2011.12.01 © Nurcombe, E.
Tomocerus minor from the USA
3rd antennomere with very fine annulation
2012.02.18 © Carr, J.F.
Pogonognathellus flavescens from Germany
Ant.3 strongly annulated
2014.09.11 © Kluck, A.
Tomocerus minor from the UK
Distinctly knobbed 'tenent hair' (*)
2015.12.02 © Horton, M.
Pogonognathellus sp. from Sweden
Longitudinally ribbed scales
2016.09.16 © Nilsson, K.
Tomocerinae from the USA
Preparing for jump with antennae bent
2019.09.19 © Wechsler, D.
Tomocerinae juv. from the USA
Hatchling
2020.05.12 © Solomon.


Tribus Tomocerini Börner C, 1906:161


Genus Tomocerina Yosii, R, 1955:394 key

Tomocerus minutus from Japan
Basal outer tooth of mucro without toothlet
After Yosii R, 1955 Abb.8D
  Syn.: Tomocerus Tomcerina(sic) Yosii, R, 1967:46 ISS


Species annamitica Yu, D-Y, Man, LC et Deharveng, L, 2016:8

Tomocerina annamitica from South Vietnam
After Yu D-H, Man LC & Deharveng L, 2016 Fig.1B

Species aokii (Yosii, R, 1972:94)

Species curta (Christiansen, K, 1964:662)

Species fuxii (Yu, D-Y, Pan, Z-X & Shi, S-D, 2017:408) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:8

            Syn.: Tomocerus fuxi(sic) Yu, D-Y, Pan, Z-X et Shi, S-D, 2017:408 IOS
Tomocerus fuxi(sic) from China
After Yu-D-Y, Pan Z-X & Shi S-D, 2017 Fig.1B

            Syn.: Tomocerina fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS
Remark by Janssens, F. 2021.07.25: Given the new combination is assigned in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given neither this electronic publication nor the recombination itself have been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, 2.the recombintions in Appendix A are unavailable for Zoology.

            Syn.: Tomocerus fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 IOS
            Syn.: Tomocerus fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 IOS
            Syn.: Tomocerus fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:7 IOS
            Syn.: Tomocerus fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS
            Syn.: Tomocerina fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 IOS
            Syn.: Tomocerina fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 IOS
            Syn.: Tomocerina fuxi(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS


Species lamellifera (Mills, HB, 1934:84)

Species liliputana (Yosii, R, 1967:18) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:8

            Syn.: Tomocerus minutus Yosii, R, 1955:394 partim
            Syn.: Tomocerina minuta Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8


             Syn.: Tomocerus (Tomocerina) lilliputanus(sic) Ma, Y-T, Chen, J-X et Christiansen, K, 2003:516 ISS
             Syn.: Tomocerus (Tomocerina) liliptanus(sic) Hasegawa, M, Suma, Y, Tanaka, S et Ichisawa, K, 2017:159 ISS
             Syn.: Tomocerus (Tomocerina) liliptanus(sic) Hasegawa, M, Suma, Y, Tanaka, S et Ichisawa, K, 2017:160 ISS

Species minuta (Tullberg, T, 1877:32) Yosii, R, 1955:394

            Syn.: Tomocerus varius Handschin, E in Dahl, F, 1929:100 =?
            Syn.: Tomocerus mixtus Gisin, 1961
            Syn.: Tomocerina simplex Yosii, 1966 =?


Tomocerina minuta from Iran
After Yahyapour E & al, 2021 Fig.15
Tomocerina minuta from Iran
Ocelli A-D in square formation
After Yahyapour E & al, 2021 Fig.19

Species nuwaae (Yu, D-Y, Pan, Z-X & Shi, S-D, 2017:404) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:8

            Syn.: Tomocerus nuwa(sic) Yu, D-Y, Pan, Z-X et Shi, S-D, 2017:404 IOS
Tomocerus nuwa(sic) from China
After Yu-D-Y, Pan Z-X & Shi S-D, 2017 Fig.1A

            Syn.: Tomocerina nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS
Remark by Janssens, F. 2021.07.25: Given the new combination is assigned in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given neither this electronic publication nor the recombination itself have been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, 2.the recombintions in Appendix A are unavailable for Zoology.

            Syn.: Tomocerus nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 IOS
            Syn.: Tomocerus nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 IOS
            Syn.: Tomocerus nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:7 IOS
            Syn.: Tomocerus nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS
            Syn.: Tomocerina nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 IOS
            Syn.: Tomocerina nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 IOS
            Syn.: Tomocerina nuwa(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8 IOS


Species simplex Yosii, 1966

Species teres (Christiansen, K, 1964:653)

Species tianshanensis (Ma, Y-T, Chen, J-X & Christiansen, K, 2003:512) Yu, D-Y, Pan, Z-X & Shi, S-D, 2017:403

Species varia (Folsom, JW, 1899:267) Yu, D-Y, Deharveng, L & Zhang, F, 2014:558

            Syn.: Tomocerus minutus Yosii, 1954
            Syn.: Tomocerus minutus Yosii, R, 1955:394 partim
            Syn.: Tomocerus minutus Yosii, R, 1956:91
            Syn.: Tomocerina minuta Yosii, R, 1956:91
            Syn.: Tomocerus minutus Salmon, JT, 1964:311
            Syn.: Tomocerus minutus Yosii, R, 1967:17
            Syn.: Tomocerus minutus Dunger, 1972
            Syn.: Tomocerus minutus Yosii, R, 1977:158
            Syn.: Tomocerus minutus Ma, Y-T, Chen, J-X et Christiansen, K, 2003:517
            Syn.: Tomocerina minuta Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:8


Tomecerina varia
Descaled
After Yu D-Y & al, 2020 Fig.5

Species violaceus (Yosii, R, 1956:91) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:5

            Syn.: Tomocerus vulgaris Japanese records =?


Species yiliensis (Ma, Y-T, Chen, J-X & Christiansen, K, 2004:43) Yu, D-Y, Pan, Z-X & Shi, S-D, 2017:403


Genus Tomocerus Nicolet, H, 1842:67 key

  Syn.: ... Poduræ(sic) Sp
  Syn.: Aptera Podura Linnæus, C, 1758:608 partim
  Syn.: Podurides(sic) Poduræ(sic) Bourlet, C, 1845:100 ISS
  Syn.: Podurides(sic) Podura Elditt, HL, 1854:42 partim
  Syn.: Podurelles(sic) Temnourus Nicolet, H in Desor, E, 1841:384 nn
  Syn.: Podures(sic) Macrotoma Bourlet, C, 1839:387 nec Laporte, 1832
  Syn.: Podurides(sic) Macrotoma Bourlet, C, 1845:168 ISS
Macrotoma sp. from France
Ocelli of left eye
Anterior 4 ocelli ABCD form square
Modified after Bourlet C, 1845 Fig.17

  Syn.: ... Palpiger Olfers, 1907



  Syn.: Collembola Toxocerus(sic) Dallas, WS in Duncan, PM, 1896:149 ISS
  Syn.: Tomocerini(sic) Tomoceri(sic) Börner, C, 1901:10 ISS
Tomocerus sp.from Europe
After Fjellberg A, 1992 Fig.1C
Remark by ICZN. 2007:821: Placed on the Official List of Names in Zoology. Opinion 239.

Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
Only 1st and 2nd antennal segments scaled
2008.12.22 © Valentine., B
Tomocerus minor from the UK
Long 3rd antennal segment subcylindrical
2009.01.26 © Robertson, A.
Tomocerus minor from Germany
Colour of 3rd antennal segment distinctly different from other segments
2009.01.26 © Peters, J.
Tomocerus sp. juv. from the UK
4th ant. segment > 1st + 2nd ant. segment
2009.04.20 © Robertson, A.
 
Tomocerus sp. from Canada
Dental spines array
2011.11.16 © Hochhalter, R.
Tomocerus sp. juv. from the UK
4th ant. segment > 1st + 2nd ant. segment
2012.09.02 © Murray, A.
Tomocerus sp. juv. from the UK
Annulated 4th antennal segment
2012.09.26 © Murray, A.
Tomocerus minor from the UK
Annulated 4th & 3rd antennal segment
2014.02.22 © Henton-Jones, D.
Tomocerus minor from Germany
Only 1st and 2nd antennal segments scaled
2014.08.21 © Kluck, A.
Tomocerus vulgaris from Holland
Eyepatch with square of 4 + 2 ocelli
2014.09.27 © van Duinen, J.
Tomocerus minor from Germany
Lateral bothriotrichum
2014.10.10 © Beck, K.
Tomocerus sp. from Czechia
Colour of ant.1 and 2 distinctly different from ant.3 and 4
2015.03.05 © Krásenský, P.
Tomocerus minor from Holland
Eyepatch with square of 4 + 2 ocelli
2015.11.03 © van Duinen, J.
Tomocerus minor from Canada
Seven macrosetae on posterior margin of abd.5 (*)
2015.11.08 © Hogue, A.
Tomocerus minor ♀ from Canada
Pointed dorsal anal valve (*)
2015.11.08 © Hogue, A.
Tomocerus minor from Belgium
Eyepatch with square of 4 + 2 ocelli
2016.02.03 © Huskens, M.-L.
Tomocerus sp. from Sweden
Eyepatch with square of 4 + 2 ocelli
2017.03.09 © Nilsson, K.
Tomocerus minor from France
Eyepatch with square of 4 + 2 ocelli
2018.01.18 © Garcelon, P.
Tomocerus minor from France
Ant.4+3 annulated
2018.01.18 © Garcelon, P.
Tomocerus sp. from Japan
2018.11.10 © Hara, Y.
Tomocerus sp. from Japan
2018.11.10 © Hara, Y.
Tomocerus sp. juv. from the UK
Ant.3 longest
Ant.4 longer then ant.1+2
2018.11.19 © Uncle, F.
Tomocerus minor from Holland
Ant.1+2 scaled, ant.3+4 not scaled
2020.01.22 © Kamsteeg, G.
Tomocerus sp. from Belgium
Left eye
Anterior ocelli ABCD form a square
2020.09.05 © Huskens, M.-L.
Tomocerus sp. from Belgium
Ocelli BDE in straight line
As in Isotomurus
2020.09.05 © Huskens, M.-L.
Tomocerus ocreatus from Taiwan
Hind tibiotarsus with 2 inner spine-like macrosetae (*)
2021.09.11 © Cheng, H.-J.
Tomocerus sp. from Taiwan
Ocelli ABCD form a square
2021.10.11 © Cheng, H.-J.
Tomocerus minor from France
Abd.5+6
2022.02.12 © Joachim, J.
Tomocerus sp. from Baltic amber
Anterior ocelli ABCD form square
2022.03.27 © Garcelon, P.
Tomocerus sp. from Baltic amber
Antenna
2022.04.02 © Garcelon, P.
Tomocerus sp. from the USA
Hind tibiotarsus with 2 inner spine-like macrosetae
2023.07.29 © Lopez, A.
Tomocerus sp. from China
Tenent macroseta of footcomplex with clavate apex
2023.07.01 © Wang, G.
Tomocerus sp. from China
Mucro apical part with 2 serial teeth
2023.07.01 © Wang, G.
Tomocerus sp. from China
Mucro basal part with 2 parallel teeth
2023.07.01 © Wang, G.
Tomocerus sp. from China
Unguis with lateral basal pseudonychia
2023.07.01 © Wang, G.
Tomocerus sp. from China
Dental spines with toothlet
2023.07.01 © Wang, G.



Species bajalis Mehra, Verma, Verma & Vyas, 2009 u

Remark by Mandal, G.P. & al. 2024:765: Excluded as the new taxon is not described properly following the ICZN - Articles no. 72 & 73.
Species neuceratus Verma & Yadav, 2019 u

Remark by Mandal, G.P. & al. 2024:765: Excluded as the new taxon is not described properly following the ICZN - Articles no. 72 & 73.


Subgenus Tomocerus Nicolet H sensu Yu, Deharveng, Lukic, Wei, Hu & Liu, 2020:5

  Syn.: Tomocerus Eutomocerus Yosii, R, 1955:394
Tomocerus minor from the USA
Basal outer tooth of mucro with toothlet (*)
After Yosii R, 1955 Abb.8C




Species sp.1 Agrell, 1939

Species sp.2 Agrell, 1939

Species sp.6 Janssens F sensu Yu D-Y et al, 2020

            Syn.: Tomocerus (Tomocerus) sp. cf. minor Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:3
            Syn.: Tomocerus (Tomocerus) sp.separate Yu D-Y et Deharveng L in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:3


Species asahinai Yosii, 1954

Tomocerus asahinai from Japan
After Yosii R, 1967 Fig.13A

Species asiaticus Martynova, 1969

Species asoka Yosii & Ashraf, 1965

Species baicalensis Martynova, 1969

Species baudoti Denis, J-R, 1932:375

            Syn.: Tomocerus Baudoti(sic) Denis, J-R, 1932:375 IOS

Species calceus (Liu, Hou & Li, 1999) Ma, Y-T, Chen, J-X & Christiansen, K, 2003:516

Species caputiviolaceus Lee, B-H, 1975:946

Species christianseni Stomp, N, 1969:695

Species cucullatus (Olfers, 1907 -) Handschin, 1926 -

Species emeicus Liu, Hou & Li, 1999

Species huoensis Sun, Y, Liang, A-P et Huang, F-S, 2006:804

Species ishibashii Yosii, 1954

            Syn.: Tomocerus katoi Uchida, 1969
            Syn.: Tomocerus katoi Yosii, R, 1977:158


Tomocerus (Tomocerus) ishibashii from Japan
After Yosii R, 1967 Fig.12A
             Syn.: Tomocerus (Tomocerus) ishibashi(sic) Yosii, R, 1967:24 ISS
             Syn.: Tomocerus ishabashii(sic) Hopkin, SP, 1997:312 ISS


Species jezoensis Matsumura S et Ishida H in Matsumura, S, 1931:1491

Species jinyunensis Liu, Y-Q, 2006:300

             Syn.: Tomocerus (Tomocerus) jinyun(sic) Liu, Y-Q, 2006:301 ISS

Species lamelliger (Börner, 1903) Bonet, F, 1931:103

             Syn.: Tomocerus lamelligerus(sic) Bonet, F, 1931:103 ISS
             Syn.: Tomocerus lamelligera(sic) Yu, D-Y, Pan, Z-X et Shi, S-D, 2017:411 ISS

Species laxalamellus Lee, B-H, 1975:953

            Syn.: Tomocerus laxalamella(sic) Lee, B-H, 1975:953 IOS
            Syn.: Tomocerus (Monolamellus) laxalamella(sic) Lee, 1983 IOS
            Syn.: Tomocerus (Monolamellus) laxalamella(sic) Lee, B-H et Park, K-H, 1992:107 IOS
            Syn.: Tomocerus laxalamella(sic) Yu, D-Y, Pan, Z-X et Shi, S-D, 2017:411 IOS

Species lividus Tullberg, T, 1877:32

Species magadanicus Martynova, 1979

            Syn.: Tomocerus magadanicus Martynova & al, 1977 nn

Species magnus Sun & Liang, 2008

Species minor (Lubbock, J, 1862:598) Schäffer, 1900, t.t., B

            Syn.: Podure(sic) Podure grise commune Geoffroy, 1762
            Syn.: Podura podure grise commune Geoffr
            Syn.: Podura plumbea von Linné, C, 1767:1013
            Syn.: Podura plumbea Fabricius, IC, 1775:301
            Syn.: Podura plumbea Müller OF, 1776
            Syn.: Podura plumbea Schrank, 1781
             Syn.: Podura plumbæa(sic) de Fourcroy, AF, 1785:524 ISS
            Syn.: Podura plumbea Latreille, 1796
            Syn.: Podura plumbea Goring, CR, 1831:246
            Syn.: Choreutes plumbeus Burmeister, 1835
            Syn.: Podura plumbea Boisduval & Lacordaire, 1835
            Syn.: Podura plumbea Bourlet, C, 1839:384
Podura plumbea from France
After Lucas M, 1840 Pl.1 Fig.2

             Syn.: Podurides(sic) Poduraplumbea(sic) Bourlet, C, 1845:102 lapsus ISS
Macrotoma plumbea from the UK
After McIntire SJ, 1867 Fig.51
After McIntire SJ, 1868


             Syn.: Choreutes plumbea(sic) Lubbock, J, 1870:284 ISS
Tomocerus plumbeus from the UK
After Beck J in Lubbock J, 1873 Pl.68

            Syn.: Tomocerus plumbeus Lubbock, J, 1873:138
Tomocerus plumbeus from the UK
After Lubbock, J., 1873 Pl.19

            Syn.: Podura gigas Fabricius O, 1783
            Syn.: Macrotoma tridentifera Tullberg, T, 1872:37
            Syn.: Tomocerus norvegicus Lie-Pettersen, 1897
             Syn.: Tomocerus tridentiferus var. orcina(sic) Absolon, K, 1901:88 ISS
             Syn.: Tomocerus tridentiferus var. orcina(sic) Houlbert, C, 1924:109 ISS
             Syn.: Tomocerus minor f. orcina(sic) Bonet, F, 1928:253 ISS
             Syn.: Tomocerus minor f. orcina(sic) Bonet, F, 1928:254 ISS
            Syn.: Tomocerus minor var. noricus Latzel, R, 1917:239
            Syn.: Macrotoma celer Houlbert, C, 1924:108



Tomocerus minor from Belgium
Tridentate dental spines
Legit Leruth, R., 1933.07.16
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
Unscaled 3rd antennal segment
Legit Leruth, R, 1933.10.03
2008.05.28 © Janssens, F.
Tomocerus minor from Belgium
2004 © Vuijlsteke, M.
Tomocerus minor from the UK
2005 © Hopkin, S.P.
Tomocerus minor from the UK
2006 © Cornwall, N.J.
Tomocerus minor from Belgium
2006 © Vuijlsteke, M.
Tomocerus minor from the UK
2007 © Barton, T.
Tomocerus minor from the USA
2007.03.03 © Murray, T.
Tomocerus minor from the USA
2007.03.03 © Murray, T.
Tomocerus minor from the USA
2007.03.11 © Murray, T.
Tomocerus minor from the USA
2007.05.04 © Moorehead, C.
Tomocerus minor from Belgium
2007.07.07 © Vuijlsteke, M.
Tomocerus minor from the UK
2007.07.22 © Valentine., B
Tomocerus minor from the USA
2007.09.21 © Moorehead, C.
Tomocerus minor from the UK
2007.12.14 © Botting, J.
Tomocerus minor from the Netherlands
2008.01.24 © Baas, A.H.
Tomocerus minor from the UK
2008.02.03 © Birch, P.
Tomocerus minor from the Netherlands
2008.02.03 © Baas, A.H.
Tomocerus minor from the UK
2008.02.27 © Spears, G.
Tomocerus minor from Belgium
2008.03.15 © Oger, P.
Tomocerus minor from the UK
2008.03.17 © Robertson, A.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.22 © Vuijlsteke, M.
Tomocerus minor from Belgium
2008.03.24 © Vuijlsteke, M.
Tomocerus minor from the USA
2008.05.29 © Nanz, S.
Tomocerus minor ♀ from Belgium
Anal valves apically elongated
2008.08.25 © Janssens, F.
Tomocerus minor from the USA
2008.09.13 © Robertson, A.
Tomocerus minor from the UK
2008.09.24 © Robertson, A.
Tomocerus minor juvenile from the UK
2008.09.24 © Robertson, A.
Tomocerus minor from the UK
2008.10.21 © Spears, G.
Tomocerus minor from the UK
2009.01.14 © Valentine., B
Tomocerus minor from Germany
2009.02.18 © Peters, J.
Tomocerus minor from the UK
2009.02.27 © Knight., B
Tomocerus minor from Belgium
2009.03.15 © Janssens, F.
Tomocerus minor from the UK
2009.04.29 © Robertson, A.
Tomocerus-minor from Belgium
2009.09.13 © Wallays, H.
Tomocerus minor, juv. from the UK
2009.10.07 © Robertson, A.
Tomocerus minor from the UK
Body scales bluish iridisation
2010.01.02 © Robertson, A.
Tomocerus minor from the UK
Macroseta with blunt apex
2010.01.02 © Robertson, A.
Tomocerus minor from the UK
Microsetae on 3rd antennal segment
2010.01.02 © Robertson, A.
Tomocerus minor from Canada
Abdominal bothriotricha
2011.09.18 © Hochhalter, R.
Tomocerus minor from Belgium
Descaled specimen with dark thorax
2011.12.28 © Menten, J.
Tomocerus minor from Belgium
Mucro with teeth and setae
2011.12.28 © Menten, J.
Tomocerus minor from Belgium
Footcomplex: unguiculus, unguis and tenant hair
2011.12.28 © Menten, J.
Tomocerus minor from Germany
Descaled specimen, revealing dark pigmented thorax
2012.03.15 © Rose, A.
Tomocerus minor from Belgium
2013.04.06 © Declercq., B
Tomocerus minor from Germany
Base of manubrium (*)
2014.10.12 © Beck, K.
Tomocerus minor from New Zealand
2014.10.21 © Kerr, S.
Tomocerus minor ♂ from the UK
Genital plate (*) on exposed extended abd.5
Trapped in ice
2014.12.06 © Henton-Jones, D.
Tomocerus minor ♂ from the UK
Furca implanted ventroposteriorly at abd.4
Trapped in ice
2014.12.06 © Henton-Jones, D.
Tomocerus minor juv. from Germany
2015.01.05 © Beck, K.
Tomocerus minor ♂ from the UK
Exposed anal valves without cerci (*)
2015.03.01 © Thomas., B
Tomocerus minor from Germany
Bothriotrichum (Bothr.)and macroseta (Mac.)
2015.09.03 © Kluck, A.
Tomocerus minor ♀ from Holland
Posterior margin of abd.5 with short blunt macrosetae
2015.11.03 © van Duinen, J.
Tomocerus minor juv. from the UK
2016.02.07 © Lewis, M.
Tomocerus minor from Hungary
Eyepatch with square of 4 +2 ocelli
2016.02.20 © Fekete, C.
Tomocerus minor from Denmark
Dental spines tridentate
2017.08.27 © Hobern, D.
Tomocerus minor from France
Square of anterior 4 ocelli
2018.03.04 © Garcelon, P.
Tomocerus minor from the UK
Dental spines formula : 1-III-4-I-1-I
2018.10.07 © Cann, A.J.
Tomocerus minor from Holland
Eye with anterior square of 4 + 2 posterior ocelli
2018.11.12 © Kamsteeg, G.
Tomocerus minor from France
Footcomplex with clavate tenent macroseta (*)
2019.01.03 © Garcelon, P.
Tomocerus minor from Belgium
2019.04.14 © Huskens, M.-L.
Tomocerus minor from Belgium
Iridiscent blue body scales
2019.04.14 © Huskens, M.-L.
Tomocerus minor from Belgium
Eye with anterior square of 4 + 2 posterior ocelli
2019.04.14 © Huskens, M.-L.
Tomocerus minor from Belgium
Footcomplex with unguiculus, unguis and sticky macroseta
2019.09.14 © Huskens, M.-L.
Tomocerus minor
Descaled
After Yu D-Y & al, 2020 Fig.5
Tomocerus minor from France
Eye with anterior square of 4 ocelli (A-D)
2020.02.23 © Joachim, J.
Tomocerus minor from France
Bothriotrichum moved by airflow
2020.03.19 © Garcelon, P.
Tomocerus minor from the USA
Rainbow iridisation of scales
2020.04.10 © Pearson, R.R.
Tomocerus minor from the UK
"Truncated" mouthcone
2020.12.19 © Atkins, W.
Tomocerus minor from France
2022.02.12 © Joachim, J.
Tomocerus minor from France
2023.03.25 © Le Monde des Arthropodes.
Tomocerus minor from France
Iridised body scales
2023.03.25 © Le Monde des Arthropodes.
Tomocerus minor from Germany
Descaled
Head + thorax darker
2023.09.12 © Cheng, H.-J.
Tomocerus minor from the UK
Cephalic chaetotaxy
2023.12.25 © Barton, T.
Tomocerus (Tomocerus) minor from India
After Mandal P & al, 2024 Fig.6A
Tomocerus (Tomocerus) minor from India
Basal outer tooth with corner toothlet
After Mandal P & al, 2024 Fig.6D
Tomocerus minor from the UK
Eye with 6 visible ocelli
Anterior 4 ocelli form square
2024.01.17 © Barton, T.

Species mitrai Prabhoo & Muraleedharan, 1980

Tomocerus mitrai from India
After Mandal GP & Hazra AK, 2009 Fig.47a

Species montanus (Oudemans, JT in Weber, M, 1890:74) Yayuk, RS (sic), 1989:9

Species monticolus Huang & Yin, 1981

Species multisetus Sun, Liang & Huang, 2007

Species nepalicus Yosii, 1971

Species nigrofasciatus Sun, Y, Liang, A-P et Huang, F-S, 2006:559

Species nodentalis Nguyen, TT, 1995:564

Tomocerus nodentalis from North Vietnam
After Nguyen TT, 1995 Hinh.5.5

Species obscurus Huang & Yin, 1981

             Syn.: Tomocerus (Tomocerus) obsculus(sic) Chen, J-X et Ma, Y-T, 1997:157 ISS
             Syn.: Tomocerus (Tomocerus) obsculus(sic) Ma, Y-T et Christiansen, KA, 1998:47 ISS
             Syn.: Tomocerus (Tomocerus) obsculus(sic) Chen, J-X et Christiansen, KA, 1998:51 ISS

Species orientalis Martynova, 1979

Species parvus Huang & Yin, 1981

Species petalospinus Salmon, 1969

             Syn.: Tomocerus petolospinus(sic) Mandal, GP et Hazra, AK, 2009:126 ISS
             Syn.: Tomocerus (Tomocerus) petalospinous(sic) Mandal, GP, Kumari, S, Mandal, P, Roy, KK, Suman, KK et Bhattacharya, KK, 2024:764 ISS

Species problematicus Cassagnau, P, 1964:741

Species purpuratus Sun, Liang & Huang, 2006

Species sachalinensis Martynova, 1979

Species serratospinus Salmon, 1969

Species setoserratus Salmon, JT, 1941:347

Species shuense Liu, 2003

Species sibiricus Reuter, 1891

Species sikkimensis Mandal, GP, Kumari, S, Mandal, P, Roy, KK, Suman, KK et Bhattacharya, KK, 2024:757

Tomocerus (Tomocerus) sikkimensis from India
After Mandal P & al, 2024 Fig.1A
Tomocerus (Tomocerus) sikkimensis from India
After Mandal P & al, 2024 Fig.1B
Tomocerus (Tomocerus) sikkimensis from India
Ocelli ABCD in square formation
Modified after Mandal P & al, 2024 Fig.2B
Tomocerus (Tomocerus) sikkimensis from India
Mucro
Basal outer tooth with corner toothlet
After Mandal P & al, 2024 Fig.4D

Species spinistriatus Lee, B-H, 1975:958

Species steinbocki Yosii, 1971

Species teldanicus Gruia, 1995

Species tridentatus Sun, Liang & Huang, 2007

Species vasconicus Bonet, F, 1928:256

Tomocerus vasconicus from Spain
Dental spines
After Bonet F, 1928 Fig.2

            Syn.: Tomocerus navaricus(lapsus) Bonet, F, 1928:254 nn



             Syn.: Tomocerus (Tomocerus) vasconius(sic) Gisin, H, 1960:254 ISS

Species viridescens Wankel, 1860

             Syn.: Macrotoma viridiscens(sic) Absolon, K, 1899:317 ISS

Species viridis Yosii, R, 1967:27



Species-group ocreatus Zhang & al, 2014:393 key
Tomocerus sp. from Taiwan
Mucro
Lateral aspect
2023.03.02 © Cheng, H.-J.
Tomocerus sp. from Taiwan
Mucro posterior aspect
With 2 basal teeth
2023.03.03 © Cheng, H.-J.
Det.: Yu, D.-Y.
Tomocerus sp. from Taiwan
Dental spines array
2023.03.03 © Cheng, H.-J.
Det.: Yu, D.-Y.
Tomocerus sp. from Taiwan
Dental spines with many denticles
2023.03.03 © Cheng, H.-J.
Det.: Yu, D.-Y.


Species sp.1 Janssens F sensu Uchida, 1953

            Syn.: Tomocerus minor Uchida, 1953
            Syn.: Tomocerus (Tomocerus) minor Uchida, 1954
            Syn.: Tomocerus minor Stach, 1964
            Syn.: Tomocerus (Tomocerus) minor? Yosii, R, 1977:158
            Syn.: Tomocerus ocreatus Stach, 1964
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.2 Janssens F sensu Yosii, 1954

            Syn.: Tomocerus kawamurai Yosii, 1954
            Syn.: Tomocerus ochreatus(sic) f. kawamurai Yosii, R, 1956:90
            Syn.: Tomocerus kawamurai Yosii, R, 1967:19
            Syn.: Tomocerus kawamurai Yosii, R, 1977:158
            Syn.: Tomocerus ochreatus(sic) f. kawamurai Yosii, R, 1977:158
            Syn.: Tomocerus (ocreatus-complex) kawamurai Yu, D-Y, Man, LC et Deharveng, L, 2016:7
            Syn.: Tomocerus kawamurai f. depicta Yosii, R, 1955:392
            Syn.: Tomocerus ochreatus(sic) f. depicta Yosii, R, 1956:90
            Syn.: Tomocerus kawamurai f. depicta Yosii, R, 1967:19
            Syn.: Tomocerus kawamurai f. depicta Yosii, R, 1977:158
             Syn.: Tomocerus ochreatus(sic) Yosii, R, 1956:90 ISS
            Syn.: Tomocerus (Tomocerus) ocreatus Yosii, R, 1967:19
            Syn.: Tomocerus ocreatus Yosii, R, 1977:158
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.3 Janssens F sensu Stach, 1964

            Syn.: Tomocerus ocreatus Stach, 1964
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.4 Janssens F sensu Stach, 1965

            Syn.: Tomocerus ocreatus Stach, 1965
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7
            Syn.: Tomocerus (ocreatus-complex) nr folsomi Yu, D-Y, Man, LC et Deharveng, L, 2016:8


Species sp.5 Janssens F sensu Chiba, 1968

            Syn.: Tomocerus ocreatus Chiba, 1968
            Syn.: Tomocerus ocreatus f.1 Chiba, 1968
            Syn.: Tomocerus ocreatus f.2 Chiba, 1968
            Syn.: Tomocerus ocreatus f.3 Chiba, 1968
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.6 Janssens F sensu Lee B-H, 1975:958

            Syn.: Tomocerus ocreatus Lee, B-H, 1975:958
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.7 Janssens F sensu Martynova, 1977

            Syn.: Tomocerus ocreatus Martynova, 1977
            Syn.: Tomocerus (ocreatus-complex) ocreatus? Yu, D-Y, Man, LC et Deharveng, L, 2016:7


Species sp.8 Janssens F sensu Suetsugu, Satoh & Koike, 2010

            Syn.: Tomocerus (Tomocerus) ocreatus Suetsugu, N, Satoh, F et Koike, T, 2010:61
Tomocerus (Tomocerus) ocreatus from Japan
Dental spines
After Suetsugu N, Satoh F & Koike T, 2010 Photo41
Tomocerus (Tomocerus) ocreatus from Japan
Dental spine with basal teeth
After Suetsugu N, Satoh F & Koike T, 2010 Photo42




Species baibungensis Sun, Y, Liang, A-P et Huang, F-S, 2006:561

Species cthulhu Yu & Li, 2016

Species deharvengi Yu & Li, 2016

Species deogyuensis Lee, B-H, 1975:949

             Syn.: Tomocerus (Tomocerus) deongyuensis(sic) Liu, Hou & Li, 1998 ISS

Species dong Yu & Li, 2016

Species folsomi Denis, 1929

Species hexipunctatus Sun, Liang & Huang, 2007

Species jiuzhaiensis Liu, Zhou & Zhang, 2013

Species wushanensis Sun, Liang & Huang, 2007


Subgenus Ciliatomurus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 nn
  Syn.: Tomocerus Ciliatomurus Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 nn
Remark by Janssens, F. 2021.07.23, 2023.03.11: Given the new taxon is described in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given this electronic publication has not been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, and 2. the taxon Ciliatomurus Yu et al, 2020 is a nomen nudum, given its description is unpublished. In addition, according to ICZN Art.21.9, the publication date of Yu & al. is 24 October, 2020, not January, 2021.

  Syn.: Tomocerus jilinensis-clade Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6



Species jesonicus Yosii, R, 1967:31, t.t.

            Syn.: Tomocerus vulgaris Yosii, 1954
            Syn.: Tomocerus vulgaris Japanese records =?
            Syn.: Tomocerus vulgaris Yosii, R, 1967:32 =?
            Syn.: Tomocerus vulgaris var. kurilensis Uchida, 1964
            Syn.: Tomocerus vulgaris var. kurilensis Yosii, R, 1967:31 =?
            Syn.: Tomocerus vulgaris var. kurilensis Yosii, R, 1977:158
            Syn.: Tomocerus jezoensis Yosii, R, 1977:158 =?


            Syn.: Tomocerus vulgaris ssp. jesonicus Yosii, 1940 nn

Species jilinensis Ma, 2011, t.t.

Tomocerus jilinensis
Descaled
After Yu D-Y & al, 2020 Fig.5

Subgenus Coloratomurus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:3 nn
  Syn.: Tomocerus Coloratomurus Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 nn
Remark by Janssens, F. 2021.07.23, 2023.03.11: Given the new taxon is described in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given this electronic publication has not been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, and 2. the taxon Coloratomurus Yu et al, 2020 is a nomen nudum, given its description is unpublished. In addition, according to ICZN Art.21.9, the publication date of Yu & al. is 24 October, 2020, not January, 2021.

  Syn.: Tomocerus bimaculatus-clade Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6




Species bimaculatus Sun, Liang & Huang, 2006, t.t.

Species cheni Ma, Y-T et Christiansen, KA, 1998:47

Species cuspidatus Börner, C, 1909:114

            Syn.: Tomocerus minor Uchida, 1954 =?
            Syn.: Tomocerus kumei Yosii, 1954
            Syn.: Tomocerus kumei Yosii, R, 1967:22
            Syn.: Tomocerus kumei Yosii, R, 1977:158
            Syn.: Tomocerus kumei Yoshii R in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:4
            Syn.: Tomocerus uenoi Yosii, 1954
            Syn.: Tomocerus uenoi Yosii, R, 1967:22
            Syn.: Tomocerus uenoi Yosii, R, 1977:158
            Syn.: Tomocerus uenoi Yoshii R in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:4


Tomocerus (Tomocerus) cuspidatus from Japan
After Yosii R, 1967 Fig.11A
             Syn.: Tomocerus cuapidatus(sic) Yosii, R, 1967:2 ISS
Tomocerus gr. cuspidatus from China
2023.06.24 © Wang, G.
Det.: Yu, D.-Y.
Tomocerus gr. cuspidatus from China
Colouration of basal antennal segments
2023.06.24 © Wang, G.
Det.: Yu, D.-Y.
Tomocerus gr. cuspidatus from China
2023.06.24 © Wang, G.
Det.: Yu, D.-Y.



Species jordanai Sun, Liang & Huang, 2006

Species nigromaculatus Sun, Y, Liang, A-P et Huang, F-S, 2006:513

Tomocerus (Tomocerus) nigromaculatus from Tibet
After Sun & al, 2006 Fig.1A

Species purpurithorus (Liu, Hou & Li, 1999) Ma, Y-T, Chen, J-X & Christiansen, K, 2003:516

Tomocerina purpurithora
Descaled
After Yu D-Y & al, 2020 Fig.5

Subgenus Striatomurus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:7 nn
  Syn.: Tomocerus Striatomurus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 nn
Remark by Janssens, F. 2021.07.23, 2023.03.11: Given the new taxon is described in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given this electronic publication has not been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, and 2. the taxon Striatomurus Yu et al, 2020 is a nomen nudum, given its description is unpublished. In addition, according to ICZN Art.21.9, the publication date of Yu & al. is 24 October, 2020, not January, 2021.

  Syn.: Tomocerus nigrus-clade Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6




  Syn.: Tomocerus Striatomurus Arbea, JI, 2021:30 nn
  Syn.: Tomocerus Striatomurus Arbea, JI, Baquero, E, Beruete, E, Pérez Fernández, T et Jordana, R, 2021:52 nn

Species botanicus Cassagnau, P, 1962:230


Tomocerus botanicus from France
Dental spines I2(3)I(2-)4/(2)3(-5)
After Cassagnau P, 1962 Fig.1AB
Tomocerus botanicus from France
Large dental spine with secondary denticle
After Cassagnau P, 1962 Fig.1C
Tomocerus botanicus from France
Footcomplex
After Cassagnau P, 1962 Fig.1D
Tomocerus botanicus from France
Mucro with 2(3) internal teeth
After Cassagnau P, 1962 Fig.1E

            Syn.: Tomocerus vulgaris Ruppel, 1953
            Syn.: Tomocerus vulgaris Ponge, J-F, 1983:367
            Syn.: Tomocerus sp. Hopkin S, 1998 =?
Tomocerus sp. from the UK
Mucro with one intermediate tooth
1998 © Hopkin, S.

            Syn.: Tomocerus catalanus Hopkin, 1998 =?
            Syn.: Tomocerus 'catalanus' Shaw, P et Benefer, CM, 2015:200 =?
Tomocerus sp.3 from the UK
2012.12.09 © Shaw, P.
Tomocerus sp.3 from the UK
Dark pigmented clypeus
2012.12.09 © Shaw, P.
Tomocerus sp.3 from the UK
Aberrant mucro with one (optional) intermediate tooth
2012.12.09 © Shaw, P.
Tomocerus sp.3 from the UK
Dental spines
2012.12.09 © Shaw, P.
Tomocerus sp.3 from the UK
Foot complex
2012.12.09 © Shaw, P.

            Syn.: Tomocerus sp.nov. Shaw, P et Benefer, CM, 2015:201 =?



Tomocerus botanicus from France
Descaled
2021.10.15 © Yu, D.-Y.
Tomocerus botanicus from France
Ocelli ABCD in square
2021.10.15 © Yu, D.-Y.

Species fopingensis Sun & Liang, 2008

Species maculatus Sun, Liang & Huang, 2007

Species nigrus Sun, Y, Liang, A-P et Huang, F-S, 2006:803

Tomocerus nigrus from China
2016.03.28 © Home, M.
Det.: Yu, D.-Y.
Tomocerus nigrus from China
2016.03.28 © Home, M.
Det.: Yu, D.-Y.
Tomocerus nigrus
Descaled
After Yu D-Y & al, 2020 Fig.5

Species punctatus Yosii, R, 1967:28

             Syn.: Tomocerus (Tomocerus) puncutatus(sic) Yosii, R, 1967:28 ISS
Tomocerus (Tomocerus) puncutatus(sic) from Japan
After Yosii R, 1967 Fig.15A
Tomocerus punctatus from China
After Sun Y & Liang A-P, 2010 Fig.1-2



Species vulgaris (Tullberg, 1871) Brook, 1883, t.t., B

            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
            Syn.: Macrotoma nigra Guthrie, JE, 1903:80
            Syn.: Tomocerus niger Guthrie, JE, 1903:80
            Syn.: Macrotoma nigra Folsom, JW, 1913:459 =?
            Syn.: Tomocerus plumbeus Packard, 1873, partim
            Syn.: Tomocerus celer Guthrie, JE, 1903:80
            Syn.: Tomocerus minutus Hammer, 1953
            Syn.: Tomocerus vulgaris var. kurilensis Uchida, H, 1965:207


Tomocerus vulgaris from Germany
2005 © Menzel, E.
Tomocerus vulgaris from Germany
2005 © Kohl, F.
Tomocerus vulgaris from Germany
2005 © Peters, J.
Tomocerus vulgaris from Germany
2006 © Peters, J.
Tomocerus vulgaris from Holland
2007 © Hautus, M.A.
Tomocerus vulgaris from Belgium
2007.07.12 © Janssens, F.
Tomocerus vulgaris from Germany
2007.04.07 © Kohl, F.
Tomocerus vulgaris from the Netherlands
2007.09.18 © Arp, X.
Tomocerus vulgaris from Germany
2008.01.25 © Peters, J.
Tomocerus vulgaris from the UK
2008.01.27 © Barton, T.
Tomocerus vulgaris from Belgium
2008.08.28 © Janssens, F.
Tomocerus vulgaris from Holland
2008.10.06 © Baas, A.H.
Tomocerus vulgaris from Holland
2008.10.07 © Baas, A.H.
Tomocerus vulgaris from Holland
2008.10.10 © Baas, A.H.
Tomocerus vulgaris from France
2008.11.03 © Ramel, A.
Tomocerus vulgaris from Germany
2008.12.07 © Peters, J.
Tomocerus vulgaris from Belgium
2009.02.21 © Vuijlsteke, M.
Tomocerus vulgaris from the UK
2009.03.04 © Ransom, T.
Tomocerus vulgaris from Belgium
2009.03.07 © Janssens, F.
Tomocerus vulgaris from France
2009.04.19 © Lefebvre, V.
Det. Ponge, J.-F.
Tomocerus vulgaris from Belgium
2009.04.19 © Doornaert, A.
Tomocerus vulgaris from Germany
2009.11.07 © Buonvolere, D.
Tomocerus vulgaris from Belgium
Dental spines
2010.01.13 © De Bruyn, L.
Tomocerus vulgaris from Slovenia
2010.04.15 © Miran, C.
Tomocerus vulgaris from Holland
2010.04.30 © van Duinen, J.
Tomocerus vulgaris from Holland
Typical dorsal banding
2011.08.17 © van Duinen, J.
Tomocerus vulgaris from Germany
2011.10.26 © Lenke, M.
Tomocerus vulgaris from Germany
Metathoracic botriotrichum: mechanoreceptive sensilla in cup-like depression of skin
2011.10.26 © Lenke, M.
Tomocerus vulgaris from the UK
2012.02.03 © Dargie, S.
Tomocerus vulgaris from Belgium
2012.02.26 © Wallays, H.
Tomocerus vulgaris from the UK
2012.05.23 © Nicholls, D.
Tomocerus vulgaris from Belgium
2013.08.09 © Wallays, H.
Tomocerus vulgaris from Belgium
The "golden Tomocerus"
2013.08.11 © Wallays, H.
Tomocerus vulgaris from Czechia
2014.05.07 © Deml, M.
Tomocerus vulgaris from Belgium
Mouthcone distinctly truncated
2016.07.15 © Huskens, M.-L.
Tomocerus vulgaris from Belgium
Clypeus, labrum and labium
2016.07.15 © Huskens, M.-L.
Tomocerus vulgaris from Belgium
Metathoracic bothriotrichum
2016.07.15 © Huskens, M.-L.
Tomocerus vulgaris from Belgium
2017.08.19 © Willems, J.
Tomocerus vulgaris from Belgium
Telescopically exposed anal valves
2017.12.07 © Huskens, M.-L.
Tomocerus vulgaris ♂ from Belgium
Male genital orifice with circumgenital setae
2018.02.04 © Huskens, M.-L.
Tomocerus vulgaris from Belgium
Eyepatch with square of 4 + 2 ocelli
2018.02.09 © Huskens, M.-L.
Tomocerus vulgaris from the UK
Abd.4-6
2018.10.23 © Smart, J.
Tomocerus vulgaris from Holland
Abd.1-3 iridisation
2020.03.01 © Kamsteeg, G.
Tomocerus vulgaris ♂ from Holland
Broadened manubrial base
2020.03.01 © Kamsteeg, G.
Tomocerus vulgaris ♂ from Holland
Genital papilla roundish
2020.03.01 © Kamsteeg, G.
Tomocerus vulgaris from the UK
Golden metallic posterior segment margins
2021.05.11 © Atkins, W.
Tomocerus vulgaris from the UK
Body scales with continuous longitudinal ribs
2021.05.11 © Atkins, W.
Tomocerus vulgaris from the UK
Abd.3+4 with lateral bothriotrichum
2021.05.11 © Atkins, W.
Tomocerus vulgaris from France
2023.02.11 © Clerc, O.
Tomocerus vulgaris from France
Ocelli ABCD form square
2023.02.11 © Clerc, O.
Tomocerus vulgaris from France
Th.2 bothriotrichum
2023.02.11 © Clerc, O.
Tomocerus vulgaris from France
Midfoot with clavate tenent macroseta
2023.02.11 © Clerc, O.
Tomocerus vulgaris from Germany
2023.04.29 © Vogl, U.
Tomocerus vulgaris from Germany
Body scale translucent
With longitudinal and transvers structure
2023.05.02 © Vogl, U.
Tomocerus vulgaris from Germany
Body scale translucent
Longitudinal rib distance about 1.5 micron
2023.05.02 © Vogl, U.
Tomocerus vulgaris from Germany
Body scale iridisation
Structural colours
2023.05.02 © Vogl, U.
Tomocerus vulgaris from Germany
Body scale with structural colours
2023.05.02 © Vogl, U.

Subgenus Ocreatomurus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 nn
  Syn.: Tomocerus Ocreatomurus Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 nn
Remark by Janssens, F. 2021.07.23, 2023.03.11: Given the new taxon is described in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given this electronic publication has not been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, and 2. the taxon Ocreatomurus Yu et al, 2020 is a nomen nudum, given its description is unpublished. In addition, according to ICZN Art.21.9, the publication date of Yu & al. is 24 October, 2020, not January, 2021.

  Syn.: Tomocerus ocreatus-clade Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6



  Syn.: Tomocerus Ocreatomurus Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:5 nn
  Syn.: Tomocerus Ocreatomurus Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:127 nn
Remark by Nguyen, A.T.T. & al. 2022:126: However, we accept these subgenera since we consider that they are sufficiently characterized in the printed version of the publication (page 5) [of Yu & al. (2021(sic))]."
Remark by Janssens, F. 2023.03.11: There is no characterisation of the subgenus Ocreatomurus on p.5 of Yu & al (2021(sic)). The name Ocreatomurus is only mentioned in Fig.2 on page 5 of both the electronic version of 24 October, 2020 as in the printed version of January, 2021.

Tomocerus (Ocreatomurus) from Taiwan
2023.03.06 © Cheng, H.-J.
Tomocerus (Ocreatomurus) from Taiwan
Dental spines with scale-like surface structures
2023.03.08 © Cheng, H.-J.




Species changbaishanensis Wang, 1999

Tomocerus changbaishanensis from China
After Yu D-Y & al, 2018 Fig.3A

Species conagensis Sun, Y, Liang, A-P et Huang, F-S, 2006:516

             Syn.: Tomocerus conagenesis(sic) Mandal, GP, Kumari, S, Mandal, P, Roy, KK, Suman, KK et Bhattacharya, KK, 2024:759 ISS
             Syn.: Tomocerus conagenesis(sic) Mandal, GP, Kumari, S, Mandal, P, Roy, KK, Suman, KK et Bhattacharya, KK, 2024:765 ISS

Species huangi Yu D-Y in Yu, D-Y, Qin, C-Y, Ding, Y-H, Hu, F, Zhang, F et Liu, M-Q, 2018:156

Tomocerus huangi from China
After Yu D-Y & al, 2018 Fig.3B

Species leyensis Yu, D-Y et Deharveng, L, 2018:55

Tomocerus leyensis from China
After Yu D-Y & Deharveng L, 2018 Fig.1B

Species maximus Liu, Hou & Li, 1999

             Syn.: Tomocerus ochreatus(sic) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6 ISS
            Syn.: Tomocerus kawamurai Yoshii R in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:6


Species nabanensis Yu, D-Y, Yang, X-D et Liu, M-Q, 2018:232

Tomocerus nabanensis from China
After Yu D-Y, Yang X-D & Liu M-Q, 2018 Fig.1C

Species nanus Yu, D-Y, Yang, X-D et Liu, M-Q, 2018:230

            Syn.: Tomocerus (ocreatus-group) nan(sic) Yu, D-Y, Yang, X-D et Liu, M-Q, 2018:230 IOS
Tomocerus nan(sic) from China
After Yu D-Y, Yang X-D & Liu M-Q, 2018 Fig.1B
Remark by Janssens, F. 2018.11.19: According to the etymology, 'nan' is a vernacular Chinese word for 'south'. ICZN Recommendation 11A: Use of vernacular names. An unmodified vernacular word should not be used as a scientific name. Appropriate latinization is the preferred means of formation of names from vernacular words.



Species ocreatus Denis, 1948, t.t.

            Syn.: Tomocerus kawamurai Lee, B-H, 1975:958


             Syn.: Tomocerus ochreatus(sic) Yosii, R et Lee, C-E, 1963:4 ISS
Tomocerus ocreatus from South Vietnam
After Yu D-H, Man LC & Deharveng L, 2016 Fig.1A
Tomocerus ocreatus from South Vietnam
Dental spines with numerous moderate denticles
After Yu D-H, Man LC & Deharveng L, 2016 Fig.3F
Tomocerus ocreatus from Taiwan
2021.08.31 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Ocelli ABCD in a square
2021.08.31 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Descaled
2021.09.10 © Cheng, H.-J.
Det.: Yu, D.-Y.
Tomocerus ocreatus from Taiwan
2021.09.11 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Ocelli ABCD form a square
2021.09.11 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Dental spines base to apex :
3 small + 4 small + 2 large
2021.09.11 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
2021.09.13 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
2021.11.18 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Dental array about half dens in length
2021.11.18 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Anterior ocelli ABCD in square formation
Typical in Tomocerus
2021.12.14 © Cheng, H.-J.
Tomocerus ocreatus from Taiwan
Distinctly long antennae
As seen in Pogonognathellus
2021.12.14 © Cheng, H.-J.



Species paraspinulus Gong, X, Qin, C-Y et Yu, D-Y, 2018:278

Tomocerus paraspinulus from China
After Gong X, Qin C-Y & Yu D-Y, 2018 Fig.1B

Species postantennalis Yu, D-Y, Zhang, F et Deharveng, L, 2014:63

Tomocerus postantennalis from Vietnam
After Yu D-Y al, 2014 Fig.1

Species pseudocreatus Yu D-Y in Yu, D-Y, Qin, C-Y, Ding, Y-H, Hu, F, Zhang, F et Liu, M-Q, 2018:157

Tomocerus pseudocreatus from China
After Yu D-Y & al, 2018 Fig.3C

Species pseudospinulus Gong, X, Qin, C-Y et Yu, D-Y, 2018:275

Tomocerus pseudospinulus from China
After Gong X, Qin C-Y & Yu D-Y, 2018 Fig.1A

Species qinae Yu, Yao & Hu, 2016

Tomocerus qinae from China
After Yu D-Y & al, 2018 Fig.2

             Syn.: Tomocerus qine(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:10 ISS
Tomocerus qinae from China
Ocelli ABCD in deformed square
2021.10.15 © Yu, D.-Y.
Tomocerus qinae from China
Axis AC longer than BD
2021.10.15 © Yu, D.-Y.
Tomocerus (Ocreatomurus) qinae from India
After Mandal P & al, 2024 Fig.8A



Species qixiaensis Yu, Yao & Hu, 2016

Species spinulus Chen, J-X et Christiansen, KA, 1998:51

Tomocerus spinulus from China
After Yu D-Y & al, 2018 Fig.3D

Species tropicus Yu, D-Y, Yang, X-D et Liu, M-Q, 2018:227

Tomocerus tropicus from China
After Yu D-Y, Yang X-D & Liu M-Q, 2018 Fig.1A

Species virgatus Yu D-Y in Yu, D-Y, Qin, C-Y, Ding, Y-H, Hu, F, Zhang, F et Liu, M-Q, 2018:161

Tomocerus virgatus from China
After Yu D-Y & al, 2018 Fig.3E

Species yueluensis Yu D-Y in Yu, D-Y, Qin, C-Y, Ding, Y-H, Hu, F, Zhang, F et Liu, M-Q, 2018:162

Tomocerus yueluensis from China
After Yu D-Y & al, 2018 Fig.3F

Species zayuensis Huang & Yin, 1981

             Syn.: Tomocerus (Tomocerus) zayensis(sic) Ma, Y-T et Christiansen, KA, 1998:47 ISS
             Syn.: Tomocerus (Tomocerus) zayensis(sic) Chen, J-X et Christiansen, KA, 1998:51 ISS
Tomocerus zayuensis from China
After Yu D-Y & al, 2018 Fig.3G



Species zhuque Yu D-Y in Yu, D-Y, Qin, C-Y, Ding, Y-H, Hu, F, Zhang, F et Liu, M-Q, 2018:166

Tomocerus zhuque from China
After Yu D-Y & al, 2018 Fig.3H


Genus Yoshiicerus Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5 nn

  Syn.: Tomocerinae Yoshiicerus Yu D-Y in Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:1 nn
Remark by Janssens, F. 2021.07.23, 2023.03.11: Given the new taxon is described in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given this electronic publication has not been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, and 2. the taxon Yoshiicerus Yu et al, 2020 is a nomen nudum, given its description is unpublished. In addition, according to ICZN Art.21.9, the publication date of Yu & al. is 24 October, 2020, not January, 2021.

  Syn.: Tomocerus kinoshitai-group Yu & al, 2017
  Syn.: Tomocerus kinoshitai-group Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:2
  Syn.: Tomocerus kinoshitai-clade Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F et Liu, M-Q, 2020:5


Species caecus (Yu & Deharveng, 2015) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:1

Species dissimilis (Yu, Ding & Ma, 2017) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:2

Tomocerus dissimilis
Descaled
After Yu D-Y & al, 2020 Fig.5

Species kinoshitai (Yosii, 1954) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:2

            Syn.: Tomocerus (Tomocerus) kinoshitai ssp. dentiferus Yosii, R, 1956:90
            Syn.: Tomocerus kinoshitai ssp. dentiferus Yosii, R, 1967:20
            Syn.: Tomocerus kinoshitai ssp. dentiferus Lee, B-H, 1974:412
            Syn.: Tomocerus kinoshitai ssp. dentiferus Yosii, R, 1977:158


             Syn.: Tomocerus kinonoshitai(sic) Lee, B-H et Park, K-H, 1991:273 ISS
             Syn.: Tomocerus kinonoshitai(sic) Lee, B-H et Park, K-H, 1991:284 ISS

  Subspecies denticulus Lee, B-H, 1975:953



Species persimilis (Yu, Ding & Ma, 2017) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:2, t.t.

Species similis (Chen, J-X & Ma, Y-T, 1997:157) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:2

Species tiani (Yu, 2016) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:2

Species troglodytes (Yu, D-Y & Deharveng, L, 2018:51) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:1

Tomocerus troglodytes from China
After Yu D-Y & Deharveng L, 2018 Fig.1A


Genus Tritomurus Frauenfeld, 1854 key

Species catalanus (Denis, J-R, 1924:198) Yu, D-Y, Deharveng, L, Lukíc, M, Wei, Y-M, Hu, F & Liu, M-Q, 2020:8

Tomocerus catalanus from Spain
Dental spines
After Bonet F, 1928 Fig.1
Remark by Janssens, F. 2021.07.25: Given the new combination is assigned in Appendix A of the original publication, given that appendix is provided as a supplementary file, in other words as an external resource, given the supplement is only available as an online document, it is not available in the printed version of the publication, so Appendix A is an electronic publication, given neither this electronic publication nor the recombination itself have been registered in Zoobank, given this online document is formatted as a Microsoft Word document, it is not in a fixed (portable) format, Appendix A has not been published in accordance to ICZN rules for published works and therefore 1. Appendix A is an unpublished work, 2.the recombintions in Appendix A are unavailable for Zoology.

Species falcifer Cassagnau, P, 1958:121

Tritomurus falcifer
After Thibaud J-M & D'Haese C, 2010 Fig.19

Species scutellatus Frauenfeld, 1854:15, t.t.

            Syn.: Tomocerus anophthalmus Absolon, K, 1901:88
            Syn.: Tomocerus anophthalmus Absolon, K, 1903:97
            Syn.: Tomocerus anophthalmus Bonet, F, 1931:100


Tritomurus scutellatus from Croatia
2010.11.19 © Lukic, M.
Tritomurus scutellatus
Descaled
After Yu D-Y & al, 2020 Fig.5

Species terrestralis Stach, 1922

            Syn.: Tomocerus gradjackae Denis, 1933


             Syn.: Tritomurus terrestrialis(sic) Denis, J-R, 1929:179 ISS
             Syn.: Tritomurus terrestrialis(sic) Bonet, F, 1931:103 ISS
             Syn.: Tomocerus terrestrialis(sic) Christiansen, K, 1964:656 ISS

Species veles Lukic, M, Houssin, C et Deharveng, L, 2010:3

Tritomurus veles from Croatia
2010.11.23 © Houssin, C.


Genus Pogonognathellus Paclt, 1944

  Syn.: Thysanoura Podura Say, 1821 partim
  Syn.: Tomocerini(sic) Pogonognathus Börner, C in Schultze, 1908:66
  Syn.: Tomoceridae Pogonognatus(sic) Yosii, R, 1967:2 ISS
  Syn.: Isotomidae Architomocerura Denis, 1931
  Syn.: ... Archisotoma(sic) Bolger, 1987 ISS
  Syn.: ... Paratomocerus Tarsia in Curia, 1938
  Syn.: Tomocerus Monodontocerus Salmon, JT, 1964:121
  Syn.: ... Maynardia Yosii, 1956, nec Dall, 1894
  Syn.: Collembola Tomerocus(sic) Haybach, G, 1959:Tab.VIII lapsus
  Syn.: Entomobryidae Tomocerus Delhez, F et Kersmaekers, M, 1973:8
  Syn.: Collembola Tomocerus Busmachiu, G, 2010:121
  Syn.: Tomoceridae Tomocerus Borodin, OI et Tsinkevich, VA, 2016:13


  Syn.: Tomoceridae Pogonognthellus(sic) Yosii, R, 1967:11 ISS
Pogonognathellus from the USA
Descaled revealing intrinsic body colour
2006 © Murray, T.
Pogonognathellus sp. from France
Furca with short dental spines array
2007.04.10 © Lebeaux, P.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.
Pogonognathellus sp. from the USA
Eyepatch with diamond of 4 + 2 ocelli
2009.02.26 © Cowen, R.
Pogonognathellus sp. from Taiwan
Tapering antenna
2010.06.23 © Chien, H.-C.
Pogonognathellus sp. from Taiwan
2010.06.30 © Chien, H.-C.
Pogonognathellus longicornis from Sweden
Scaled 3rd antennal segment
2011.11.25 © Hallmén, J.
Pogonognathellus sp. from the UK
Eyepatch with diamond of 4 + 2 ocelli
2012.01.23 © Kent, J.J.

  Syn.: Tomoceridae Pogonathellus(sic) Smith, CL, 2015:3 u ISS
Pogonognathellus longicornis from France
Base of ant.3 scaled (*)
2015.05.16 © Ramel, A.
Pogonognathellus longicornis from France
Distinctly long macrosetae at posterior margin of abd.5
2015.05.16 © Ramel, A.
Pogonognathellus longicornis from the UK
Abd.5 with very long bent dorsal macrosetae
2015.09.29 © Horton, M.
Pogonognathellus longicornis from the UK
Eyepatch with diamond of 4 + 2 ocelli
2015.09.29 © Horton, M.
Pogonognathellus sp. from Slovenia
Abd.5 with very long bent dorsal macrosetae
2015.11.15 © Pisec, N.
Pogonognathellus sp. from the UK
Eyepatch with diamond of 4 + 2 ocelli
2016.02.10 © Phillips, E.
Pogonognathellus longicornis from the UK
Abd.5 with very long bent dorsal macrosetae
2016.02.23 © Branch, A.
Pogonognathellus sp. from Sweden
Eyepatch with diamond of 4 + 2 ocelli
2016.09.16 © Nilsson, K.
Pogonognathellus longicornis from the UK
Abd.5 with very long bent dorsal macrosetae
2016.10.29 © Henton-Jones, D.
Pogonognathellus longicornis from the UK
Third tibiotarsus with distinctly long spiky macrosetae (*)
2016.10.29 © Henton-Jones, D.
Pogonognathellus sp. from Canada
Anterior 4 ocelli form a diamond
2018.01.12 © Barbin, T.
Pogonognathellus sp. from Canada
Scales present on ant.3
2018.01.12 © Barbin, T.
Pogonognathellus sp. from Baltic amber
Ant.3 distinctly long and curled-up
2019.02.28 © Klug, G.
Pogonognathellus sp. from Baltic amber
Ant.3 distinctly annulated
2019.02.28 © Klug, G.
Pogonognathellus sp. from France
Abd.5+6
2019.12.19 © Dejoux, P.
Pogonognathellus sp. from Belgium
Early juvenile instar
2021.01.03 © Van Elegem, Be.
Pogonognathellus dubius s.l. from the USA
Ant.3 apically tapering
Ant.4 shorter then ant.2
2021.06.14 © Pavy, R.

  Syn.: Tomoceridae Pogonognathelus(sic) Bellini, BC, Zhang, F, Souza, PGC, dos Santos-Costa, RC, Medeiros, GdS et Godeiro, NN, 2022:3 ISS
  Syn.: Tomoceridae Pogonognathelus(sic) Bellini, BC, Zhang, F, Souza, PGC, dos Santos-Costa, RC, Medeiros, GdS et Godeiro, NN, 2022:5 ISS
Pogonognathellus dubius s.l. from the USA
Antennae curling up before jump
2022.02.02 © Pavy, R.
Pogonognathellus sp. from Taiwan
2022.12.23 © Cheng, H.-J.
Pogonognathellus sp. from Taiwan
Eye : anterior 4 ocelli form a diamond
2022.12.23 © Cheng, H.-J.
Pogonognathellus sp. from Taiwan
2022.12.23 © Cheng, H.-J.
Pogonognathellus sp. ♀ from Taiwan
Anal valve elongate pointed
2022.12.30 © Cheng, H.-J.
Pogonognathellus sp. from the USA
Hind tibiotarsus with 1 inner spine-like macroseta
(Descaled specimen)
2023.07.29 © Moore, J.
Pogonognathellus sp. from the USA
Anus telescoped out
Genital segment exposed
2024.03.24 © Miller, B.T.



Species sp.1 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475

Species beckeri (Börner, C, 1909:115) Yosii, 1946

            Syn.: Tomocerus flavescens Gisin, 1944 =?
            Syn.: Pogonognathellus flavescens Yosii, 1954
            Syn.: Pogonognathellus flavescens Yosii, R, 1956:89
            Syn.: Pogonognathellus flavescens Selga, 1961
            Syn.: Pogonognathellus flavescens Yosii, R, 1967:10
            Syn.: Pogonognathellus flavescens Yosii, R, 1977:157


             Syn.: Tomocerus backeri(sic) Selga, 1961 ISS
             Syn.: Pogonognathellus bockeri(sic) Yosii, R, 1967:6 ISS
Pogonognathellus beckeri from Japan
Antennae mutilated
2008.01.20 © Tsukiji, T.
Pogonognathellus beckeri from Japan
Right antenna partly recovered
2008.12.19 © Tsukiji, T.
Pogonognathellus beckeri from Japan
Eyepatch with diamond of 4 + 2 ocelli
2016.07.18 © Hara, Y.
Pogonognathellus beckeri from Japan
Ant.3 annulated, scales present
2016.07.18 © Hara, Y.
Pogonognathellus beckeri juv. from Japan
2016.07.30 © Hara, Y.
Pogonognathellus beckeri from Japan
2018.10.16 © Hara, Y.
Pogonognathellus beckeri from Japan
2018.10.16 © Hara, Y.
Pogonognathellus beckeri from Japan
Eyepatch with diamond of 4 anterior + 2 posterior ocelli
2018.10.16 © Hara, Y.



Species belmontorum Park, K-H, Bernard, EC et Moulton, JK, 2011:2

Pogonognathellus belmontorum from the USA
After Park K-H & al., 2011 Fig.1A
            Syn.: Pogonognathellus flavescens Nearctic records partim


Species bidentatus (Folsom, JW, 1913:463) Yosii, R, 1967:7

            Syn.: Tomocerus bidentatus ssp. Jeanneli(sic) Bonet, 1934
            Syn.: Tritomurus henroti Delamare Deboutteville, 1949
             Syn.: Tritomurus Henroti(sic) Cassagnau, P, 1958:121 ISS


             Syn.: Tomocerus (Pogonognathellus) dibentatus(sic) Christiansen, K, 1964:668 ISS
Pogonognathellus bidentatus from the USA
After Felderhoff KL et al., 2010 Fig.2A
Antennae typically curved backwards in capricorn fashion
Pogonognathellus bidentatus from the USA
After Felderhoff KL et al., 2010 Fig.5A
Descaled; cuticular pattern
Pogonognathellus bidentatus? from the USA
Ant.4 distinctly very short (*)
2017.03.14 © Dobbs, Br.
Pogonognathellus bidentatus? from the USA
Ant.4 distinctly very short (*)
2017.05.27 © Montgomery, G.
Pogonognathellus bidentatus? from the USA
Ant.4 distinctly very short
2017.10.11 © Dobbs, Br.
Pogonognathellus bidentatus? from the USA
Dark spots mark bothriotricha
2017.10.11 © Dobbs, Br.
Pogonognathellus bidentatus? from the USA
Diamond of 4 anterior ocelli
2017.10.11 © Dobbs, Br.
Pogonognathellus bidentatus? from Canada
Distinctly short ant.4
2017.12.05 © Barbin, T.
Pogonognathellus bidentatus? from the USA
Distinctly short ant.4 (*)
2017.12.24 © Fleming, K.
Pogonognathellus bidentatus? from Canada
Distinctly short ant.4 (*)
2018.07.01 © Barbin, T.
Pogonognathellus bidentatus? from the USA
Distinctly short ant.4 (*)
2019.01.03 © Pearson, R.R.



Species borealis Yosii, R, 1967:11

            Syn.: Tomocerus flavescens ssp. arcticus Yosii, R, 1956:89
            Syn.: Tomocerus flavescens var. arcticus Yosii, R, 1967:11
            Syn.: Pogonognathellus flavescens var. arcticus Yosii, R, 1977:157


Pogonognathellus ?borealis? from Japan
2005.09.07 © Keresztes, G.

Species brevifulvus Park, K-H, Bernard, EC et Moulton, JK, 2011:6

Pogonognathellus brevifulvus from the USA
After Park K-H & al., 2011 Fig.3A
Pogonognathellus brevifulvus from the USA
2013.07.04 © Liberta, A.

Species celsus (Christiansen, K, 1964:670) Yosii, R, 1967:46

Species danieli Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488

Pogonognathellus danieli from the USA
After Felderhoff KL et al., 2010 Fig.5B
Descaled; cuticular pattern

Species dubius (Christiansen, K, 1964:671) Yosii, R, 1967:12

            Syn.: Pogonognathellus grayspeciescomplex Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488
Remark by Felderhoff, K.L. et al. 2010:488: The taxa included in this group are united by cuticular pigmentation, scale color, and alignment of the basal and lateral mucronal teeth, but the sequenced specimens do not seem to form a monophyletic group. Instead, they clustered in four separate groups scattered among members of the "pale complex." Additional collections, including specimens from the P. dubius type locality, will be necessary to begin to satisfactorily resolve these taxa.

            Syn.: Pogonognathellus sp.4 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.9 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475


Pogonognathellus dubius s.l. from the USA
2005 © DiTerlizzi, T.
Pogonognathellus dubius s.l. from the USA
2005 © Murray, T.
Pogonognathellus dubius s.l. from the USA
2006 © Cresswell, S.
Pogonognathellus dubius s.l. from the USA
2007 © Murray, T.
Pogonognathellus dubius s.l. from Canada
2007.12.17 © McCann, S.
Pogonognathellus dubius s.l. from the USA
2008.09.09 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2008.10.08 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2008.12.06 © Houston, S.
Pogonognathellus dubius s.l. from the USA
2009.06.18 © Bradford, A.
Pogonognathellus dubius s.l. from the USA
2009.10.25 © Bergen, L.
Pogonognathellus dubius s.l. from the USA
2010.06.26 © Ueda, K.
Pogonognathellus dubius s.l. from the USA
2016.08.20 © Schulz, K.
Pogonognathellus dubius s.l. from Canada
2017.10.29 © Barbin, T.
Pogonognathellus dubius s.l. from Canada
Body scales in 2 colours
2017.10.29 © Barbin, T.
Pogonognathellus dubius s.l. from the USA
2020.09.16 © Rose, J.S.

Species elongatus (Maynard, EA, 1951:126) Yosii, R, 1967:3

            Syn.: Tomocerus (Pogonognathus) nigritus Christiansen, K, 1964:649 =?,is


Pogonognathellus elongatus from the USA
2006 © Cresswell, S.
Pogonognathellus elongatus from the USA
Body scales removed
2008.08.12 © Trently, D.
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3A
 
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3B
Posterior dorsal macrochaetotaxy
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3B
Specimen with projected mesotergum
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.3D
Specimen with projected mesotergum
Pogonognathellus elongatus from the USA
After Felderhoff KL et al., 2010 Fig.5C
Descaled; cuticular pattern
Pogonognathellus elongatus from the USA
2012.07.01 © Cresswell, S.
Pogonognathellus elongatus from the USA
2014.05.14 © Gallagher, J.
Pogonognathellus elongatus from the USA
2014.09.14 © Schrayer, G.
Pogonognathellus elongatus from the USA
2015.07.13 © Cicanese, M.
Pogonognathellus elongatus from the USA
Abd.3 with large apically rounded scales
2015.07.13 © Cicanese, M.
Pogonognathellus elongatus from the USA
With asymmetry in antennae
2015.08.07 © Cicanese, M.
Pogonognathellus elongatus ♂ from the USA
With raised genital papilla (*)
2016.04.30 © Schulz, K.
Pogonognathellus elongatus from the USA
Eyepatch with diamond of 4 anterior ocelli
2016.06.25 © Schulz, K.
Pogonognathellus elongatus from the USA
2016.08.05 © Eiseman, C.
Pogonognathellus elongatus from the USA
2016.09.03 © Schulz, K.
Pogonognathellus elongatus from the USA
2022.09.11 © Wilcox, T.

Species flavescens (Tullberg, 1871) Stach, 1929, B

            Syn.: Podura plumbea Linnæus, C, 1758:609
            Syn.: Podura plumbea Nicolet, H, 1842:69
            Syn.: Podura plumbea Templeton, R in Templeton, R et Westwood, JO, 1836:93
Podura plumbea from Ireland
After Templeton R, 1836 Pl.XI Fig.4,4.1

             Syn.: Podura Plumbea(sic) Bourlet, C, 1845:153 ISS
             Syn.: Podura Plumbea(sic) Bourlet, C, 1845:155 ISS
             Syn.: Podura Plumbea(sic) Bourlet, C, 1845:160 ISS
             Syn.: Macrotoma Plumbea(sic) Bourlet, C, 1845:160 ISS
             Syn.: Macrotoma Plumbea(sic) Bourlet, C, 1845:168 ISS
Macrotoma Plumbea(sic) from France
Scaled
After Bourlet C, 1845 Fig.1
Macrotoma Plumbea(sic) from France
Ant.1+2 distinctly longer then head
After Bourlet C, 1845 Fig.1
Macrotoma Plumbea(sic) from France
Descaled
After Bourlet C, 1845 Fig.2
Macrotoma Plumbea(sic) from France
Ventral aspect
Furca extended
After Bourlet C, 1845 Fig.3
Macrotoma Plumbea(sic) from France
Ventral aspect
Furca flexed
After Bourlet C, 1845 Fig.4

            Syn.: Tomocerus plumbeus Willem, V, 1900:44,B
Tomocerus plumbeus ♂ from Belgium
Abd.5 elongated
Modified after Willem V, 1900 Planche IX Fig.1
Tomocerus plumbeus ♀ from Belgium
Anal valves with elongated "cerci" (*)
Modified after Willem V, 1900 Planche IX Fig.2

            Syn.: ... plumbea Ågren, 1903, nec Templeton, 1835
             Syn.: Pogonognathus plumbaeus(sic) Bonet, F, 1931:99 ISS
            Syn.: Podura violacea Geoffroy
            Syn.: Podura longicornis Müller OF, 1776
            Syn.: Podura gigas Fabricius O, 1783
            Syn.: Podura communis Fourcroy, 1785
            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
            Syn.: Macrotoma nigra Bourlet, C, 1839:390
            Syn.: Tomocerus celer Nicolet, H, 1842:69
Tomocerus celer from Switzerland
After Nicolet H, 1842 Pl.7 Fig.9
Macrotome agile from Europe
Macrotoma celer
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.7

            Syn.: Tomocerus celer Bourlet, C, 1845:68
            Syn.: Tomocerus celer Bourlet, C in Erichson, WP, 1846:304
            Syn.: Tomocerus celer Nicolet, H, 1847:381
            Syn.: Tomocerus celer Folsom, JW, 1902:97
            Syn.: Macrotoma lepida Gervais in Walckenaër, 1844
             Syn.: Tomocerus lepida(sic) Folsom, JW, 1902:97 ISS
             Syn.: Macrotoma Spiricornis(sic) Bourlet, C, 1845:168 ISS
Macrotoma Spiricornis(sic) from France
With antennae curled in spiral
After Bourlet C, 1845 Fig.21

            Syn.: Tomocerus lubbocki Schäffer, 1900
            Syn.: Pogonognathus beckeri Börner, C, 1909:134?
            Syn.: Pogonognathus beckeri Salmon, JT, 1964:300
            Syn.: Architomocerura crassicauda Denis, 1931 tt
Architomocerura crassicauda from Britain
After Bagnall RS, 1940 Fig.1
=P. flavescens first instar

            Syn.: Paratomocerus pierantonii Tarsia in Curia, 1938,tt




Pogonognathellus flavescens from Belgium
2004 © Vuijlsteke, M.

             Syn.: Tomocerus Flavencens(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:365 ISS
             Syn.: Tomocerus flavencens(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:366 ISS
Pogonognathellus flavescens from the UK
2006.02.17 © Valentine., B
Pogonognathellus flavescens from Belgium
2006.03.19 © Vuijlsteke, M.
Pogonognathellus flavescens from Belgium
Black basal antennal rings
2006.10.25 © Janssens, F.
Pogonognathellus flavescens from the UK
2007.02.13 © Jones, D.
Pogonognathellus flavescens from the UK
2007.02.18 © Barton, T.
Pogonognathellus flavescens from Czechia
2007.06.23 © Krásenský, P.
Pogonognathellus flavescens from Czechia
2007.07.22 © Krásenský, P.
Pogonognathellus flavescens from the Netherlands
2007.11.05 © Baas, A.H.
Pogonognathellus flavescens from the UK
2007.11.14 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.18 © Campbell, A.
 
Pogonognathellus flavescens from the UK
2007.11.25 © Warner, P.
Pogonognathellus flavescens from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus flavescens from the Netherlands
2008.02.10 © Baas, A.H.
Pogonognathellus flavescens from the UK
2008.02.10 © Campbell, A.
Pogonognathellus flavescens from France
2008.02.11 © Lavoué., B
Pogonognathellus flavescens from the UK
Juvenile, descaled
2008.03.31 © Williams., B
Pogonognathellus flavescens from the UK
2008.05.01 © Farrell, S.
Pogonognathellus flavescens from Belgium
Scaled 3rd antennal segment
Legit Leruth, R, 1933.09.30
2008.05.28 © Janssens, F.
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
 
Pogonognathellus flavescens juv.
from the Netherlands
2008.10.15 © Baas, A.H.
Pogonognathellus flavescens
from the Netherlands
2008.11.03 © Baas, A.H.
Pogonognathellus flavescens from France
2008.11.08 © Jacquemin, G.
 
Pogonognathellus flavescens from the UK
2009.01.31 © Ransom, T.
Pogonognathellus flavescens from Belgium
2009.02.21 © Vuijlsteke, M.
Pogonognathellus flavescens (bar=2µm)
Hexagonal epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.2A'
Pogonognathellus flavescens from the UK
2012.09.12 © Jones, A.
Pogonognathellus flavescens from Belgium
2013.04.06 © Declercq., B
Pogonognathellus flavescens from Holland
Descaled specimen revealing intrinsic yellow body colour
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Moulting, final phase: redrawing furca from old cuticula
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Just moulted, dorsal aspect
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Just moulted, lateral aspect: thorax bulged
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from Holland
Moulting complete
2013.10.26 © Baas, A.H.
Pogonognathellus flavescens from the UK
Descaled specimen with partly recovered damaged left antenna
2013.12.24 © Evans, M.
Pogonognathellus flavescens from Germany
Dark clypeus and antennal bases
2015.02.22 © Kluck, A.
Pogonognathellus flavescens ♀ from the UK
Long spikes apically on anal valves (*)
2015.04.02 © Thomas., B
Pogonognathellus flavescens juv. from the UK
Abd.5 with long dorsal macrosetae
2015.05.05 © Henton-Jones, D.
Pogonognathellus flavescens from the UK
Antenna with black basal ring
2015.05.05 © Henton-Jones, D.
Pogonognathellus flavescens from Holland
Eyepatch
2015.10.24 © van Duinen, J.
Pogonognathellus flavescens ♀ from Holland
Abd.6 with dorsal short blunt macrosetae (*)
2015.11.08 © van Duinen, J.
Pogonognathellus flavescens ♀ from the UK
Curled antennae: ready to jump
2015.11.25 © David W.
Pogonognathellus flavescens ♀ from the UK
Long spike apically on anal valve (*)
2015.11.25 © David W.
Pogonognathellus flavescens from the UK
Black ring around antennal base
2015.12.08 © Thomas, Br.
Pogonognathellus flavescens juv. from Belgium
Black ring around antennal base
2016.03.28 © Huskens, M.-L.
Pogonognathellus flavescens juv. from Belgium
Abd.6 with telescopic anal valves (*)
2018.01.22 © Huskens, M.-L.
Pogonognathellus flavescens from the UK
Curled up antennae indicate specimen is preparing for a jump
2018.11.04 © Featherstone, A.W.
Pogonognathellus flavescens from Germany
Preparing for an escape jump
2019.10.17 © Fischer, A.
Pogonognathellus flavescens
Descaled
After Yu D-Y & al, 2020 Fig.5
Pogonognathellus flavescens juv. from Belgium
2020.01.05 © Huskens, M.-L.
Pogonognathellus flavescens juv. from Belgium
Footcomplex with distinctly triangular clavate tenent macroseta (*)
2020.12.03 © Huskens, M.-L.
Pogonognathellus flavescens juv. from Belgium
Dark antennal basal ring
Anterior 4 ocelli form a diamond
2020.12.03 © Huskens, M.-L.
Pogonognathellus flavescens from Belgium
Left antenna partly recovered
After Vanthournout B & al, 2021 Fig.p.7

             Syn.: Pogonognathelus(sic) flavensis(sic) Bellini, BC, Zhang, F, Souza, PGC, dos Santos-Costa, RC, Medeiros, GdS et Godeiro, NN, 2022:3 ISS
             Syn.: Pogonognathelus(sic) flavensis(sic) Bellini, BC, Zhang, F, Souza, PGC, dos Santos-Costa, RC, Medeiros, GdS et Godeiro, NN, 2022:5 ISS
Pogonognathellus flavescens from Belgium
Anterior 4 ocelli ABCD form diamond
2023.02.02 © Huskens, M.-L.
Pogonognathellus flavescens from the UK
Using antennae to sense environment
2023.10.02 © B.S.C.G.
Pogonognathellus flavescens from Germany
Dark purple partly pigmented body scale
Dorsal at midsegment
2024.06.04 © Vogl, U.
Pogonognathellus flavescens from Germany
Translucent unpigmented body scale
Dorsal at posterior segment margin
2024.06.04 © Vogl, U.
Pogonognathellus flavescens from Germany
Body scale with longitudinal ribs
forming comb at apical scale margin
2024.06.04 © Vogl, U.
Pogonognathellus flavescens from Germany
Body scale with longitudinal ribs
interconnected by transvers structures
2024.06.04 © Vogl, U.
Pogonognathellus flavescens from Germany
Epicuticular ultrastructure
forming hexagonal pattern
2024.06.04 © Vogl, U.
Pogonognathellus flavescens from Germany
Scale body matrix formed by longitudinal surface ribs and transverse internal structures
2024.06.10 © Vogl, U.
Pogonognathellus flavescens from Germany
Scales with structural colours
Each colour matches another scale thickness
2024.06.10 © Vogl, U.



Species flavescens2 Lee, B-H et Park, K-H, 1992:102 nec Tullberg, 1871

Species heterochrosatus Wang, TL, Yu, DY et Zhang, F, 2013:1244

            Syn.: Pogonognathellus heterochros(sic) Wang, TL, Yu, DY et Zhang, F, 2013:1244 IOS
Pogonognathellus heterochros(sic) from China
After Wang TL, Yu YD & Zhang F, 2013 Fig.1A

             Syn.: Pogonognathellus heterochros(sic) Yu, D-Y, Deharveng, L et Zhang, F, 2014:559 ISS
             Syn.: Pogonognathellus heterochros(sic) Yu, D-Y, Zhang, F, Stevens, MI, Yan, Q-B, Liu, M-Q et Hu, F, 2016:288 ISS




Species litoralis Kuprin, AV et Potapov, MB, 2019:2

Pogonognathellus litoralis ♀ from Far East Russia
After Kuprin AV & Potapov M, 2019 Fig.1

Species longicornis (Müller OF, 1776) Paclt, 1944, t.t., B

            Syn.: Podura plumbea Linnæus, 1761
            Syn.: Podura plumbea von Linné, C, 1767:1013
            Syn.: Podura plumbea Villers, 1789
            Syn.: Podura plumbea Templeton, 1835
            Syn.: Tomocerus plumbeus Nicolet, H, 1842:68
Tomocerus plumbeus from Switzerland
After Nicolet H, 1842 Pl.7 Fig.8

             Syn.: Tomocerus plombeus(sic) Nicolet, H, 1847:380 ISS
Tomocerus plombeus(sic) from Europe
After Nicolet, H, 1847 Pl.6 Fig.12

             Syn.: Tomocerus plombeus(sic) Frauenfeld, 1854 ISS
             Syn.: Tomocerus plumbea(sic) Lubbock, J, 1873:138 ISS
            Syn.: Pogonognathus plumbeus Boldori, 1938
            Syn.: Podura bicolor Say, 1821
             Syn.: Podura Bicolor(sic) Bourlet, C, 1845:158 ISS
            Syn.: Podura iricolor Say, 1821
             Syn.: Podura Iricolor(sic) Bourlet, C, 1845:158 ISS
            Syn.: Macrotoma ferruginosa Bourlet, C, 1839:390
             Syn.: Macrotoma ferruginea(sic) Elditt, HL, 1854:37 ISS
            Syn.: Desoria (Première division) tigrina Nicolet, H, 1842:59
Desoria tigrina from Switzerland
After Nicolet H, 1842 Pl.5 Fig.11
= juvenile Pogonognathellus longicornis
Desoria tigrina from Switzerland
Modified after Nicolet H, 1842 Pl.5 Fig.11
Compared with body of juvenile Tomocerinae
Top: Tomocerinae; bottom: Desoria tigrina Nicolet
Desoria tigrina from Switzerland
Modified after Nicolet H, 1842 Pl.5 Fig.11
Comparative segmental body division
Top: adult Tomocerus; bottom: adult Desoria
Desoria tigrina from Switzerland
Modified after Nicolet H, 1842 Pl.5 Fig.11
Comparative segmental body division
Top: juvenile Tomocerinae; bottom: adult Desoria


            Syn.: Macrotoma spiricornis Bourlet, 1843
             Syn.: Macrotoma Spiricornis(sic) Bourlet, C, 1845:67 ISS
             Syn.: Macrotoma Spiricornis(sic) Bourlet, C, 1845:101 ISS
            Syn.: Macrotoma rufescens Tullberg, 1871
            Syn.: Architomocerura litsteriana Bagnall, RS, 1940:173



             Syn.: Podura Longicornis(sic) Bourlet, C, 1845:153 ISS
Tomocerus longicornis from the UK
After Beck J in Lubbock J, 1873 Pl.67
Tomocerus longicornis from the UK
After Lubbock J, 1873 Pl.17


             Syn.: Tomocerus longicornes(sic) Noble-Nesbitt, J, 1963:253 ISS
             Syn.: Pogonognathellus logicornis(sic) Yosii, R, 1967:9 ISS
Pogonognathellus longicornis from the UK
2002 © Hopkin, S.P.
Pogonognathellus longicornis
2003 © Pettersson., B
Pogonognathellus longicornis from the UK
Large leaf-like basal dental scales
2004 © Hopkin, S.P.
Pogonognathellus longicornis from Sweden
2005 © Hall, K.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from the UK
2006 © Cornwall, N.J.
Pogonognathellus longicornis from Russia
2006 © Macroclub.ru.
Pogonognathellus longicornis from Germany
2006.09.30 © Kohl, F.
Pogonognathellus longicornis from Belgium
Black interantennal spot
2006.10.25 © Janssens, F.
Pogonognathellus longicornis from Germany
2007 © Peters, J.
Pogonognathellus longicornis from Belgium
2007.02.07 © Lindsley, J.
Pogonognathellus longicornis from France
2007.06.09 © Lebeaux, P.
Pogonognathellus longicornis from Belgium
2007.09.08 © Oger, P.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.10.13 © Brian.
Pogonognathellus longicornis from the UK
2007.11.10 © Campbell, A.
Pogonognathellus longicornis from Belgium
2007.11.24 © Carlet, G.
Pogonognathellus longicornis from Sweden
2007.12.04 © Hallmén, J.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the Netherlands
2007.12.08 © Heemskerk, R.
Pogonognathellus longicornis from the UK
2008.04.03 © Spears, G.
Pogonognathellus longicornis from the UK
2008.07.27 © Michaelson, J.
Pogonognathellus longicornis from the UK
2008.09.01 © Talbot, M.
Pogonognathellus longicornis from the UK
2009 © Chaplin, M.
Cleaning the antenna with maxillae
Pogonognathellus longicornis from the UK
2009 © Chaplin, M.
Everting vesicles of collophore
Pogonognathellus longicornis from the UK
2009.02.17 © Ransom, T.
Pogonognathellus longicornis from the UK
Unguis (U) with lateral teeth (LT)
Unguicular filament (uf) longer than unguis
2009.04.03 © Robertson, A.
Pogonognathellus longicornis from Sweden
5th and 6th abdominal segments
2011.11.25 © Hallmén, J.
Pogonognathellus longicornis from Sweden
2011.11.25 © Hallmén, J.
Pogonognathellus longicornis from the UK
Spirally curled antennae, ready to jump
2012.08.09 © Lindsay., B
Pogonognathellus longicornis from Germany
Eyepatch with 6 ocelli
2012.09.12 © Schmidt, R.
Pogonognathellus longicornis from the UK
Abdominal dorsal bothriotricha
2012.11.06 © Nurcombe, E.
Pogonognathellus longicornis from France
2014.07.14 © Quintin, C.
Pogonognathellus longicornis from the UK
Unguiculus with apical filament as long as unguis (*)
2015.11.30 © Boardman, P.
Pogonognathellus longicornis from the UK
Eyepatch with diamond of 4 +2 ocelli
2016.02.27 © David W.
Pogonognathellus longicornis from the UK
Antenna curled-up spirally
2017.08.28 © Henton-Jones, D.
Pogonognathellus longicornis from the UK
Descaled revealing darker 4 anterior segments
2019.02.16 © Jeffery, J.
Pogonognathellus longicornis from the UK
Footcomplex
Unguiculus longer then unguis
2019.06.09 © Cann, A.J.
Pogonognathellus longicornis from the UK
2021.02.02 © Atkins, W.
Pogonognathellus longicornis from the UK
With frontal macrosetae
With collar of macrosetae
2021.02.02 © Atkins, W.
Pogonognathellus longicornis from the UK
Antennae longer than body
2021.03.02 © Atkins, W.
Pogonognathellus longicornis from the UK
2021.03.02 © Atkins, W.
Pogonognathellus longicornis from the UK
Ant.1+2 longer than head
2021.03.02 © Atkins, W.
Pogonognathellus longicornis from the UK
Scales of mesothorax arranged in longitudinal lines
2021.03.02 © Atkins, W.
Pogonognathellus longicornis from the UK
Eye with 6 visible ocelli
Anterior 4 ocelli form a diamond
2021.03.02 © Atkins, W.



Species mai Wang, TL, Yu, DY et Zhang, F, 2013:1249

Pogonognathellus mai from China
After Wang TL, Yu YD & Zhang F, 2013 Fig.1B

Species magnibrunneus Park, K-H, Bernard, EC et Moulton, JK, 2011:9

Pogonognathellus magnibrunneus from the USA
After Park K-H & al., 2011 Fig.5A

Species mystax Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:488

Pogonognathellus n. sp. from the USA
2009.09.01 © Hodge, K.
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2B
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2C
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.2D
Patterned clypeus
Pogonognathellus mystax from the USA
After Felderhoff KL et al., 2010 Fig.5E
Descaled; cuticular pattern
Pogonognathellus mystax from the USA
2016.11.03 © Dobbs, Br.
Pogonognathellus mystax from the USA
2016.11.03 © Dobbs, Br.
Pogonognathellus mystax from the USA
Begin of moulting
2017.11.10 © Dobbs, Br.
Pogonognathellus mystax from the USA
New scale cover during moult
2017.11.10 © Dobbs, Br.
Pogonognathellus mystax from the USA
Moulted
2017.11.10 © Dobbs, Br.
Pogonognathellus ?mystax? from Canada
2019.09.15 © Barbin, T.

Species nigritus (Maynard, EA, 1951:130) Felderhoff, KL, Bernard, EC & Moulton, JK, 2010:484

            Syn.: Tomocerus (Pogonognathellus) elongatus Christiansen, K, 1964:672 partim


Pogonognathellus nigritus from the USA
2009.12.05 © Guzman, D.
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.4A
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.4C
Dorsal macrochaetotaxy
Pogonognathellus nigritus from the USA
After Felderhoff KL et al., 2010 Fig.5G
Descaled; cuticular pattern
Pogonognathellus nigritus from the USA
2011.10.09 © Bradford, A.
Pogonognathellus nigritus from the USA
2011.10.09 © Bradford, A.

Pogonognathellus nigritus from the USA
2013.07.04 © Liberta, A.

             Syn.: Pogonognathellus nigirtus(sic) Soto-Adames, FN et Taylor, SJ, 2013:158 ISS
Pogonognathellus nigritus from the USA
Ant.3 basally pale, distally darker
2014.10.02 © Rosenfeld, J.
Pogonognathellus nigritus from the USA
2015.06.22 © Dankowicz, E.
Pogonognathellus nigritus from the USA
Ant.3 basally pale, distally darker
2015.06.22 © Dankowicz, E.
Pogonognathellus nigritus from the USA
2016.07.16 © Schulz, K.
Pogonognathellus nigritus from the USA
2019.07.24 © Keim, Bi.
Pogonognathellus nigritus from the USA
2021.12.25 © Childs, K.
Pogonognathellus nigritus from the USA
2021.12.25 © Childs, K.
Pogonognathellus nigritus from the USA
Antenna
2021.12.25 © Childs, K.
Pogonognathellus nigritus from the USA
2022.05.25 © Hendrix, S.



Species palespeciesgroup Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:487

Remark by Felderhoff, K.L. et al. 2010:488: We prefer the term "pale species group" to denote this group for convenience only, rather than "flavescens complex," to separate these North American taxa from the European species.
            Syn.: Tomocerus plumbea(sic) Packard, 1871 IOS
            Syn.: Tomocerus alba(sic) Packard, 1877 IOS
            Syn.: Tomocerus pallidus Packard, 1888
            Syn.: Tomocerus arcticus Schött, H, 1893:43
            Syn.: Pogonognathellus arcticus Edwards, 1972
            Syn.: Pogonognathellus flavescens ssp. arcticus Danks, 1981
            Syn.: Tomocerus americanus Schött, 1896
Tomocerus flavescens var. americanus from the USA
After Folsom JW, 1913 Fig.2
Tomocerus flavescens var. americanus ♂ from the USA
Genital segment not extended
Modified after Folsom JW, 1913 Fig.6
Tomocerus flavescens var. americanus ♂ from the USA
Genital segment extended
Modified after Folsom JW, 1913 Fig.7
Tomocerus flavescens var. americanus ♀ from the USA
Anal segment with long cerci
After Folsom JW, 1913 Fig.8
Remark by Folsom, J.W. 1913:455: "Fig.7.-Tomocerus flavescens, var. americanus. Posterior segments of abdomen of male, showing protruded genital segment. f. fold. X 30." [In error long cerci of female are shown]
Remark by Folsom, J.W. 1913:455: "Fig.8.-Tomocerus flavescens, var. americanus. Anal segment of male, showing cerci. X 102." [In error "of male"; should be "of female", given long cerci are shown]

            Syn.: Pogonognathellus flavescens var. americanus Hammer, 1953
            Syn.: Pogonognathellus flavescens var. americanus Mills & Richards, 1953
            Syn.: Tomocerus niger Folsom, JW, 1902:97
            Syn.: Macrotoma nigra Folsom, JW, 1904:95
            Syn.: Tomocerus celer Folsom, JW, 1904:95
            Syn.: Macrotoma celer Folsom, JW, 1904:95
            Syn.: Macrotoma ferruginosa Folsom, JW, 1904:95
            Syn.: Macrotoma flavescens Folsom, JW, 1904:95
            Syn.: Tomocerus flavescens Folsom, JW, 1904:95
            Syn.: Macrotoma lepida Folsom, JW, 1904:95
             Syn.: Tomocerus lepida(sic) Folsom, JW, 1904:95 ISS
            Syn.: Tomocerus flavescens var. separatus Folsom, JW, 1913:460
            Syn.: Pogonognathellus flavescens ssp. separatus Stach, 1966
            Syn.: Tomocerus flavescens Bird, 1930
            Syn.: Tomocerus flavescens James, 1933
            Syn.: Tomocerus flavescens Brown, 1934
            Syn.: Tomocerus flavescens Mills, 1934
            Syn.: Tomocerus flavescens Spencer, 1948
            Syn.: Tomocerus flavescens Weber, 1950
            Syn.: Tomocerus flavescens Maynard, 1951
            Syn.: Tomocerus flavescens Judd, 1961
            Syn.: Tomocerus (Pogonognathellus) flavescens Christiansen, K, 1964:673
Remark by Christiansen, K. 1964:675: It is apparent that the species here classified as T. flavescens represents a vast complex of forms.

            Syn.: Pogonognathellus flavescens Marshall, 1964
            Syn.: Pogonognathellus flavescens Judd, 1965
            Syn.: Pogonognathellus flavescens Martin, 1965
            Syn.: Pogonognathellus flavescens Marshall, 1967
            Syn.: Pogonognathellus flavescens Powell, 1971
            Syn.: Pogonognathellus flavescens Judd, 1972
            Syn.: Pogonognathellus flavescens Powell & Skaley, 1975
            Syn.: Pogonognathellus flavescens Aitchison, 1979
            Syn.: Pogonognathellus flavescens Christiansen & Bellinger, 1980
            Syn.: Pogonognathellus flavescens Aitchison, 1984
            Syn.: Pogonognathellus flavescens Bird & Chatarpaul, 1986
            Syn.: Pogonognathellus flavescens Peck, 1988
            Syn.: Pogonognathellus flavescens Battigelli & Marshall, 1993
            Syn.: Pogonognathellus flavescens Setälä & Marshall, 1994
            Syn.: Pogonognathellus flavescens Arulnayagam, 1995
            Syn.: Pogonognathellus flavescens Rusek & Marshall, 1995
            Syn.: Pogonognathellus flavescens Setälä & al, 1995
            Syn.: Pogonognathellus flavescens Skidmore, 1995
            Syn.: Pogonognathellus flavescens Cannings & Cannings, 1997
            Syn.: Pogonognathellus flavescens Puvanendran & al, 1997
            Syn.: Pogonognathellus flavescens Christiansen & Bellinger, 1998
            Syn.: Pogonognathellus flavescens Therrien & al, 1999
            Syn.: Pogonognathellus flavescens Chagnon & al, 2000
            Syn.: Pogonognathellus flavescens Chagnon & al, 2001
            Syn.: Pogonognathellus flavescens Addison & al, 2003
            Syn.: Pogonognathellus flavescens Babenko & Fjellberg, 2006
            Syn.: Pogonognathellus flavescens Cannings, 2010
            Syn.: Pogonognathellus flavescens Park & al, 2011
            Syn.: Pogonognathellus flavescens Huebner & al, 2012
            Syn.: Pogonognathellus flavescens Lindo, 2014
            Syn.: Pogonognathellus flavescens Turnbull, 2014
            Syn.: Tomocerus flavescens Glime, JM, 2015:3
            Syn.: Pogonognathellus flavescens Babenko, A, Stebaeva, S et Turnbull, MS, 2019:78
            Syn.: Architomocerura crassicauda Mills, HB, 1932:265
Architomocerura crassicauda from the USA
After Mills HB, 1932 PlateI Fig.11
=P. palespeciesgroup second instar
Architomocerura crassicauda from the USA
Left eye
After Mills HB, 1932 PlateI Fig.9

            Syn.: Pogonognathellus sp.2 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.3 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.5 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.6 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.7 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475
            Syn.: Pogonognathellus sp.8 Felderhoff, KL, Bernard, EC et Moulton, JK, 2010:475



Pogonognathellus "flavescens" from the USA
2006 © Murray, T.
Pogonognathellus "flavescens" from the USA
2009 © Bernard, E.C.
Pogonognathellus "flavescens" from the USA
2010.05.24 © Taft, M.
2+4 frontal setae



Species taeniatus (Koch, CL & Berendt, GC in Berendt, GC, 1854:119, -) -

            Syn.: Podura taenita(sic) Koch, CL et Berendt, GC in Berendt, GC, 1854:119,- IOS
Podura taeniata from Baltic amber
After Koch CL & Berendt GC, 1854 Tab.XV Fig.126

            Syn.: Catastylus calcaratus von Olfers, 1907 -
            Syn.: Orchesella concolor von Olfers, 1907 -
            Syn.: Palpiger cucullatus von Olfers, 1907 -
            Syn.: Lepidocyrtus curvicollis von Olfers, 1907 -
            Syn.: Lepidocyrtus gibbulus von Olfers, 1907 -
            Syn.: Catastylus nasicornis von Olfers, 1907 -
            Syn.: Tomocerus plumbeus von Olfers, 1907 -






Genus Tomolonus Mills, HB, 1949:353

Species reductus Mills, HB, 1949:353, t.t.

Tomolonus reductus from the USA
Reduced eyepatch with post-antennal organ (PAO)
After Mills HB, 1949 Fig.2
Tomolonus reductus from the USA
Eyepatch with 2 ocelli; post-antennal organ (PAO) with 4 lobes
After Mills HB, 1949 Fig.1
Tomolonus reductus from the USA
Reduced eyepatch with post-antennal organ (PAO)
After Yosii R, 1967 Fig.27I,G
Tomolonus reductus from the USA
2023.12.23 © Wang, G.


Genus Monodontocerus Yosii, R, 1955:394 key

Monodontocerus modificatus from Tokara Island
Basal outer tooth of mucro absent
After Yosii R, 1955 Abb.8E
  Syn.: Tomoceridae Mondontocerus(sic) Yosii, R, 1977:157 ISS
Monodontocerus sp. from Taiwan
2022.06.18 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Ocelli ABCD form a diamond?
2022.06.18 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
2022.06.18 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Dental spines array
2022.06.19 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Dental spines tridentate
2022.06.19 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Mucro (apical tooth broken)
2022.06.19 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Mucro with single basal tooth
2022.06.19 © Cheng, H.-J.
Ident.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Ant.3 subcylindrical
2022.06.20 © Cheng, H.-J.
Det.: Yu, D.-Y.
Monodontocerus sp. from Taiwan
Hind foot
2022.06.20 © Cheng, H.-J.
Det.: Yu, D.-Y.



Species absens Yu, D-Y, Deharveng, L et Zhang, F, 2014:559

Monodontocerus absens from China
Pigmented sockets of macrosetae (arrows)
After Yu-D-Y, Deharveng L & Zhang F, 2014 Fig.1

Species cinereus Yu, 2016

Species leqingensis Sun & Liang, 2009

Monodontocerus leqingensis
Descaled
After Yu D-Y & al, 2020 Fig.5

Species modificatus Yosii, R, 1955:394, t.t.

            Syn.: Monodontocerus modificatus ssp. satsumensis Yosii, R, 1956:89
            Syn.: Tomocerus (Monodontocerus) modificatus ssp. satsumensis Yosii, R, 1967:14
            Syn.: Tomocerus (Monodontocerus) modificatus ab. satsumensis Yosii, R, 1967:15
            Syn.: Mondontocerus(sic) modificatus ssp. satsumensis Yosii, R, 1977:157


             Syn.: Tomocerus (Monodontocerus) mocificatus(sic) Yosii, R, 1967:15 ISS

Species mulunensis Yu, D-Y, Deharveng, L et Zhang, F, 2014:565

Species odongnyeoensis (Park, K-H & Lee, B-H, 1995:437) Yu, D-Y, Deharveng, L & Zhang, F, 2014:557

             Syn.: Tomocerus odongyeoensis(sic) Park, K-H et Lee, B-H, 1995:436 ISS

Species trigrandis Yu, D-Y, Deharveng, L et Zhang, F, 2014:568


Genus Aphaenomurus Yosii, R, 1956:86

Species interpositus Yosii, 1954, t.t.

            Syn.: Tomocerus (Aphaenomurus) vicinus Yosii, R, 1966:556
            Syn.: Aphaenomurus vicinus Yosii, R, 1967:34
            Syn.: Aphaenomurus vicinus Yosii, R, 1977:158


             Syn.: Aphaenomurus intespositus(sic) Yosii, R, 1967:36 ISS
  Subspecies denticulatus Yosii, R, 1956:87

            Syn.: Aphaenomurus interpositus ssp. dentiferus(sic) Yosii, R, 1967:35 lapsus



Genus Plutomurus Yosii, R, 1956:78 key  key  key  key  key  key

  Syn.: Tomoceridae Tritomurus Salmon, JT, 1964:121


  Syn.: Tomoceridae Plutomulus(sic) Yosii, R, 1967:47 ISS
  Syn.: Tomoceridae Plutomorus(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:154 sp ISS
  Syn.: Tomoceridae Plutomorus(sic) Palacios-Vargas, JG, Castaño-Meneses, G et Valladares, L, 2021:155 sp ISS
Plutomurus sp. from the Causasus
After Barjadze S & al, 2022 cover
Plutomurus sp. from the USA
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
Antenna
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
Antennae 'folded'
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
Hindfoot with acuminate tenent macroseta
Unguis with 5 inner teeth
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
1st foot with serrate inner unguicular edge
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
Bothriotrichum socket semi-surrounded by 7 microsetae
2023.10.28 © Wang, G.
Plutomurus sp. from the USA
2023.12.26 © Temmi, N.



Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20

Species sp.2 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:894

Species abchasicus Martynova, 1969

             Syn.: Plutomurus abchasic.(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:30 ISS

Species baschkiricus (Skorikov, 1900) Yosii, 1956 tt

Species belozerovi Martynova, 1979

Species birsteini Djanashvili, R et Barjadze, S, 2011:28

Species borealis Suma, Y, 1981:502

Species brevimucronatus (Denis, J-R, 1929:176) Yosii, R, 1967:47

Tomocerus brevimucronatus from the USA
After Denis J-R, 1929 Fig.VI.1
             Syn.: Tomocerus (Plutomurus) brevimucromatus(sic) Skidmore, RE, 1995:60 ISS


Species californicus (Folsom, JW, 1913:469) Yosii, R, 1967:47

            Syn.: Tritomurus oregonensis Denis, J-R, 1929:178
Tritomurus oregonensis from the USA
After Denis J-R, 1929 Fig.VII.1




             Syn.: Tomocerus cilifornicus(sic) Yosii, R, 1967:47 ISS

Species carpaticus Rusek, J et Weiner, WM, 1978:742

            Syn.: Tomocerus terrestralis Paclt, 1957, nec Stach, 1922

Species danialensis Barjadze, S, Mehrafrooz Mayvan, M, Parimuchová, A, Maghradze, E et Kovác, L, 2024:?

Species diversispinus (Yosii, R, 1966:555) Yosii, R, 1967:36

Species edaphicus Yosii, R, 1967:37

Species ehimensis Yosii, R, 1956:83

             Syn.: Plutomurus suzukaensis ssp. ehimesis(sic) Yosii, R, 1977:158 ISS

Species eristoi Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:87

            Syn.: Plutomurus sp.7 Barjadze & al, 2015


Species grahami (Christiansen, KA, 1980:121) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:78

Species gul (Yosii, R, 1966:552) Yosii, R, 1967:36

Plutomurus gul
Descaled
After Yu D-Y & al, 2020 Fig.5
Remark by Janssens, F. 2023.10.27: "Gul" is a vernacular Korean word that means "cave". The ICZN Rec.11A discourages the use of vernacular words as scientific names. Given appropriate latinisation is the prefered means of formation of names from vernacular words, "cavus" (= pertaining to the cave) would be an example of an appropriate name in line with Rec.11A.

Species iwatensis Yoshii, 1991

Species jangamensis Chang & Park, 2020

Species jeleznovodskii Kniss & Thibaud, 1999

            Syn.: Plutomurus jeleznovodski(sic) Kniss & Thibaud, 1999 IOS
             Syn.: Plutomurus jeleznovodski(sic) Djanashvili, R et Barjadze, S, 2011:28 ISS
             Syn.: Plutomurus jeleznovozski(sic) Djanashvili, R et Barjadze, S, 2011:29 ISS
             Syn.: Plutomurus jeleznovodski(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:78 ISS
             Syn.: Plutomurus jeleznovodski(sic) Barjadze, S et Djanashvili, R, 2014:1 ISS
             Syn.: Plutomurus jeleznovodski(sic) Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:78 ISS



Species jordanai Barjadze, S, Giordana, R et Soto-Adames, F, 2020:?

Species kawasawai Yosii, R, 1956:85

  Subspecies kyushuensis Yosii, R, 1956:85

Species kelasuricus Martynova, 1969

Species kharagauliensis Barjadze, S, Parimuchova, A, Raschmanova, N, Maghradze, E et Kovác, L, 2022:?

Species leei (Yosii, R, 1966:554) Yosii, R, 1967:36

Species marmorarius Yosii, R, 1967:45

Species ortobalaganensis Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:77

Plutomurus ortobalaganensis from Abkazia
The deepest terrestrial animal living about 2 km below surface
2012 © Jordana R & Baquero E.

             Syn.: Plutomurus ortobelaganensis(sic) Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:92 ISS
             Syn.: Plutomurus orthobal.(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:30 ISS
             Syn.: Plutomurus orthobalaganesis(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:39 ISS


Species pichkhaiai Barjadze S, Jordana R et Soto-Adames FN in Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:36

Plutomurus pichkhaiai from Georgia
After Barjadze S & al, 2018 Fig.23

            Syn.: Plutomurus sp.4 Barjadze & al, 2015
            Syn.: Plutomurus sp.1 Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:79



             Syn.: Plutomurus pichkaiai(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:38 ISS

Species revazi Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:81

Plutomurus revazi from Georgia
After Barjadze S & al, 2016 Fig.5

            Syn.: Plutomurus sp.2 Barjadze & al, 2015


Species riugadoensis (Yosii, 1939) Yosii, R, 1956:78, t.t.

            Syn.: Tritomurus ishikawai Uchida, 1940
            Syn.: Tritomurus ishikawai Yosii, R, 1977:159


             Syn.: Tritomurus ruigadoensis(sic) Mills, HB, 1949:358 ISS
             Syn.: Plutomurus ryugadoensis(sic) Yosii, R, 1967:4 ISS

Species ruseki Barjadze, S, Parimuchova, A, Raschmanova, N, Maghradze, E et Kovác, L, 2022:?

Species shurubumuensis Barjadze S, Jordana R et Soto-Adames FN in Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:32

            Syn.: Plutomurus sp.8 Barjadze & al, 2015
            Syn.: Plutomurus sp.2 Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:79


             Syn.: Plutomurus shurubu.(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:30 ISS
             Syn.: Plutomurus shurubumensis(sic) Barjadze, S, Jordana, R, Baquero, E, Giordano, R et Soto-Adames, FN, 2018:38 ISS

Species sorosii Kniss & Thibaud, 1999

            Syn.: Plutomurus sorosi(sic) Kniss & Thibaud, 1999 IOS
             Syn.: Plutomurus sorosi(sic) Djanashvili, R et Barjadze, S, 2011:28 ISS
             Syn.: Plutomurus sorosi(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:79 ISS
             Syn.: Plutomurus sorosi(sic) Barjadze, S et Djanashvili, R, 2014:4 ISS
             Syn.: Plutomurus sorosi(sic) Barjadze, S, Baquero, E, Soto-Adames, FN, Giordano, R et Jordana, R, 2016:78 ISS



Species suzukaensis (Yosii, 1939) Yosii, R, 1956:81

            Syn.: Plutomurus suzukaensis ssp. naikaiensis Yosii, R, 1956:83
            Syn.: Plutomurus suzukaensis ssp. naikaiensis Yosii, R, 1967:39
            Syn.: Plutomurus suzukaensis ssp. naikaiensis Yosii, R, 1977:159
            Syn.: Plutomurus suzukaensis ssp. ohminensis Yosii, R, 1956:82
            Syn.: Plutomurus suzukaensis ssp. ohminensis Yosii, R, 1967:39
            Syn.: Plutomurus suzukaensis ssp. ohminensis Yosii, R, 1977:159


Plutomurus suzukaensis from Japan
Head
After Yosii R, 1967 Fig.22A

Species unidentatus (Börner, 1901) Dobat, 1979, B

            Syn.: Pogonognathellus flavescens Dethier, M et Hubart, J-M, 2000:29,B


             Syn.: Plutomurus unidentaus(sic) Stomp, N et Weiner, WM, 1994:359 ISS

Species vigintiferispinus (Lee, B-H, 1974:410) Thibaud, 1976

            Syn.: Tomocerus (Plutomurus) vigintiferispina(sic) Lee, B-H, 1974:410 IOS
            Syn.: Tomocerus (Plutomurus) vigintiferispina(sic) Lee, B-H et Park, K-H, 1992:106 IOS
            Syn.: Tomocerus (Plutomurus) vigintiferispina(sic) Park, K-H et Lee, B-H, 1995:440 IOS
            Syn.: Plutomurus vigintiferispina(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:79 IOS
            Syn.: Plutomurus vigintiferispina(sic) Yu, D-Y, Zhang, F et Deharveng, L, 2014:69 IOS
            Syn.: Plutomurus vigintiferispina(sic) Potapov, M, 2020:il IOS
Remark by Janssens, F. 2024.05.24: The gender of the species name "vigentiferispina" does not match with the gender of the genus name "Tomocerus". In ICZN nomenclature, Latin adjectives take the declension -us, -a, -um and must agree in gender, number and case with the noun they modify. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). Given the stem "vigintiferispin-", "vigintiferispinus" is the adjective form agreeing with the gender of "Tomocerus" and in line with the ICZN code.


Species weinerae Barjadze, S, Mehrafrooz Mayvan, M, Parimuchová, A, Maghradze, E et Kovác, L, 2024:?

Species wilkeyi (Christiansen, K, 1964:659) Yosii, R, 1967:47

Plutomurus wilkeyi from the USA
Dental spines array
I-4-I-2-I-4/III-9
2023.10.14 © Wang, G.
Plutomurus wilkeyi from the USA
Mucro with 2 basal teeth
2023.10.14 © Wang, G.
Plutomurus wilkeyi from the USA
Mucro with 5 intermediate teeth
2023.10.14 © Wang, G.
Plutomurus wilkeyi from the USA
Mucro with small apical + large subapical tooth
2023.10.14 © Wang, G.
Plutomurus wilkeyi from the USA
First foot with 5 inner teeth
2023.10.14 © Wang, G.
Plutomurus wilkeyi from the USA
Bothriotrichal socket with semi-circle of 6 microsetae
2023.10.14 © Wang, G.

Species yamatensis Yosii, R, 1956:84

  Forma albida Yosii R, 1956:85


Genus Lethemurus Yosii, R, 1970:4

Species sp.1 Pomorski, RJ et Steinmann, D, 2004:20

Species finitimus Yosii, R, 1970:6

Species missus (Mills, HB, 1949:356) Yosii, 1970

             Syn.: Tomocerus misus(sic) Cutz-Pool, LQ, Palacios-Vargas, JG et Vázquez, M, 2003:97 ISS



Genus Entomocerus Christiansen, KA et Pike, E, 2002:170, -

Species mirus (Christiansen, KA & Pike, E, 2002:170, -) Christiansen, KA & Pike, E, 2002:170, -, t.t.


References