http://www.collembola.org/taxa/anurinae.htm - Last updated on 2017.06.20 by Frans Janssens
Checklist of the Collembola: Anurophorinae

Subfamilia Anurophorinae Börner C, 1901:42
Genus Number
of species
Aggressopygus2
Antarctophorus1
Anurophorus53
Archisotoma28
Arlea8
Blissia2
Cryptopygus88
Cylindropygus1
Dagamaea4
Dimorphacanthella2
Folsomia179
Gnathofolsomia2
Gressittacantha1
Haploisotoma1
Hemisotoma8
Isotomiella53
Isotominella2
Isotomodella6
Isotomodes35
Jesenikia2
Martynovella1
Micranurophorus2
Micrisotoma1
Mucracanthus2
Mucrosomia4
Neocryptopygus1
Octodontophora1
Pauropygus3
Pectenisotoma1
Proctostephanus6
Proisotomodes2
Pseudanurophorus12
Pseudofolsomia2
Rhodanella1
Sahacanthella1
Secotomodes2
Sibiracanthella3
Stachanorema3
Tetracanthella99
Tiancanthella1
Tuvia2
Uzelia12
Weberacantha7
Womersleyella1
Yosiiella1
Anurophorinae649
Genera with number of species
Anurophorinae from Australia
Anus opens ventrally
2015.11.07 © Murray, A.

Remark by Melville, R.V. & Smith, J.D.D. 1987:5: Anurophorinae Börner, 1901. Placed on the Official List of Family-Group Names in Zoology. Type genus: Anurophorus Nicolet, [1842]. Opinion 439.
Remark by Melville, R.V. & Smith, J.D.D. 1987:8: Bourletiini Börner in Brohmer, 1932. Placed on the Official Index of Rejected and Invalid Family-Group Names in Zoology. Type genus: Bourletia MacGillivray, 1893. Junior objective synonym of Anurophorinae Börner, 1901, the respective type genera of the two families having the same nominal species as type species. Opinion 439.



Genus Proctostephanus Börner, C, 1902:606 key
Species cidi Selga, 1958
Species dalii Arbea, JI, 2003:5
Species madeirensis da Gama, 1959
Species provincialis Poinsot & Dallai, 1970
Species sanctiaugustini Cassagnau, 1963
Species stuckeni Börner, C, 1902:607, t.t.
            Syn.: Proctostephanus cidi Selga, 1958 =?

Genus Uzelia Absolon, K, 1901:209
Uzelia sp. from the USA
2006 © Bernard, E.
Uzelia sp. from Russia
2006 © Boldsystems.org

Species badakschanica Martynova, 1971
Species clavata Jordana, R et Ardanaz, A, 1981:37
Species dahli (Börner, 1903) Gisin, 1960:?, s.d.
Species furcata Grinbergs, A, 1962:61
  Forma lutea Grinbergs, A, 1962:63
Species hansoni Mills & Richards, 1953
Species japonica Yosii, 1961
Species kondarensis Martynova, 1971
Species kuehnelti Cassagnau, P, 1954:613
            Syn.: Uzelia kühnelti(sic) Cassagnau, P, 1954:613 IOS
Species mongolica Martynova, 1975
Species oxoniensis (Bagnall, 1914) Kloet, GS & Hincks, WD, 1945:6
Species setifera Absolon, K, 1901:210, t.t.
            Syn.: Protanurophorus pearmani Womersley & Bagnall in Womersley, 1925
             Syn.: Anurophorus Pearmani(sic) Cassagnau, P, 1953:156 ISS
Uzelia setifera from Poland
After Stach, J., 1947 Plate VI Fig.1.

Species tadjika Martynova, 1971

Genus Tetracanthella Schött, 1891 key
  Syn.: ... Lubbockia Haller, 1880 nec Claus, 1862
  Syn.: ... Deuterolubbockia Dalla-Torre, 1895
  Syn.: Isotomidae Tetracantella(sic) Fiera, C, Konikiewicz, M et Skarzynski, D, 2013:468 ISS
Species acuminata Cassagnau, P, 1959:223
Species afurcata Handschin, 1919
Species albanordica Peja, 1985
Species alpina Carl, J, 1901:255
Species alticola Deharveng, 1987
Species andalusiaca Deharveng, 1987
Tetracanthella andalusiaca from Spain
After Fiera C & al, 2013 Fig.1

Species antoni Potapov, 1997
Species apenninica Parisi, V, 2007:24
Species arctica Cassagnau, P, 1959:234
            Syn.: Tetracanthella Wahlgreni(sic) Hammer, 1944, nec Linnaniemi, 1911
            Syn.: Tetracanthella iberica Steiner, 1958
Tetracanthella arctica from Norway
2011 © Fjellberg, A.

  Subspecies estaranhensis Cassagnau, P, 1959:236
Species ariegica Deharveng & Bedos, 1997

Tetracanthella bellingeri from the USA
2006 © Bernard, E.
Species bellingeri Deharveng, 1978

Species bichaeta Potapov, MB et Deharveng, L, 2005:87
Species bipartita Cassagnau, P, 1959:231
Species borsa Dányi, L et Traser, G, 2008:213
Species bosnia Palissa & Zivadinovic, 1974

Tetracanthella brachyura
2005 © Brocklehurst, K.
Species brachyura Bagnall, 1949

Species brevempodialis Gisin, 1963
Species brevifurca Stach, J, 1930:287
Species caerulea (Haller, 1880) Schäffer, 1900
            Syn.: Lubbockia cærulea(sic) Haller, 1880 IOS
             Syn.: Lubbockia cærulea(sic) MacGillivray, AD, 1893:318 ISS
             Syn.: Lubbockia Coerulea(sic) Carl, J, 1899:274,sd ISS
             Syn.: Tetracanthella coerulea(sic) Schäffer, 1900 ISS
Species calcarata Deharveng, 1987
Species californica Deharveng, 1978
Species cantabrica Deharveng, 1987
Species carpatica Stach, 1947
Species cassagnaui Gisin, H, 1962:19
Species caucasica Stach, J, 1947:32
Species christianseni Cassagnau, P, 1959:217
            Syn.: Tetracanthella Christianseni(sic) Cassagnau, P, 1959:217 IOS
Species clandestina Deharveng, 1987
Species coiffaiti Deharveng, 1987
Species corsica Potapov, MB et Deharveng, L, 2005:89
Species czernovae Kutyreva, 1980
Species dallaii Deharveng, 1987
Species deficiens Steiner, 1958
Species deharvengi Potapov, 1997
Species doftana Fiera, C, Konikiewicz, M et Skarzynski, D, 2013:468
Tetracanthella doftana from Romania
After Fiera C & al, 2013 Fig.2

Species elevata Cassagnau, P, 1959:237
Species emucronata Deharveng, 1987
Species ethelae Wray, 1945
            Syn.: Tetracanthella Ethelae(sic) Cassagnau, P, 1959:219 IOS
Species fjellbergi Deharveng, 1987
            Syn.: Tetracanthella arctica auct.
            Syn.: Tetracanthella wahlgreni auct.
Species fluorina Raynal, 1972
Species franzi Cassagnau, P, 1959:240
            Syn.: Tetracanthella Franzi(sic) Cassagnau, P, 1959:240 IOS
            Syn.: Tetracanthella montana Stach, 1947, a.p.
Species fusca Potapov, MB et Deharveng, L, 2005:91
Species gallica Deharveng, 1987
Species gamae Deharveng, 1987
Species gisini Cassagnau, P, 1959:239
            Syn.: Tetracanthella Gisini(sic) Cassagnau, P, 1959:239 IOS
Species grindbergsi Deharveng, 1987
Species gruiae Rusek, 1979
Species hatipari Potapov & Kuchiev, 1993
Species hellenica Deharveng, 1987
Species hygropetrica Cassagnau, P, 1954:615
            Syn.: Tetracanthella matthesii da Gama, 1959
             Syn.: Tetracanthella matthesi(sic) Paclt, J, 1961:222 ISS
Species hystrix Cassagnau, P, 1959:242
            Syn.: Tetracanthella montana Stach, 1947
Species ignisiana Dányi, L et Traser, G, 2008:216
Tetracanthella ignisiana from Romania
After Dányi L & Traser G, 2008 Fig.20

Species intermedia Palissa, 1968
Species irregularis Deharveng, 1987
Species juneaui Potapov, 1997
Species kendalli Bagnall, 1939
             Syn.: Tetracanthella Kendalli(sic) Cassagnau, P, 1953:154 ISS
             Syn.: Tetracanthella Kendalli(sic) Cassagnau, P, 1959:244 ISS
Species ksenemani Nosek, 1964
Species luxemburgensis Stomp, 1968
Tetracanthella luxemburgensis from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.4

Species mansurica Kutyreva, 1980
Species martini Simón-Benito, JC et Luciáñez Sánchez, MJ, 1998:234
Species martynovae Potapov, 1997
Species matthesii da Gama, 1959
Species montana Stach, 1947, a.p.
            Syn.: Tetracanthella bescidica Potapov, MB et Karpus, IJ, 1993:32
Species nitida Deharveng, 1987
Species orientalis Martynova, 1977
Species osetica Potapov & Kuchiev, 1993
Species pacifica Rusek & Marshall, 1977
Species perezi Delamare Deboutteville, 1943
            Syn.: Tetracanthella reticulata Cassagnau, P, 1953:151
             Syn.: Tetracanthella Perezi(sic) Delamare Deboutteville, C, 1946:67 ISS
             Syn.: Tetracanthella Perezi(sic) Cassagnau, P, 1953:154 ISS
             Syn.: Tetracanthella Perezi(sic) Cassagnau, P, 1959:223 ISS
Species pericarpathica Kaprus, IJ et Tsalan, JV, 2009:176
Tetracanthella pericarpathica from Ukraine
After Kaprus' IJ & Tsalan JV, 2009 Fig.2.1
             Syn.: Tetracanthella pericarpatica(sic) Fiera, C, Konikiewicz, M et Skarzynski, D, 2013:471 ISS
Species pilosa Schött, 1891, t.t.
            Syn.: ... coerulea Haller, 1880
            Syn.: Tetracanthella Delamarei(sic) Cassagnau, P, 1953:150 IOS
            Syn.: Tetracanthella Delamarei(sic) Cassagnau, P, 1959:221 IOS
Tetrachantella pilosa from Norway
After Schött H, 1893 Taf.VII Fig.2

Species proxima Steiner, 1955
Species pseudomontana Cassagnau, P, 1953:152
Species pyrenaica Cassagnau, P, 1953:152
Species quinqueoculata Simón-Benito, JC et Luciáñez Sánchez, MJ, 1998:231
Species raynalae Deharveng, 1987
Species recta Deharveng, 1987
            Syn.: Tetracanthella tubercolata(sic) Parisi, V, 2007:20
Species reducta von Törne, 1955
Species religiosa Deharveng, 1987
Species schalleri Hemmer, 1992
Species selgae Deharveng, 1987
Species septemsetosa Martynova, 1971
Species serrana Steiner, 1955
Species sexsetosa Martynova, 1971
Species sibirica Deharveng, 1987
Species sigillata Deharveng, 1987
Species similis Deharveng, 1987
Species specifica Palissa, 1968
Species stachi Cassagnau, P, 1959:240
            Syn.: Tetracanthella Stachi(sic) Cassagnau, P, 1959:240 IOS
Species stebevae Deharveng, 1987
Species steineri Deharveng, 1987
Species strenzkei Gisin, 1949
            Syn.: Tetracanthella lichnidis Bagnall, 1949
             Syn.: Tetracanthella Strenzkei(sic) Cassagnau, P, 1959:225 ISS
Tetracanthella strenzkei from the UK
2013.04.25 © Shaw, P.

Species subdeficiens Deharveng, 1987
Species suursarensis Linnaniemi, 1912
Species sylvatica Yosii, 1939
Species tarbae Potapov & Kuchiev, 1993
Species transylvanica Cassagnau, P, 1959:243
             Syn.: Tetracanthella transsylvanica(sic) Gisin, 1960 ISS
Species travei Deharveng, 1987
Species tuberculata Cassagnau, P, 1954:617
             Syn.: Tetracanthella tubercolata(sic) Parisi, V, 2007:20 ISS
  Subspecies orbaicetensis Cassagnau, P, 1959:230
Species uniseta Deharveng, 1987
Species wahlgreni Axelson, 1907
            Syn.: Tetracanthella coerulea Schäffer, 1900
            Syn.: Tetracanthella pilosa Schött, 1891, partim
Tetracanthella wahlgreni from Arctica
After Folsom JW, 1919 Plate 4 Fig.33
             Syn.: Tetracanthella Wahlgreni(sic) Cassagnau, P, 1953:154 ISS
             Syn.: Tetracanthella Wahlgreni(sic) Cassagnau, P, 1959:234 ISS
Tetracantella wahlgreni from the UK
2012.12.16 © Murray, A.
Tetracantella wahlgreni from the UK
2012.12.16 © Murray, A.
Tetracantella wahlgreni from the UK
Four anal spines
2012.12.16 © Murray, A.


Genus Blissia Rusek, 1985
Species glabra Rusek, 1985
Species robusta Fjellberg, A et Potapov, MB, 1998:87

Genus Sibiracanthella Potapov & Stebaeva, 1995
  Syn.: Isotomidae Sibirocanthella(sic) Potapov, MB, Bu, Y, Huang, C-W, Gao, Y et Luan, Y-X, 2010:15 ISS
Species nuda Stebaeva & Potapov, 1995
Species rara (Dunger, WG, 1982:38)
Species sohondo Potapov & Stebaeva, 1995

Genus Dimorphacanthella Potapov, MB, Bu, Y, Huang, C-W, Gao, Y et Luan, Y-X, 2010:14
  Syn.: Isotomidae Dimorphocanthella(sic) Potapov, MB, Bu, Y, Huang, C-W, Gao, Y et Luan, Y-X, 2010:15 ISS
Species anommatos (Chen & Yin, 1984 tt) Potapov, MB, Bu, Y, Huang, C-W, Gao, Y & Luan, Y-X, 2010:16
Dimorphacanthella anommatos from China
Adult specimen with 4 anal spines
Juvenile specimen with 2 anal spines
After Potapov MB et al, 2010 Fig.3,4
            Syn.: Uzelia anommatos Yue & Yin, 1999
Species mediaseta Potapov, MB, Bu, Y, Huang, C-W, Gao, Y et Luan, Y-X, 2010:19

Genus Martynovella Deharveng, 1979
Species nana (Martynova, 1967)
  Subspecies kirgisica Martynova, 1978

Genus Tuvia Grinbergs, A, 1962:61
Species chinensis Chen, B-R et Yin, W-Y, 1983:173
Species prima Grinbergs, A, 1962:61, t.t.

Genus Womersleyella Salmon, JT, 1944:142
Species niveata Salmon, JT, 1944:142, t.t.

Genus Stachanorema Wray, 1957
Species arnaudi Wray, 1957
Species contra Christiansen & Bellinger, 1980
Species tolerans Babenko, 1994

Genus Antarctophorus Potapov, 1992
Species subpolaris (Salmon, 1962) Block, 1984
             Syn.: Antarctophorus sudpolaris(sic) Hopkin, SP, 1998:118 ISS
             Syn.: Antarctophorus sudpolaris(sic) Hopkin, SP, 2002:208 ISS
Anurophorus subpolaris from Antarctica
2008 © Univ of Waikato.



Genus Neocryptopygus Salmon, 1965
Species nivicolus Salmon, 1965
Neocryptopygus nivicolus from Antarctica
2008 © Univ of Waikato.


Genus Cryptopygus Willem, V, 1901:261
  Syn.: Isotomidae Cryptpygos(sic) Christiansen, 1951 ISS
Cryptopygus sp. from Antarctica
2003 © Stevens, M.

Species sp.1 Convey, P, Greenslade, P, Richard, KJ et Block, W, 1996:1
Species sp.2 Potapov, 2001
Species sp.3 Potapov, 2001
Species sp.4 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.5 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.6 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.7 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.8 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.9 Stevens, MI, Greenslade, P, Hogg, ID et Sunnucks, P, 2006:876
Species sp.10 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:141
Species sp.11 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:141
Species sp.12 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:141
Species sp.13 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:141
Species sp.14 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:141
Species sp.15 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:142
Species sp.16 Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:142
Species agreni (Börner, 1903), t.t.
             Syn.: Isotoma (Isotomina) Ågreni(sic) Ågren, H, 1904:19 ISS
Species albus Yosii, 1939
Species albaredai Selga, 1962
Species ambus Christiansen & Bellinger, 1980
Cryptopygus ambus from the USA
2007.09.07 © Snider, R.

Species andinus Díaz & Najt, 1995
Species annobonensis Selga, 1962
Species anomala Linnaniemi, 1912
Species antarcticus Willem, V, 1901:261, t.t.
            Syn.: Cryptopygus crassus Carpenter, GH, 1907:476
Cryptopygus crassus from Antarctica
After Carpenter GH, 1907 Plate II Fig.14

Cryptopygus antarcticus from Antarctica
After Willem V, 1902 Pl.III Fig.7
Cryptopygus antarcticus from the Falkland Islands
Stamp of 13 P, 1982
Cryptopygus antarcticus from Antarctica
After Rusek, J, 2002 Fig.1A
Cryptopygus antarcticus from Antarctica
After Davis, G. in Greenslade, P., 2006 Fig.63

  Subspecies maximus Deharveng, 1981
  Subspecies reagens Enderlein, 1909
            Syn.: Isotoma reagens Vanhöffen, 1908 nn
            Syn.: Cryptopygus reagens Vanhöffen, 1908 nn
            Syn.: Cryptopygus reagens Enderlein, 1908 nn
            Syn.: Cryptopygus reagens Enderlein, 1909 nn
  Subspecies travei Deharveng, 1981
Species aquae (Bacon, 1914) Poole, RW in Poole, RW & Gentili, P, 1997:50
Species araucanus Massoud & Rapoport, 1968
Species arcticus Christiansen & Bellinger, 1980
Species axayacatl Palacios-Vargas, JG et Thibaud, J-M, 2001:164
             Syn.: Cryptopygus axayacalt(sic) Thibaud, J-M, 2007:185 ISS
Species badasa Greenslade, P, 1995:312
Species beijiangensis Hao & Huang, 1995
Species benhami Christiansen & Bellinger, 1980
Species binoculatus Deharveng, 1981
Species campbellensis Wise, 1964
Species cardusi Selga, 1962
Species cinctus Wahlgren, E, 1906:11
Species cisantarcticus Wise, 1967
Cryptopygus cisantarcticus from Antarctica
2008 © Univ of Waikato.

Species clavatus (Schött, H, 1893:73) Fjellberg, A, 2007:67
Species constrictus (Folsom, JW, 1937:64) Zorrilla, 1985
Species decemoculatus (Salmon, JT, 1949:20)
Species delamarei Poinsot, 1970
Species dubius Deharveng, 1981
Species elegans (Rapoport & Izarra, 1962) Mari Mutt & Bellinger, 1990:70
Species elegans (Cardoso MA, 1973:16, nec Rapoport & Izarra, 1962)
Species exilis (Gisin, 1960) Bernard, EC, 1977:76
Species hirsutus (Denis, 1931) Rapoport, 1971
            Syn.: Isotomina neomuelleri Winter, 1963 nn
Species indecisus Massoud & Rapoport, 1968
Species indicus Brown, 1932
Species insignis Massoud & Rapoport, 1968
Species interruptus Schött, 1927
Species kahuziensis Martynova, 1978

Isotomina lamellata from New Zealand
After Salmon, J.T. 1941 Pl.53 Fig.221.
Species lamellatus (Salmon, JT, 1941:334)
Species lapponicus (Brown, 1931)
Species lawrencei Deharveng, 1981
Species loftyensis (Womersley, 1934) Womersley, H, 1936:320
Species mauretanica Handschin, 1925
Species minimus Salmon, 1941
Species nanjiensis Shao, H, Zhang, Y, KE, X, Yue, Q et Yin, W, 2000:589
Species novaezealandiae (Salmon, JT, 1943:74) Wise, 1974
            Syn.: Folsomia novae-zealandiae(sic) Salmon, JT, 1943:74 IOS
             Syn.: Folsomia novae-zealandiae(sic) Stach, 1947 ISS
             Syn.: Parafolsomia novae-zealandiae(sic) Salmon, JT, 1949:20 ISS
             Syn.: Folsomia novae-zealandiae(sic) Salmon, JT, 1958:707 ISS
             Syn.: Spinurosomia novae-zealandiae(sic) Salmon, JT, 1964:332 ISS
             Syn.: Parafolsomia novae-zealandiae(sic) Salmon, JT, 1964:359 ISS
Species novazealandia (Salmon, JT, 1941:334) Bellinger, PF & Christiansen, KA, 2000:1
             Syn.: Zealandotoma nova-zealandia(sic) Salmon, JT, 1964:359 ISS
Species oeensis (Caroli, 1914)
Species parallelus (Wahlgren, 1901) Wahlgren, 1907
Species parasiticus (Salmon, JT, 1943:380)
Species patagonicus Izarra, 1972
Species pentatomus (Börner, C, 1906:172) Mari Mutt & Bellinger, 1990:71
Species perisi Selga, 1960
Species pilosus (Womersley, 1934) Greenslade P in Harris, Reeves & Sympson, 1982
            Syn.: Cryptopygus sp. cf. cisantarcticus Greenslade P, 1974
Species pseudominuta Schött, 1927
Species quadrioculatus (Rapoport, 1963) Mari Mutt & Bellinger, 1990:71
Species quadrioculatus Martynova, 1967, nec Rapoport, 1963
Species quadrioculatus Yoshii, 1995, nec Rapoport, 1963
Species quinqueoculatus Izarra, 1970
Species roberti (Fjellberg, 1991) Potapov, M, Janion-Scheepers, C & Deharveng, L, 2017:132
Species scapelliferus (Gisin, 1955)
             Syn.: Isotomina scalpellifera(sic) Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:8 ISS
             Syn.: Isotomina scapelifera(sic) Jordana, R, 1980:101 ISS
             Syn.: Cryptopigus(sic) scapelifera(sic) Jordana, R, 1980:101 ISS
             Syn.: Isotomia(sic) scapelifera(sic) Jordana, R, 1980:105 ISS
             Syn.: Hemisotoma scapelliferus(sic) Potapov, M et Babenko, A, 2014:15 ISS
Species separatus (Denis, 1931) Mari Mutt & Bellinger, 1990:71
             Syn.: Cryptopygus separata(sic) Loranger, G, Ponge, J-F, Blanchart, E et Lavelle, P, 1998:24 ISS
Species sphagneticola (Linnaniemi, WM, 1912:171)
Species subalpinus (Salmon, JT, 1944:147)
Species subantarcticus Wise, KAJ, 1970:192
Species sverdrupi Lawrence, 1978
Crytopygus (sic) sverdrupi from Antarctica
After Thibaud J-M & D'Haese C, 2010 Fig.36
2009 © D'Haese, C.

Species tasmaniensis Womersley, H, 1942:25
Cryptopygus tasmaniensis from Australia
After Womersley H, 1942 Fig.2A

Species tingus Queiroz & de Mendonça, 2010
Species tricuspis Enderlein, 1909
            Syn.: Parafolsomia quadrioculata Wise, KAJ, 1970:194
Species tridentatus (Handschin, E, 1929:237)
Cryptopygus tridentatus from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573

Species triglenus Ellis, WN, 1976:287
Species trioculatus Izarra, 1972
Species ulrikeae (Najt & Thibaud, 1987) Potapov, M, Janion-Scheepers, C & Deharveng, L, 2017:141
Species vtorovi Martynova, 1978
Species yosiii Izarra, 1965
            Syn.: Cryptopygus yosii(sic) Izarra, 1965 IOS
             Syn.: Cryptopygus yosii(sic) Izarra, 1970 ISS
             Syn.: Cryptopygus yosii(sic) Mari Mutt, JA et Bellinger, PF, 1990:73 ISS
             Syn.: Cryptopygus yosii(sic) Potapov, 2001 ISS
             Syn.: Cryptopygus yosii(sic) Potapov, M, Babenko, A et Fjellberg, A, 2006:73 ISS
             Syn.: Cryptopygus yosii(sic) Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:143 ISS
Species zenderi (Winter, 1967) Hazelton, 1978

Genus Pauropygus Potapov, Gao & Deharveng, 2013
Species caussaneli (Thibaud, 1996) Potapov, Gao & Deharveng, 2013
             Syn.: Cryptopygus caussanelli(sic) Thibaud, J-M, 2007:185 ISS
            Syn.: Cryptopygus riebi Barra, J-A, 1997:470
Species pacificus Potapov, Gao & Deharveng, 2013
Species projectus Potapov, Gao & Deharveng, 2013

Genus Hemisotoma Bagnall, 1949
  Syn.: Isotomidae Cyrptopygus(sic) Kahrarian, M et Arbea, J, 2013:92 ISS
Species bibasiosetis (Potapov & Stebaeva, 1999) Rusek, J, 2002:307
Species bituberculata (Wahlgren, E, 1906:4) Jordana, R & Baquero, E, 2010:364
             Syn.: Hemisotoma bituberculatus(sic) Jordana, R et Baquero, E, 2010:364 ISS
Species orientalis (Stach, J, 1947:275) Cox, 1982
Isotomina orientalis from Poland
After Stach, J., 1947 Plate XXXI Fig.1.

Species pontica (Stach, 1947) Kahrarian, M & Arbea, J, 2013:92
Hemisotoma pontica from Iran
After Kahrarian M & Arbea J, 2013 Fig.6

Species posteroculata (Stach, 1947) Rusek, J, 2002:307
Species subpontica Thibaud, 2010
            Syn.: Hemisotoma subponticus(sic) Thibaud, 2010 IOS
Species thermophila (Axelson, 1900:114) Bagnall, 1949, t.t., B
            Syn.: Isotoma lombokensis Schött, 1901
            Syn.: Proisotoma (Isotomina) thermophila var. anomala Linnanieimi, 1912
            Syn.: Proisotoma (Isotomina) thermophila var. impunctata Linnanieimi, 1912
            Syn.: Isotoma bituberculata Wahlgren, 1912
            Syn.: Isotomina thermophila f. C.R.I Denis, 1931 IOS
             Syn.: Isotomina thermophila f. C.R.I Denis, 1933 ISS
             Syn.: Isotomina thermophila f. Costa Rica Winter, 1963 ISS
            Syn.: Isotoma linnaniemia Womersley, 1934
             Syn.: Isotoma linnaniemi(sic) Womersley, H, 1936:321 ISS
            Syn.: Proisotoma (Isotomina) sexoculata Womersley, 1934
            Syn.: Isotoma decemoculata Womersley, H, 1935:216
            Syn.: Proisotoma (Proisotoma) cognata Folsom, JW, 1937:65
            Syn.: Proisotoma constructua Folsom, 1937
            Syn.: Proisotoma (Proisotoma) tenelloides Folsom, JW, 1937:62
            Syn.: Isotoma (Isotoma) pentomma Womersley, H, 1939:165
            Syn.: Isotomina martiniquae Stach, 1947
            Syn.: Isotomina salaymehi Christiansen, K, 1958:72
Isotomina salaymehi from Lebanon
After Christiansen, K., 1958 Pl.9 Fig.14.

            Syn.: Isotomina thermophila ssp. peruensis Winter, 1963,nn
            Syn.: Isotomina thermophila ssp. costaricensis Winter, 1967
            Syn.: Cryptopygus orientalis Poinsot, 1971
            Syn.: Cryptopygus constrictus Bellinger, PF et Christiansen, KA, 1974:34
            Syn.: Isotomina thermophila ssp. galapagoensis Jacquemart, 1975 in Peck SB & Jacquemart J, 2011:3 u
Isotomina thermophila from Poland
After Stach, J., 1947 Plate XXX Fig.1.
             Syn.: Isotomina termophila(sic) Cardoso, MA, 1966:90 ISS
             Syn.: Cryptopigus(sic) termophilus(sic) Miranda, RA, Palacios-Vargas, JG, Garcia, CN et Pescador, RA, 1995:103 ISS
             Syn.: Cryptopygus thermophila(sic) Skidmore, RE, 1995:55 ISS
             Syn.: Cryptopygus thermophila(sic) Poole, RW in Poole, RW et Gentili, P, 1997:50 ISS
             Syn.: Cryptopygus termophilus(sic) Palacios-Vargas, JG, 2000:30 ISS
             Syn.: Cryptopygus thermophylus(sic) Arbea, JI et Blasco-Zumeta, J, 2001:35 ISS
             Syn.: Cryphopygus(sic) thermophilus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:22 ISS
             Syn.: Cryptopyhus(sic) thermophilus Bonte, D, Van Heuverswyn, F et Mertens, J, 2002:24 ISS
Hemisotoma thermophila from Europe
With foil seta
After Rusek, J, 2002 Fig.3

             Syn.: Cryptopygus thermophylus(sic) Cutz-Pool, LQ, Palacios-Vargas, JG, Castaño-Meneses, G et Carcía-Calderón, N, 2003:191 ISS
             Syn.: Cryptopygus thermophila(sic) Shaw, P, 2003:7 ISS
             Syn.: Cryptopygus thermophylus(sic) Cutz-Pool, LQ, Palacios-Vargas, JG et Castaño-Meneses, G, 2005:172 ISS
Cryptopygus thermophilus from the UK
Mucro
2005 © Hopkin, S.P.

             Syn.: Cryptopyguscomplex thermophila(sic) Potapov, M, Babenko, A et Fjellberg, A, 2006:73 ISS
             Syn.: Hemisotoma thermophyla(sic) Fiera, C, 2007:20 ISS
             Syn.: Cryptopygus thermophylus(sic) Cutz-Pool, LQ, Palacios-Vargas, JG et Castaño-Meneses, G, 2008:4 ISS
Hemisotoma thermophila from Australia
2008.11.19 © PADIL
Det. Greenslade, P.
Cryptopygus thermophilus from India
After Mandal GP & Hazra AK, 2009 Fig.24g

             Syn.: Hemisotoma thermophilus(sic) Zeppelini, D, Bellini, BC, Creao-Duarte, AJ et Hernández, MIM, 2009:1165 ISS
             Syn.: Hemisotoma termopila(sic) Magaña-Martinez,CS et Palacios-Vargas, JG, 2010:76 ISS
             Syn.: Hemisotoma thermophilus(sic) Jordana, R et Baquero, E, 2010:364 ISS
Hemisotoma thermophila from Holland
2011.03.05 © van Duinen, J.
Hemisotoma thermophila from the USA
2011.12.11 © Kouri, J.


             Syn.: Cryptopygus thermophylus(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:110 ISS
Hemisotoma thermophila from Holland
2012.10.18 © van Duinen, J.
Hemisotoma thermophila from the UK
2012.12.26 © Wilkinson, K.
Hemisotoma thermophila from Holland
2013.02.25 © van Duinen, J.
Hemisotoma thermophila from France
2013.03.31 © Sandra.
Hemisotoma thermophila from the UK
2013.04.10 © Jones, A.
Hemisotoma thermophila from Belgium
2016.10.17 © Janssens, F.

Species tribasiosetis (Potapov & Stebaeva, 1999) Rusek, J, 2002:307

Genus Micrisotoma Bellinger, PF, 1952:20
  Syn.: Anurophorinae Microisotoma(sic) Potapov, M et Babenko, A, 2014:9 ISS
Species achromata Bellinger, PF, 1952:20
Microisotoma(sic) achromata from Russia
After Potapov M & Babenko A, 2014 Fig.4C


Genus Proisotomodes Bagnall, 1949
Species bipunctatus (Axelson, WM, 1903:9) Rusek, J, 2002:307, t.t., B
Isotoma bipunctata from Poland
After Stach, J., 1947 Plate XXXIX Fig.1.

             Syn.: Cryptopygus bipunctata(sic) Poole, RW in Poole, RW et Gentili, P, 1997:50 ISS
Proisotomodes bipunctatus from Europe
With flattened sensilla
After Rusek, J, 2002 Fig.4
Proisotomodes bipunctatus from the UK
2012.11.15 © Murray, A.

Species debilis (Cassagnau, P, 1959:84) Potapov, M & Babenko, A, 2014:7

Genus Cylindropygus Deharveng, Potapov & Bedos, 2005
Species ferox Deharveng, Potapov & Bedos, 2005

Genus Isotominella Delamare Deboutteville, 1948
Species geophila Delamare Deboutteville, 1948
Isotominella-geophila from Algeria
After Jordana R, Hamra-Kroua S & Baquero E, 2009 Fig.2A

Species pilosa Delamare Deboutteville, 1948

Genus Dagamaea Yosii, 1965
Species danieli Nosek, J, 1974:397
Species fragilis Yoshii, 1995
Species morei Tamura, 2002
Species tenuis (Folsom JW in Mills, HB, 1934:46) habitus lateral (fig. B)
            Syn.: Dagamaea japonica Yosii, 1965
            Syn.: Dagamaea japonica Yosii, R, 1977:154
Dagamaea tenuis from the USA
2016.12.29 © Abela, A.


Genus Aggressopygus Potapov, M et Babenko, A, 2014:7
Species armatus Potapov, M et Babenko, A, 2014:11
Aggressopygus armatus from Russia
After Potapov M & Babenko A, 2014 Fig.4B
            Syn.: Micrisotoma sp.1 Potapov, 2002

Species sibiricus Potapov, M et Babenko, A, 2014:8, t.t.
Aggressopygus sibiricus from Russia
After Potapov M & Babenko A, 2014 Fig.4A


Genus Arlea Womersley, H, 1939:319 key
  Syn.: ... Falcisotomina Stach, 1947
Species adetolai de Mendonça, MC, Abrantes, EA et Fernandes, LH, 2006:58
Species arenicola Abrantes, EA et de Mendonça, MC, 2005:936
Species caeca Rapoport & Rubio, 1968
Species judithnajtae Bernard, EC, 2017:88
Arlea judithnajtae from the USA
After Bernard EC, 2017 Fig.1A

Species lucifuga (Arlé, 1939) Womersley, H, 1939:319, t.t.
Species psammophila de Mendonça, MC, Abrantes, EA et Fernandes, LH, 2006:57
             Syn.: Arlea psamophila(sic) Abrantes, EA et de Mendonça, MC, 2007:55 ISS
Species spinisetis Mendonça & Arlé, 1987
Species tridens Barra, J-A, 1997:472

Genus Gressittacantha Wise, 1967
Species terranova Wise, 1967
Gressittacantha terranova from Antarctica
2008 © Univ of Waikato.


Genus Rhodanella Salmon, JT, 1945:69
Species fasciata (Carpenter, 1912), t.t.
            Syn.: Isotomina XII-oculata(sic) Carpenter, 1919 IOS
             Syn.: Isotomina 12-oculata(sic) Stach, 1947 ISS
             Syn.: Parafolsomia duodecemoculata(sic) Salmon, JT, 1964:359 ISS
             Syn.: Isotoma (Isotomina xii-oculata(sic) Lawrence, PN, 1978:266 ISS
            Syn.: Vertagopus minos Denis, 1928
Rhodonella minos ♂
After Chimitova A & Potapov M, 2011 Fig.7

            Syn.: Proisotoma stachi Goto, 1957


Genus Isotomodes Axelson, 1907 key
Species alavensis Simón, Luciáñez, Ruiz & Martin, 1995
Species alexius Palacios-Vargas, JG et Kovác, L, 1995:502
            Syn.: Isotomodes sp. Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:10
Species armatus Naglitsch, 1962
Species bahiensis Rapoport, 1962
Species bisetosus Cassagnau, P, 1959:84
Species buchardi Thibaud, J-M, 2008:516
Species cariocus Thibaud, J-M et Palacios-Vargas, JG, 1999:29
            Syn.: Isotomodes carioca(sic) Thibaud, J-M et Palacios-Vargas, JG, 1999:29 IOS
             Syn.: Isotomodes carioca(sic) Culik, MP et Zeppelini, DF, 2003:1125 ISS
             Syn.: Isotomodes carioca(sic) Abrantes, EA et de Mendonça, MC, 2007:55 ISS
             Syn.: Isotomodes carioca(sic) Thibaud, J-M, 2007:185 ISS
             Syn.: Isotomurus carioca(sic) da Rocha Neves, AC et de Mendonça, MC, 2014:1 ISS
             Syn.: Isotomodes carioca(sic) Arbea, JI et Kahrarian, M, 2015:82 ISS
Species cassagnaui Izarra, 1969
Species ccoccatirensis Winter, 1963
Species cuzcoensis Winter, 1963
Species dagamae Prabhoo, 1971
Species denisi Folsom, JW, 1932:59
Species falsus Christiansen & Bellinger, 1980
Species fernandesae Abrantes, EA et de Mendonça, MC, 2007:?
Species fiscus Christiansen & Bellinger, 1980
Species gamae Izarra, 1971
             Syn.: Isotomodes gammae(sic) Mari Mutt & Bellinger, 1990 ISS
Species gisini da Gama, 1963
Species gredensis Lucianez & Simón, 1989
Species ibanezi Simón & Lucianez, 1989
Species klostermani Bernard, EC, 1971:125
Species korkorensis Arbea, JI et Kahrarian, M, 2015:75
Isotomodes korkorensis from Iran
After Arbea JI & Kahrarian M, 2015 Fig.17-18

Species maroccanus Stach, 1947
Species martae Simón, Luciáñez, Ruiz & Martin, 1995
Species productus (Axelson, WM, 1906:11) Linnaniemi, 1907, t.t., B
            Syn.: Isotoma elongata Axelson, WM, 1903:6
            Syn.: Isotomodes britannicus Bagnall, 1939
            Syn.: ... maroccanus Stach, 1947 =?
Isotomodes productus from New Zealand
After Salmon, J.T., 1941 Pl.51 Fig.189.
Isotomodes productus from Poland
After Stach, J., 1947 Plate XIII Fig.1.

             Syn.: Isotomodes producta(sic) Poole, RW in Poole, RW et Gentili, P, 1997:52 ISS
Species pseudoproductus Stach, J, 1965:352
Species quadrisetosus da Gama, 1963
Species rafaeli Arbea, JI, 2006:48
Species rosae Arbea, JI, 2006:46
Species sexsetosus da Gama, 1963
  Subspecies provincialis Poinsot, 1966
Species sotoensis Simón, Luciáñez, Ruiz & Martin, 1995
Species subarmatus Jordana, R et Arbea, JI, 1990:423
Species templetoni Bagnall, 1939
Species trisetosus Axelson, 1907
            Syn.: Isotomodes plurisetosus Denis, 1923
Species venezuelensis Rapoport & Maño, 1969
             Syn.: Isotomodes vanezuelensis(sic) González, V, Díaz, M et Palacios-Vargas, JG, 1997:46 ISS
Species xishaensis Chen, 1986

Genus Jesenikia Rusek, J, 1997:115
Species filiformis Rusek, J, 1997:117, t.t.
Jesenikia filiformis from Czechia
After Rusek J, 1997 Fig.3A

Species sensibilis (Cassagnau, P, 1959:80) Potapov, 2001

Genus Pseudanurophorus Stach, 1922
Species acuta Denis, 1932
Species arcticus Christiansen, KA, 1952:131
            Syn.: Pseudanurophorus arctica(sic) Christiansen, KA, 1952:131 IOS
Pseudanurophorus arctica(sic) from Alaska
After Christiansen K, 1951 Pl.11 Fig.3

             Syn.: Pseudanurophorus arctica(sic) Skidmore, RE, 1995:54 ISS
Species binoculatus (Kseneman, 1934) Christiansen, 1951:134, B
Species boerneri Stach, 1922, t.t.
Species cassagnaui Winter, 1963
Species hissaricus Martynova, 1971
Species isotoma (Börner, 1903) Kseneman, 1934
  Forma acuta Denis, 1932
Species lapponicus Agrell, 1939
Species mobilis Potapov, 1997
Species montanus Martynova, 1971
Species octoculatus Martynova, 1971
Species quadrioculatus von Törne, 1955

Genus Isotomodella Martynova, 1968
Species alticola (Bagnall, 1949) Potapov, MB & Stebaeva, SK, 2002:?
            Syn.: Pseudanurophorus binoculatus var. inoculatus Bödvarsson, H, 1957:19
            Syn.: ... sensibilis Cassagnau, 1959 =?
             Syn.: Isotomodella alticolus(sic) Potapov, MB et Stebaeva, SK, 2002:?,gr ISS
             Syn.: Isotomodella articola(sic) Potapov, MB et Stebaeva, SK, 2002:? ISS
Species asiatica (Martynova, 1968) Potapov, MB & Stebaeva, SK, 2002:?
Species microlobata Potapov, MB et Stebaeva, SK, 2002:?
             Syn.: Isotomodella microlabata(sic) Potapov, MB et Stebaeva, SK, 2002:? ISS
Species psammophila Potapov, MB et Stebaeva, SK, 2002:?
Species pusilla Martynova, 1968
Species zhejiangensis (Chen, 1985) Potapov, MB & Stebaeva, SK, 2002:?

Genus Anurophorus Nicolet, 1842, n.c.
  Syn.: Aphoruridæ(sic) Bourletia MacGillivray, AD, 1893:313
  Syn.: Isotomidae Anrophorus(sic) Christiansen, KA in Dindal, DL, 1990:987 ISS
Remark by ICZN. 2007:52: Placed on the Official List of Names in Zoology. Opinion 439.

Anurophorus sp. from the USA
2009.11.21 © Thombs, D.
Anurophorus sp. from the USA
2013.05.12 © Gorring, P.

Species alpinus Potapov & Stebaeva, 1990
Species altus Christiansen & Bellinger, 1980

Anurophorus asfouri from Lebanon
After Christiansen, K., 1958 Pl.8 Fig.1.
Species asfouri Christiansen, 1958:60

Species atlanticus Fjellberg, 1974
Species balcellsi Selga, 1959
Species barroudensis Cassagnau, P, 1959:78
Anurophorus barroudensis from France
2008.02.07 © Lavoué, Br.
Det.: Arnaud, J.-P.

Species bimus Christiansen & Bellinger, 1980
Species changjiensis Hao & Huang, 1995
Species chukoticus Potapov & Stebaeva, 1990
Species cinereus Potapov, MB, 1997:18
Species clavipilus Stach, 1947
Species coiffaiti Cassagnau, P et Delamare Deboutteville, C, 1955:377
            Syn.: Anurophorus Coiffaiti(sic) Cassagnau, P et Delamare Deboutteville, C, 1955:377 IOS
            Syn.: Anurophorus silvaticus from Azerbaijan =?
             Syn.: Anurophorus sylvaticus(sic) Falahati HosseinAbad, A, Potapov, M, Sarailoo, MH, Mehr, MS et Yazdanian, M, 2013:237 ISS
Anurophorus coiffaiti (bar=2µm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.4A

Species continentalis Dunger, WG, 1982:43
Species cuspidatus Stach, 1920
Species duodecimoculatus Steiner, 1958
Species elongatus Fjellberg, 1984
Species eximius Potapov, MB, 1997:12
Species fulvus Fjellberg, A, 1988:39
Species fjellbergi Potapov, MB, 1997:27
Species ganghwaensis Lee, B-H, 1977:155
             Syn.: Anurophorus ganghwaiensis(sic) Lee, B-H, Kim, B-J et Kim, J-T, 1993:290 ISS
Species konseli Kseneman, 1936
             Syn.: Anurophorus Konseli(sic) Cassagnau, P, 1953:156 ISS
Species koreanus Potapov, MB, 1997:8
Species laricis Nicolet, H, 1842:53, t.t., B
Anurophorus laricis from Switzerland
After Nicolet, H., 1842 Pl.4 Fig.3.

            Syn.: Adicranus corticinus Bourlet, 1842
Lipura corticina from the UK
After Lubbock J, 1873 Pl.45 Fig.1

            Syn.: Anurophorus laricis var. pallida Absolon, K, 1901:216
             Syn.: Anurophorus Laricis(sic) Oudemans, JT, 1888:151 ISS
             Syn.: Anurophorus Laricis(sic) Lameere, A, 1895:595,B ISS
Anurophorus laricis from Poland
After Stach, J., 1947 Plate VII Fig.1.
Anurophorus laricis from the UK
2006.04.28 © Storey, M.
Anurophorus laricis from Germany
2008.03.14 © Peters, J.
Anurophorus laricis from France
2009.04.01 © Ramel, A.
Anurophorus laricis from the UK
2011.03.22 © Michaelson, J.
Anurophorus laricis from Holland
2011.07.09 © van Duinen, J.
Anurophorus laricis from Holland
Abdominal segment 6 very short
2011.07.09 © van Duinen, J.
Anurophorus laricis from the UK
2013.04.23 © Michaelson, J.
Anurophorus laricis from the UK
2013.04.23 © Michaelson, J.
Anurophorus laricis from Holland
2014.02.04 © van Duinen, J.
Anurophorus laricis from Holland
2014.03.22 © Baas, A.H.
Anurophorus laricis from Germany
2015.02.15 © Kluck, A.
Anurophorus laricis from Germany
2015.02.22 © Kluck, A.
Anurophorus laricis from the UK
2015.12.26 © Barton, T.

Species lohi Christiansen, K et Bellinger, P, 1992:117
Species lydiae Lucianez & Simón, 1989
Species massoudi Poinsot, 1970
Species mongolicus Dunger, WG, 1982:45
Species montanus Martynova, 1968
Species nitrophilus Potapov, MB, 1997:29
Species olympicus Potapov, MB, 1997:5
Species oredonensis Cassagnau, P, 1953:155
Species orientalis Potapov & Stebaeva, 1990
Species pacificus Potapov, MB, 1997:22
Species palearcticus Potapov, MB, 1997:32
Species pseudolaricis Loksa, 1978
Species racovitzai Denis, 1932
Species rarus (Yosii, 1939)
Species satchelli Goto, 1956
Species scheueri Christiansen & Bellinger, 1980
Species senex Fjellberg, 1984
Species sensibilis Potapov, MB, 1997:7
Species septentrionalis Palissa, 1966
Species septentrionalis2 Christiansen, K et Bellinger, P, 1980:535
Anurophorus (Anurophorus) septentrionalis from the USA
After Christiansen K & Bellinger P, 1980 Fig.462A
            Syn.: Anurophorus laricis Guthrie, JE, 1903:98
            Syn.: Anurophorus laricis Folsom, 1937
Anurophorus laricis from the USA
After Maynard EA PlateI Fig.6
Anurophorus laricis from the USA
After Maynard EA PlateIX Fig.146

Anurophorus nr septentrionalis from the USA
2016.06.12 © Eiseman, C.
Anurophorus nr septentrionalis from the USA
Abd.5+6 fused
2016.06.12 © Eiseman, C.
Anurophorus nr septentrionalis from the USA
Eyepatch with 8 ocelli
2016.06.12 © Eiseman, C.

Species serratus Deharveng, 1976
Species silvaticus Potapov & Stebaeva, 1990
Species sorosi Potapov, MB, 1997:9
Species spinosus Dallai, R, 1971:342
Species stepposus Potapov & Stebaeva, 1990
Species szeptyckii Potapov, MB, 1997:18
Species trisensillus Potapov, MB, 1997:11
Species unguiculus Bagnall, 1940
Species ursi Potapov & Stebaeva, 1990
Species utahensis (Wray, 1958) Poole, RW in Poole, RW & Gentili, P, 1997:50

Genus Micranurophorus Bernard, EC, 1977:75 key
  Syn.: Isotomidae Micraurophorus(sic) Petersen, H, 2011:475 ISS
Species musci Bernard, EC, 1977:75, t.t.
            Syn.: Micranurophorus schalleri Christian, E, 1986:121
Species valdelatensis Simón-Benito, JC, Luciáñez Sánchez, MJ et M Ruiz Ortega, 1996:48

Genus Mucrosomia Bagnall, 1949
  Syn.: Anurophorinae Bathyterra Rusek, J, 1996:207
Species alticola de Mendonça, MC et Queiroz, GC, 2013:217
Species bipartita (Rusek, J, 1996:207) Potapov, M, 2001:359
Species caeca (Wahlgren, E, 1906:12) Potapov, M, 2001:359
            Syn.: Folsomia fimetaroides Womersley, 1934 excluded from Australian fauna
            Syn.: Parafolsomia litorea Salmon, JT, 1949:20
            Syn.: Isotomina nerudai Rapoport, EH et Rubio, IR, 1963:120
Isotomina nerudai from Chile
After Rapoport EH & Rubio IR, 1963 Fig.75
Isotomina nerudai from Chile
Chaetotaxy of head + mesothorax
After Rapoport EH & Rubio IR, 1963 Fig.76

            Syn.: Isotomina zenderi Winter, 1963 nn

Species garretti (Bagnall, 1939 tt) Bagnall, 1949, B
            Syn.: Isotomina caeca Gisin, 1960, nec Wahlgren, 1906
             Syn.: Cryptopygus coecus(sic) Fratello & al., 1989 ISS
            Syn.: Cryptopygus gisini Massoud & Rapoport, 1968
             Syn.: Cryptopygus garetti(sic) Dunger, W et Zivadinovic, J, 1989:1 ISS
Cryptopygus garretti from the UK
Post-antennal organ
2005 © Hopkin, S.P.


Genus Haploisotoma Izarra, 1965
Species ventanensis Izarra, 1965, t.t.

Genus Pectenisotoma Gruia, M, 1983:193
Species theodori Gruia, M, 1983:193, t.t.

Genus Yosiiella Hüther, 1967
Species mira Hüther, 1967, t.t.

Genus Archisotoma Linnaniemi, WM, 1912:118 key
  Syn.: Collembola Isotama(sic) Childs, GH, 1915:39 ISS
Species sp.1 Arbea, JI et Ariza, E, 2012:206
Species sp.2 Arbea, JI et Ariza, E, 2012:206
Species arariboia da Rocha Neves, AC et de Mendonça, MC, 2014:2
Species besselsi (Packard, AS, 1877:52) Linnaniemi, WM, 1912:119, t.t.
            Syn.: Isotoma Besselsii(sic) Packard, AS, 1877:52,tt IOS
            Syn.: Isotoma pulchella Moniez, 1891
            Syn.: Isotoma spitzbergensis(sic) Lubbock, 1898 IOS
             Syn.: Isotoma spitzbergensis(sic) Carpenter, GH, 1907:474 ISS
            Syn.: Isotoma arctica Scherbakow, 1899
            Syn.: Isotoma janmayensis Wahlgren, 1900
            Syn.: Isotoma bidenticula Bacon, 1912
            Syn.: Isotoma klovstadi Enderlein, 1912
            Syn.: Proisotoma laguna Folsom, JW, 1937:118
             Syn.: Isotoma Besselsii(sic) MacGillivray, 1891 ISS
             Syn.: Isotoma Besselsii(sic) MacGillivray, 1896 ISS
             Syn.: Isotoma Besselsi(sic) Willem, 1902 ISS
             Syn.: Isotoma Besselsii(sic) Davenport, 1903 ISS
             Syn.: Isotoma Beselsii(sic) Imms, AD in Herdman, WA, 1906:77 ISS
             Syn.: Isotoma Beselsii(sic) Carpenter, GH, 1907:474 ISS
             Syn.: Isotoma besselsii(sic) Nielsen, 1909 ISS
             Syn.: Isotoma besselsii(sic) Bacon, 1912 ISS
             Syn.: Isotoma besselsii(sic) Bacon, 1914 ISS
             Syn.: Isotama(sic) besselsii(sic) Childs, GH, 1915:39 ISS
             Syn.: Archisotoma beselsi(sic) Carpenter & Philips, 1922 ISS
             Syn.: Archisotoma Besselsi(sic) Denis, JR, 1924:197 ISS
             Syn.: Archisotoma Besselsi(sic) Willem, V, 1925:617 ISS
             Syn.: Archisotoma beselsi(sic) Carpenter, 1927 ISS
             Syn.: Archisotoma Besselsi(sic) van den Bruel, WE, 1945:38 ISS
             Syn.: Archistoma(sic) lesselsi(sic) Obenberger, J, 1955:95 ISS
             Syn.: Archisotoma besselsii(sic) Strenzke, 1958 ISS
             Syn.: Archisotoma besselsii(sic) Arlé, 1959 ISS
             Syn.: Archisotoma besselsii(sic) Hepburn & Ross, 1964 ISS
             Syn.: Archisotoma besselsii(sic) Salmon, JT, 1964:347 ISS
             Syn.: Archisotoma besselsii(sic) Palacios-Vargas, 1983 ISS
             Syn.: Archisotoma besselsii(sic) Christiansen & Bellinger, 1988 ISS
             Syn.: Archisotoma besselsii(sic) Mari Mutt, JA et Bellinger, PF, 1990:73,is ISS
             Syn.: Archisotoma besselsii(sic) Palacios-Vargas & Gómez-Anaya, 1994 ISS
  Subspecies uniseriata Strenzke, 1955
Species bothrilongaequalis Thibaud & Ndiaye, 2006
Species brucei (Carpenter, GH, 1907:474) Womersley, H, 1936:321
            Syn.: Isotoma Brucei(sic) Carpenter, GH, 1907:474 IOS
Isotoma Brucei(sic) from Antarctica
After Carpenter GH, 1907 Plate I Fig.1

             Syn.: Archisotoma Brucei(sic) Delamare Deboutteville, C, 1948:40 ISS
             Syn.: Archisotoma Brucei(sic) Cassagnau, P, 1957:394 ISS
Archisotoma brucei from Antarctica
After Carpenter, 1907 in Greenslade, P., 2006 Fig.61.1.

Species catiae Abrantes, EA et de Mendonça, MC, 2007:55
Species gourbaultae Thibaud, 1993
            Syn.: Archisotoma interstialis(sic) Palacios-Vargas, 1997
             Syn.: Archisotoma gorbaultae(sic) da Rocha Neves, AC et de Mendonça, MC, 2014:2 ISS
Species heraultensis Christian, E et Thibaud, J-M, 1996:94
             Syn.: Archisotoma heraultae(sic) Christian & Thibaud, 1996 ISS
Species interstitialis Delamare Deboutteville, 1953
            Syn.: Archisotoma Renaudi(sic) Delamare Deboutteville, 1956 IOS
            Syn.: Archisotoma xavieri Cardoso, MA, 1966:84
            Syn.: Archisotoma utinomii Yosii, R, 1977:154 =?
             Syn.: Archisotoma intertialis(sic) Selga, 1960 ISS
             Syn.: Archisotoma interstialis(sic) Poinsot, N, 1972:677 ISS
Species litoralis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:81
Species madagascariensis Thibaud, J-M, 2008:513
Species martae Fjellberg, A et Jucevica, E, 2000:21
Species mayotti Thibaud, 2011
Species megalops Bagnall, 1939
            Syn.: Archisotoma subbrucei Delamare Deboutteville, 1953
Species nigricans Bagnall, 1939
Species pauliani Delamare Deboutteville, 1953
             Syn.: Archisotoma pauliana(sic) Salmon, JT, 1964:348 ISS
Species poinsotae da Gama, 1968
Species polaris Fjellberg & Poinsot, 1975
Species pulchella (Moniez, 1890) Denis, 1923, B
             Syn.: Archisotoma pullchella(sic) Thibaud, J-M et Palacios-Vargas, JG, 2001:350 ISS
Species quadrioculata Fjellberg, A, 1988:40
Archisotoma quadrioculata from Norway
After Fjellberg, A, 1988 Fig.4A

Species sabulosa Barra, J-A, 1997:474
Species senegalensis Thibaud & Ndiaye, 2006
Species subtheae Thibaud & Boumezzough, 2006
Species theae Fjellberg, 1980
Species utinomii Yosii, 1971
            Syn.: Archisotoma tokiokai Yosii, 1971
Species vaoensis Thibaud, J-M et Weiner, WM in Najt, J et Matile, L, 1997:81
Species vareli Sterzynska & Ehrnsberger, 2000

Genus Folsomia Willem, 1902 key
  Syn.: Isotomidae Folsoma(sic) Poinsot, N, 1972:682 ISS
Remark by ICZN. 2007:322: Placed on the Official List of Names in Zoology. Opinion 1431.

Folsomia sp. from Holland
2012.08.31 © van Duinen, J.
Folsomia sp. from the UK
2012.11.27 © Murray, A.
Folsomia candida from the UK
Abdominal segments 4-6 fused
2014.01.25 © Scyrene., B
Folsomia sp. from Canada
Narrow elliptic post-antennal organ (PAO)
2016.02.06 © Hochhalter, R.
Folsomia sp. from Spain
2016.02.18 © Baquero, E.

Species sp.1 Potapov, MB, 2000:235
Species sp.2 Potapov, MB, 2000:235
Species sp.3 Potapov, MB, 2000:235
Species sp.4 Potapov, MB, 2000:235
Species sp.5 Potapov, MB, 2000:235
Species sp.6 Potapov, MB, 2000:235
Species sp.7 Potapov, MB, 2000:235
Species sp.8 Potapov, MB, 2000:236
Species sp.9 Potapov, MB, 2000:236
Species sp.10 Potapov, MB, 2000:236
Species sp.11 Potapov, MB, 2000:236
Species sp.12 Potapov, MB, 2000:236
Species sp.13 Potapov, MB, 2000:236
Species sp.14 Potapov, MB, 2000:236
Species sp.15 Potapov, MB, 2000:236
Species sp.16 Potapov, MB, 2000:236
Species sp.17 Potapov, MB, 2000:236
Species sp.18 Potapov, MB et Chimitova, A, 2009:55
Species sp.19 Potapov, MB et Chimitova, A, 2009:55
Species sp.20 Potapov, MB et Chimitova, A, 2009:55
Species sp.21 Potapov, MB et Chimitova, A, 2009:55
Species sp.22 Kahrarian, M et Arbea, J, 2013:92
Folsomia n.sp. from Iran
After Kahrarian M & Arbea J, 2013 Fig.5

Species sp.23 Potapov, M et Gulgenova, A, 2013:319
Species abrupta Potapov & Deharveng, 2000
Species achaeta Bagnall, 1939
Species agrelli Gisin, 1944
            Syn.: Folsomia quadrioculata f. 2-punctata(sic) Agrell, I, 1939:5 IOS
Species albens Kaprus, IJ et Potapov, MB, 1999:355
Species alpigena Stach, 1947
Species alpina Kseneman, 1936
Species altaica Martynova, 1974:18
Species altamontana Yosii, 1971
             Syn.: Folsomia altamomtana(sic) Berezina, OG et Potapov, MB, 2006:9 ISS
Species amazonae Winter, 1963
Species americana Stach, 1947
Species amplissima Potapov & Babenko, 2000
Species ancestor Potapov & Babenko, 2000
Species annamariae Winter, 1963
Species antricola Loksa, I, 1959:37
Folsomia antricola from Hungary
After Loksa I, 1959 Fig.1A

Species aquatica Wang & Wu, 2012
Species arena Potapov MB et Babenko A in Potapov, MB, 2006:16
Folsomia arena from Russia
After Potapov MB, 2006 Fig.19

Species asiatica Martynova, 1971
Folsomia asiatica from Iran
After Arbea JI & Kahrarian M, 2015 Fig.1

Species atropolaris Potapov & Babenko, 2000
Species australica Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:137
            Syn.: Folsomia loftyensis Potapov & Greenslade, 2010
Species baicalica Stebaeva & Potapov, 1998
Species baida Potapov, MB, 2006:12
Folsomia baida from Russia
After Potapov MB, 2006 Fig.2

Species baijali Prabhoo, 1971
Species bashkira Potapov MB et Kuznetsova N in Potapov, MB, 2006:14
Folsomia bashkira from Russia
After Potapov MB, 2006 Fig.4

Species bidentata Lee, 1974
             Syn.: Folsomia bidenetata(sic) Lee, B-H, Kim, B-J et Kim, J-T, 1993:290 ISS
Species binoculata (Wahlgren, 1899)
Species biparis Potapov, M et Gulgenova, A, 2013:305
Species bisetosa Gisin, 1953
Species bisetosella Fjellberg, A, 2005:111
Species bogojevicae Dunger W in Dunger, W et Zivadinovic, J, 1989:3
            Syn.: Folsomia diplophthalma Simon JC in Arbea, JI, 2016:426
Folsomia bogojevicae ♂ from Spain
After Arbea JI, 2016 Fig.1
Folsomia bogojevicae juv. from Spain
After Arbea JI, 2016 Fig.7

Species borealis Potapov & Babenko, 2000
Species brevicauda Agrell, 1939
Species brevifurca (Bagnall, 1949) Goto, HE & Lawrence, PN in Kloet, GS & Hincks, WD, 1964:7
             Syn.: Folsomia brevifurcata(sic) Loksa, I, 1964:96 ISS
Remark by Potapov, M. 2001:186: As was shown by Lawrence (1973), the species was described from a single juvenile, so it became a species dubia.

Species brevisensilla Potapov & Babenko, 2000
Species britannica Stach, 1947
Species candida Willem, V, 1902:280, t.t., B
            Syn.: Isotoma fimetaria var. caldaria Axelson, WM, 1905:790
            Syn.: Folsomia dubia Kinoshita, 1917
            Syn.: Folsomia dubia Kinoshita, 1932
            Syn.: Folsomia dubia Yosii, R, 1977:153
            Syn.: Folsomia fimetaria var. dentata Folsom, JW, 1927:7
            Syn.: Folsomia fimetaria Stach, 1929
            Syn.: Folsomia fimetaria Geyer & Mann, 1940
            Syn.: Folsomia fimetaria Bonet, F, 1931:380,B
            Syn.: Folsomia fimetaria Wolf, B, 1938:120,B
            Syn.: Folsomia fimetaria Leruth, R, 1939:199,B
            Syn.: Folsomia fimetaria Folsom, 1932
            Syn.: Folsomia fimetaria Womersley, 1934
            Syn.: Folsomia fimetaria Womersley, H, 1939:145
            Syn.: Folsomia fimetaria Yosii, 1939
            Syn.: Folsomia fimetaria Uchida, 1940
            Syn.: Folsomia fimetaria Uchida, 1952
            Syn.: Folsomia fimetaria Yosii, 1955
            Syn.: Entomobrya emeraldica Rayment, 1937
            Syn.: Folsomia distincta Bagnall, 1939
            Syn.: Folsomia kingi Bagnall, 1939
            Syn.: Folsomia litsteri Bagnall, 1939
             Syn.: Folsomia listeri(sic) auctt. ISS
             Syn.: Folsomia listeri(sic) Christiansen & Tucker, 1977 ISS
             Syn.: Folsomia listeri(sic) Szeptycki & Weiner in Razowski, 1990 ISS
             Syn.: Folsomia litseri(sic) Hopkin, 1997 ISS
             Syn.: Folsomia listeri(sic) Irmler, U, 2006:54 ISS
             Syn.: Folsomia listeri(sic) Auclerc, A, Ponge, JF, Barot, S et Dubs, F, 2009:1599 ISS
Remark by Potapov, M. 2001:188: According to the accurate study of types by Lawrence (1973), F. litsteri, F. distincta and F. kingi (described as three independent species) are the growth stages of this large-sized species.


            Syn.: Folsomia fontis Yosii, 1953
            Syn.: Folsomia fontis Yosii, R, 1977:153
            Syn.: Folsomia cavicola Cassagnau, P et Delamare Deboutteville, C, 1955:381
            Syn.: Folsomia abchasica Martynova, 1964
            Syn.: Folsomia candida var. indistincta Green, 1964
            Syn.: Folsomia lawrencei Rusek, 1984
Folsomia candida from the UK
2003 © Hopkin, S.P.
Folsomia candida from the UK
2005 © CEH.
Folsomia candida from the UK
2005 © CEH.
Folsomia candida from the UK
2006.04.08 © Read, D.
Folsomia candida from Germany
2006.12.26 © Kohl, F.
Folsomia candida from Germany
2007.12.14 © Pageler, J.
Folsomia candida from Germany
2007.12.14 © Pageler, J.
Folsomia candida from France, juvenile
2007 © Raveton, M.
Remark by Dányi, L. & Traser, G. 2008:47: Data of Stach (1929) and Geyer & Mann (1940) were corrected by Loksa (1960:49).

Folsomia candida from France
After Tully, T., 2008 ©
First instar 0.35mm
Folsomia candida from Spain
After Baquero, E., 2009.05.22 ©
In laboratory culture
Folsomia candida from the USA
2011.03.02 © Justis, S.
Folsomia candida from the UK
2012.09.05 © Murray, A.
Folsomia candida from the UK
With brown gut
2013.01.26 © Murray, A.
Folsomia candida from the UK
Antenna
2013.01.26 © Murray, A.
Folsomia candida from the UK
2014.01.25 © Scyrene., B
Folsomia candida from Holland
2014.10.21 © van Duinen, J.

Species checae Lucianez & Simon, 2003
Species ciliata Babenko & Bulavintsev, 1993
Species clavipila Cassagnau, P, 1954:619
Species coeruleogrisea (Hammer, 1938) Fjellberg, A, 2007:35
             Syn.: Cryptopygus coeruleogrisea(sic) Poole, RW in Poole, RW et Gentili, P, 1997:50 ISS
Species continentalis Potapov & Dunger, 2000
Species cryptophila Potapov & Babenko, 2000
             Syn.: Folsomia (fimetaria-group) cryptopila(sic) Potapov, MB et Chimitova, A, 2009:55 ISS
Species culter Potapov, MB et Chimitova, A, 2009:49
Folsomia culter from Russia
After Potapov MB & Chimitova A, 2009 Fig.1

Species decaxiophthalma Ford, 1962
Species decemoculata Stach, J, 1946:29
             Syn.: Folsomia 10-oculata(sic) Parisi, V, 2007:22 ISS
Species decophthalma Steiner, 1958
Species decopsis Steiner, 1958
Species diplophthalma (Axelson, 1902) Axelson, 1907, B
            Syn.: Isotoma cacuminum Wahlgren, 1906
            Syn.: Isotoma binoculata Collinge & Shoebotham, 1909, nec Wahlgren, 1899
            Syn.: Folsomia diplophthalma var. caeca Kseneman, M, 1935:12
            Syn.: Folsomia diplophthalma var. pallida Kseneman, M, 1938:462
            Syn.: Folsomia diplophthalma f. 4-punctata(sic) Agrell, I, 1939:13 IOS
            Syn.: Folsomia diplophthalma ab. 2-punctata(sic) Agrell, I, 1939:7 IOS
            Syn.: Folsomia diplophthalma f. anocellata Maynard, EA, 1951:87
             Syn.: Folsomia diplopthalma(sic) Salmon, JT, 1943:73 ISS
             Syn.: Folsomia diplophtalma(sic) Jacobs, W, 1997:71,B ISS
Folsomia diplophthalma from Japan
After Suetsugu N, Satoh F & Koike T, 2010 Photo16

Species dovrensis Fjellberg, 1976
Species duodecimoculata Ford, 1962
            Syn.: Folsomia hoffi Scott, HG, 1961:264 partim holotype
             Syn.: Folsomia duodecimaculata(sic) Christiansen & Tucker, 1977 ISS
Species echinosetosa Stebaeva, 1978
Species elongata (MacGillivray, 1896) James, 1933
            Syn.: Isotoma bidenticula Guthrie, JE, 1903:66
             Syn.: Folsomia elongate(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:110 ISS
Species evansi Fjellberg, 1984
Species fimetaria (Linnæus, C, 1758:609) Willem, V, 1925:626, B
            Syn.: Podura terrestris alba Linnæus, C, 1746:343
            Syn.: Podura fimaria Linnæus, C, 1748:67
             Syn.: Podura Terrestris(sic) Scopoli, JA, 1763:379 ISS
            Syn.: Isotoma alba Tullberg, 1871
            Syn.: ... nitida Becker, 1902
            Syn.: Isotoma splendes Becker, 1902
            Syn.: Folsomia dubia Kinoshita, 1917
Isotoma fimetaria from Europe
After Börner in Escherich K, 1923 Abb.1a
Folsomia fimetaria from the UK
Feeding instar showing gut contents
2012.12.09 © Murray, A.
Folsomia fimetaria from Czechia
Feeding instar showing gut contents
2015.02.13 © Deml, M.

Species fimetarioides (Axelson, WM, 1903:8) Handschin, 1929:59, B
             Syn.: Folsomia fimetariodes(sic) Hopkin, 1997 ISS
Species fumosa Potapov, M et Gulgenova, A, 2013:313
Folsomia fumosa from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.11

Species gebhardti Loksa, I, 1964:95
Species giustii Dallai, R, 1970:451
Species hasegawai Yosii, R, 1959:117
Species heterocellata Stebaeva & Potapov, 1998
Species heterothorax Potapov, M et Gulgenova, A, 2013:309
Folsomia heterothorax from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.2

Species hexasetosa Lee, B-H, 1977:157
             Syn.: Folsomia hexasetasa(sic) Lee, B-H, Kim, B-J et Kim, J-T, 1993:290 ISS
Species hidakana Uchida, H et Tamura, H, 1968:7
Species highlandia Wray, DL, 1950:95
Species hissarica Martynova, 1971
Species hodgei Maynard, EA, 1951:86
Species hoffi Scott, HG, 1961:264 partim paratypes
            Syn.: Proisotoma hoffi Scott, 1962
Species hrabei Rusek, 1984
Species hubeiensis Ding, Huang & Chen, 2006
Species imadatei Tamura & Zhao, 2000
Species inoculata Stach, J, 1946:34
            Syn.: Folsomia ezoensis Yosii, 1965
            Syn.: Folsomia ezoensis Yosii, R, 1977:154
            Syn.: Folsomia spinosa Zivadinovic, 1973
Folsomia inoculata from the UK
Post-antennal organ
2004 © Hopkin, S.P.
Folsomia inoculata from the UK
Basal dental ridges toothlike extended
2008.04.04 © Shaw, P.
Folsomia inoculata from the UK
2+2 anterior manubrial setae
2008.04.04 © Shaw, P.

Species kerni Gisin, 1948
Species kirgisica Martynova, 1973
Species ksenemani Stach, 1947
            Syn.: Folsomia pseudodiplophthalma Stach, 1947
             Syn.: Folsomia pseudo(sic) Poinsot, N, 1972:680 ISS
Species kuramotoi Yoshii, 1995
Species kurushica Potapov, MB, Abdurakhmanov, GM et Aitekova, MM, 2001:?
Species kuznetsovae Potapov MB in Potapov, MB et Taskaeva, AA, 2009:1
Folsomia kuznetsovae from Russia
After Potapov MB, 2009 Fig.14
             Syn.: Folsomia kuznetsove(sic) Schulz, H-J et Potapov, M, 2010:63 ISS
Species longidens Potapov & Babenko, 2000
Species lunata Salmon, JT, 1943:73
Species macrochaetosa Martynova, 1977
Species macroseta Ford, 1962
Species magadani Martynova, 1977
Species manolachei Bagnall, 1939, Bhabitus lateral
            Syn.: Folsomia anophthalma Axelson, 1902
            Syn.: Isotoma quadrioculata var. pallida Axelson, WM, 1905:791
            Syn.: Folsomia nana Gisin, 1957
            Syn.: Folsomia diplophthalma Selga, 1971
            Syn.: Folsomia diplophthalma Selga D in Arbea, JI, 2016:426
            Syn.: Folsomia diplophthalma Steiner W in Arbea, JI, 2016:426
            Syn.: Folsomia spinosa Zivadinovic, 1973
             Syn.: Folsomia manobechei(sic) Bagnall, 1939 ISS
             Syn.: Folsomia manobechei(sic) Gisin, H, 1960:178 ISS
             Syn.: Folsomia monobechei(sic) Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:7 ISS
             Syn.: Folsomia monolachei(sic) Jordana, R, 1980:105 ISS
             Syn.: Folsomia monolachei(sic) Christian, E, 1985:143 ISS
             Syn.: Folsomia manolacheri(sic) Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:897 ISS
Remark by Dányi, L. & Traser, G. 2008:48: The [Hungarian] data of Folsomia quadrioculata (Tullberg) reported by Dunger (1975) included "Folsomia nana Gisin" [= Folsomia manolachei] as well.

Folsomia manolachei from the UK
2012.12.08 © Murray, A.
Folsomia manolachei from the UK
2013.02.02 © Nurcombe, E.
Folsomia manolachei from the UK
2013.02.02 © Nurcombe, E.
Folsomia manolachei from the UK
Eye with 2 distinctly separated ocelli
2013.02.02 © Nurcombe, E.

Species martynovae Potapov, 2001
Folsomia duodecimoculata from Ukraine
After Kaprus IJ, 1997 Fig.1A

Species maximovi Potapov, M et Gulgenova, A, 2013:314
Species microchaeta Agrell, I, 1939:5
            Syn.: Folsomia microchaeta ab. 2-punctata(sic) Agrell, I, 1939:6 IOS
            Syn.: Folsomia microchaeta f. 4-punctata(sic) Agrell, I, 1939:5 IOS
Species mikhaili Jordana R et Baquero E in Baquero, E, Arbea, J et Jordana, R, 2010:299
Species minipunctata Zhao & Tamura, 1992
Species minorae Potapov, M, Janion-Scheepers, C et Deharveng, L, 2017:133
Folsomia minorae from New Zealand
After Potapov M, Janion-Scheepers C & Deharveng L, 2017 Fig.1

Species minuta Lee, B-H et Kim, J-T, 1994:211
Species miradentata Salmon, JT, 1943:74
Species mofettophila Schulz, H-J et Potapov, M, 2010:60
Folsomia mofettophila from Czechia
After Schulz H-J & Potapov M, 2010 Fig.5

Species monoculata (Bagnall, 1949) Goto & Lawrence in Kloet & Hincks, 1964:7
Species monophthalma Bagnall, 1939
Species montana Martynova, 1971
Species montigena Stach, J, 1946:32
Species najtae Potapov, MB, Fjellberg, A et Bokova, A, 2017:96
Folsomia najtae ♀ from Far East Russia
After Potapov MB, Fjellberg A & Bokova A, 2017 Fig.1A
Folsomia najtae juv. from Far East Russia
After Potapov MB, Fjellberg A & Bokova A, 2017 Fig.2A
Folsomia najtae juv. from Far East Russia
Mobile form
After Potapov MB, Fjellberg A & Bokova A, 2017 Fig.2B

Species nakajimai Yosii, R, 1959:116
Species nanjingensis Ding, Ji & Chen, 2006
Species nigromaculata Najt, 1981
Species nivalis (Packard, AS, 1873:31) Scott, HG, 1961:265
            Syn.: Folsomia duodecimsetosa Hammer, 1953
Species obscurocellata Potapov & Cassagnau, 2000
Species ocellata Jordana, R, 1980:101
Species octoculata Handschin, 1925
            Syn.: Folsomia uenoea Uchida, 1943
            Syn.: Folsomia uenoea Yosii, R, 1977:154
            Syn.: Parafolsomia trioculata Salmon, 1956
             Syn.: Folsomia octooculata(sic) Potapov, MB, 2000:235 ISS
             Syn.: Folsomia octooculata Potapov, MB, Fjellberg, A et Bokova, A, 2017:96 ISS
Species orientalis Martynova, 1977
Species ozeana Yosii, 1954
            Syn.: Folsomia diplophthalma Yosii, 1939
             Syn.: Folsomia oézana(sic) Lee, B-H et Kim, J-T, 1994:214 ISS
Species palaearctica Potapov & Babenko, 2000
            Syn.: Folsomia diplophthalma auct., nec Axelson, 1902
Species paoinflata Potapov MB et Stebaeva S in Potapov, MB, 2006:18
Species penicula Bagnall, 1939
            Syn.: Folsomia diplophthalma var. tetrophthalma Kseneman, 1934
            Syn.: Folsomia tetrophthalma Gisin, 1946
            Syn.: Folsomia multiseta Stach, 1947
            Syn.: Folsomia multiseta ssp. dives Stach, 1947
            Syn.: Folsomia ksenemani Christiansen, 1959
Species picea Christiansen & Tucker, 1977:374
Species postsensilis Potapov, M et Gao, Y, 2012:54
Folsomia postsensilis from China
After Potapov M & Gao Y, 2012 Fig.12

Species potapovi Babenko, AB, 2007:375
Species prima Mills, HB, 1931:3
            Syn.: Folsomia quadrioculata Folsom, 1928, partim
            Syn.: Folsomia silvestrii Folsom, JW, 1937:20
             Syn.: Folsomia silvestri(sic) Stach, 1947 ISS
             Syn.: Folsomia silvestri(sic) Scott, HG, 1961:266 ISS
Folsomia prima from the USA
2009.03.07 © Bergen, L.
Folsomia prima from the USA
2009.03.07 © Bergen, L.

Species pseudodecemoculata Stebaeva, 1971
             Syn.: Folsomia pseudodecimoculata(sic) Potapov, MB, 2000:236 ISS
Species pseudodiplophthalma Stach, 1947
Species pseudovtorovi Potapov, M et Gulgenova, A, 2013:321
Species pusilla Salmon, JT, 1944:143
Species pyrenaea Cassagnau, P, 1954:622
Species quadrioculata (Tullberg, 1871) Agrell, I, 1939:4, B
            Syn.: Folsomia anophthalma Axelson, 1902
            Syn.: Isotoma quadrioculata var. pallida Axelson, WM, 1905:791
            Syn.: Folsomia gracilis Latzel, 1922
            Syn.: Folsomia quadrioculata f. 4-punctata(sic) Agrell, I, 1939:2 IOS
            Syn.: Folsomia diplophthalma f. 4-punctata(sic) Agrell, I, 1939:6 IOS
            Syn.: Folsomia quadrioculata ab. 2-punctata(sic) Agrell, I, 1939:5 IOS
            Syn.: Folsomia manolachei Bagnall, 1939
            Syn.: Folsomia quadrioculata ssp. americana Stach, 1947
            Syn.: Folsomia nana Gisin, 1957
            Syn.: Folsomia monosetosa Rusek, 1966
            Syn.: Folsomia sp. De Bie, P et Dop, H, 1980:A63 map
            Syn.: Folsomia spec. De Bie, P et Dop, H, 1980:7
             Syn.: Isotoma 4-oculata(sic) Absolon, K, 1901:33 ISS
Folsomia quadrioculata from New Zealand
After Salmon JT, 1941 Pl.49 Fig.154

             Syn.: Folsomia 4-oculata(sic) Cassagnau, P in Vandel, A, 1958:3379 ISS
             Syn.: Folsomia 4-oculata(sic) Delamare Deboutteville & Jacquemart, 1961 ISS
             Syn.: Folsomia 4-oculata(sic) Poole, 1961 ISS
             Syn.: Folsomia 4-oculata(sic) Betsch, J-M et Cassagnau, P, 1966:125 ISS
             Syn.: Folsomia 4-oculata(sic) Peterson, AJ, 1971:82 ISS
             Syn.: Folsomia quadriculata(sic) Martynova, EF, 1974:19 ISS
             Syn.: Folsomia 4-oculata(sic) Deharveng, L, 1982:613,gr ISS
Folsomia quadrioculata from the UK
2005 © Brocklehurst, K.

             Syn.: Folsomia 4-oculata(sic) Parisi, V, 2007:22 ISS
             Syn.: Folsomia quadrioculala(sic) Busmachiu, G, 2010:154 ISS
Folsomia quadrioculata (bar=2µm)
Rhombic epicuticular ultrastructure
After Nickerl J & al, 2012 Fig.4C

Folsomia quadrioculata from Holland
2012.10.30 © van Duinen, J.
Folsomia quadrioculata from Holland
Fused 4th-6th abdominal segments
2012.10.30 © van Duinen, J.
Folsomia quadrioculata from Holland
Eye with 2 distinctly separated ocelli
2012.10.30 © van Duinen, J.
Folsomia quadrioculata juv. from Switzerland
2012.11.30 © Murray, A.

             Syn.: Folsomia qudrioculata(sic) Zhang, F, Chen, Z, Dong, R-R, Deharveng, L, Stevens, MI, Huang, Y-H et Zhu, CD, 2014:233 ISS
             Syn.: Folsomia qudrioculata(sic) Zhang, F, Chen, Z, Dong, R-R, Deharveng, L, Stevens, MI, Huang, Y-H et Zhu, CD, 2014:235 ISS
             Syn.: Folsomia qudrioculata(sic) Zhang, F, Chen, Z, Dong, R-R, Deharveng, L, Stevens, MI, Huang, Y-H et Zhu, CD, 2014:236 ISS
             Syn.: Folsomia qudrioculata(sic) Zhang, F, Chen, Z, Dong, R-R, Deharveng, L, Stevens, MI, Huang, Y-H et Zhu, CD, 2014:237 ISS
Folsomia quadrioculata from Holland
2014.10.28 © van Duinen, J.
Folsomia quadrioculata from Germany
2015.01.07 © Mattew, M.
Folsomia quadrioculata from Germany
Four distinctly separated ocelli
2015.01.07 © Mattew, M.
Folsomia quadrioculata from Germany
2015.02.15 © Mattew, M.

             Syn.: Folsomia quadriocullata(sic) Raschmanová, N, Miklisová, D et Kovác, L, 2016:189 ISS
             Syn.: Folsomia quadriocula(sic) Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:599 ISS
             Syn.: Folsomia qudrioculata(sic) Zhang, F, Pan, Z-X, Wu, J, Ding, Y-H, Yu, D-Y et Wang, B-X, 2016:615 ISS
Species regularis Hammer, 1953
            Syn.: Folsomia ozeana Yosii, R, 1977:154
             Syn.: Folsomia regulars(sic) Suetsugu, N, Satoh, F et Koike, T, 2010:59 ISS
Folsomia regularis from Canada
2014.02.02 © Hochhalter, R.
Folsomia regularis from Canada
2014.02.02 © Hochhalter, R.

Species riozoyoshiii Potapov & Cassagnau, 2000
Species rossica Potapov & Dunger, 2000
Species salmoni Stach, 1947
Species sanjiangensis Wang & Wu, 2012
Species santokhi (Baijal, HN, 1958:351)
Bagnallela santokhi from the Himalaya
After Baijal HN, 1958 Fig.3A.
"Folsomia santokhi"(sic) from India
After Santeshwari, Raghuraman M & Singh J, 2015 p.573
See note by Janssens F, 2016.11.02
Remark by Janssens, F. 2016.11.02: Folsomia is to be excluded for the record of Santeshwari et al., 2015:573 from India given abd.4-6 are not fused. Cryptopygus sp. might be a better match given only abd.5-6 are fused.

Species sedecimoculata Salmon, JT, 1943:75
Species sensibilis Kseneman, 1934
            Syn.: Isotomina pazae Selga, 1966
Species sensivillosa Potapov, M et Gulgenova, A, 2013:317
Folsomia sensivillosa from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.20

Species setifrontalis Potapov & Marusik, 2000
Species setosa Gisin, 1953
Species setula Christiansen, KA et Tucker, BE, 1977:376
Species sexoculata (Tullberg, 1871) Handschin, 1929:57, B
            Syn.: ... sexoculata ssp. alpigena Stach, 1947
            Syn.: Folsomia anglicana Paclt, 1952
            Syn.: Folsomia sexoculata ssp. britannica Stach, 1947
            Syn.: ... folsomiella Bagnall, 1949
            Syn.: ... sexoculata var. pyrenaea Cassagnau, 1954
             Syn.: Folsomia 6-oculata(sic) Parisi, V, 2007:22 ISS
             Syn.: Tetracanthella(sic) 6-oculata(sic) Parisi, V, 2007:23 lapsus ISS
Folsomia sexoculata from Norway
2010.03.24 © Haraldseide, H.
Folsomia sexoculata from Norway
2010.03.24 © Haraldseide, H.
Folsomia sexoculata from the UK
2012.01.10 © Jones, A.
Folsomia sexoculata from the UK
2012.01.10 © Jones, A.
Folsomia sexoculata from the UK
2012.01.10 © Jones, A.
Folsomia sexoculata from the UK
2012.01.12 © Jones, A.
Folsomia sexoculata from Canada
2012.02.09 © Hochhalter, R.
Folsomia sexoculata from the UK
2012.09.12 © Jones, A.
Folsomia sexoculata from France
2012.11.03 © Quintin, C.
Folsomia sexoculata from the UK
2012.11.27 © Murray, A.
Folsomia sexoculata from the USA
2016.12.07 © Dobbs, Br.
Folsomia sexoculata from the USA
2016.12.07 © Dobbs, Br.

Species similis Bagnall, 1939:57, B
             Syn.: Folsomia simile(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:258 ISS
Folsomia similis from Iran
After Arbea JI & Kahrarian M, 2015 Fig.7

Species sparsosetosa Potapov, M et Stebaeva, S, 1997:130
Folsomia sparsosetosa from Russia
After Potapov M & Stebaeva S, 1997 Fig.27

Species spinosa Kseneman, 1936, B
            Syn.: Folsomia aspinosa Bartosova, 1958
            Syn.: Folsomia montigena Stach, 1946
Folsomia spinosa from the UK
PAO longer than width of ant.1
2012.12.25 © Murray, A.
Folsomia spinosa from the UK
2012.12.25 © Murray, A.

Species stebaevi Potapov MB et Stevaeva S in Potapov, MB, 2006:19
Folsomia stebaevi from Russia
After Potapov MB, 2006 Fig.43

Species stella Christiansen & Tucker, 1977:379
            Syn.: Folsomia fimetaria auctt
            Syn.: Isotoma manubriata Macnamara
Species strenzkei Nosek, 1963
Species sylvia Wray, DL, 1953:148
             Syn.: Folsomia sylviae(sic) Mari Mutt & Bellinger, 1990 ISS
Species taigicola Stebaeva & Potapov, 1998
Species taimyrica Martynova & al., 1973
Species tatarica Martynova, 1964
Folsomia tatarica from Russia
After Potapov MB, 2006 Fig.3

Species tesari Dunger, 1970
Species thalassophila Bagnall, 1940
Species tianshanica Martynova, 1969
            Syn.: Folsomia tianschanica(sic) Martynova, 1969 IOS
             Syn.: Folsomia tianschanica(sic) Rusek, 1973 ISS
             Syn.: Folsomia tianschanica(sic) Potapov, M et Stebaeva, S, 1997:130 ISS
Species torpeda Potapov MB et Taskaeva A in Potapov, MB, 2006:11
Folsomia torpeda from Russia
After Potapov MB, 2006 Fig.1

Species trisetata Jordana, R et Ardanaz, A, 1981:41
Species uniparis Potapov, M et Gulgenova, A, 2013:322
Folsomia uniparis from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.31

Species uniramia Potapov, M et Gulgenova, A, 2013:323
Folsomia uniramia ♂ from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.32

Species variabilis Fjellberg, 1984
Species vasilyevi Berezina, OG et Potapov, MB, 2006:9
Species villosa Potapov & Marusik, 2000
Species vitimica Potapov, M et Gulgenova, A, 2013:325
Folsomia vitimica from Buryat Republic
After Potapov M & Gulgenova A, 2013 Fig.39

Species volgensis Martynova, 1967
Species vtorovi Martynova, 1971
Species wanxianensis Ji, M, Ding, Y-F et Chen, J-X, 2007:59
            Syn.: Folsomia wanxianesis(sic) Ji, M, Ding, Y-F et Chen, J-X, 2007:59 IOS
Species wellingdae Potapov, M et Culik, MP, 2002:69

Genus Gnathofolsomia Deharveng, L et Christian, E, 1984:97
Species contrapunctata (Kseneman, 1950) Potapov, MB, 2000:236
Species palpata Deharveng, L et Christian, E, 1984:98, t.t.

Genus Isotomiella Bagnall, 1939 key
  Syn.: Isotominae Denisia Folsom, JW, 1932:61
  Syn.: Isotomidae Isotoma Leleup, N, 1948:10,B
  Syn.: Isotomidae Isotomellia(sic) Greenslade, P in Houston, WWK, 1994:78 ISS
  Syn.: Isotomiella minor King, Hutchinson & Greenslade P, 1976
  Syn.: Isotomiella minor Greenslade P & Thompson, 1981
  Syn.: Isotomiella minor King, Greenslade P & Hutchinson, 1985
  Syn.: Isotomiella minor Greenslade P & Greenslade PJM in Striganova, 1987
  Syn.: Isotominae Isotominella(sic) Snider, RJ, 1967:216 ISS
  Syn.: Isotomidae Isotomiela(sic) Zeppelini Filho, D et Castaño Meneses, G, 1995:10 ISS
  Syn.: Isotomidae Isotomiela(sic) Cutz-Pool, LQ et Vázquez-González, M, 2012:106 ISS
Species alulu Christiansen, K et Bellinger, P, 1992:147
Species aluluminor Deharveng & Suhardjono, 1995
Species annae Deharveng, L et Suhardjono, YR, 1994:313
Species amazonica Oliveira & Deharveng, 1990
Species arlei Oliveira & Deharveng, 1990
Species barisan Deharveng, L et Suhardjono, YR, 1994:315
Species barivierai Deharveng, 1988
Species barrai Deharveng & Oliveira, 1990
Species barrana de Mendonça, MC et Abrantes, EA, 2007:?
Species bidentata Delamare Deboutteville, 1950
Species brevidens Bedos & Deharveng, 1994
Species canina de Mendonça, MC et Fernandes, LH, 2003:2
Species ciliata Cardoso, MA, 1969:115
Species cribrata Deharveng, L et Suhardjono, YR, 1994:320
Species deforestai Deharveng, L et Suhardjono, YR, 1994:316
Species deharvengi Gao & Potapov, 2011
Species delamarei Barra, 1968
Species digitata Deharveng & Oliveira, 1990
Species distincta de Mendonça, MC et Fernandes, LH, 2003:113
Species dubia Deharveng, L et Suhardjono, YR, 1994:318
Species dupliseta Deharveng & Oliveira, 1990
Species edaphica Bedos & Deharveng, 1994
Species falcata de Mendonça, MC et Fernandes, LH, 2003:113
Species fellina de Mendonça, MC et Fernandes, LH, 2003:5
Species gracilimucronata Rusek, 1981
Species granulata Oliveira & Deharveng, 1990
Species hirsuta Bedos & Deharveng, 1994
Species hygrophila Sterzynska, M et Kaprus, I, 2001:275
Species insulae Barra, J-A, 2006:65
Species inthanonensis Bedos & Deharveng, 1994
Species leksawasdii Bedos & Deharveng, 1994
Species longisensilla Gao & Potapov, 2011
Species louisi de Mendonça, MC et Queiroz, GC, 2016:1
Species macedoi de Mendonça, MC, Abrantes, EA et Neves, ACR, 2012:22
Species madeirensis da Gama, 1959
Species michonae Deharveng, L et Suhardjono, YR, 1994:322
Species minor (Schäffer, 1896) Yosii, 1939, t.t., B
            Syn.: Isotoma muscorum Schäffer, 1900 =?
            Syn.: Denisia falcata Folsom, JW, 1932:61
            Syn.: Isotomiella distinguenda Bagnall, 1939
            Syn.: Isotomiella media Gisin, 1948
            Syn.: Isotomiella sp. De Bie, P et Dop, H, 1980:A62 map
            Syn.: Isotomiella spec. De Bie, P et Dop, H, 1980:7
Isotoma minor from Russia
After Scherbakov AM, 1898 Table II Fig.14
Isotomiella minor from Belgium
2009.03.09 © Janssens, F.
Isotomiella minor from the UK
2012.08.21 © Murray, A.
Isotomiella minor from the UK
2012.08.21 © Murray, A.
Isotomiella minor from Holland
2012.10.31 © van Duinen, J.
Isotomiella minor from Holland
2012.10.31 © van Duinen, J.
Isotomiella minor from Holland
2012.10.31 © van Duinen, J.
Isotomiella minor from the UK
2012.12.25 © Murray, A.

Species muscorum (Schäffer, 1900) Kovác, L & Palacios-Vargas, JG, 2008:11
Species nummulifer Deharveng & Oliveira, 1990
Species paraminor Gisin, 1942
            Syn.: Isotomiella minor ssp. bulgarica Stach, J, 1947:305
            Syn.: Isotomiella paraminor var. madeirensis da Gama, 1959
Species proxima de Mendonça, MC et Fernandes, LH, 2003:111
Species quadriseta Deharveng & Oliveira, 1990
Species sensillata Deharveng & Oliveira, 1990
Species similis Oliveira & Deharveng, 1990
Species sodwana Barra, J-A, 1997:466
Species spinifer Deharveng & Oliveira, 1990
Species spinosa Deharveng, L et Fjellberg, A, 1993:121
Species symetrimucronata (Najt & Thibaud, 1987) Mari Mutt, JA & Bellinger, PF, 1990:82
            Syn.: Isotomiella minor Greenslade P & Thompson, 1981
            Syn.: Isotomiella prussianae Oliveira & Deharveng, 1990
             Syn.: Isotomiella symmetrimucronata(sic) Culik, MP, de Souza, JL et Ventura, JA, 2002:52 ISS
             Syn.: Isotomiella symmetrimucronata(sic) Culik, MP, Martins, DS et Ventura, JA, 2006:5 ISS
             Syn.: Isotomiella symmetrimucronata(sic) Greenslade, P et Potapov, M, 2008:91 ISS
Species tamurai Tanaka & Niijima, 2009
Species thiollayi Deharveng, L et Suhardjono, YR, 1994:319
             Syn.: Isotomiella thiollay(sic) Kovác, L et Palacios-Vargas, JG, 2008:12 ISS
Species uai de Mendonça, MC, Abrantes, EA et Neves, ACR, 2012:26
Species unguiculata Deharveng, 1989
Species yinae Tamura & Zhao, 2000

Genus Mucracanthus Stebaeva, 1976
Species altaicus Stebaeva, 1976
Species ischimicus Stebaeva, 1984

Genus Octodontophora Tshelnokov, 1990
Species ornata Tshelnokov, 1990, t.t.

Genus Pseudofolsomia Martynova, 1967
Species acanthella Martynova, 1967
Species spinata (Martynova, 1964)

Genus Sahacanthella Potapov & Stebaeva, 1995
Species kele Potapov & Stebaeva, 1995

Genus Secotomodes Potapov, 1988
Species caucasicus Potapov, 1988
Species sibiricus Potapov, 1988

Genus Tiancanthella Rusek, 1979
Species martynovae Rusek, 1979

Genus Weberacantha Christiansen, KA, 1951:24 key
Species beckeri (Stebaeva, 1966) Potapov, M, Babenko, A & Fjellberg, A, 2006:58
Species cylindrica Huang & Potapov, 2012
Species echinodermata Potapov, M, Babenko, A et Fjellberg, A, 2006:58
Weberacantha echinodermata from Russia
After Potapov M et al., 2006 Fig.168

Species janetscheki (Yosii, 1971) Potapov, M, Babenko, A & Fjellberg, A, 2006:58
Species magnomucrella Potapov, M, Babenko, A et Fjellberg, A, 2006:61
Weberacantha magnomucrella from Mongolia
After Potapov M et al., 2006 Fig.176

Species octa Christiansen, KA, 1951:25, t.t.
Weberacantha octa from Alaska
After Christiansen K, 1951 Fig.1

Species striganovae Babenko & Potapov, 2012



References