http://www.collembola.org/taxa/troginae.htm - Last updated on 2021.01.10 by Frans Janssens
Checklist of the Collembola: Troglopedetinae

Subfamilia Troglopedetinae Börner C, 1913:321

Genus Number
of species
Cyphoderopsis17
Troglopedetes33
Troglopedetes21
Trogolaphysa78
Troglopedetinae129
Genera with number of species
  Syn.: Cyphoderinae Troglopedetini Börner, C, 1913:321
  Syn.: Entomobryoidea Troglopedetidae Delamare Deboutteville, C, 1947:37
  Syn.: ... Troglopedetinae Lukic, M, Delic, T, Zagmajster, M et Deharveng, L, 2018:1159
  Syn.: Paronellidae Troglopedetinae Jantarit, S, Surakhamhaeng, K et Deharveng, L, 2020:5


Genus Troglopedetes Joseph, 1872 M

  Syn.: Paronellidae Cyphoderopsis Bonet, 1931
  Syn.: Paronellidae Cyphoderopsis Wilson, JM, 1982:211
  Syn.: Paronellidae Cypherodopsis(sic) Wilson, JM, 1982:214 ISS


  Syn.: Paronellidae Troglopedetes Wilson, JM, 1982:211 nec Joseph, 1872
Remark by Paclt, 1947, cited from Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Joseph, 1872 is a nomen nudum.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:57: The opinion of Paclt, 1947, that Troglopedetes Joseph is a nomen nudum cannot be upheld, however meagre the description may be.



Species sp.1 Deharveng, L, 1990:281

Species absoloni Bonet, F, 1931:363

            Syn.: Troglopedetes sp. Arbea & Baena, 2004
            Syn.: Troglopedetes sp. Pérez Fernández & Tinaut, 2005
            Syn.: Troglopedetes sp. Pérez Fernández & Pérez Ruiz, 2006


             Syn.: Troglopedetes Absoloni(sic) Absolon, K et Ksenemann, M, 1942:17 ISS
             Syn.: Troglopedetes Absoloni(sic) Delamare Deboutteville, C, 1945:46 ISS
Troglopedetes absoloni from Spain
After Arbea JI, 2013 Fig.15
Troglopedetes absoloni from Spain
Ant.4 subdivided in 2 parts
After Soto-Adames & al., 2014 Fig.6



Species albus Joseph, 1872, t.t.

            Syn.: Troglopedetes pallidus Paclt, 1946 =?
            Syn.: Troglopedetes pallidus Wilson, JM, 1982:211
            Syn.: Troglopedetes pallidus ssp. distinctus Wilson, JM, 1982:211


Species caeruleus (Carpenter, GH, 1916:43) Deharveng, L & Bedos, A, 1995:22

             Syn.: Paronella coerulea(sic) Handschin, 1929 ISS
             Syn.: Microparonella caeruleus(sic) Yoshii, R et Suhardjno, YR, 1992:48 ISS

Species calvus Deharveng, L et Gers, C, 1993:112

Species canis Christiansen, K, 1957:86

Species cavernicolus Delamare Deboutteville, 1944

             Syn.: Troglopedetes cavernicola(sic) Delamare Deboutteville, 1944 ISS
            Syn.: Troglopedetes wichmanni Delamare Deboutteville, 1950 nn
            Syn.: Troglopedetes wichmanni Wilson, JM, 1982:214 nn
            Syn.: Troglopedetes cretensis Wilson, JM, 1982:214 =?


             Syn.: Troglopedetes cavernicola(sic) Thibaud, J-M et Najt, J, 1988:720 ISS
             Syn.: Troglopedetes cavernicola(sic) da Gama & Ferreira, 2000 ISS
             Syn.: Troglopedetes cavernicola(sic) Wilson, JM, 1982:214 ISS

Species centralis Deharveng, L et Gers, C, 1993:109

Species churchillatus Wilson in Durrant, Smart, Turner & Wilson, 1979

Troglopedetes churchillatus from Nepal
After Wilson JM, 1982 Fig.1A

Species convergens Deharveng, L et Gers, C, 1993:111

Species cretensis Ellis, WN, 1976:306

            Syn.: Troglopedetes wichmanni Delamare Deboutteville, 1950 =?
            Syn.: Troglopedetes wichmanni Wilson, JM, 1982:214 =?


Species dispersus Deharveng, L et Gers, C, 1993:110

Species fredstonei Deharveng, L, 1988:95

Species ildumensis Soto-Adames, FN, Jordana, R et Baquero, E, 2014:8

Species jeanneli (Delamare Deboutteville, C, 1945:41) Thibaud, J-M & Najt, J, 1988:720

            Syn.: Troglopedetina Jeanneli(sic) Jeannel, R, 1945:15 IOS

Species kae Jantarit, S, Surakhamhaeng, K et Deharveng, L, 2020:7

Troglopedetes kae from Thailand
After Jantarit S, Surkahamhaeng K & Deharveng L, 2020 Fig.1 inset
Remark by Janssens, F. 2020.12.21: The name "Troglopedetes kae" is confusing and misleading, especially so when the specific name is wrongly given an upper case initial letter. In order to avoid the appearance that the specific name is a citation of the authorship of the generic name, the specific name should not be formed as a noun in apposition (ICZN Recommendation 31A). To avoid confusion with an author bearing the name "Kae", the new specific name which is based on a geographical name (a toponym) should preferably be formed with the latin -ensis suffix. Therefore "Troglopedetes kaeensis" would have been the recommended name.

Species leclerci Deharveng, L, 1990:279

Species longicornis Deharveng, L et Gers, C, 1993:109

Species machadoi Delamare Deboutteville, 1946

             Syn.: Troglopedetes rnachadoi(sic) da Gama, MM, 1988:85 ISS

Species maffrei Deharveng, L et Gers, C, 1993:108

Species maungonensis Deharveng, L et Gers, C, 1993:108

Species meridionalis Jantarit, S, Surakhamhaeng, K et Deharveng, L, 2020:20

Troglopedetes meridionalis from Thailand
After Jantarit S, Surkahamhaeng K & Deharveng L, 2020 Fig.11A

Species microps Deharveng, L et Gers, C, 1993:108

Species minor Hüther, 1983

Species multispinosus Deharveng, L et Gers, C, 1993:110

Species orientalis Cassagnau, P et Delamare Deboutteville, C, 1955:385

Species paucisetosus Deharveng, L et Gers, C, 1993:111

Species rasendrans Bhattacharjee, 1985

Species ruffoi Delamare Deboutteville, 1951

Species schalleri (Hüther, 1962) Thibaud, J-M & Najt, J, 1988:720

Species tridentatus (Salmon, 1954) Thibaud, J-M & Najt, J, 1988:720

             Syn.: Troglopedetes tridentata(sic) Thibaud, J-M et Najt, J, 1988:720 ISS

Species vandeli Cassagnau, P et Delamare Deboutteville, C, 1955:387

            Syn.: Troglopedetes Vandeli(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:387 IOS

Species vanharteni Barra, J-A, 2006:69


Genus Troglopedetes2

  Syn.: ... Troglopedetes2 Absolon, 1907 nec Joseph, 1872
  Syn.: Paronellini Microparonella Carpenter, GH, 1916:43 partim
  Syn.: Troglopedetinae Cyphoderopsis Bonet, F, 1931:362
  Syn.: Paronellidae Cyphoderopsis Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5
  Syn.: Paronellinae Trogolaphysa Bonet, F, 1942:58
  Syn.: Paronellidae Trogolaphysa Salmon, 1964
  Syn.: Paronellidae Trogolaphysa Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5
  Syn.: Troglopedetini Troglopedetina Delamare Deboutteville, C, 1945:40
  Syn.: Paronellidae Dicranocentruga Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:5


Remark by Paclt, 1947, cited from Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Absolon, 1907 might be identical with Troglopedetes Joseph, 1872.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:57: Troglopedetes Absolon, 1907 is a junior homonym of Troglopedetes Joseph, 1872.

  Syn.: Paronellidae Trolgopedetes(sic) Greenslade, P et Yoshii, R, 2000:139 ISS

Species pallidus Absolon, 1907 tt

             Syn.: Troglopedetes pallidus var. distincta(sic) Absolon, K et Ksenemann, M, 1942:6 ISS


Troglopedetes pallidus from Croatia
2010.11.19 © Lukic, M.
Troglopedetes pallidus from Croatia
2010.11.19 © Lukic, M.
Troglopedetes pallidus from Croatia
2019.03.dd © Lukic, M.


Genus Cyphoderopsis Carpenter, 1917

  Syn.: ... Trogonella Delamare Deboutteville, 1951
  Syn.: Paronellidae Trogonella Thibaud, J-M et Najt, J, 1988:721
  Syn.: Paronellinae Trogonella Soto-Adames, FN, Jordana, R et Baquero, E, 2014:1



Species-group to be defined

Species sp. Deharveng & Bedos, 2000

Species lindbergi (Stach, J, 1960:546) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetina Lindbergi(sic) Stach, J, 1960:546 IOS

Species-group ceylonica Yosii, 1985

Species ceylonica Yosii, 1966

             Syn.: Troglopedetes ceylonica(sic) Wilson, JM, 1982:214 ISS

Species gorumaraensis Mandal, GP, Suman, KK et Bhattacharya, KK, 2019:148

Cyphoderopsis gorumaraensis from India
After Mandal GP & al, 2019 Fig.5

Species nayakensis (Stach, J, 1960:543) Thibaud, J-M & Najt, J, 1988:721

Species nepalensis (Wilson, JM, 1982:221) Thibaud, J-M & Najt, J, 1988:721

Troglopedetes nepalensis from Nepal
After Wilson JM, 1982 Fig.5A

Species sexocellatus Yosii, 1966

            Syn.: Cyphoderopsis 6-ocellata(sic) Yosii, 1966 IOS
             Syn.: Troglopedetes sexoculata(sic) Wilson, JM, 1982:214 ISS
             Syn.: Cyphoderopsis (ceylonica-group) sexoculata(sic) Thibaud, J-M et Najt, J, 1988:721 ISS
             Syn.: Cyphoderopsis sixocellata(sic) Mandal, GP, 2018:124 ISS

Species-group kempi Yosii, 1985



Species cavicola Jantarit , Satasook & Deharveng, 2013

Species decemoculatus Prabhoo, 1971

             Syn.: Troglopedetes decemoculata(sic) Wilson, JM, 1982:214 ISS
             Syn.: Troglopedetes decemoculata(sic) Wilson, JM, 1982:219 ISS

Species gracilis Carpenter, 1924

Cyphoderopsis gracilis from India
After Mandal GP & Hazra AK, 2009 Fig.42a
             Syn.: Cyphoderopsis gracilils(sic) Mandal, GP et Hazra, AK, 2009:41 ISS


Species kempi Carpenter, 1917 tt

             Syn.: Cyphoderopsis Kempi(sic) Delamare Deboutteville, C, 1945:46 ISS
Cyphoderopsis kempi from India
After Mandal GP & Hazra AK, 2009 Fig.44a



Species khaophang Jantarit , Satasook & Deharveng, 2013

Species lamottei Delamare Deboutteville, 1950

Trogolaphysa lamottei from Guinea
After Nguyen M & Soto-Adames FN, 2018 Fig.5I

Species madagascarensis (Wilson, JM, 1982:222) Thibaud, J-M & Najt, J, 1988:721

Troglopedetes madagascarensis from Madagascar
After Wilson JM, 1982 Fig.6

Species pauliani (Delamare Deboutteville, 1951) Thibaud, J-M & Najt, J, 1988:721

Species phangnga Jantarit , Satasook & Deharveng, 2013

Species thachana Jantarit , Satasook & Deharveng, 2013


Genus Trogolaphysa Mills, HB in Pearse, AS, 1938:184 key  key

  Syn.: Paronellinae Dicranocentruga Wray, DL, 1953:141
  Syn.: ... Dicranocentrus(sic) Mari Mutt, JA, 1979:60 ISS
  Syn.: Paronellini Diranocentruga(sic) Mitra, SK, 1993:42 ISS
Dicranocentruga
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.31

  Syn.: Paronellini Dicraanocentruga(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellinae Dicranocentuga(sic) Mitra, SK, 2002:109 ISS
  Syn.: Paronellinae Metaparonella Mitra, 1972 u



Trogolaphysa
Cephalic chaetotaxy
After Mitra SK, 1993 Fig.40

  Syn.: Troglopedetini Trogolaphyusa(sic) Mitra, SK, 1993:66 ISS
  Syn.: Paronellidae Troglolaphysa(sic) Palacios-Vargas, JG et Zeppelini, D, 1995:173 ISS
  Syn.: Paronellini Trogolpahysa Soto-Adames, FN, 2015:23 ISS
  Syn.: Collembola Troglolaphysa(sic) Duarte Núñez, DA, Pinzón Florián, OP et Palacios-Vargas, JG, 2020:1205 ISS


Species sp.1 (Mari Mutt, JA, 1987:403)

Species sp.2 Palacios-Vargas, JG et Thibaud, J-M, 1997:328

Species sp.3 Bellini, BC et Zeppelini, D, 2004:588

Species sp.4 (Bellini, BC et Zeppelini, D, 2004:588)

Species sp.5 Palacios-Vargas, JG, 2008:46

Species sp.6 Bellini, 2014

Species sp.7 de Oliveira, MPA, de Oliveira Bernardi, LF, Zeppelini, D et Ferreira, RL, 2016:133

Trogolaphysa new sp. from Brazil
Parasitised by mite (Parasitengona, Microtrombidiidae)
After de Oliveira MPA & al, 2016 Fig.2A

Species sp.8 Mari Mutt & al, 2016

Species sp.9 Soto-Adames, FN et Anderson, EW, 2017:40

Species aelleni Yoshii, R, 1988:453

             Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:36 ISS
             Syn.: Trogolaphysa alleni(sic) Soto-Adames, FN, 2015:39 ISS

Species balteata Soto-Adames, FN, 2015:27

Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4C
Trogolaphysa balteata from Jamaica
After Soto-Adames FN, 2015 Fig.4D

Species beguei (Delamare Deboutteville, 1951) Nguyen, M & Soto-Adames, FN, 2018:182

            Syn.: Paronella beguei Delamare Deboutteville, 1948 nn
Trogolaphysa beguei from the Ivory Coast
After Nguyen M & Soto-Adames FN, 2018 Fig.5J



Species belizeana Palacios-Vargas, JG et Thibaud, J-M, 1997:326

Species berlandi (Denis, 1925) Thibaud, J-M & Najt, J, 1988:726

             Syn.: Paronella Berlandi(sic) Cassagnau, P in Delamare Deboutteville, C et Rapoport, E, 1963:134 ISS

Species bessoni Thibaud, J-M et Najt, J, 1988:724

Species caripensis (Gruia, M, 1987:155) Thibaud, J-M & Najt, J, 1988:723

Species carpenteri (Denis, 1925) Yoshii, 1988

            Syn.: Dicranocentruga geminata Mari Mutt, JA, 1987:402 =?
            Syn.: Dicranocentruga jataca Wray, DL, 1953:143 =?


             Syn.: Paronella Carpenteri(sic) Cassagnau, 1963 ISS

Species clarencei Soto-Adames, FN, 2015:21

Trogolaphysa clarencei from Mexico
After Soto-Adames FN, 2015 Fig.3E

Species cotopaxiana Thibaud, J-M et Najt, J, 1988:726

Species dimorphica Soto-Adames, FN, 2015:14

Trogolaphysa dimorphica from Mexico
After Soto-Adames FN, 2015 Fig.3B

Species distinguenda (Denis, 1931) Thibaud, J-M & Najt, J, 1988:726

Species ecuatorica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:22) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa ecuatoricus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa ecuatoricus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS
             Syn.: Trogolaphysa ecuatoriana(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 ISS

Species entreriosensis Soto-Adames, FN, 2015:23

Trogolaphysa entreriosensis from Argentina
After Soto-Adames FN, 2015 Fig.3F

Species ernesti Cipola NG et Bellini BC in Bellini, BC et Cipola, NG, 2017:170

Trogolaphysa ernesti from Brazil
After Bellini BC & Cipola NG, 2017 Fig.62

Species formosensis da Silva & Bellini, 2015

Species fuelleborni (Börner, 1903) Nguyen, M & Soto-Adames, FN, 2018:182

            Syn.: Paronella fülleborni(sic) Börner, 1903 IOS
             Syn.: Paronella fülleborni(sic) Wahlgren, 1908 ISS
             Syn.: Paronella fülleborni(sic) Handschin, 1929 ISS
             Syn.: Paronella fülleborni(sic) Denis, 1933 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:441 ISS
             Syn.: Paronella fülleborni(sic) Womersley, H, 1934:471 ISS
             Syn.: Paronella fülleborni(sic) Absolon, K et Ksenemann, M, 1942:13 ISS



Species geminata (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species ghesquierei (Marlier, 1945) Nguyen, M & Soto-Adames, FN, 2018:182

             Syn.: Dicranocentruga ghesourieri(sic) Mitra, SK, 2002:114 ISS
             Syn.: Trogolaphysa ghesquieri(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS
             Syn.: Dicranocentruga ghesouerei(sic) Nguyen, M et Soto-Adames, FN, 2018:183 ISS

Species giordanoae Soto-Adames, FN et Taylor, SJ, 2013:40

Trogolaphysa giordanoae from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.1

Species grassei (Delamare Deboutteville, 1952) Nguyen, M & Soto-Adames, FN, 2018:183

            Syn.: Paronella Grassei(sic) Delamare Deboutteville, 1951 nn IOS

Species guacharo Yoshii, R, 1988:455

Species guayabambensis (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species haitica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:16) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa haiticus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa haiticus(sic) Palacios-Vargas, JG et Thibaud, J-M, 1997:328 ISS

Species hauseri Yoshii, R, 1988:457

Species hirtipes (Handschin, 1924) Thibaud, J-M & Najt, J, 1988:725

            Syn.: Paronella carpenteri Denis, 1925 =?


Trogolaphysa cf. hirtipes from Brazil
After Cipola NG & al, 2019 Fig.10

Species hondurasensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa honduraensis(sic) Soto-Adames, FN, 2015:36 ISS

Species innominata (Denis, 1933) Thibaud, J-M & Najt, J, 1988:726

            Syn.: Paronella innominata f. I Denis, 1933
            Syn.: Paronella innominata f. II Denis, 1933
            Syn.: Paronella innominata f. III Denis, 1933


Remark by Mitra, S.K. 2009.07.31:in litt.: Originally, Denis (1933) described 3 forms of "innominata": C.R.I, II, and III. They were considered as different species by Salmon (1964:544), but are infraspecific morphs or varieties.

Species jacobyae Soto-Adames, FN et Taylor, SJ, 2013:47

            Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN et Taylor, SJ, 2013:47 IOS


Trogolaphysa jacobyi(sic) from Belize
After Soto-Adames FN & Taylor SJ, 2013 Fig.24
Trogolaphysa jacobyi(sic) from Belize
Ant.4 with constriction
After Soto-Adames FN & Taylor SJ, 2013 Fig.25

             Syn.: Trogolaphysa jacobyi(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:2 ISS
             Syn.: Trogolaphysa jacobyi(sic) Zeppelini, D et Oliveira, JVLC, 2016:569 ISS




Species jamaicana (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:19) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa jamaicanus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species jataca (Wray, DL, 1953:143) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Dicranocentruga jacata(sic) Mari Mutt, JA, 1979:60 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:28 ISS
             Syn.: Troglopedetes jataca(sic) Palacios-Vargas et al., 1986 ISS

Species judithnajtae Nguyen, M et Soto-Adames, FN, 2018:184

Trogolaphysa judithnajtae from Madagascar
After Nguyen M & Soto-Adames FN, 2018 Fig.1

Species laterolineata Soto-Adames, FN, 2015:16

Trogolaphysa laterolineata from Mexico
After Soto-Adames FN, 2015 Fig.3C

Species luquillensis (Mari Mutt, JA, 1987:402) Mari Mutt, JA & Bellinger, PF, 1996:171

Species marielouiseae Soto-Adames, FN, 2015:19

             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:1 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:2 ISS
             Syn.: Trogolaphysa marieloiseae(sic) Soto-Adames, FN, 2015:20 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:10 ISS
             Syn.: Trogolaphysa marielouisea(sic) Soto-Adames, FN, 2015:17 ISS
Trogolaphysa marielouisea(sic) from Mexico
After Soto-Adames FN, 2015 Fig.3D



Species marimutti (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:24) Thibaud, J-M & Najt, J, 1988:723

Species maya Mills, HB in Pearse, AS, 1938:184, t.t.

            Syn.: Troglopedetes delamarei Massoud, Z et Gruia, M, 1973:339
Troglopedetes delamarei from Cuba
After Massoud Z & Gruia M, 1973 Fig.7E




Species millsi Arlé, 1939

Species monomaculata (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

Species nacionalica (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa nacionalicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species neoannulicornis (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species nigromaculata (Schött, H, 1903:VII) Nguyen, M & Soto-Adames, FN, 2018:189

Paronella nigromaculata from Kamerun
Schematic outline of habitus
After Schött H, 1903 Taf.I Fig.10

             Syn.: Paronella fülleborni(sic) Philiptschenko, 1926 ISS
             Syn.: Paronella fülleborni(sic) Martynova, 1961 ISS
            Syn.: Neophorella dubia Paclt, 1959



  Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Pauliani, 1952

            Syn.: Paronella nigromaculata ssp. pallida Delamare Deboutteville, 1951 nn

Species ocellata Soto-Adames, FN, 2015:25

Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4A
Trogolaphysa ocellata from Mexico
After Soto-Adames FN, 2015 Fig.4B

Species octosetosa Soto-Adames, FN, 2015:32

Trogolaphysa octosetosa from Mexico
After Soto-Adames FN, 2015 Fig.4F

Species oztotlica (Ojeda, M & Palacios-Vargas, JG, 1984:16) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetes sp. Palacios-Vargas, 1982


             Syn.: Trogolaphysa oztotlicus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species palaciosi Soto-Adames, FN, 2015:32

Species paracarpenteri Soto-Adames, FN, 2015:29

Trogolaphysa paracarpenteri from Mexico
After Soto-Adames FN, 2015 Fig.4E
             Syn.: Trogolaphysa paracapenteri(sic) Soto-Adames, FN, 2015:31 ISS


Species penicillata (Schött, 1926) Nguyen, M & Soto-Adames, FN, 2018:190

  Subspecies obscura Delamare Deboutteville, 1950

            Syn.: Paronella penicillata ssp. obscura Delamare Deboutteville, 1951 nn

  Subspecies pallida Delamare Deboutteville in Delamare Deboutteville & Paulian, 1952



Species piracurucaensis Nunes, RC et Bellini, BC, 2018:87

Trogolaphysa-piracurucaensis from Brazil
After Nunes RC & Bellini BC, 2018 Fig.44

Species quinquesetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species quisqueyana Soto-Adames, FN, Jordana, R et Baquero, E, 2014:10

Trogolaphysa quisqueyana from the Dominican Republic
After Soto-Adames & al., 2014 Fig.43

Species relicta (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:26) Thibaud, J-M & Najt, J, 1988:723

             Syn.: Trogolaphysa relictus(sic) Thibaud, J-M et Najt, J, 1988:723 ISS

Species riopedrensis (Mari Mutt, JA, 1987:403)

Species sauroni Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12

            Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, Jordana, R et Baquero, E, 2014:12 IOS
Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 left
Trogolaphysa sauron(sic) from Martinique
After Soto-Adames & al., 2014 Fig.51 right
Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 left
Trogolaphysa sauron(sic) from Martinique
Pale form
After Soto-Adames & al., 2014 Fig.52 right

             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:17 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN, 2015:29 ISS
             Syn.: Trogolaphysa sauron(sic) Soto-Adames, FN et Anderson, EW, 2017:40 ISS




Species separata (Denis, 1933) Thibaud, J-M & Najt, J, 1988:726

            Syn.: Dicranocentruga riopedrensis Mari Mutt, JA, 1987:403 =?


Species sexsetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species stannardi Soto-Adames, FN, 2015:8

Trogolaphysa stannardi from Mexico
After Soto-Adames FN, 2015 Fig.3A

Species strinatii Yoshii, R, 1988:452

Species subterranea (Mari Mutt, JA, 1987:403) Mari Mutt, JA & Bellinger, PF, 1996:171

            Syn.: Dicranocentruga subterranean(sic) Mari Mutt, JA, 1987:403 IOS

Species tijucana (Arlé & Guimaraes, 1979) Mari Mutt, JA & Bellinger, PF, 1990:124

             Syn.: Troglopedetes tijucana(sic) Palacios-Vargas et al., 1989 ISS
             Syn.: Trogolaphysa tijuacana Soto-Adames, FN, 2015:39 ISS

Species toroi (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:12) Thibaud, J-M & Najt, J, 1988:723

Species trioculata Soto-Adames, FN, 2015:35

Trogolaphysa trioculata from Mexico
After Soto-Adames FN, 2015 Fig.3G

Species trisetosa (Winter, 1963) Mari Mutt & Bellinger, 1990 nn

Species variabilis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:18) Thibaud, J-M & Najt, J, 1988:723

Species villiersi (Delamare Deboutteville, 1950) Soto-Adames, FN, 2014:il

            Syn.: Paronella villiersi Delamare Deboutteville & Paulian, 1947 nn
             Syn.: Paronella villersi(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

  Subspecies anomala Delamare Deboutteville, 1951

  Subspecies pallida Delamare Deboutteville, 1950



Species vivieni (Barra, 1969) Soto-Adames, FN & Bellini, BC, 2015:337

             Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:181 ISS
             Syn.: Trogolaphysa viviani(sic) Nguyen, M et Soto-Adames, FN, 2018:182 ISS

Species wahlgreni (Mitra, SK, 2002:109) Nguyen, M & Soto-Adames, FN, 2018:192

            Syn.: Paronella nigromaculata Wahlgren, 1908


Dicranocentruga wahlgreni from Kenia
After Mitra SK, 2002 Fig.1A
             Syn.: Dicranocentruga whalgreni(sic) Nguyen, M et Soto-Adames, FN, 2018:189 ISS
             Syn.: Trogolaphysa wahlbergi(sic) Nguyen, M et Soto-Adames, FN, 2018:194 lapsus ISS


Species xtolokensis (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:10) Thibaud, J-M & Najt, J, 1988:723

Species yoshiii (Palacios-Vargas, JG, Ojeda, M & Christiansen, KA, 1985:14) Thibaud, J-M & Najt, J, 1988:723

            Syn.: Troglopedetes yoshiius(sic) Palacios-Vargas, JG, Ojeda, M et Christiansen, KA, 1985:14 IOS


             Syn.: Trogolaphysa yoshius(sic) Thibaud, J-M et Najt, J, 1988:723 ISS
             Syn.: Trogolaphysa yoshiia(sic) Mari Mutt, JA et Bellinger, PF, 1990:124 ISS
             Syn.: Trogolaphysa yoshiia(sic) Palacios-Vargas, JG, 2006:10 ISS
             Syn.: Trogolaphysa yoshiia(sic) Soto-Adames, FN et Taylor, SJ, 2013:37 ISS

References