http://www.collembola.org/taxa/onycinae.htm - Last updated on 2019.02.23 by Frans Janssens
Checklist of the Collembola: Onychiurinae

Subfamilia Onychiurinae Börner, 1901 key

Genus Number
of species
Absolonia1
Agraphorura11
Allonychiurus23
Argonychiurus7
Arneria2
Bionychiurus6
Chribellphorura1
Cribrochiurus2
Deharvengiurus10
Detriturus4
Deuteraphorura83
Dinochiurus1
Dungeraphorura1
Formosanonychiurus2
Heteraphorura19
Hymenaphorura46
IncertaeSedis9
Jacekaphorura1
Kalaphorura10
Kennethia1
Leeonychiurus8
Megaphorura2
Micronychiurus8
Oligaphorura56
Ongulonychiurus1
Onychiuroides25
Onychiurus48
Orthonychiurus28
Paronychiurus6
Pilonychiurus1
Probolaphorura2
Protaphorura138
Protaphorurodes1
Psyllaphorura10
Reducturus1
Sacaphorura1
Sensillonychiurus13
Similonychiurus1
Spelaphorura1
Spinonychiurus13
Supraphorura1
Thalassaphorura67
Uralaphorura6
Vexaphorura1
Vibronychiurus5
Wandaphorura1
Yoshiiphorura1
Onychiurinae689
Genera with number of species
Onychiurinae from China
Pigment and eyes absent
2008.07.17 © NCode, A.
Onychiurinae from Canada
Pigment and eyes absent
2009.01.09 © Weeta, W.
Onychiurinae from Greece
Ventral furca absent
2012.11.14 © Henderickx, H.
Onychiurinae from France
Cave species
2014.05.07 © Alonso, C.


Genus Dungeraphorura Gulgenova & Potapov, 2012

Species yaruna Gulgenova & Potapov, 2012


Tribus Hymenaphorurini Pomorski RJ, 1996:99 key


Genus Kalaphorura Absolon, K, 1901:388habitus dorsal

  Syn.: Onychiuridae (Onychiurinae) Kalaphorua(sic) Salmon, JT, 1964:156 ISS

Species bearei (Bagnall, 1937) Salmon, JT, 1959:151

Species bermani (Martynova, 1976) Weiner, WM, 1996:179

Species burmeisteri (Lubbock, J, 1873:190) Handschin, 1920, t.t., B

            Syn.: Lipura Burmeisteri(sic) Lubbock, J, 1873:190,tt IOS
            Syn.: Lipura tuberculata Moniez, 1890


Lipura Burmeisteri(sic) from the UK
After Lubbock, J., 1873 Pl.44.
             Syn.: Aphorura (Kalaphorura?) Burmeisteri(sic) Absolon, K, 1901:388 ISS
Remark by Dányi, L. & Traser, G. 2008:22: According to Dunger (1975) the early Hungarian data of Kalaphorura burmeisteri could include Kalaphorura paradoxus (Schäffer, 1900) as well.

Kalaphorura burmeisteri from the UK
Vestigial furca
2002 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2004 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2004 © Hopkin, S.P.
Kalaphorura burmeisteri from the UK
2006 © Valentine., B
Kalaphorura burmeisteri from the UK
2006 © Barton, T.
Kalaphorura burmeisteri from the UK
2006 © Cornwall, N.J.
Kalaphorura burmeisteri from Belgium
2007 © Doornaert, A.
Kalaphorura burmeisteri from the UK
2007 © Tonsbeek, M.
Kalaphorura burmeisteri from the UK
2007 © Tonsbeek, M.
Kalaphorura burmeisteri from the UK
2007.11.28 © Oakshott, A.
Kalaphorura burmeisteri from the UK
2007.11.28 © Oakshott, A.
Kalaphorura burmeisteri from the UK
2008.12.24 © Barton, T.
Kalaphorura burmeisteri from the UK
Large anals spines
2008.12.24 © Barton, T.
Kalaphorura burmeisteri from the UK
Tuberculate cuticula
2009.01.01 © Barton, T.
Kalaphorura burmeisteri from the UK
Groove marking post-antennal organ
2009.10.18 © Barton, T.
Kalaphorura burmeisteri from the UK
Juvenile
2011.07.16 © Barton, T.
Kalaphorura burmeisteri from the UK
Granulated skin
2012.11.22 © Murray, A.
Kalaphorura burmeisteri from the UK
Moulting
2013.05.10 © Murray, A.
Kalaphorura burmeisteri from the UK
Aged creamy specimen
2013.05.12 © Murray, A.
Kalaphorura burmeisteri from the UK
2013.06.06 © Murray, A.
Kalaphorura burmeisteri from the UK
Large anal spines
2013.09.15 © Murray, A.
Kalaphorura burmeisteri from the UK
Large anal spines
2013.11.13 © Phillips, E.
Kalaphorura burmeisteri from the UK
Club-shaped antenna
2013.11.13 © Phillips, E.
Kalaphorura burmeisteri from the UK
Short tibiotarsus with long unguis (U) and unguiculus (u)
2013.12.18 © Phillips, E.
Kalaphorura burmeisteri from the UK
Tuberculate skin
2013.12.18 © Phillips, E.
Kalaphorura burmeisteri from the UK
2014.01.05 © Barton, T.
Kalaphorura burmeisteri from the UK
2016.12.18
Body segments with broad posterior margin
© Barton, T.





Species carpenteri (Stach, 1919) Handschin, 1924

Kalaphorura carpenteri from Romania
Seta a0 above p-setae
After Fiera C & Weiner WM, 2013 Fig.2B

Species granulata (Börner, C, 1909:103) Handschin, 1924

Species heterodoxa (Gisin, H, 1964:655) Weiner, WM, 1996:179

Kalaphorura heterodoxa (bar=2µm)
Hexagonal epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.5A

Species ludzaki Weiner, WM et Stomp, N, 2005:96

Species paradoxa (Schäffer, 1900) Ellis, WN & Bellinger, PF, 1973:26, B

            Syn.: Lipura tuberculata Schäffer, 1900 =?
            Syn.: Lipura tuberculata Börner, 1901 =?
            Syn.: Onychiurus (Kalaphorura) burmeisteri Kovács, 1953 partim
            Syn.: Kalaphorura burmeisteri Loksa, 1959 partim


Remark by Dányi, L. & Traser, G. 2008:22: Loksa's "Onychiurus burmeisteri" data were corrected by Traser (1999) as Kalaphorura paradoxa.

Kalaphorura paradoxa (bar=2µm)
Hexagonal epicuticular ultrastructure with secondary granules
After Nickerl J & al, 2012 Fig.5E
Kalaphorura paradoxa from Romania
Seta a0 in line with p-setae
After Fiera C & Weiner WM, 2013 Fig.2A

Species tuberculata (Moniez, 1890) Gruia, 1964

            Syn.: Lipura burmeisteri Stach, J, 1954:113 =?
            Syn.: Onychiurus burmeisteri Denis, 1937
            Syn.: Aphorura paradoxa Stach, J, 1954:117


  Subspecies clavipila Stach, 1920

Species vannieri (Massoud, 1968) Weiner, WM, 1996:179


Genus Hymenaphorura Bagnall, 1948 key

  Syn.: ... Hymenaphorurodes Bagnall, 1949
  Syn.: Onychiurinae Hymenophorurodes(sic) Stach, J, 1954:22 ISS
  Syn.: Onychiurinae Sibirichiurus Stach, J, 1954:25 i

Species sp. Arbea, JI et Kahrarian, M, 2017:147

            Syn.: Hymenaphorura sibirica Cox, 1982
            Syn.: Hymenaphorura sibirica Daghighi & al, 2013
             Syn.: Onychiurus sibricus(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:260 ISS
            Syn.: Hymenaphorura sibirica Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:78
            Syn.: Hymenaphorura sibirica Yoosefi-Lafooraki & Shayanmehr, 2015
            Syn.: Hymenaphorura sibirica Arbea, JI et Kahrarian, M, 2017:147


Species alaskana Pomorski, RJ, 2001:451

Species alpina (Stach, J, 1946:8) Salmon, JT, 1964:159

Species alticola (Bagnall, 1935) Salmon, JT, 1959:153

            Syn.: Onychiurus sibiricus Handschin, 1921


             Syn.: Onychiurus alticola(sic) Gisin, 1960 ISS
             Syn.: Onychiurus (Protaphorura) alticola(sic) Gisin, H, 1960:127 ISS

Species anatolii Pomorski, RJ, 2001:467

Species arantiana Weiner & Stomp, 2001

             Syn.: Hymenaphorura aurantiana(sic) Pomorski, RJ, 2001:440 ISS
Hymenaphorura arantiana from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.6



Species californica (Coleman, 1941, i.s.) Salmon, 1964:159

Species cocklei (Folsom, 1908) Bagnall, 1948

            Syn.: Aphorura lutea Bacon, 1913
            Syn.: Aphorura montis Bacon, 1913
            Syn.: Onychiurus irregularis Chamberlain, 1943


             Syn.: Hymenaphorura cockleyi(sic) Salmon, 1974 ISS
Hymenaphorura cocklei from the USA
2008.07.07 © Wagner., B
Hymenaphorura cocklei from the USA
2008.12.28 © Hampel, P.
Hymenaphorura cocklei from the USA
2008.12.28 © Hampel, P.
Hymenaphorura cocklei from the USA
2009.01.13 © Davis, J.
Hymenaphorura cocklei from the USA
2009.01.13 © Davis, J.
Hymenaphorura cocklei from Canada
2009.06.18 © McIvor M. & McIvor D.
Hymenaphorura cocklei from Canada
Mass aggregation
2012.07.10 © Walker, I.
Hymenaphorura cocklei from the USA
2016.10.06 © Johnson, S.T.
Hymenaphorura cocklei from the USA
Distinct anal spines
2016.10.06 © Johnson, S.T.
Hymenaphorura cocklei from Canada
2018.01.12 © Barbin, T.



Species deca (Christiansen, K & Bellinger, P, 1980:431) Rusek & Marshall, 1996

             Syn.: Hymenaphorura decus(sic) Rusek & Marshall, 1996 ISS
             Syn.: Hymenaphorura decus(sic) Fjellberg, 1999 ISS
             Syn.: Hymenaphorura decus(sic) Pomorski, RJ, 2001:447 ISS

Species dentifera (Stach, 1934) Salmon, 1964:161

Remark by Dányi, L. & Traser, G. 2008:21: Hymenaphorura dentifera (Stach, 1934) is a "species inquirenda according to Gisin (1960)". [Janssens, F., 2008.11.17: Contrary to what Dányi & Traser (2008:21) claim, Gisin (1960:151) refers to fimetarius var. dentifera Kos, 1939, not to sibiricus f. dentifera Stach, 1934. And the first is considered as a species dubia by Gisin (1960:151), not as species inquirenda.]

Species gamae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:50

Species granulata Pomorski, RJ, 2001:453

Species hispanica Pomorski, 1992

Species improvisa Pomorski & Skarzynski, 2000

Species inopinata Babenko, Potapov & Taskaeva, 2017

Species ioni Busmachiu, G, Popa, I et Weiner, WM, 2014:550

Hymenaphorura ioni from Romania
After Busmachiu G, Popa I & Weiner WM, 2014 Fig.2

Species jugoslavica (Gisin, 1963) Pomorski, RJ, 2002:il

Species liberta Pomorski, RJ, 1990:219

Species maiteae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:51

Species maoerensis Sun, X, 2014:58

Hymenaphorura maoerensis from China
After Sun X, 2014 Fig.1

Species minuta Sun, X, 2014:60

Hymenaphorura minuta from China
After Sun X, 2014 Fig.11

Species montana (Handschin, 1921) Salmon, JT, 1964:159

Species mystica Pomorski, RJ, 2001:462

Species nearctica Pomorski, RJ, 2001:460

Species nicolae Barra, J-A, 1998:522

Species nova Pomorski, RJ, 1990:210

Species palaearctica Pomorski, RJ, 2001:467

             Syn.: Hymenaphorura palearctica(sic) Pomorski, RJ, 2001:473 ISS

Species parva Skarzynski, D et Pomorski, RJ, 1996:319

Species polonica Pomorski, RJ, 1990:213

            Syn.: Onychiurus sibiricus Axelson, 1905
            Syn.: Onychiurus pseudosibiricus Rusek, 1966


Species pseudosibirica (Stach, 1954:101) Salmon, JT, 1964:160

Species rafalskii Weiner, WM et Szeptycki, A, 1997:193

Species reducta Pomorski, RJ, 2001:442

Species reisingeri (Neuherz, 1979) Pomorski, RJ, 2002:il

Species ridibunda Pomorski, RJ, 2001:451

Species sensitiva Pomorski, RJ, 2001:457

Species sibirica (Tullberg, T, 1877:40) Bagnall, 1948:637, t.t., B

            Syn.: Aphorura watanabei Matsumoto & Saito, 1926
            Syn.: Onychiurus watanabei Matsumoto, 1929
            Syn.: Onychiurus watanabei Yosii, R, 1977:151
            Syn.: Onychiurus alpinus Gisin, H, 1960:130
            Syn.: Onychiurus montanus Gisin, H, 1960:130
            Syn.: Hymenaphorura troglodytes Gisin, H, 1960:130


             Syn.: Aphorura siberica(sic) Schäffer, 1900 ISS
             Syn.: Protaphorura sibiricus(sic) Skidmore, RE, 1995:53 ISS
             Syn.: Onychiurus sibirica(sic) Sun, X, 2014:57,gr ISS

Species similis (Folsom, 1917) Bagnall, 1948

Species strasseri (Stach, J, 1934:147) Salmon, 1964:161

            Syn.: Onychiurus strasseri f. principalis Kos, F, 1939:297


Species submontana (Denis, 1926) Salmon, 1964:161

Species subsimilis (Bagnall, 1948) Salmon, JT, 1959:145

Species superba Pomorski, RJ, 2001:457

Species teretis Pomorski, RJ, 2001:464

Species troglodytes Bagnall, 1948

Species uzicensis Curcic, BPM, Lucic, LR, Curcic, SB et Curcic, NB, 2005:176

Species valdegranulata (Stach, J, 1954:103) Salmon, 1964:161

Species wusuliensis Sun & Wu, 2011


Genus Heteraphorura Bagnall, 1948 key  key  key


Species-group nearctic Arbea JI, 2014:3
Species bima (Christiansen, K & Bellinger, P, 1980:438) Pomorski, RJ, 2002:461

Species casa (Christiansen, K & Bellinger, P, 1980:440) Pomorski, RJ, 2002:463

             Syn.: Onychiurus cassus(sic) Pomorski, RJ, 2002:457 ISS
             Syn.: Heteraphorura cassa(sic) Pomorski, RJ, 2002:463 ISS

Species intricata Pomorski, RJ, 2002:468

Species justynae Pomorski, RJ, 2002:465

Species magnina (Wray, 1950) Bernard, EC, 2015:330

Species subtenuis (Folsom, 1917) Pomorski, RJ, 2002:459

            Syn.: Hymenaphorura similis Bagnall, 1948


Onychiurus (Protaphorura) subtenuis from the USA
After Christiansen K & Bellinger P, 1998 Fig.343A

             Syn.: Heteraphorura subtenua(sic) Soto-Adames, FN et Taylor, SJ, 2013:157 ISS
Heteraphorura subtenuis from the USA
2015.03.28 © Himmler, K.
Heteraphorura subtenuis from the USA
2018.02.24 © Murray, A.



Species tala (Christiansen, K & Bellinger, P, 1980:444) Pomorski, RJ, 2002:461


Species-group euroasiatic Arbea JI, 2014:4
Species carpatica (Stach in Kseneman, 1938:468) Weiner, WM, 1996:177

Species edentata (Kos, F, 1939:300) Pomorski, RJ, 2002:459

Onychiurus strasseri var. edentatus from Yugoslavia
After Kos F, 1939 Fig.57




             Syn.: Onychiurus strasseri var. edenticulatus(sic) Kos, F, 1939:297 ISS
Remark by Arbea, J.I. 2014:4: For the presence of 2+2 pseudocelli on abdominal tergum III, this species departs from Heteraphorura.


Species orientalis (Martynova, 1976) Pomorski, RJ, 2002:459

Species steineri Arbea, JI, 2014:5

Heteraphorura steineri from Italy
After Arbea JI, 2014 Fig.1




            Syn.: Onychiurus apuanus Steiner in Franz, 1975 nn

Species variotuberculata (Stach, J, 1934:161) Bagnall, 1948 tt

            Syn.: Onychiurus variotuberculatus var. terricolus Kos, 1940
            Syn.: Onychiurus setiventris Butschek, 1948


Heteraphorura variotuberculata from Turkey
After Özata MA, Sevgili H & Kaprus IJ, 2017 Fig.9A

Species-group oriental Arbea JI, 2014:4
            Syn.: ... Japonychiurus Yoshii, 1996


Species conjungens (Börner, C, 1909:104) Pomorski, RJ, 2002:459

            Syn.: Onychiurus (Paronychiurus) conjungens f. ezoensis Uchida, H et Tamura, H, 1968:5


             Syn.: Onychiurus congugens(sic) Mills, HB, 1935:134 ISS

Species imadatei (Yosii, R, 1956:58) Arbea, JI, 2014:3

            Syn.: Onychiurus (Paronychiurus) Imadatéi(sic) Yosii, R, 1956:58 IOS

Species iranica Kaprus I, Shayanmehr M et Kahrarian M in Kaprus, I, Shayanmehr, M, Kahrarian, M et Lafooraki, EY, 2017:336

Heteraphorura iranica from Iran
After Kaprus I & al, 2017 Fig.1
            Syn.: Heteraphorura sp. Yoosefi-Lafooraki & Shayanmehr, 2015
            Syn.: Heteraphorura sp. Arbea, JI et Kahrarian, M, 2017:142
            Syn.: Heteraphorura japonica Mayvan & al, 2015 cf
            Syn.: Heteraphorura japonica Hosseini & al, 2016 cf
            Syn.: Heteraphorura japonica Qazy & Shayanmehr, 2016a cf
            Syn.: Heteraphorura japonica Qazy & Shayanmehr, 2016b cf
            Syn.: Heteraphorura japonica Kaprus, I, Shayanmehr, M, Kahrarian, M et Lafooraki, EY, 2017:335,cf
            Syn.: Heteraphorura japonica Arbea, JI et Kahrarian, M, 2017:142,cf



Heteraphorura iranica from Iran
After Arbea JI & Kahrarian M, 2017 Fig.1

Species japonica (Yosii, 1967) Pomorski, RJ, 2002:459

  Subspecies kyotensis Yoshii, 1996

Species longisetosa (Lee & Park, 1986) Pomorski, RJ, 2002:459

             Syn.: Heteraphorura longisetosus(sic) Pomorski, RJ, 2002:459 ISS
             Syn.: Heteraphorura (oriental-group) longisetosus(sic) Arbea, JI, 2014:4 ISS

Species pseudoseolagensis (Martynova, 1981) Pomorski, RJ, 2002:459

Species seolagensis (Lee, 1974) Pomorski, RJ, 2002:459


Genus Kennethia Smolis, A et Skarzynski, D, 2013:42

Species pomorskii Smolis, A et Skarzynski, D, 2013:43, t.t.

Kennethia pomorskii from the USA
After Smolis A & Skarzynski D, 2003 Fig.1


Genus Arneria Pomorski, 2000

Species alinae Pomorski, 2000

Species filiformis Pomorski, 2000


Genus Paronychiurus Bagnall, 1948 key

  Syn.: Onychiurinae Parnychiurus(sic) Pomorski, RJ, 2003:187 ISS

Species dentatus (Folsom, JW, 1902:91), i.s.

             Syn.: Onychiurus dentata(sic) Poole, RW in Poole, RW et Gentili, P, 1997:53 ISS
Paronychiurus dentatus? from the USA
2017.02.14 © Dobbs, Br.
Paronychiurus dentatus? from the USA
Abdomen with small Abd.6
2017.02.14 © Dobbs, Br.
Paronychiurus dentatus? from the USA
Abd.6 with small anal spines
2017.02.14 © Dobbs, Br.



Species eous (Christiansen, K & Bellinger, P, 1980:449) Pomorski, RJ, 2003:190

Species hubbardi Pomorski, RJ, 2003:192

Species japonicus (Yosii, R, 1956:57) Salmon, JT, 1964:186

Species probus Pomorski, RJ, 2003:191

Species ramosus (Folsom, 1917) Bagnall, 1948, t.t.

            Syn.: Onychiurus millsianus Bagnall, 1934,nn
            Syn.: Paronychiurus millsi Salmon, 1959, nec Chamberlain, 1943


Paronychiurus ramosus from the USA
2018.01.13 © Abbott, J.C.
Det.: Bernard, E.C., 2018.01.14


Genus Psyllaphorura Bagnall, 1948

Species bashkirica (Khanislamova, 1986) Weiner, WM, 1996:181

Species jiangsuensis Yan, HJ, Huang, C et Chen, J-X, 2007:?

Species martynovae (Stebaeva, 1985, s.d.) Weiner, WM, 1996:181

Species obesa (Mills, HB, 1934:35) Bagnall, 1948, t.t.

Species okafujii (Yosii, 1967) Weiner, WM, 1996:181

             Syn.: Psyllaphorura okafuji(sic) Weiner, WM, 1996:181 ISS

Species olga Babenko & Fjellberg, 2016

Species raoheensis Sun & Wu, 2011

Species ryozoyoshii Weiner & Najt, 2000

Species sensillifera (Martynova, 1981) Weiner, WM, 1996:181

Species uenoi (Yosii, R, 1954:62) Weiner, WM, 1996:181

             Syn.: Onychiurus uénoi(sic) Yosii, R, 1954:62 ISS
             Syn.: Onychiurus Uénoi(sic) Yosii, R, 1956:49 ISS
             Syn.: Onychiurus ueno(sic) Yosii, R, 1956:52 ISS



Genus Dinochiurus Pomorski, RJ et Steinmann, D, 2004:15

Species reus (Christiansen, K & Bellinger, P, 1980:433) Pomorski, RJ & Steinmann, D, 2004:17, t.t.

Dinochiurus reus from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.1


Genus Reducturus Pomorski, RJ et Steinmann, D, 2004:19

Species christianseni Pomorski, RJ et Steinmann, D, 2004:19, t.t.

Reducturus christianseni from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.15


Genus Sacaphorura Pomorski, RJ et Steinmann, D, 2004:20

Species parapseudocellata Pomorski, RJ et Steinmann, D, 2004:23, t.t.

Sacaphorura parapseudocellata from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.19-20


Genus Vexaphorura Pomorski, RJ et Steinmann, D, 2004:23

Species robustiseta Pomorski, RJ et Steinmann, D, 2004:23, t.t.

Vexaphorura robustiseta from the USA
After Pomorski RJ & Steinmann D, 2004 Fig.29


Genus Wandaphorura Pomorski, RJ, 2007:53

Species millsi (Chamberlain, 1943) Pomorski, RJ, 2007:53, t.t.




Onychiurus (Onychiurus) millsi from the USA
After Christiansen K & Bellinger P, 1998 Fig.349A


Genus Probolaphorura Dunger, W, 1977:1

Remark by Pomorski, R.J. & Steinmann, D. 2004:26: The tribe Hymenaphorurini probably also comprises the genus Probolaphorura, but it needs redescription.

Species plumosetosa Martynova, 1978

Species sachalinensis Dunger, W, 1977:2, t.t.


Genus Protaphorurodes Bagnall, 1949

Species tomuraushensis (Yosii, 1940) Pomorski, RJ, 2002:il


Tribus Oligaphorurini Bagnall, 1949 key  key  key


Genus Oligaphorura Bagnall, 1949 key  key

  Syn.: Oligaphorurinae Dimorphaphorura Bagnall, 1949
  Syn.: Oligaphorurinae Micraphorura Bagnall, 1949
  Syn.: Onychiurinae Affinichiurus Stach, J, 1954:25 i
  Syn.: Oligaphorurinae Archaphorura Bagnall, 1949
  Syn.: Onychiurinae Arcaphorura(sic) Salmon, 1964 ISS
  Syn.: Onychiurus Arcaphorura(sic) Curcic, BPM et Lucic, LR, 1999:189 ISS


  Syn.: Onychiurus Olygaphorura(sic)) Loksa, I, 1978:51 ISS
  Syn.: Onychiurus Olygaphorura(sic)) Loksa, I, 1978:56 ISS
  Syn.: Onychiurus Olygaphorura(sic)) Loksa, I, 1978:64 ISS

Species aborigensis (Fjellberg, 1987) Pomorski, RJ, 2002:il

Species absoloni (Börner, 1901) Dallai & Malatesta in Calderini, 1995:7, t.t., B

            Syn.: Aphorura Absoloni(sic) Börner, 1901 IOS
            Syn.: Lipura cirrigera Moniez, 1893 =?
             Syn.: Lipura cirragera(sic) Salmon, JT, 1964:189,sd ISS
            Syn.: Aphorura minor Carl, J, 1899:281
            Syn.: Onychiurus groenlandicus ssp. affinis Ågren, 1903
            Syn.: Onychiurus quadrituberculatus Denis, 1931
            Syn.: Onychiurus schötti(sic) Denis, 1931
            Syn.: Micraphorura handschini Bagnall, 1949




Species alavensis (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:73) Pasnik, G & Weiner, WM, 2017:628

            Syn.: Arcaphorura(sic) serratotuberculata Pozo, 1983
            Syn.: Arcaphorura(sic) serratotuberculata Pozo, Selga & Simon, 1986


Species alna (Fjellberg, 1987) Potapov, M & Stebaeva, S, 1997:121

             Syn.: Micraphorura alnus(sic) Weiner, WM, 1996:175 ISS
             Syn.: Oligaphorura alnus(sic) Potapov, M et Stebaeva, S, 1997:121 ISS
             Syn.: Dimorphaphorura alnus(sic) Weiner & Kaprus, 2014 ISS
             Syn.: Dimorphaphorura alnus(sic) Weiner, WM et Kaprus, IJ, 2014:6 ISS
             Syn.: Oligaphorura alnus(sic) Pasnik, G et Weiner, WM, 2017:629 ISS

Species ambigua Babenko & Fjellberg, 2015

Species changbaiensis (Sun, X & Wu, D, 2012:18) Pasnik, G & Weiner, WM, 2017:622

Species chankaensis Sun & Wu, 2012

Species chatyrdagi (Kaprus, IJ, Weiner, WM & Pomorski, RJ, 2002:356) Pasnik, G & Weiner, WM, 2017:628

Micraphorura chatyrdagi from Ukraine
After Kaprus I.J., Weiner, W.M. & Pomorski, R.J, 2002 Fig.16

Species caucasica (Weiner, WM & Kaprus, IJ, 2014:7) Pasnik, G & Weiner, WM, 2017:622

             Syn.: Dimorphaphorura caucasisa(sic) Weiner, WM et Kaprus, IJ, 2014:5 ISS
Dimorphaphorura caucasica from Russia
After Weiner WM & Kaprus IJ, 2014 Fig.11



Species daii (Pomorski, RJ, Skarzynski, D & Kaprus, I, 1998:253) Pasnik, G & Weiner, WM, 2017:622

Species differens (Bagnall, 1949) Pasnik, G & Weiner, WM, 2017:622

            Syn.: Onychiurus quadrituberculatus Stach, 1947


Dimorphaphorura differens from Austria
After Weiner WM & Kaprus IJ, 2014 Fig.4

Species duocellata Babenko & Fjellberg, 2015

Species eremia (Kaprus, IJ, Weiner, WM & Pomorski, RJ, 2002:354) Pasnik, G & Weiner, WM, 2017:622

Micraphorura eremia from Ukraine
After Kaprus IJ, Weiner WM & Pomorski RJ, 2002 Fig.9

Species gamae (Busmachiu & Weiner, 2013) Pasnik, G & Weiner, WM, 2017:622

Species gela (Christiansen, K & Bellinger, P, 1980:407) Pomorski, RJ, 2002:il

Species groenlandica (Tullberg, T, 1877:41) Pomorski, 1996:31, t.t.

            Syn.: Lipura schötti(sic) Lie-Pettersen, 1896 IOS
             Syn.: Onychiurus schötti(sic) Ågren, 1903 ISS
             Syn.: Onychiurus schötti(sic) Wahlgren, 1906 ISS
            Syn.: Aphorura quadrituberculata Folsom, JW, 1919:274 =?
            Syn.: Onychiurus affinis Ågren, 1903



Onychiurus groenlandicus from Greenland
After Folsom JW, 1919 Fig.14

             Syn.: Onychiurus grönlandica(sic) Handschin, E in Dahl, F, 1929:43 ISS
             Syn.: Onychiurus groenlandica(sic) Poole, RW in Poole, RW et Gentili, P, 1997:53 ISS
Oligaphorura groenlandica from Norway
2011 © Fjellberg, A.





Species hackeri (Christian, E, 1986:177) Pomorski, RJ, Skarzynski, D & Kaprus, I, 1998:259

Species humicola Shvejonkova & Potapov, 2011

Species interrupta (Fjellberg, 1987) Pasnik, G & Weiner, WM, 2017:628

Species irinae (Thibaud & Taraschuk, 1997) Pomorski, RJ, 2002:il

Species inya (Weiner, WM & Kaprus, IJ, 2014:10) Pasnik, G & Weiner, WM, 2017:622

             Syn.: Dimorphaphorura inyae(sic) Weiner, WM et Kaprus, IJ, 2014:3 ISS
             Syn.: Dimorphaphorura inyae(sic) Weiner, WM et Kaprus, IJ, 2014:11 ISS
             Syn.: Dimorphaphorura inyae(sic) Weiner, WM et Kaprus, IJ, 2014:18 ISS
             Syn.: Dimorphaphorura inyae(sic) Weiner, WM et Kaprus, IJ, 2014:27 ISS
Dimorphaphorura inyae(sic) from Russia
After Weiner WM & Kaprus IJ, 2014 Fig.19

             Syn.: Dimorphaphorura inyae(sic) Pasnik, G et Weiner, WM, 2017:622 ISS
             Syn.: Dimorphaphorura inyae(sic) Pasnik, G et Weiner, WM, 2017:624 ISS
             Syn.: Dimorphaphorura inyae(sic) Pasnik, G et Weiner, WM, 2017:625 ISS
             Syn.: Oligaphorura (clade-C) inyae(sic) Pasnik, G et Weiner, WM, 2017:622 ISS


Species jingyueensis (Sun & Wu, 2012) Pasnik, G & Weiner, WM, 2017:622

Species judithae (Weiner, 1994) Pomorski, RJ, 1996:91

Species judithnajtae Weiner, WM et Pasnik, G, 2017:70

Oligaphorura judithnajtae from Japan
After Weiner WM & Pasnik G, 2017 Fig.1A

Species koreana (Weiner, 1994) Weiner, WM, 1996:176

Species kremenitsai Shvejonkova & Potapov, 2011

Species kurtshevae Martynova, 1981

Species linderae (Weiner, 1994) Weiner, WM, 1996:176

Species marcuzzii (Cassagnau, 1968) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:7

Species melittae (Christian, E, 1993:163) Kaprus, IJ & Weiner, WM, 2009:il

Species montana Weiner, 1994

Species multiperforata (Gruia, M, 1973:173) Pasnik, G & Weiner, WM, 2017:629

             Syn.: Onychiurus (Oligophorura) multiperforata(sic) Weiner, WM et Pasnik, G, 2017:73 ISS

Species nataliae (Fjellberg, 1987) Pasnik, G & Weiner, WM, 2017:622

             Syn.: Micraphorura natalie(sic) Kaprus, IJ et Weiner, WM, 2009:il ISS

Species nuda (Fjellberg, 1987) Pomorski, RJ, 2002:il

Species olenae (Weiner, WM & Kaprus, IJ, 2014:13) Pasnik, G & Weiner, WM, 2017:622

Dimorphaphorura olenae from Ukraine
After Weiner WM & Kaprus IJ, 2014 Fig.29

Species palissai (Yosii, 1971) Pomorski, RJ, 2002:il

Species pieninensis (Weiner, 1988) Potapov, M & Stebaeva, S, 1997:121

Species pingicola (Fjellberg, 1987) Pomorski, RJ, 2002:il

Species pseudoinya (Weiner, WM & Kaprus, IJ, 2014:14) Pasnik, G & Weiner, WM, 2017:622

             Syn.: Dimorphaphorura pseudoinyae(sic) Weiner, WM et Kaprus, IJ, 2014:3 ISS
             Syn.: Dimorphaphorura pseudoinaye(sic) Weiner, WM et Kaprus, IJ, 2014:11 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Weiner, WM et Kaprus, IJ, 2014:11 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Weiner, WM et Kaprus, IJ, 2014:18 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Weiner, WM et Kaprus, IJ, 2014:27 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Pasnik, G et Weiner, WM, 2017:622 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Pasnik, G et Weiner, WM, 2017:624 ISS
             Syn.: Dimorphaphorura pseudoinyae(sic) Pasnik, G et Weiner, WM, 2017:625 ISS
             Syn.: Oligaphorura (clade-C) pseudoinyae(sic) Pasnik, G et Weiner, WM, 2017:622 ISS

Species pseudomontana Sun, X et Wu, D, 2012:21

Species pseudoraxensis (Nosek, J & Christian, E, 1983:397) Pasnik, G & Weiner, WM, 2017:629

             Syn.: Dimorphaphorura pseuraxensis(sic) Weiner, WM et Kaprus, IJ, 2014:19 ISS

Species quadrituberculata (Börner, C, 1901:4) Pasnik, G & Weiner, WM, 2017:629

             Syn.: Aphorura 4-tuberculata(sic) Absolon, K, 1901:84 ISS
             Syn.: Aphorura 4-tuberculata(sic) Absolon, K, 1901:380 ISS
             Syn.: Aphorura 4-tuberculata(sic) Absolon, K, 1901:386 ISS
             Syn.: Onychiurus quadrituberculata(sic) Frenzel, 1937 ISS
             Syn.: Arcaphorura(sic) quadrituberculatus(sic) Salmon, JT, 1959:154 ISS
             Syn.: Onychiurus (Oligaphorura) quadrituberculata(sic) Gisin, H, 1960:112 ISS
             Syn.: Onychiurus (Oligophorura) quadrituberculata(sic) Gruia, 1973 ISS

Species raxensis (Gisin, 1961) Pasnik, G & Weiner, WM, 2017:622

Species reversa (Fjellberg, 1987) Pomorski, RJ, 2002:il

Species sabulosa Babenko, AB, 2007:366

Species sanjiangensis (Sun & Wu, 2012) Pasnik, G & Weiner, WM, 2017:622

Species schoetti (Lie-Pettersen, 1896) Bagnall, 1949

            Syn.: Lipura Schötti(sic) Lie-Pettersen, 1896 IOS
             Syn.: Onychiurus grönlandicus(sic) Folsom, 1919 ISS
             Syn.: Onychiurus grönlandicus(sic) Stach, 1921 ISS
            Syn.: Aphorura quadrituberculata Salmon, JT, 1964:159


             Syn.: Onychiurus schötti(sic) Wahlgren, 1906 ISS
             Syn.: Onychiurus schötti(sic) Denis, 1931 ISS
             Syn.: Archaphorura schötti(sic) Salmon, 1964 ISS
             Syn.: Onychiurus (Oligophorura) schotti(sic) Gruia, 1973 ISS
Oligaphorura schoetti from the UK
2002 © Hopkin, S.P.





Species serratotuberculata (Stach, 1933) Pasnik, G & Weiner, WM, 2017:628, t.t.

Species sibirica (Weiner, WM & Kaprus, IJ, 2014:16) Pasnik, G & Weiner, WM, 2017:622

Dimorphaphorura sibirica from Russia
After Weiner WM & Kaprus IJ, 2014 Fig.37

Species sophyae (Weiner, WM & Kaprus, IJ, 2014:18) Pasnik, G & Weiner, WM, 2017:622

Dimorphaphorura sophyae from Russia
After Weiner WM & Kaprus IJ, 2014 Fig.45

Species steposa (Kaprus, IJ, Weiner, WM & Pomorski, RJ, 2002:353) Pasnik, G & Weiner, WM, 2017:622

Micraphorura steposa from Ukraine
After Kaprus I.J., Weiner, W.M. & Pomorski, R.J, 2002 Fig.1

Species stojkoae (Shvejonkova & Potapov, 2011) Pasnik, G & Weiner, WM, 2017:628

Species tottabetsuensis (Yosii, R, 1972:83) Pomorski, RJ, 2002:il

Species tuvinica Potapov, M et Stebaeva, S, 1997:121

Oligaphorura tuvinica from Russia
After Potapov M & Stebaeva S, 1997 Fig.4

Species uralica (Khanislamova, 1986) Kaprus, IJ & Weiner, WM, [1995]:59

             Syn.: Onychiurus (Oligophorura) uralica(sic) Weiner, WM et Pasnik, G, 2017:73 ISS

Species ursi Fjellberg, 1984


Genus Chribellphorura Weiner, WM, 1996:174

  Syn.: Oligaphorurini Chribellaphorura(sic) Weiner, WM, 1996:197 ISS

Species allanae (Christiansen, K & Bellinger, P, 1980:405) Weiner, WM, 1996:174, t.t.


Tribus Protaphorurini Bagnall, 1949


Genus Supraphorura Stach, J, 1954:26

Species furcifera (Börner, C, 1901:3) Pomorski, 1996:27, B

            Syn.: Aphorura armata var. parumpunctata Schäffer, 1896
            Syn.: Onychiurus celticus Bagnall, 1935
            Syn.: Onychiurus scoticus Bagnall, 1935
            Syn.: Onychiurus subfurcifer Jacquemart, S, 1967:1,B


             Syn.: Aphorura furcifer(sic) Börner, 1901 ISS
             Syn.: Onychiurus furcifer(sic) Axelson, 1905 ISS
             Syn.: Onychiurus furcifer(sic) Axelson, 1905 ISS
             Syn.: Onychiurus furcifer(sic) Axelson, 1907 ISS
             Syn.: Onychiurus furcifer(sic) Axelson, 1912 ISS
             Syn.: Onychiurus furcifer(sic) Denis, 1925 ISS
             Syn.: Onychiurus furcifer(sic) Handschin, 1925 ISS
             Syn.: Onychiurus furcifer(sic) Handschin, E in Dahl, F, 1929:45 ISS
             Syn.: Onychiurus furcifer(sic) Nordberg, 1936 ISS
             Syn.: Onychiurus furcifer(sic) Tarsia in Curia, 1936 ISS
             Syn.: Onychiurus furcifer(sic) Agrell, 1939 ISS
             Syn.: Onychiurus furcifer(sic) Agrell, 1943 ISS
             Syn.: Onychiurus furcifer(sic) Agrell, 1945 ISS
             Syn.: Onychiurus furcifer(sic) Harvey, 1952 ISS
             Syn.: Onychiurus furcifer(sic) MacFadyen, 1952 ISS
             Syn.: Onychiurus furcifer(sic) Gisin, 1953 ISS
             Syn.: Onychiurus furcifer(sic) Strenzke, 1955 ISS
             Syn.: Onychiurus furcifer(sic) Bockemuhl, 1956 ISS
             Syn.: Onychiurus furcifer(sic) von Törne, 1956 ISS
             Syn.: Onychiurus (Protaphorura) furcifer(sic) Gisin, H, 1960:112 ISS
             Syn.: Onychiurus (Protaphorura) furcifer(sic) Grinbergs, 1960 ISS
             Syn.: Onychiurus furcifer(sic) Milne, 1960 ISS
             Syn.: Onychiurus furcifer(sic) Jacquemart, S, 1967:1 ISS
             Syn.: Onychiurus furcifer(sic) Rusek, 1977 ISS
             Syn.: Protaphorura furcifer(sic) Thibaud, J-M et Massoud, Z, 1977:6,B ISS
             Syn.: Onychiurus furcifer(sic) Walsh & Bolger, 1990 ISS
             Syn.: Protaphorura furcifer(sic) Dallai & Malatesta in Calderini, 1995 ISS
             Syn.: Supraphorura surcifera(sic) Pomorski, 1996 ISS
             Syn.: Onychiurus furcifer(sic) Dethier, M, 1998:56,B ISS
             Syn.: Onychiurus furcifer(sic) Delhez, F(+), Dethier, M et Hubart, J-M, 1999:34,B ISS
             Syn.: Onychiurus furcifer(sic) Ponge, J-F, 1999:56 ISS


Supraphorura furcifera from the UK
Vestigial furca
2002 © Hopkin, S.P.
Supraphorura furcifera from the UK
2013.06.08 © Murray, A.
Supraphorura furcifera from the UK
Post-antennal organ
2013.06.08 © Murray, A.
Supraphorura furcifera from the UK
Vestigial furca
2013.06.08 © Murray, A.


Genus Protaphorura Absolon, K, 1901:387 key  key  key  key

  Syn.: ... Allaphorura Bagnall, 1949
  Syn.: ... Heteronychiurus Bagnall, 1949
  Syn.: ... Protaphorurodes Bagnall, 1949
  Syn.: ... Stenonychiurus Bagnall, 1949


  Syn.: Onychiuridae Protophorura(sic) Greenslade P, [1996] ISS
Protaphorura sp. from Germany
2006.12.30 © Kohl, F.

  Syn.: Onychiuridae Protophorura(sic) Greenslade, P, 2010:235 ISS
Protaphorura sp. from the USA
2011.11.06 © Murray, T.

  Syn.: Onychiuridae Protahorura(sic) Gulgenova, A et Potapov, M, 2013:211 ISS


Species sp.1 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:893

Species sp.2 Babenko, AB et Kaprus, IJ, 2014:595

            Syn.: Onychiurus stachi Bagnall, 1935
Remark by Pitkin.R., B 1980:423: Type specimen not adult.


            Syn.: Onychiurus paucisetosus Hammer, 1942

Species abscondita Kaprus, I, Weiner, W et Pasnik, G, 2016:121

Protaphorura abscondita from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.1

Species aconae Arbea, JI et Jordana, R in Jordana, R et Ariño, AH, 1994:43

Species ajudagi Pomorski, RJ, Skarzynski, D et Kaprus, I, 1998:259

Species aksuensis (Martynova, 1972, s.d.) Pomorski, RJ, 2002:il

Species armata (Tullberg, 1869) Börner, 1909, t.t., B

            Syn.: Pediculus terrestris Linnæus, 1746
            Syn.: Podura ambulans Linnæus, 1758, partim Salmon, 1964
            Syn.: Podura fimetaria Mohr, 1786
            Syn.: Adicranus fimetarius Bourlet, 1842
            Syn.: Anurophorus gracilis Müller, 1859
            Syn.: Lipura arctica Tullberg, 1876
            Syn.: Lipura aurantiaca Ridley, 1880
             Syn.: Onychiurus aurantiaca(sic) Turk, 1933 ISS
            Syn.: Lipura incerta Moniez, 1893
            Syn.: Drepanura(sic) incerta Schött, H, 1917:2
            Syn.: Lipura ambulans Meinert, 1896
            Syn.: Aphorura ambulans Guthrie, 1903
            Syn.: Lipura quadrispina Lie-Pettersen, 1896
             Syn.: Aphorura 4-spina(sic) Absolon, K, 1901:380 ISS
             Syn.: Aphorura (Protaphorura) 4-spina(sic) Absolon, K, 1901:388 ISS
            Syn.: Aphorura armata ssp. stalagmitorum Absolon, 1900
            Syn.: Onychiurus pseudarmatus Folsom, 1917
            Syn.: Onychiurus daviesi Bagnall, 1935
            Syn.: Onychiurus makarensis Salmon, JT, 1937:352
            Syn.: Onychiurus waterstoni Bagnall, 1937
             Syn.: Onychiurus watersoni(sic) Parimuchová, A et Kovác, L, 2016:262 ISS
            Syn.: Onychiurus campatus Gisin, 1952
            Syn.: Onychiurus illaboratus Gisin, 1952
            Syn.: Onychiurus nemoratus Gisin, 1952
            Syn.: Onychiurus uliginatus Bödvarsson, H, 1970:30 =?
            Syn.: Onychiurus subuliginatus Bödvarsson, H, 1970:150
            Syn.: Onychiurus vanderdrifti Gisin, 1952
            Syn.: Onychiurus tricampatus Gisin, 1956
            Syn.: Onychiurus pseudovanderdrifti Gisin, 1957
            Syn.: Onychiurus subarmatus Bödvarsson, H, 1970:150
            Syn.: Onychiurus sublatus Bödvarsson, H, 1970:30 =?
            Syn.: Onychiurus flavescens Salmon, 1959
            Syn.: Onychiurus flavidulus Salmon, 1959
            Syn.: Onychiurus pannonicus Haybach, 1960
            Syn.: Onychiurus franconianus Gisin, 1961
            Syn.: Onychiurus subhumatus Selga, 1963
            Syn.: Lipura aurantiaca Salmon, JT, 1964:164
            Syn.: Onychiurus flavescens Salmon, JT, 1964:164
            Syn.: Onychiurus flavescens Salmon, JT, 1964:166
            Syn.: Onychiurus flavidulus Salmon, JT, 1964:164
            Syn.: Onychiurus flavidulus Salmon, JT, 1964:166
            Syn.: Lipura volvator Salmon, JT, 1964:163 partim
            Syn.: Protaphorura bagnalli Shaw, P, Faria, C et Emerson, B, 2013:165 =?


             Syn.: Onychiurus Armatus(sic) Kos, F, 1939:278 ISS
Onychiurus armatus from New Zealand
After Salmon, J.T., 1941 Pl.47 Fig.125.

             Syn.: Protaphorura armatus(sic) Snider, RJ, 1967:195 ISS
             Syn.: Protaphorura armatus(sic) Grégoire-Wibo, C, 1978:22 ISS
Onychiurus sp. armatus gr. from Tasmania
Post-antennal organ with single lobes
After Ireson JE, 1990 Fig.72
Onychiurus sp. armatus gr. from Tasmania
Ant.3 sensory organ
After Ireson JE, 1990 Fig.74

             Syn.: Protaphorura armatus(sic) Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:163 ISS
             Syn.: Protaphorura armatus(sic) Skidmore, 1995 ISS
             Syn.: Onychiurus armata(sic) Poole, RW in Poole, RW et Gentili, P, 1997:53 ISS
             Syn.: Protaphorura armatus(sic) Peck, SB, 2001:103,gr ISS
             Syn.: Protaphorura armatus(sic) Peck & al, 2001 ISS
Protaphorura armata from the UK
Vestigial furca
2002 © Hopkin, S.P.
Protaphorura armata from Sweden
2005 © Hall, K.
Protaphorura armata from Sweden
2005 © Hall, K.
 
Protaphorura armata from Denmark
2005.10.02 © Ipsen, K.
Protaphorura armata from Denmark
2007.02.09 © Larsen, T.
Protaphorura armata from the UK
2007.03.18 © Cornwall, N.J.
Protaphorura armata from France
2008.mm.dd © Lebeaux, P.
Protaphorura armata from the UK
2008.04.07 © Farrell, S.
Protaphorura armata from the UK
2008.04.08 © Farrell, S.
Protaphorura armata from Tasmania
2009.08.16 © Daley, A.

             Syn.: Protaphorura armatus(sic) Peck, SB et Jacquemart, J, 2011:5 ISS
Protaphorura armata from Tasmania
2011.06.17 © Daley, A.
Protaphorura armata from Tasmania
Ant.3 organ and post-antennal organ
2011.06.17 © Daley, A.
Protaphorura armata from the UK
2012.01.27 © Nurcombe, E.
Protaphorura armata from Holland
2012.10.31 © van Duinen, J.
Protaphorura armata juv. from the UK
2012.11.21 © Murray, A.
Protaphorura armata juv. from Switzerland
Post-antennal organ and antenna 3 organ
2012.12.01 © Murray, A.
Protaphorura armata from Spain
After Arbea JI, 2013 Fig.6
Protaphorura armata from the UK
Linea ventralis
2013.04.15 © Nurcombe, E.
Protaphorura armata from the UK
Anal spines
2014.01.18 © Garlick, T.
Protaphorura armata from the UK
Leg with long unguis
2015.12.29 © Robson, S.
Protaphorura armata from Slovenia
2016.02.26 © Pisec, N.
Protaphorura armata from Czechia
2018.10.08 © Huskens, M.-L.

  Subspecies denticulata Handschin, 1924

            Syn.: Onychiurus armatus var. denticulata(sic) Handschin, 1924 IOS
            Syn.: Onychiurus subaequalis Bagnall, 1937


             Syn.: Onychiurus armatus var. denticulata(sic) Handschin, 1929 ISS
             Syn.: Onychiurus armatus var. denticulata(sic) Kseneman, M, 1932:14 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Stach, 1934 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Frenzel, 1937 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Kseneman, 1937 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Kseneman, 1938 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Kseneman, 1938 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Strebel, 1938 ISS
             Syn.: Onychiurus armatus var. denticulata(sic) Kos, F, 1939:272 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Kos, 1941 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Muhlmann, 1942 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Strenzke, 1949 ISS
             Syn.: Onychiurus armatus ssp. denticulata(sic) Grinbergs, 1958 ISS
  Subspecies furcifera Schött, 1924


            Syn.: Onychiurus armatus ssp. furcifera(sic) Schött, 1924 IOS
             Syn.: Onychiurus armatus ssp. furcifera(sic) Stach, 1934 ISS
  Subspecies inermis Axelson, WM, 1905:790

  Subspecies multituberculata Stach, J, 1934:127,sd

  Subspecies trispina Axelson, 1935

  Forma scanica Bödvarsson, H, 1970:6





Species asensitiva (Stach, 1954) Salmon, JT, 1964:170

Species aurantiaca (Ridley, 1880) Salmon, JT, 1964:176

            Syn.: ... alborufescens Waterston, 1930, nec Vögler, 1893
            Syn.: Onychiurus flavescens Bagnall, 1935 nec Kinoshita, 1916
            Syn.: Onychiurus flavidulus Bagnall, 1939
             Syn.: Onychiurus flavidus(sic) Stach, 1954 ISS
            Syn.: Onychiurus magnicornis Bagnall, 1937
            Syn.: Onychiurus austriacus Butschek, 1948
            Syn.: Onychiurus uliginatus Gisin, 1952
            Syn.: Onychiurus latus Gisin, H, 1956:341
            Syn.: Onychiurus prolatus Gisin, H, 1956:340
            Syn.: Onychiurus sublatus Gisin, 1957
            Syn.: Onychiurus vontoernei Gisin, 1957
             Syn.: Protaphorura vontörnei(sic) Salmon, JT, 1964:176 ISS
            Syn.: Onychiurus subparallatus Selga, 1962
            Syn.: Onychiurus prolatus ssp. conlatus Gisin, H, 1962:3
            Syn.: Onychiurus (Protaphorura) subcancellatus Gisin, 1963
            Syn.: Onychiurus prolatus ssp. trilatus Gisin, 1963
            Syn.: Onychiurus gisini Ellis, WN, 1976:232
            Syn.: Onychiurus gisini Arbea, JI et Jordana, R, 1991:135
            Syn.: Onychiurus gisini da Gama, 1964 :
            Syn.: Onychiurus cancellatus Jordana & Beruete, 1983
            Syn.: Protaphorura meridiata Arbea & al., 1986
            Syn.: Onychiurus tetragrammatus Pomorski, 1990
            Syn.: Onychiurus tetragrammatus Parimuchová, A et Kovác, L, 2016:262


Onychiurus aurantiacus from the UK
After Pitkin, B.R., 1980 Fig.1
Protaphorura aurantiaca from the UK
2006.01.02 © Botting, J.
Protaphorura aurantiaca from the UK
2008.02.26 © Botting, J.
Protaphorura aurantiaca from the UK
2008.04.11 © Robertson, A.
Protaphorura aurantiaca from the UK
2009.01.24 © Robertson, A.
Protaphorura aurantiaca from the UK
2009.04.12© Robertson, A.
Protaphorura aurantiaca from the UK
2010.02.01© Robertson, A.
Protaphorura aurantiaca from the UK
2015.02.10 © Murray, A.
Protaphorura aurantiaca from the UK
Short 6th abdominal segment
2015.02.10 © Murray, A.
Protaphorura aurantiaca from the UK
2015.02.10 © Murray, A.
Protaphorura aurantiaca from the UK
Body yellowish; antennae white
2015.04.01 © Phillips, E.
Protaphorura aurantiaca from the UK
Ant.3 with constriction (*)
2015.04.01 © Phillips, E.
Protaphorura aurantiaca from the UK
2015.12.16 © Evans, K.
Protaphorura aurantiaca from the UK
2016.03.12 © Evans, K.
Protaphorura aurantiaca from the UK
Groove of post-antennal organ (*)
2016.03.12 © David W.
Protaphorura aurantiaca from the UK
2017.01.14 © Doogue, M.
Protaphorura aurantiaca from the UK
2019.02.04 © David W.

Species bakhchisaraica Kaprus, IJ, Pasnik, G et Weiner, WM, 2014:466

Protaphorura bakhchisaraica from Ukraine
After Kaprus IJ, Pasnik G & Weiner WM, 2014 Fig.1

Species bicampata (Gisin, H, 1956:344) Salmon, JT, 1964:170

            Syn.: Onychiurus islandicus Pomorski, 1990
            Syn.: Protaphorura islandica Pomorski, 1998


Species borealis (Martynova in Martynova, Gorodkov & Tshelnokov, 1973) Pomorski, RJ, 2002:il

Species borinensis Parimuchová, A et Kovác, L, 2016:256

Protaphorura borinensis from Slovakia
After Parimuchová A & Kovác L, 2016 Fig.1

Species borzica (Gruia, 1999) Pomorski, Leithner & Bruckner, 2003

Species brevispinata (Yosii, R, 1966:548) Pomorski, RJ, 2002:il

Species buryatica Gulgenova, A et Potapov, M, 2013:207

Protaphorura buryatica from Russia
After Gulgenova A & Potapov M, 2013 Fig.8

Species caerulea (Börner, 1901) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:7

Species caledonica (Bagnall, 1935) Kloet, GS & Hincks, WD, 1945:5

Species campata (Gisin, 1952) Salmon, JT, 1964:170, B

            Syn.: Onychiurus procampatus Gisin, H, 1956:343
Protaphorura procampata from the UK
After Shaw P, 2009 ©

            Syn.: Onychiurus stachi Murphy, 1960
            Syn.: Protaphorura cancellata Luciáñez & Simón, 1989



             Syn.: Protaphorura camapata(sic) Pomorski, RJ, 1996:98 ISS

Species cancellata (Gisin, H, 1956:338) Salmon, JT, 1964:170

Protaphorura cancellata from Holland
20110.17 © Wieringa, P.
Det.: Berg, M.

Species changbaiensis Sun, Zhang & Wu, 2013

             Syn.: Protaphorura changbainensis(sic) Parimuchová, A et Kovác, L, 2016:264 ISS

Species chilensis (Loksa & Rubio, 1966) Pomorski, RJ, 2002:il

Species christiani Pomorski, RJ, Leithner, C et Bruckner, A, 2003:307

             Syn.: Protaphorura christianseni(sic) Kaprus, IJ, Pasnik, G et Weiner, WM, 2014:470 ISS

Species churchilliana (Hammer, 1953) Salmon, JT, 1964:170

             Syn.: Onychiurus churchilliana(sic) Poole, RW in Poole, RW et Gentili, P, 1997:53 ISS

Species crassa (Kos, 1940, s.d.) Salmon, JT, 1964:169

Species cykini Parimuchova A et Kovac L in Parimuchová, A, Kovác, L, Zurovcová, M et Kadebskaya, OI, 2017:187

Protaphorura cykini from Siberia
After Parimuchová A & al, 2017 Fig.1c

Species dallaii (Nosek & Paoletti, 1981, s.d.) Nosek & Paoletti, 1981

Species daviesi (Bagnall, 1935) Kloet, GS & Hincks, WD, 1945:5

            Syn.: Onychiurus octopunctatus ssp. edenticulata Davies, 1934 nec Wahlgren, 1900


Species decempunctata (Kos, F, 1939:292) Salmon, JT, 1964:170

Species diplosensillata (Dunger, 1978)

Species discrepans (Bagnall, 1948) Salmon, JT, 1959:154

            Syn.: Onychiurus octopunctatus Handschin, 1920, nec Tullberg, 1876
            Syn.: Onychiurus octopunctatus Womersley, 1930, nec Tullberg, 1876
            Syn.: Onychiurus octopunctatus Bagnall, 1948, nec Tullberg, 1876


             Syn.: Protaphorura discrepens(sic) Salmon, JT, 1964:171 ISS

Species dorzhievi Gulgenova, A et Potapov, M, 2013:204

Protaphorura dorzhievi from Russia
After Gulgenova A & Potapov M, 2013 Fig.1

Species dudichi (Loksa, I & Rubio, I, 1966:148, s.d.) Dallai, R, Malatesta, E & Ramellini, P in Minelli, A, Ruffo, S & La Posta, S, 1995:7

Species duodecimpunctata (Folsom, JW, 1919:6A) Salmon, JT, 1964:171

Remark by Folsom, J.W. 1919:6A: Pseudocelli of antennal bases 6+6 (two specimens) or 5+5 (one specimen).
Onychiurus duodecimpunctatus from Arctica
After Folsom JW, 1919 Plate 3 Fig.29
             Syn.: Onychiurus 12-punctatus(sic) Hammer, M, 1953:36 ISS
Remark by Hammer, M. 1953:36-37: In specimens from Northern Canada, the number of pseudocelli at the antennal base of Onychiurus 12-punctatus(sic) varies between 4 to 6. Of 23 specimens, 10 were found with 4+4, 11 with 5+5, 1 with 6+6 and 1 with a bilateral asymmetric number of 5+4 pseudocelli.


             Syn.: Protaphorura duodecimpunctatus(sic) Skidmore, RE, 1995:53 ISS
             Syn.: Protaphorura duodecimpunctatus(sic) Bowser, M, 2008:il ISS
Remark by Janssens, F. 2008.08.15: Given the description of 'decimpunctatus Skidmore, 1995:53' is lacking it is a nomen nudum, therefore an invalid name. The name implicitly refers to specimens of duodecimpunctatus having 5 pseudocelli at the antennal base in steadt of the 6 pseudocelli of the nominal form. Such specimens are considered to be an intraspecific variation of duodecimpunctatus. Therefore, 'decimpunctatus Skidmore, 1995:53' is a partial junior synonym of duodecimpunctatus Folsom, 1919:6A.




Species dzherga Gulgenova, A et Potapov, M, 2013:208

Protaphorura dzherga from Russia
After Gulgenova A & Potapov M, 2013 Fig.16

Species edentata (Kos, F, 1939:276) Salmon, JT, 1964:174

Species eichhorni (Gisin, 1954) Salmon, JT, 1964:171, B

             Syn.: Protaphorura eichorni(sic) Salmon, JT, 1964:171 ISS
Protaphorura eichhorni from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.11



Species elenae Kaprus, IJ et Pomorski, RJ, 2008:668

Protaphorura elenae from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.1

Species federicoi Arbea, JI, 2003:93

            Syn.: Protaphorura armata Lawrence, 1979 =?


Species fimata (Gisin, 1952) Salmon, JT, 1964:171

            Syn.: Onychiurus (Protaphorura) armatus Christiansen, K et Bellinger, P, 1980:411 partim
            Syn.: Protaphorura armata Greenslade P, 1990 :
            Syn.: Onychiurus (Protaphorura) armatus Christiansen, K et Bellinger, P, 1998:452 partim
            Syn.: Onychiurus sp. Greenslade & Wise, 1984
            Syn.: Onychiurus sp. Greenslade, 1990


Species fistania Thibaud & Peja, 1994

Species florae Simón-Benito, JC et Luciáñez, MJ in Jordana, R et Ariño, AH, 1994:70

Species genheensis Sun, Chang & Wu, 2015

Species gisini (Haybach, 1960) Selga & al., 1977

Species glebata (Gisin, 1952) Arbea & Jordana, 1985

Protaphorura glebata from Romania
After Radwanski JM, Fiera C & Weiner WM, 2006 Fig.8

Species golestanica Kaprus I, Shayanmehr M et Kahrarian M in Kaprus, I, Shayanmehr, M, Kahrarian, M et Lafooraki, EY, 2017:339

Protaphorura golestanica from Iran
After Kaprus I & al, 2017 Fig.9

Species ianstachi Yosii, 1972

            Syn.: Onychiurus octopunctatus Stach, 1934 partim


             Syn.: Protaphorura janstachi(sic) Pomorski, RJ et Kaprus, I, 2007:138 ISS
             Syn.: Protaphorura ianstachia(sic) Kaprus, IJ, Pasnik, G et Weiner, WM, 2014:465 ISS

Species ionescui Radwanski, JM, Fiera, C Weiner, WM, 2006:449

Protaphorura ionescui from Romania
After Radwanski JM, Fiera C & Weiner WM, 2006 Fig.1

Species islandica (Bödvarsson, 1959) Salmon, JT, 1964:169

            Syn.: Protaphorura bicampata Pomorski, 1998


Species jacutica Martynova, 1976

Species janosik Weiner, 1990

Protaphorura janosik from Slovakia
2014.07.dd © Kovác L & Parimuchová A.

Species januarii (Weiner, 1977) Pomorski, RJ, Leithner, C & Bruckner, A, 2003:309

Species jernika Kaprus, I, Weiner, W et Pasnik, G, 2016:124

Protaphorura jernika from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.10
             Syn.: Protaphorura jernica(sic) Kaprus, I, Weiner, W et Pasnik, G, 2016:143 ISS


Species jiamusiensis Sun, Wu & Gao, 2013

Species kadici (Loksa, I, 1967:290, s.d.) Pomorski, RJ, 2002:il

  Varietas geminiocellatus Loksa, I, 1967:292

Species karadzicae (Lucic, Dimitrijevic & Mihajlova, 2003) Parimuchová, A & Kovác, L, 2016:261

Species kaszabi Dunger, 1978

Species kolenatii (Hemmer, 2000) Parimuchová, A & Kovác, L, 2016:261

  Forma alba Hemmer, 2000

Onychiurus kolentatii f. alba from Austria
After Kopeszki H, 2000 Abb.2

  Forma rubra Hemmer, 2000

Onychiurus kolentatii var. rubra from Austria
After Kopeszki H, 2000 Abb.1

Species kopetdagi Pomorski, RJ, 1994:193

Species kultia (Singh, Baijal & Mathew, 1956) Salmon, JT, 1964:172

            Syn.: Onychiurus kultia(sic) Singh, Baijal & Mathew, 1956 IOS

Species latosimilis (Loksa, I et Rubio, I, 1966:149), s.d.

Species levantina (Christiansen, K, 1956:123) Salmon, JT, 1964:172

             Syn.: Protaphorura levantica(sic) Kahrarian, M, Kaprus, I, Vafaei-Shoushtari, R et Shayanmehr, M, 2016:224 ISS

Species licheniphila Kaprus, IJ et Pomorski, RJ, 2008:670

Protaphorura licheniphila from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.6
            Syn.: Protaphorura sp.B Babenko & Fjellberg, 2006



Species longisensillata (Yosii, 1969)

Species lucensis Arbea, JI, 2015:236

            Syn.: Protaphorura (octopunctata-group) lucense(sic) Arbea, JI, 2015:236 IOS
Protaphorura lucense(sic) from Spain
After Arbea JI, 2015 Fig.1-2



Species macfadyeni (Gisin, 1953) Salmon, JT, 1964:172

Protaphorura macfadyeni from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.11
Protaphorura macfadyeni from Norway
2011 © Fjellberg, A.

Species macrodentata (Hammer, 1953) Salmon, JT, 1964:172

             Syn.: Protaphorura macrodentatus(sic) Skidmore, RE, 1995:53 ISS

Species macrotuberculata (Kos, F, 1939:309, i.s.) Salmon, JT, 1964:172

Onychiurus macrotuberculata from Yugoslavia
After Kos F ©, 1939 Fig.70

Species maglicensis (Zivadinovic, 1970, s.d.) Zivadinovic, 1970

Species maoerensis Sun, Wu & Gao, 2013

Species matsumotoi (Kinoshita, 1923) Pomorski, RJ, 2002:il

Species meridiata (Gisin, 1952, s.d.) Arbea, JI, 2003:95

            Syn.: Onychiurus subnemoratus Gisin, 1957
            Syn.: Onychiurus paranemoratus Selga, 1962
            Syn.: Onychiurus (Protaphorura) pulvinatus ssp. metapulvinatus Loksa, I, 1964:91
             Syn.: Onychiurus (Protaphorura) pulvinatus ssp. matapulvinatus(sic) Loksa, I, 1964:92 ISS
            Syn.: Protaphorura troglophila Nosek, 1975
             Syn.: Protaphorura troglofila(sic) Arbea, JI, 2003:95 ISS
            Syn.: Protaphorura cancellata Pomorski, 1990
            Syn.: Protaphorura gisini Pomorski, 1990
            Syn.: Onychiurus pulvinatus Pomorski, 1990
            Syn.: Protaphorura pulvinata Hopkin, SP, 1997:94
            Syn.: Onychiurus pulvinatus Parimuchová, A et Kovác, L, 2016:262


Species merita Kaprus, IJ et Pomorski, RJ, 2008:672

Protaphorura merita from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.16-17

Species microcellata (Dunger, 1978) Pomorski, RJ, 2002:il

Protaphorura cf. microcellata from Russia
After Babenko AB & Kaprus IJ, 2014 Fig.7.1

Species microcellata2 Babenko, AB et Kaprus, IJ, 2014:587

Species microtica Dunger, 1978

Species minima Sun, Zhang & Wu, 2013

Species mongolica (Martynova, 1975) Pomorski, RJ & Kaprus, I, 2007:137

Protaphorura mongolica from Mongalia
After Kaprus IJ & Pomorski RJ, 2008 Fig.24

Species montana Mateos Frias, EM et Arbea, JI, 1986:283

Species multipunctata (Absolon, K, 1900:406, i.s., s.d.) Salmon, JT, 1964:169

             Syn.: Onychiurus armatus var. multipunctata(sic) Kos, F, 1939:275 ISS

Species nazarovensis Kaprus, IJ et Pomorski, RJ, 2008:678

Protaphorura nazarovensis from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.28

Species neriensis (Martynova, 1976, s.d.) Martynova, 1976

Species nikolai Kaprus, I, Weiner, W et Pasnik, G, 2016:126

Protaphorura nikolai from Russia Far East
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.18

Species nutak (Yosii, R, 1972:86) Pomorski, RJ, 2002:il

Protaphorura nutak from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.32

Species octopunctata (Tullberg, T, 1877:40) Kloet, GS & Hincks, WD, 1945:5, B

            Syn.: Lipura octo-punctata(sic) Tullberg, T, 1877:40 IOS
            Syn.: Lipura disjuncta Moniez, 1894
            Syn.: Aphorura octopunctata var. edenticulata Wahlgren, 1900
             Syn.: Onychiurus octopunctata(sic) ssp. edenticulata(sic) Handschin, 1921 ISS
             Syn.: Onychiurus octopunctata(sic) var. edenticulata(sic) Handschin, E in Dahl, F, 1929:47 ISS
             Syn.: Onychiurus octopunctatus var. edenticulata(sic) Kos, F, 1939:285 ISS
             Syn.: Onychiurus octopunctatus ssp. edenticulata(sic) Kos, 1941 ISS
            Syn.: Onychiurus daviesi Stach, J, 1954:137,sd
            Syn.: Onychiurus octopunctatus f. daviesi Gisin, 1943 =?
            Syn.: Onychiurus armatus ssp. quadriocellatus Gisin, 1952
            Syn.: Onychiurus discrepans Stach, 1954
            Syn.: Onychiurus (Protaphorura) flavorufulus Martynova, 1976
            Syn.: Protaphorura glebata Arbea & Jordana, 1985, partim
            Syn.: Protaphorura quadriocellata Arbea & Jordana, 1985


             Syn.: Lipura octo-punctata(sic) Schött, 1893 ISS
             Syn.: Aphorura 8-punctata(sic) Absolon, K, 1901:381 ISS
             Syn.: Onychiurus octopuntata(sic) Jacquemart, S, 1967:4,B ISS
Protaphorura octopunctata from Russia
After Pomorski RJ & Kaprus IJ, 2007 Fig.1

             Syn.: Protaphorura octopunctatus(sic) Kaprus, IJ et Pomorski, RJ, 2008:667 ISS




Species okaensis Becker, 1905, i.s.

            Syn.: Onychiurus okaënsis(sic) Becker, 1905 IOS
             Syn.: Onychiurus (Protaphorura) okaënsis(sic) Gisin, H, 1960:112 ISS
             Syn.: Onychiurus (Protaphorura) okaënsis(sic) Grinbergs, 1960 ISS
             Syn.: Protaphorura okaënsis(sic) Salmon, JT, 1964:173 ISS



Species oligopseudocellata Kaprus, I, Weiner, W et Pasnik, G, 2016:129

Protaphorura oligopseudocellata from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.26

Species ombrophila (Stach, J, 1960:509) Yosii, 1966

Species orthacantha (Handschin, 1920, s.d.) Salmon, JT, 1964:173, B

Species pannonica (Haybach, 1960) Pomorski, 1990

            Syn.: Onychiurus trinotatus Gisin, 1961
             Syn.: Onychiurus trinotatatus(sic) Parimuchová, A et Kovác, L, 2016:263 ISS
            Syn.: Onychiurus pseudocellatus Szeptycki, A et Weiner, WM in Razowski J, 1990:21
            Syn.: Onychiurus pseudocellatus Fjellberg, A, 1998:116


Species parallata (Gisin, 1952, s.d.) Salmon, JT, 1964:174

Species paucisetosa (Hammer, 1942) Skidmore, 1995:53

             Syn.: Protaphorura paucisetosus(sic) Skidmore, 1995 ISS

Species persica Arbea, JI et Kahrarian, M, 2017:144

Protaphorura persica from Iran
After Arbea JI & Kahrarian M, 2017 Fig.14

Species pjasinae (Martynova, 1976) Pomorski, RJ, 2002:il

            Syn.: Onychiurus armatus f. scanica Bödvarsson, 1970 =?
            Syn.: Protaphorura boedvarssoni Pomorski, RJ, 1993:125


Protaphorura pjasinae from Russia
After Babenko AB & Kaprus IJ, 2014 Fig.2.1

Species pseudarmata (Folsom, 1917, s.d.) Salmon, JT, 1964:174

             Syn.: Onychiurus pseudoarmatus(sic) Aitchison, CW, 1984:299 ISS
             Syn.: Protaphorura pseudarmatus(sic) Skidmore, RE, 1995:53 ISS

Species pseudocellata (Naglitsch, 1962)

Species pseudoglebata Arbea, JI et Jordana, R, 1989:11

Species pseudostyriaca (Loksa, I, 1964:88, s.d.) Pomorski, RJ, Leithner, C & Bruckner, A, 2003:307

             Syn.: Protaphorura pseudostiriaca(sic) Gulgenova, A et Potapov, M, 2013:204 ISS
             Syn.: Protaphorura pseudostiriaca(sic) Parimuchová, A et Kovác, L, 2016:267,sd ISS

Species pseudovanderdrifti (Gisin, 1957) Salmon, JT, 1964:174

Species pulvinata (Gisin, 1954) Salmon, JT, 1964:174

Species quadriocellata (Gisin, 1947) Salmon, JT, 1964:170, B

             Syn.: Onychiurus 4-ocellatus(sic) Bockemuhl, 1956 ISS
             Syn.: Onychiurus 4-ocellatus(sic) Haybach, 1959 ISS
Protaphorura quadriocellata from Europe
After Pomorski RJ & Kaprus I, 2007 Fig.11



Species quercetana Mateos Frias, EM et Arbea, JI, 1986:285

Species sakatoi (Yosii, 1966)

            Syn.: Onychiurus octopunctatus Stach, 1934 partim
            Syn.: Protaphorura octopunctata Skarzynski, D, Pomorski, RJ, Smolis, A, Weiner, WM, Szeptycki, A, Slawska, M et Sterzynska, M, 2002:
            Syn.: Protaphorura rectopunctata Busmachiu, 1996
            Syn.: Onychiurus (Protaphorura) tetragrammatus ssp. serbicus Loksa & Bogojevic, 1967


Species salsa Kaprus, IJ, Pasnik, G et Weiner, WM, 2014:468

Protaphorura salsa from Russia
After Kaprus IJ, Pasnik G & Weiner WM, 2014 Fig.9

Species saltuaria Pomorski, RJ et Kaprus, I, 2007:135

Protaphorura saltuaria from Ukraine
After Pomorski RJ & Kaprus I, 2007 Fig.16

Species sayanica Kaprus, I, Weiner, W et Pasnik, G, 2016:134

Protaphorura sayanica from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.37

Species sensillata (Khanislamova, 1986, s.d.) Pomorski, RJ, 2002:il

Species septempapillata Palissa, 1986

Species spinoidea (Steiner, 1955) Salmon, JT, 1964:175

Species stalagmitorum (Absolon, K, 1900:407, s.d.) Pomorski, RJ, 2002:il

Species stiriaca (Stach, J, 1946:13) Salmon, JT, 1964:175

Species stogovi Pomorski, RJ, 1993:121

Species subarmata (Gisin, 1957) Salmon, JT, 1964:175

            Syn.: Onychiurus subarmatus Gisin, 1957
            Syn.: Protaphorura subarmata f. occidentalis Pomorski, RJ, 1998:95
            Syn.: Protaphorura subarmata f. silesciaca Pomorski, RJ, 1998:95


             Syn.: Protaphorura subaramata(sic) Pomorski, RJ, 1996:98 ISS

Species subaequalis (Bagnall, 1937, s.d.) Kloet, GS & Hincks, WD, 1945:5

Remark by Pitkin.R., B 1980:423: Type specimen not adult.

Species subarctica (Martynova, 1976) Pomorski, RJ, 2002:il

Protaphorura subarctica from Russia
After Babenko AB & Kaprus IJ, 2014 Fig.3.1

Species subfimata Thibaud & Christian, 1986

Species submersa Kaprus, IJ et Pomorski, RJ, 2008:681

Protaphorura submersa from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.37

Species suboctopunctata Khanislamova, 1986

Species subuliginata (Gisin, H, 1956:339, s.d.) Salmon, JT, 1964:175, B

            Syn.: Onychiurus subuliginatus Gisin, H, 1956:339
             Syn.: Protaphorura trivontörnei(sic) Salmon, JT, 1964:175 ISS
             Syn.: Onychiurus (Protaphorura) trivortoernei(sic) Loksa, I et Rubio, I, 1966:150 ISS
            Syn.: Onychiurus mediovanderdrifti Haybach, 1960
            Syn.: Protaphorura subarmata Pomorski, 1990
            Syn.: Protaphorura subarmata Hopkin, SP, 1997:94
            Syn.: Protaphorura subarmata Fjellberg, 1998


             Syn.: Protaphorura subluliginata(sic) Zinkler, 1971 ISS
             Syn.: Onychiurus subliginatus(sic) Pitkin, BR, 1980:423 ISS
             Syn.: Protaphorura subluliginata(sic) Saur, E et Ponge, J-F, 1988:371 ISS

Species svontoernei (Gisin, 1957, s.d.) Salmon, JT, 1964:175

            Syn.: Onychiurus s-vontoernei(sic) Gisin, 1957 IOS


             Syn.: Onychiurus s-vontornei(sic) Haybach, 1959 ISS
             Syn.: Onychiurus (Protaphorura) s-vontoernei(sic) Gisin, H, 1960:121 ISS
             Syn.: Onychiurus (Protaphorura) s-vontoerneri(sic) Goto, HE et Lawrence, PN in Kloet, GS et Hincks, WD, 1964:6 ISS
             Syn.: Protaphorura s-vontörnei(sic) Salmon, JT, 1964:175 ISS
             Syn.: Onychiurus armatus f. s-vontoernei(sic) Bödvarsson, H, 1970:8 ISS
             Syn.: Onychiurus s-vontoernei(sic) Rusek, J in Dlabola, J, 1977:13 ISS
             Syn.: Onychiurus svontornei(sic) Pitkin, BR, 1980:423 ISS
             Syn.: Protaphorura s-vontoerneri(sic) Hopkin, SP, 1997:95 ISS
Remark by Nicolson, D. 2007.mm.dd:in litt.: Based on art.NN the hyphen in the original spelling is removed.


             Syn.: Protaphorura s-vontoernei Parimuchová, A et Kovác, L, 2016:260,sd ISS



Species taimyrica (Martynova, 1976) Pomorski, RJ, 2002:il

Species teres (Yosii, R, 1956:53, s.d.) Salmon, JT, 1964:174

Species tetragrammata (Gisin, 1961) Kaprus, IJ & Pomorski, RJ, 2008:684

Protaphorura tetragrammata from Russia
After Kaprus IJ & Pomorski RJ, 2008 Fig.47
             Syn.: Protaphorura tetragramata(sic) Fiera, C et Weiner, WM, 2013:3 ISS
             Syn.: Protaphorura (octopunctata-group) tetragramata(sic) Fiera, C et Weiner, WM, 2013:4 ISS
             Syn.: Protaphorura (octopunctata-group) tetragramata(sic) Fiera, C et Weiner, WM, 2013:5 ISS


Species tolae Pomorski, RJ et Kaprus, I, 2007:131

Protaphorura tolae from Russia
After Pomorski RJ & Kaprus I, 2007 Fig.6

Species tricampata (Gisin, H, 1956:343) Salmon, JT, 1964:175

Species tschernovi (Martynova, 1976) Pomorski, RJ, 2002:il

Species tullbergi (Bagnall, 1937, s.d.) Kloet, GS & Hincks, WD, 1945:5

Species tundricola (Martynova, 1976) Pomorski, RJ, 2002:il

            Syn.: Onychiurus (Protaphorura) tundricola(sic) Martynova, 1976 IOS


Species tuvinica Kaprus, I, Weiner, W et Pasnik, G, 2016:137

Protaphorura tuvinica from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.45

Species unari Rusek, 1995

Species uniparis Gulgenova, A et Potapov, M, 2013:211

Protaphorura uniparis from Russia
After Gulgenova A & Potapov M, 2013 Fig.23

Species ussurica (Martynova, 1981, s.d.) Pomorski, RJ, 2002:il

Species valsainensis Acón, 1981

Species vasilinae Kaprus, I, Weiner, W et Pasnik, G, 2016:140

Protaphorura vasilinae from South Siberia
After Kaprus I, Weiner W & Pasnik G, 2016 Fig.51

Species yagii (Miyoshi, 1923) Salmon, JT, 1964:174

Species zlatiborensis Lucic, Curcic, Pavkovic-Lucic & Tomic, 2008

Species zloti (Curcic & Lucic, 1997) Parimuchová, A & Kovác, L, 2016:254

Species zori (Martynova in Martynova & Chelnokov, 1975, s.d.) Pomorski, RJ, 2002:il


Genus Spelaphorura Bagnall, 1948

Species glenniei Bagnall, 1948, t.t.


Genus Jacekaphorura Pomorski, RJ et Babenko, A, 2010:65

Species sakhaensis Pomorski, RJ et Babenko, A, 2010:67, t.t.

Jacekaphorura sakhaensis from Siberia
After Pomorski RJ & Babenko A, 2010 Fig.6


Genus Yoshiiphorura Jordana R et Martínez M in Martínez, M, Baquero, E, Barranco, P, Ariño, AH et Jordana, R, 2004:3

Species bellingeri Jordana R et Martínez M in Martínez, M, Baquero, E, Barranco, P, Ariño, AH et Jordana, R, 2004:7, t.t.


Genus Megaphorura Fjellberg, A, 1998:108

Species arctica (Tullberg, T, 1877:39) Fjellberg, A, 1998:108, t.t.

            Syn.: Lipura incerta Moniez, 1893
            Syn.: Onychiurus pseudarmatus Folsom, 1917


             Syn.: Onychiurus armatus var. arctica(sic) Denis, J-R, 1924:224 ISS
             Syn.: Onychiurus armatus var. arctica(sic) Denis, J-R, 1924:225 ISS
             Syn.: Onychiurus armata(sic) ssp. arctica(sic) Schött, 1924 ISS
             Syn.: Onychiurus armatus ssp. arctica(sic) Stach, 1934 ISS
             Syn.: Protaphorura artica(sic) Hopkin, 1997 ISS
             Syn.: Onychiurus antarcticus(sic) Worland, Grubor-Lajsic & Montiel, 1998 ISS
Megaphorura arctica from the UK
2002 © Hopkin, S.P.
Megaphorura arctica from the Arctic region
2003.03.06 © Holmstrups, M.
Megaphorura arctica from the Arctic region, dehydrated
2003.03.06 © Holmstrups, M.
Megaphorura arctica from the Arctic region, hydrated
2007 © Borland, R.
Megaphorura arctica from the Arctic region, dehydrated
2007 © Borland, R.
Megaphorura arctica from Sweden
2007.12.03 © Hall, K.

             Syn.: Onychiurus antarcticus(sic) Zettel, J et Zettel, U, 2008:104 ISS
Megaphorura arctica from Norway
2011 © Hermansen, P.



Species evansi Bagnall, 1935

Remark by Pitkin.R., B 1980:423: This species lacks a rudimentary furca and belongs to the arcticus group.


Tribus Thalassaphorurini Pomorski RJ, 1998:134 sensu Babenko AB, Chimitova AB & Stebaeva SK, 2011:3 key


Genus IncertaeSedis Sun, X, Chen, J-X et Deharveng, L, 2011:32

Species caprariae (Dallai, 1969) Sun, X, Chen, J-X & Deharveng, L, 2011:32, i.s.

Species michelbacheri (Bagnall, 1948) Sun, X, Chen, J-X & Deharveng, L, 2011:32, i.s.

Species sensitivus (Handschin, 1928) Sun, X, Chen, J-X & Deharveng, L, 2011:32, i.s.


Genus Spinonychiurus Weiner, WM, 1996:188

Spinonychiurus epaphius from Ukraine
After Kaprus' IJ & Tsalan JV, 2009 Fig.1.6
Abdominal sternum III divided into 2 subsegments

Species arabelensis Pomorski, RJ et Kaprus, IJ, 2015:118

Spinonychiurus arabelensis from Kyrgystan
After Pomorski RJ & Kaprus IJ, 2015 Fig.65
             Syn.: Spinonychiurus alabelensis(sic) Pomorski, RJ et Kaprus, IJ, 2015:101 ISS


Species alaskensis Pomorski, RJ et Kaprus, IJ, 2015:115

Spinonychiurus alaskensis from Alaska
After Pomorski RJ & Kaprus IJ, 2015 Fig.58

Species edinensis (Bagnall, 1935 tt) Weiner, WM, 1996:188, t.t.

            Syn.: Onychiuroides spinosus Bagnall, 1949
            Syn.: Onychiurus spinularius Murphy, 1960


Allonychiurus edinensis from the UK
Anal spines; 4 stout setae on abd.5
2002 © Hopkin, S.P.
Onychiurus edinensis from Holland
2014.03.14 © van Duinen, J.
Det.: Berg, M.




             Syn.: Spinonychiurus edinesis(sic) Pomorski, RJ et Kaprus, IJ, 2015:104 ISS

Species epaphius Kaprus, IJ et Tsalan, JV, 2009:174

Spinonychiurus epaphius from Ukraine
After Kaprus' IJ & Tsalan JV, 2009 Fig.1.1

Species issykkulensis Pomorski, RJ et Kaprus, IJ, 2015:111

Spinonychiurus issykkulensis from Kyrgystan
After Pomorski RJ & Kaprus IJ, 2015 Fig.36

Species natashae Pomorski, RJ et Kaprus, IJ, 2015:115

Spinonychiurus natashae from Kazakhstan
After Pomorski RJ & Kaprus IJ, 2015 Fig.57
             Syn.: Spinonychiurus natashe(sic) Pomorski, RJ et Kaprus, IJ, 2015:120 ISS


Species nazguli Pomorski, RJ et Kaprus, IJ, 2015:113

Spinonychiurus nazguli from Kyrgystan
After Pomorski RJ & Kaprus IJ, 2015 Fig.44

Species pamirensis (Martynova, 1975) Pomorski, RJ & Kaprus, IJ, 2015:110

Species persicus Kaprus I, Shayanmehr M et Kahrarian M in Kaprus, I, Shayanmehr, M, Kahrarian, M et Lafooraki, EY, 2017:342

Spinonychiurus persicus from Iran
After Kaprus I & al, 2017 Fig.16

Species spinularius (Gisin, 1952) Pomorski, RJ & Kaprus, IJ, 2015:104

Spinonychiurus spinularius from Switzerland
After Pomorski RJ & Kaprus IJ, 2015 Fig.7

Species subedinensis (Arbea, JI & Jordana, R, 1985:66) Sun & al, 2009

Spinonychiurus subedinensis from Spain
After Pomorski RJ & Kaprus IJ, 2015 Fig.13




             Syn.: Spinonychiurus subedinesis(sic) Pomorski, RJ et Kaprus, IJ, 2015:106 ISS
             Syn.: Spinonychiurus subedinesis(sic) Pomorski, RJ et Kaprus, IJ, 2015:118 ISS

Species tianshanicus (Martynova, 1971) Pomorski, RJ & Kaprus, IJ, 2015:106

Spinonychiurus tianshanicus from Kyrgystan
After Pomorski RJ & Kaprus IJ, 2015 Fig.21

Species vandeli (Cassagnau, P, 1960:405) Sun, X, Chen, J-X & Deharveng, L, 2011:33

            Syn.: Onychiurus Vandeli(sic) Cassagnau, P, 1960:405 IOS





Genus Detriturus Pomorski, RJ, 1998:142

Species jubilarius (Gisin, 1957) Pomorski, RJ, 1998:143, t.t.

Species lotius (Palacios-Vargas, JG & Díaz, M, 1996:5) Pomorski, RJ, 2002:il

Species cassagnaui (Gers, 1980) Pomorski, RJ, 2002:il

Species trilobatus (Palacios-Vargas, JG & Díaz, M, 1996:7) Pomorski, RJ, 2002:il


Genus Sensillonychiurus Pomorski, RJ et Sveenkova, YB, 2006:191 key

Species sp. Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:23

Species amuricus Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:19

Species changchunensis Sun & Wu, 2012

Species eisi (Rusek, 1976) Babenko, AB, Chimitova, AB & Stebaeva, SK, 2011:1

             Syn.: Onychiurus eisei(sic) Palacios-Vargas, JG et Deharveng, L, 1982:6 ISS

Species geminus Pomorski, RJ et Sveenkova, YB, 2006:195

Species minusculus Pomorski, RJ et Sveenkova, YB, 2006:193, t.t.

Species mirus Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:10

Species pseudoreductus Sun, X et Wu, D-H, 2012:565

            Syn.: Sensillonychiurus pseudoreducta(sic) Sun, X et Wu, D-H, 2012:565 IOS
Sensillonychiurus pseudoreducta(sic) from China
After Sun X & Wu D-H, 2012 Fig.13




             Syn.: Sensillonychiurus pseudoreducta(sic) Sun, X, Deharveng, L et Wu, D-H, 2013:2 ISS



Species reductus Sun, X et Wu, D-H, 2012:564

            Syn.: Sensillonychiurus reducta(sic) Sun, X et Wu, D-H, 2012:564 IOS
Sensillonychiurus reducta(sic) from China
After Sun X & Wu D-H, 2012 Fig.1




             Syn.: Sensillonychiurus reducta(sic) Sun, X, Deharveng, L et Wu, D-H, 2013:2 ISS



Species taimyrensis Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:14

            Syn.: Tantulonychiurus volinensis Babenko, 2003
            Syn.: Tantulonychiurus volinensis Babenko & Fjellberg, 2006 partim

Species vegae Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:16

            Syn.: Tantulonychiurus volinensis Babenko & Fjellberg, 2006 partim

Species virginis Pomorski, RJ et Sveenkova, YB, 2006:193

Species vitimicus Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:21


Genus Thalassaphorura Bagnall, 1949 key

  Syn.: Podurida(sic) Anurophorus Kolenati, FA, 1858:131 partim
  Syn.: Onychiuridae Allaphorura Bagnall, 1949
  Syn.: Onychiuridae Handschiniella Bagnall, 1949
  Syn.: Onychiurus Parthenaphorura Yoshii, R et Suhardjono, YR, 1989:31
  Syn.: Onychiurus Jailolaphorura Yoshii, R et Suhardjno, YR, 1992:9


  Syn.: Onychiurinae Talassaphorura(sic) Stach, J, 1954:7 ISS
  Syn.: Onychiurinae Talassaphorura(sic) Stach, J, 1954:22 ISS

Species alborufescens (Vogler, 1895) Arbea, JI, 2017:323

            Syn.: Anurophorus Kollarii(sic) Kolenati, 1858
             Syn.: Anurophorus Kollarii(sic) Kolenati, FA, 1858:131 ISS
             Syn.: Lipura Kollarii(sic) Lubbock, J, 1873:195 ISS
             Syn.: Aphorura Kollarii(sic) Absolon, K, 1901:213 ISS
             Syn.: Aphorura Kollarii(sic) Absolon, K, 1901:381 ISS


             Syn.: Aphorura albo-rufescens(sic) Latzel, 1907 ISS
             Syn.: Protaphorura albo-rufescens(sic) Salmon, 1964 ISS
             Syn.: Protaphorura albrufescens(sic) Salmon, 1964 ISS

Species bagnalli (Salmon, JT, 1959:155) Parimuchová, A & Kovác, L, 2016:261

            Syn.: Onychiurus differens Bagnall, 1934 nn


Species bapen Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species biquaternata Sun & Li, 2015

Species bisetosa Sun & Wu, 2013

Species brevisetosa Sun, X, Gao, Y et Potapov, MB, 2014:576

Species butrosi (Christiansen, K, 1956:126) Pomorski, RJ, 2002:il

Species debilis (Moniez, 1889) Pomorski, RJ, 2002:il

            Syn.: Aphorura neglecta Schäffer, 1896
            Syn.: Onychiurus litoreus Folsom, 1917
            Syn.: Onychiurus evansi Bagnall, 1935
            Syn.: Onychiurus littoralis Dürkop, 1935
             Syn.: Protaphorura litoralis(sic) Salmon, JT, 1964:172 ISS
            Syn.: Onychiurus imminutus Bagnall, 1937


Thalassaphorura debilis from the UK
2002 © Hopkin, S.P.
Thalassaphorura debilis from Holland
2018.09.30 © van Leeuwen, J.

Species duplopunctata (Strenzke, 1954) Pomorski, RJ, 2002:il

            Syn.: Onychiurus debilis Gisin, 1953, nec Moniez, 1890

Species encarpata (Denis, 1931) Fjellberg, A, 1998:111

            Syn.: Onychiurus hortensis Gisin, 1949
            Syn.: Onychiurus parthenogeneticus Choudhuri, DK, 1958:155
            Syn.: Protaphorura prosensillata Najt, J, Thibaud, J-M et Jacquemart, S (+), 1991:163


Species ghatensis (Prabhoo, 1971) Pomorski, RJ, 2002:il

Species grandis Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species guangdongensis Sun & Li, 2015

Species guangxiensis Sun, Bedos & Deharveng, 2017

Species guineaensis (Salmon, 1965) Pomorski, RJ, 2002:il

Species hainanica Sun, X, Gao, Y et Potapov, MB, 2014:580

             Syn.: Thalassaphorura hainamica(sic) Kaprus, I et Pasnik, G, 2017:243 ISS

Species halophila (Bagnall, 1937) Pomorski, RJ, 2002:il

            Syn.: Onychiurus littoralis Bagnall, 1935 nec Dürkop, 1935


Thalassaphorura halophila from the UK
2002 © Hopkin, S.P.

Species ishikawai (Yosii, 1956) Pomorski, RJ, 2002:il

Species jailolonis (Yoshii, R & Suhardjno, YR, 1992:9) Pomorski, RJ, 2002:il

Species kirgisica (Martynova, 1971) Arbea, JI, 2017:323

Species kwona (Thibaud & Lee, 1994) Pomorski, RJ, 2002:il

Species lagunensis (Palacios-Vargas, JG & Díaz, M, 1996:2) Pomorski, RJ, 2002:il

            Syn.: Onychiurus encarpatus Vázquez & Palacios-Vargas, 1990 ca


Species lifouensis (Thibaud, J-M & Weiner, WM in Najt, J & Matile, L, 1997:72) Pomorski, RJ, 2002:il

Species linzhiensis Sun & Li, 2015

Species lolae Arbea, JI, 2017:322

Thalassaphorura lolae from Spain
After Arbea JI, 2017 Fig.1

Species macrospinata Sun & Wu, 2012

Species microspinata Sun, Bedos & Deharveng, 2017

Species moniezi (Bagnall, 1935) Pomorski, RJ, 2002:il

            Syn.: Onychiurus evansi Bagnall, 1935


Remark by Pitkin.R., B 1980:423: Type specimen not adult.

Species orientalis (Stach, J, 1964:9) Pomorski, RJ, 2002:il

Species petaloides (Rusek, 1981) Pomorski, RJ, 2002:il

Species petiti Sun & Wu, 2013

Species pomorskii Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species problematica Sun, X, Deharveng, L et Wu, D-H, 2013:3

Species qinlingensis Sun & Wu, 2013

Species qixiaensis Yan, Shi & Chen, 2006

Species reducta Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species sensilata (Thibaud & Massoud, 1980) Sun, X, Chen, J-X & Deharveng, L, 2011:32

             Syn.: Onychiurus (Protaphorura) sensillata(sic) Palacios-Vargas, 1990 ISS

Species tamdaona (Nguyen, 1995) Parimuchová, A & Kovác, L, 2016:261

Species tamurai (Yue, Q-Y & Yin, W-Y, 2000:44) Pomorski, RJ, 2002:il

Species tenuis Babenko, AB, 2007:369

Species thaibinhensis (Nguyen, 2001) Kaprus, I & Pasnik, G, 2017:242

Species thalassophila (Bagnall, 1937) Bagnall, 1949

            Syn.: Thalassaphorura debilis Fjellberg, A, 1998:109


             Syn.: Spelaphorura thalassophilus(sic) Salmon, JT, 1959:149 ISS
             Syn.: Onychiurus (Protaphorura) thalassophila(sic) Gisin, H, 1960:123,si ISS

Species tiani Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species tibiotarsalis Sun, X, Chen, J-X et Deharveng, L, 2010:?

Species tovtrensis (Kaprus, IJ & Weiner, WM, [1995]:61) Thibaud & al, 1999

Allaphorura tovtrensis from Ukraine
After Kaprus I.J. & Weiner W.M., [1995] Fig.9

Species xihuensis Sun & Li, 2015

Species yodai (Yosii, 1966) Pomorski, RJ, 2002:il

Species yolandae (Izarra, 1971) Kaprus, I & Pasnik, G, 2017:242

Species zschokkei (Handschin, 1919) Pomorski, RJ, 1998:139

            Syn.: Onychiurus zschokkei Handschin, 1919, partim Handschin, 1924


             Syn.: Onychiurus Zschokkei(sic) Handschin, 1926 ISS
  Subspecies armata Handschin, E, 1924:102

            Syn.: Protaphorura zschokkei Handschin, 1919, partim Handschin, 1924



Species-group devoid-of-anal-spines
Species aspinata Kaprus, I et Pasnik, G, 2017:226

Thalassaphorura aspinata from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.1

Species cryptopyga (Denis, 1931) Pomorski, RJ, 2002:il

            Syn.: Onychiurus (Parthenaphorura) bogoriensis Yoshii, R et Suhardjono, YR, 1989:31


Species franzi (Stach, J, 1946:11) Pomorski, RJ, 2002:il, t.t.

Species hoguei (Palacios-Vargas, JG & Díaz, M, 1996:10) Pomorski, RJ, 2002:il

Species sayana Kaprus, I et Pasnik, G, 2017:228

Thalassaphorura sayana from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.9

Species sibirica Kaprus, I et Pasnik, G, 2017:230

Thalassaphorura sibirica from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.16

Species stebaevae Kaprus, I et Pasnik, G, 2017:232

Thalassaphorura stebaevae from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.24

Species stepensis Kaprus, I et Pasnik, G, 2017:234

Thalassaphorura stepensis from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.31

Species weinerae Kaprus, I et Pasnik, G, 2017:236

            Syn.: Thalassaphorura (devoid-of-anal-spines-group) weineri(sic) Kaprus, I et Pasnik, G, 2017:236 IOS
Thalassaphorura weineri(sic) from South Siberia
After Kaprus I & Pasnik G, 2017 Fig.38


Species-group insufficiently-described


Species boskovae (Lucic, Dimitrijevic & Mihajlova, 2003) Parimuchová, A & Kovác, L, 2016:261

Species clayae (Salmon, 1958) Pomorski, RJ, 2002:il

Species dinghuensis (Lin & Xia, 1985) Pomorski, RJ, 2002:il

Species hera (Christiansen, K & Bellinger, P, 1980:419) Pomorski, RJ, 2002:il

             Syn.: Allaphorura herus(sic) Weiner, WM, 1996:176 ISS

Species martynovae (Dunger, 1978) Pomorski, RJ, 2002:il

Species parvicornis (Mills, HB, 1934:33) Pomorski, RJ, 2002:il

Species petallata (Salmon, JT, 1958:709) Pomorski, RJ, 2002:il

Species saccardyi (Denis, JR, 1935:230) Arbea, JI, 2017:323

            Syn.: Onychiurus Saccardyi(sic) Denis, JR, 1935:230 IOS

Species tridentata (Stebaeva, 1982) Parimuchová, A & Kovác, L, 2016:261


Genus Micronychiurus Bagnall, 1949

Species aguzouensis (Deharveng, 1978) Weiner, WM, 1996:185

Species aranzadii (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:29) Weiner, WM, 1996:186

Species bernardoi (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:22) Weiner, WM, 1996:185

Species borensis (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:30) Weiner, WM, 1996:186

            Syn.: Onychiurus aguzouensis da Gama, 1985

Species duprei (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:26) Weiner, WM, 1996:186

Species maiteae (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:28) Weiner, WM, 1996:186

Species minutus (Denis, 1932) Weiner, WM, 1996:185, t.t.

Species sorogainensis (Beruete, E, Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:24) Weiner, WM, 1996:185


Genus Agraphorura Pomorski, RJ, 1998:150 key

Species acuitlapanensis (Palacios-Vargas, JG & Deharveng, L, 1982:3) Pomorski, RJ, 2004:76

             Syn.: Agraphorura acuitlapansis(sic) Palacios-Vargas, JG, 2006:10 ISS

Species calvoi Arbea, JI, 2005:21

Species fernandae (Oliveira & Thibaud, 1992) Pomorski, RJ, 2004:76

Species gambiaria (Murphy, 1965) Pomorski, RJ, 2004:76

Species mariapetrae (Thibaud, 1993) Pomorski, RJ, 2004:76

Species naglitschi (Gisin, 1960) Pomorski, RJ, 1998:150, t.t.

             Syn.: Agraphorura naglitshi(sic) Pomorski, RJ, 1998:150,tt ISS
             Syn.: Agraphorura naglitshi(sic) Pomorski, RJ, 2004:76 ISS

Species portucalensis (da Gama, 1964) Pomorski, RJ, 2004:76

Species pseudojusti (Thibaud & Massoud, 1980) Pomorski, RJ, 2004:76

Species sangelensis Kaprus, IJ et Stebaeva, SK, 2006:351

Agraphorura sangelensis from Southern Siberia
After Kaprus I.J. & Stebaeva S.K, 2006 Fig.1

Species spelaea Pomorski, RJ, 2004:76

Species xenonis (Ellis, WN, 1976:234) Pomorski, RJ, 2004:76


Genus Allonychiurus Yoshii, 1995 key

  Syn.: ... Pseudonychiurus Lin S-X, 1980, nec Bagnall, 1948
  Syn.: Onychiurini Tantulonychiurus Pomorski, RJ, 1996:94
  Syn.: Onychiurini Tantul-onychiurus(sic) Pomorski, RJ, 1996:100 ISS
  Syn.: Onychiurini Thibaudichiurus Weiner, WM, 1996:189
  Syn.: Onychiurinae Onychurus(sic) Yadav, RS, 2017:11057 partim ISS


Species sp. Qazi & Shayanmehr, 2016

Species antennalis Sun, X, Chen, J-X et Deharveng, L, 2011:35

Species asiaticus (Babenko, AB, 2007:366) Babenko, AB, Chimitova, AB & Stebaeva, SK, 2011:35

Species completus Babenko & Fjellberg, 2016

Species donjiensis (Lee, B-H & Kim, J-T, 1994:208) Babenko, AB, 2007:369

Species elikonius Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:26

Species flavescens (Kinoshita, 1916) Yoshii, 1995 tt

            Syn.: Onychiurus oreadis Christiansen, K et Bellinger, P, 1980:404
            Syn.: Onychiurus oreadis Poole, RW in Poole, RW et Gentili, P, 1997:53
            Syn.: Onychiurus oreadis Christiansen & Bellinger, 1998


Species foliatus (Rusek, J, 1967:189) Babenko, AB, 2007:369

Species hangchowensis (Stach, J, 1964:8) Xiong, Y, Gao, Y, Yin, W-Y & Luan, Y-X, 2008:730

Species indicus (Choudhuri & Roy, 1965) Sun, X, Chen, J-X & Deharveng, L, 2011:31

Onychiurus indicus from India
After Mandal GP & Hazra AK, 2009 Fig.14a
             Syn.: Onychiurus indica(sic) Mandal, GP et Hazra, AK, 2009:115 ISS


Species jindoensis (Lee, B-H & Kim, J-T, 1994:211) Babenko, AB, 2007:369

Species jongaksanensis (Weiner, 1989) Weiner, WM, 1996:182

Species kimi (Lee, 1973) Yoshii, 1995

Species mariangeae (Thibaud & Lee, 1994) Babenko, AB, 2007:369

Species mediaseta (Lee, 1974) Weiner, WM, 1996:182

             Syn.: Allonychiurus mediasetus(sic) Sun, X, Chen, J-X et Deharveng, L, 2011:30 ISS

Species megasomus Sun, Yan & Chen, 2009

Species pseudocellitriadis (Lee, 1974) Weiner, WM, 1996:182

Species shanghaiensis (Rusek, 1971) Sun & al, 2009

Species shinbugensis (Lee, 1974) Weiner, WM, 1996:182

             Syn.: Allonychiurus shibougensis(sic) Weiner, WM, 1996:182 ISS

Species songi Sun & Wu, 2012

Species subvolinensis Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:30

Species unisetosus Babenko, AB, Chimitova, AB et Stebaeva, SK, 2011:33

            Syn.: Onychiurus (Onychiurus) sp. Stebaeva & al, 2001

Species volinensis (Szeptycki, 1964) Babenko, AB, Chimitova, AB & Stebaeva, SK, 2011:29, t.t.


Tribus Onychiurini Börner, 1906


Genus IncertaeSedis Arbea J, 2012:35

Species cadaverinus (Handschin, 1921) Arbea, JI, 2012:35

Remark by Arbea, J. 2012:35: For the absence of anal spines, this species departs from Onychiurus.

Species certus (Nicolet, H, 1847:386) Arbea, JI, 2012:35

Anurophorus certus from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.4.
Remark by Stach, J. 1947:?: Onychiurus indeterminable.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.


Species dubius (Nicolet, H, 1847:386) Arbea, JI, 2012:35

Remark by Stach, J. 1947:?: Onychiurus indeterminable.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species fimetarioides (Denis, 1938) Arbea, JI, 2012:35

            Syn.: Onychiurus pseudofimetarius Handschin, 1921, nec Folsom, 1912
            Syn.: Onychiurus fimetarioides var. talpae Delamare Deboutteville, C, 1947:239


Remark by Arbea, J. 2012:35: For the absence of anal spines, this species departs from Onychiurus.

Species justi (Denis, 1938) Arbea, JI, 2012:35

            Syn.: Onychiurus Justi(sic) Denis, 1938 IOS
            Syn.: Onychiurus fimetarius Folsom, 1917, nec von Linné, 1767
            Syn.: Onychiurus justi ssp. porteri Tomlin, 1975


Remark by Christiansen, K. & Bellinger, P.F. 1998:?: The name justi is not based on any real population and should be abandoned.

Remark by Arbea, J. 2012:35: For the absence of anal spines, this species departs from Onychiurus.



Species justiprovisorius (Denis, 1938) Arbea, JI, 2012:36

            Syn.: Onychiurus justi-provisorius(sic) Denis, 1938 IOS
            Syn.: Onychiurus justi Deharveng & al, 2011


             Syn.: Onychiurus justi-provisorius(sic) Salmon, JT, 1964:183 ISS
             Syn.: IncertaeSedis justi-provisorius(sic) Arbea, JI, 2012:36 ISS



Species pancici (Curcic, BPM & Lucic, LR, 1999:190) Arbea, JI, 2012:36

Remark by Arbea, J. 2012:36: For its granulated sense clubs on antennal organ III, the postantennal organ of "sibiricus" type and the absence of anal spines, this species departs from Onychiurus.

Species ravanicae (Curcic, BPM & Lucic, LR, 1998:71) Arbea, JI, 2012:36

Remark by Arbea, J. 2012:36: For its paddle-shaped clubs on antennal organ III and the absence of anal spines, this species departs from Onychiurus. A redescription of this species would probably result in its reallocation to Orthonychiurus.

Species trojan (Curcic, BPM & Lucic, LR, 1998:70) Arbea, JI, 2012:36

Remark by Arbea, J. 2012:36: For its granulated sense clubs on antennal organ III and the absence of anal spines, this species departs from Onychiurus. A redescrition of the species would probably result in its reallocation to Onychiuroides.


Genus Similonychiurus Pomorski, RJ, 2007:49

Species voegtlini (Christiansen, K & Bellinger, P, 1980:446) Pomorski, RJ, 2007:50, t.t.

Similonychiurus voegtlini from the USA
After Pomorski RJ, 2007 Fig.9


Genus Onychiurus Gervais, 1841 key  key  key

  Syn.: ... Lipura Burmeister, 1838 nec Illiger, 1811
  Syn.: ... Anurophorus auctt. partim nec Nicolet, 1841
  Syn.: ... Adicranus Bourlet, 1843
  Syn.: Aphoruridæ(sic) Aphorura MacGillivray, AD, 1893:313


  Syn.: Tullbergia neomexicana Scott, 1961 partim paratypes
Remark by ICZN. 2007:575: Placed on the Official List of Names in Zoology. Opinion 291.

Onychiurus sp. from Japan
2008 © Tsukiji, T.
Onychiurus sp. from Portugal
Post-antennal organ: double array of globular vesicles in long groove
2012.10.13 © Murray, A.

  Syn.: Onychiurinae Onychurus(sic) Yadav, RS, 2017:11056 partim ISS
  Syn.: Onychiurinae Onychurus(sic) Yadav, RS, 2017:11057 partim ISS


Species sp.1 Absolon, K, 1900:191

Species sp.2 Leruth, R, 1939:198, B

Species sp.3 Palacios-Vargas, JG et Díaz, M, 1996:168

Species sp.4 Pomorski, RJ et Steinmann, D, 2004:20

Species sp.5 Kaprus, IJ, Pomorski, RJ, Skarzynski, D et Potapov, MB, 2005:893

Species sp.6 Elliott, WR, 2007:143

Species sp.7 Elliott, WR, 2007:143

Species ambulans (Linnæus, C, 1758:609) Gervais, 1841, t.t., B

Remark by Linnæus, C. 1758:609: [Podura] ambulans. 10. P. alba, furca extensa. Fn. suec. 1170. Habitat in Europa.' [Note that, while the diagnosis is in conflict with that of the refered species 1170 of the first edition of the Fauna Suecica (1746), Linnaeus, paradoxically, refers to the correct furca lacking onychiurid species.]

            Syn.: Pediculus terrestris niveus Linnæus, C, 1746:341
Remark by Linnæus, C. 1746:341 cited from Bellinger, P.F., 1997:(in litt.): 1170. Pediculus terrestris niveus. Habitat in terra subhumente calida, proinfrequens in terra prægnante et allis in hypocerusto per totem hyemem(sic). Descr. Fascies omnino Poduræ, teres oblongus albus, sed furca carens. Copiosa et gregarium vivit, mox ab adfusa aqua accedit ad superficiem terræ, fissa vero terra debet descendere.

Podura terrestris nivea from Sweden
After de Geer, C., 1778 Tom.VII Pl.3 Fig.5.

            Syn.: Anurophorus fimetarius Nicolet, H, 1842:53
Anurophorus fimetarius from Switzerland
After Nicolet, H., 1842 Pl.5 Fig.2.

            Syn.: Lipura agilis Moniez, 1889
            Syn.: Lipura noctiluca Dubois, 1896
            Syn.: Onychiurus laminatipes Bagnall, 1937
            Syn.: Onychiurus helveticus Denis, 1938
            Syn.: Lipura volvator Salmon, JT, 1964:176 partim



Remark by Linnæus, C. 1761 cited from Lubbock, 1873:5: P. ambulans alba, furca extensa.

Remark by von Linné, C. 1767 cited from Lubbock, 1873:5: P. alba, cauda bifida extensa obtusa. Corpus cylindricum, reliquis longius, album, molle. Punctum fuscum supra anum. Cauda setis 2 extensis, longitudine 1/2 corporis. Non salit.

Lipure ambulante from Europe
Lipura ambulans
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.2
Anurophorus ambulans from Europe
After Nicolet, H., 1847 Pl.6 Fig.14.
Remark by Nicolet, H. 1847:384-385: Anurophorus ambulans Nicolet, 1847:384 is based on Lipura ambulans Burmeister. [Note that Nicolet, based on the erroneous interpretation of the "cauda extensa" of Linnaeus as "deux petits crochets recourbés en dessus", synonymises in error the Podura ambulans of Linnaeus.]

             Syn.: Anurophorus Ambulans(sic) Kolenati, FA, 1858:131 ISS
Lipura ambulans from the UK
After Lubbock J, 1873 Pl.43 Fig.1
Lipura ambulans from the UK
After Lubbock J, 1873 Pl.43 Fig.2
Remark by Lubbock, J. 1873:5: Podura ambulans de Geer, 1782 does neither answer to the description of Linnæus (1761) nor to that of von Linné (1767), but appears to be Lipura ambulans Lubbock, 1873:189. 'It does not seem to have been noticed by any writer that Lipura ambulans is mentioned in the first edition of the "Faun. Suec.," though not among the Poduræ, being described as a Pediculus, although Linnæus adds "facies omnino Poduræ."'

Remark by Lubbock, J. 1873:7,190,213: Podura ambulans von Linné (1767) =? Campodea staphylinus. The Podura ambulans of Linnæus, both of, 1761 and, 1767 is described as having long caudal appendages. Campodea staphylinus Westwood, 1842 appears to be the species described by Linnæus under the name Podura ambulans in the "Systema Naturæ," ed. xii, [1767], though it is not the Podura ambulans of the Fauna Suecica [first ed. of, 1746].

Remark by Stach, J. 1954:60,157: Onychiurus ambulans Stach, 1954:60,157 is based on Anurophorus ambulans Nicolet, 1847:384, while reference is made to the redescription of Stach (1934). Lipura ambulans Burmeister, 1838 is listed as doubthfull synonym.

Remark by ICZN. 1958:25 cited from Salmon, 1964:89,176,178: Podura (Onychiurus) ambulans has been placed on the Official List of Specific Names as proposed by Gisin, Opinion 291 (1954:104).

Remark by Gisin, H. 1960:148: Refers to Stach (1934:174) and (1954:157) for Onychiurus (Onychiurus) ambulans, while doubthing the synonymy with Podura ambulans Linnæus, 1758:609.

Onychiurus ambulans from the UK
2002 © Hopkin, S.P.
Remark by Fjellberg, A. 2004.10.04(in litt.): A new neotype should be designated for Podura ambulans, as Pomorski's selection of a neotype from a German cave does not meet the "qualifying conditions" for such actions set by the ICZN (neotypes should be selected from material as nearly as practicable possible from the original type locality - which is Sweden).

Remark by Dányi, L. & Traser, G. 2008:23: All data published before description of Onychiurus circulans Gisin, 1952 are dubious.

Remark by Arbea, J. 2012:34: Fjellberg (1998) did not include O. ambulans in his Fennoscandian catalogue and, perhaps, it might well be the same as his northern O. circulans. All records are dubious until redescriptions of O. ambulans and O. circulans will be made.

Remark by Arbea, J. 2012:44: Both O. circulans and O. ambulans sensu Pomorski, 1998 key out together [suggesting being synonymous].


  Subspecies inermis Ågren, 1903


             Syn.: Onychiurus ambulans-inermis(sic) Denis, JR, 1924:197 ISS
  Subspecies intercedens Latzel, R, 1917:237



Species ambulans2 Stach, 1934

Remark by Arbea, J. 2012:44: O. ambulans sensu Stach, 1934 and Gisin, 1956 key out differently from O. ambulans sensu Pomorski, 1998.

Species arans Gisin, 1952, B

Species ariegicus Deharveng, 1978

Species bhattii Yosii, 1963

Remark by Arbea, J. 2012:34: A redescription of the species would probably result in its reallocation to another Onychiurini genus.

Species boldorii Denis, 1937

Species circulans Gisin, 1952, B

Remark by Dányi, L. & Traser, G. 2008:23: According to Pomorsky(sic) (1998) Onychiurus circulans is a junior synonym of Onychiurus ambulans (Linnaeus), but in our opinion it is a valid, separate taxon.

Remark by Arbea, J. 2012:34: A redescription of the type material would be needed.

Species circulans2 Gisin, 1958

Remark by Arbea, J. 2012:34: The populations of Madeira island might represent a closely related species (Gisin, 1958, Gama, 1961).

Species confugiens da Gama, 1962

Species darii Pomorski, 1998

Species decemsetosus Yosii, 1966

Species dissimulans Gisin, 1952, B

Species fimetarius Caroli, 1914

Remark by Gisin, H. 1960:140: Species inquirenda.

Species fraterculi Babenko & Fjellberg, 2016

Species furcisetosus Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1040

Onychiurus furcisetosus from the USA
After Pomorski et. al, 2009 Fig.10-11

Species gevorum Arbea, JI, 2012:36

            Syn.: Onychiurus gevi(sic) Arbea, JI, 2012:36 IOS
Onychiurus gevorum from Spain
After Wynne JJ & al, 2018 Fig.4C
Onychiurus gevorum from Spain
After Wynne JJ & al, 2018 Fig.4D



Species heilongjiangensis Sun, X et Wu, D-H, 2014:106

Species insinuans Gisin, 1952

Species jaceki Kaprus, IJ, 2009:60

Onychiurus jaceki from Kyrgyzstan
After Kaprus IJ, 2009 Fig.4A

Species longipes von Olfers, 1907

Species movilae Gruia, M, 1989:103

             Syn.: Onychiurus movile(sic) Fiera, C, 2007:16 ISS

Species nathanieli Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1040

Onychiurus nathanieli from the USA
After Pomorski et. al, 2009 Fig.20-21
             Syn.: Onychiurus nathanielli(sic) Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1042 ISS


Species obsiones Cassagnau, 1963

Species orienteuropeus Kaprus, IJ, 2009:58

Onychiurus orienteuropeus from Ukraine
After Kaprus IJ, 2009 Fig.3A

Species ortus Denis, 1935

Species paolettii Dallai, 1975

             Syn.: Onychiurus paoletti(sic) Arbea, JI, 2012:42 ISS

Species penetrans Gisin, 1952

Species pipistrellae Soto-Adames, FN et Taylor, SJ, 2013:153

Species rectospinatus Stach, 1922

            Syn.: Onychiurus arvensis Rusek, 1979


Species reluctoides Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1043

Onychiurus reluctoides from the USA
After Pomorski et. al, 2009 Fig.28-29

Species reluctus Christiansen, K, 1961:61

Onychiurus reluctus from the USA
After Pomorski et. al, 2009 Fig.1-2

Species sarmaticus Kaprus, IJ, 2009:62

Onychiurus sarmaticus from Ukraine
After Kaprus IJ, 2009 Fig.5A

Species steinmanni Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1045

Onychiurus steinmanni from the USA
After Pomorski et. al, 2009 Fig.35-37

Species stillicidii (Schiödte, 1849) Latzel, 1922

            Syn.: Aphorura Hamanni(sic) Schäffer, 1900 IOS


             Syn.: Anurophorus Stillicidii(sic) Schiödte, JC, 1851:14 ISS
             Syn.: Aphorura (Deuteraphorura) stilicidii(sic) Absolon, K, 1901:388 ISS
             Syn.: Aphorura stilicidii(sic) Absolon, K, 1901:579 ISS
             Syn.: Onychiurus stillicidi(sic) Frenzel, 1937 ISS



Species subambulans Denis, 1935

Species subcirculans Gisin, H, 1962:17

Species sublegans Gisin, 1960, B

Species szeptyckii Smolis, A et Skarzynski, D, 2009:18

            Syn.: Allonychiurus sp. Smolis & Skarzynski, 2006


Species vtorovi Martynova, 1976 in Vtorov & Martynova, 1976

Onychiurus vtorovi from Kyrgyzstan
After Kaprus IJ, 2009 Fig.1A

Species wilchi Wray, DL, 1950:98

Onychiurus wilchi from the USA
After Wray DL, 1950 Pl.5 Fig.18
            Syn.: Onychiurus (Onychiurus) reluctus Christiansen, K et Bellinger, P, 1980:455 =?
            Syn.: Onychiurus steinmanni Pomorski, RJ, Furgol, M et Christiansen, K, 2009:1045=?
            Syn.: Onychiurus formosanus Bernard, EC, 2015:335 =?



Remark by Pomorski et al. 2009:1049: O. wilchi will key out as steinmanni, but the collection sites make it improbable that they are the same species.
Remark by Pomorski et al. 2009:?: O. wilchi almost certainly belongs in this genus, but the type specimens are in such poor condition and Wray's description is clearly erroneous in some respects and inadequate in others such that certain placement will have to await further material study.
Species willemi (Börner, 1901) Börner, 1902

            Syn.: Aphorura Willemi(sic) Börner, 1901 IOS





Genus Deuteraphorura Absolon, K, 1901:387 key  key

  Syn.: Lipuridæ(sic) Lipura Schött, H, 1891:24
  Syn.: Onychiurus fimetarius-group Börner, 1909
  Syn.: Onychiuridae Aphorurra(sic) Kos, F, 1939:323 ISS
  Syn.: Collembola Onychiums(sic) da Gama, MM, 1988:76 ISS


  Syn.: Onychiuridae Deutheraphorura(sic) Weiner, WM et Fiera, C, 2014:248 ISS

Species sp.1 Parimuchová, A et Kovác, L, 2016:335

            Syn.: Deuteraphorura closanica Kovac & al, 2014 cf


Species sp.2 Parimuchová, A et Kovác, L, 2016:336

Species acicindelia (Salmon, JT, 1958:710) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:65

Deuteraphorura acicindelia from New Zealand
2014.02.06 © Murray, A.

Species akelaris (Jordana & Beruete, 1983) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:65

Species ameskoana (Beruete, E, Arbea, JI & Jordana, R, 2001:22) Pomorski, RJ, 2002:il

Species angelieri (Izarra, 1968) Pomorski, RJ in Christiansen, K, 2001:il

Species antheuili (Denis, 1936) Pomorski, RJ in Christiansen, K, 2001:il

  Subspecies aellenii Gisin, 1960

  Subspecies thesauri Gisin, 1960

Species apuanica (Dallai, R, 1970:446) Pomorski, RJ, 2002:il

Species aralarensis (Beruete, E, Arbea, JI & Jordana, R, 2001:17) Pomorski, RJ, 2002:il

Species arminiaria (Gisin, 1961) Christian, E, 2002:264

Species banatica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il

Species banii Dallai & Fanciulli, 2009

Species bergamaria (Gisin, H, 1956:350) Pomorski, RJ in Christiansen, K, 2001:il

Species boneti (Gisin, 1953) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus handschini Bonet, 1931, nec Denis, 1925


Species bosnaria (Gisin, H, 1964:394) Pomorski, RJ in Christiansen, K, 2001:il

Species caprelleana Fanciulli, Loreti & Dallai, 2010

Species cebennaria (Gisin, H, 1956:348) Pomorski, RJ, 1996:52

Remark by Janssens, F 2000.04.30: "cebennaria" is presently depriciated. In this checklist, "inermis" is established as the tentatively valid name of "cebennaria", in line with the proposal of Bellinger (2000:in litt.) to resolve the invalid state of "Deuteraphorura".

Remark by Fjellberg, A. 2004.10.04:in litt.: But until a neotype for "inermis" is designated, there is no published evidence that "cebennaria" is a junior synonym and according to ICZN rulings, until the appropriate actions are taken, "cebennaria" should be used in taxonomic publications.

Species closanica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il

Species defensaria (Gisin, H, 1964:397) Pomorski, RJ in Christiansen, K, 2001:il

Species diaelleni (Neuherz & Nosek, 1976) Pomorski, RJ, 2002:il

Species dianae Weiner, WM et Fiera, C, 2014:248

Deuteraphorura dianae from Romania
After Weiner WM & Fiera C, 2014 Fig.1

Species difficilis (Dallai, 1969) Pomorski, RJ in Christiansen, K, 2001:il

Species doftana Weiner, WM et Fiera, C, 2014:250

Deuteraphorura doftana from Romania
After Weiner WM & Fiera C, 2014 Fig.8

Species doneztebensis (Beruete, E, Arbea, JI & Jordana, R, 2001:19) Pomorski, RJ, 2002:il

Species dunaria (Gisin, H, 1956:349) Pomorski, RJ in Christiansen, K, 2001:il

Species eduardi (Denis, 1937) Pomorski, RJ in Christiansen, K, 2001:il

Species fazii (Christiansen, K, 1956:130) Pomorski, RJ in Christiansen, K, 2001:il

Onychiurus fazii from Lebanon
After Christiansen K, 1956 Fig.4

Species frasassii (Fanciulli, 1999) Pomorski, RJ in Christiansen, K, 2001:il

             Syn.: Deuteraphorura frasassi(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:71 ISS
             Syn.: Deuteraphorura frassasii(sic) Weiner, WM et Fiera, C, 2014:250 ISS

Species galani (Beruete, E, Arbea, JI & Jordana, R, 2001:16) Pomorski, RJ, 2002:il

Species gangjinensis (Lee, B-H & Kim, J-T, 1995:138) Pomorski, RJ, 2002:il

Species gemae (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:61) Pomorski, RJ, 2002:il

            Syn.: Onychiurus cavernicolus Pozo, 1983
            Syn.: Onychiurus fimetarius Selga, 1971

Species ghidinii (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Lipura bipunctata Moniez, 1889 nn
            Syn.: Onychiurus fimetarius Denis, 1923
            Syn.: Onychiurus ghidinii-varians(sic) Denis, 1938


             Syn.: Onychiurus ghidini(sic) Gridelli, 1955 ISS
             Syn.: Onychiurus ghidini(sic) Haybach, G, 1962:94 ISS

Species gigoni (Gisin, H, 1962:6) Pomorski, RJ in Christiansen, K, 2001:il

Species gridellii (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus stachianus Bagnall, 1939


             Syn.: Protaphorura gridelli(sic) Salmon, JT, 1964:171 ISS
             Syn.: Deuteraphorura gridelli(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:70 ISS

Species handschini (Denis, JR, 1924:197) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus Handschini(sic) Denis, JR, 1924:197 IOS


  Subspecies strinatii Gisin, 1963

             Syn.: Deuteraphorura handschini ssp. strinati(sic) Weiner, WM et Fiera, C, 2014:253 ISS
             Syn.: Deuteraphorura handschini ssp. strinati(sic) Parimuchová, A et Kovác, L, 2016:337 ISS



Species harrobiensis (Beruete, E, Arbea, JI & Jordana, R, 2001:15) Pomorski, RJ, 2002:il

Species hategana (Gruia, M, 1971:283) Pomorski, RJ, 2002:il

Species haybachae (Gisin, H, 1962:8) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus austriarius Gisin, H, 1962:14


Species hussoni (Denis, 1935) Pomorski, RJ in Christiansen, K, 2001:il

             Syn.: Onychiurus handschini ssp. hussini(sic) Jacquemart, S, 1976:157 ISS

Species imperfecta (Denis, 1938) Pomorski, RJ in Christiansen, K, 2001:il, B

Species inermis (Tullberg, 1869) Absolon, 1901, t.t., B

            Syn.: Podura fimetaria von Linné, C, 1767:1014
             Syn.: Podura fimitaria(sic) Herbst, JFW, 1787:130 ISS
             Syn.: Podura Fimetaria(sic) Herbst, JFW, 1787:350 ISS
Podura Fimetaria(sic) from Germany
After Herbst JFW, 1787 Taf.LXXV Fig.2

            Syn.: Anurophorus fimetarius Nicolet, H, 1847:385
Anurophorus fimetarius from Europe
After Nicolet,, 1847 Pl.6 Fig.15.
Lipura fimetaria from the UK
After Lubbock J, 1870 Pl.XXII Fig.27
Lipura fimetaria from the UK
Posterior foot
After Lubbock J, 1870 Pl.XXII Fig.28
            Syn.: Lipura fimetaria Lubbock, J, 1873:191
Lipura fimetaria from the UK
After Lubbock J, 1873 Pl.46

             Syn.: Lipura fimentaria(sic) Parfitt, 1891 ISS
             Syn.: Onychiurus funetarius(sic) Lie-Pettersen, 1906 ISS
             Syn.: Aphorura fimetarius(sic) Navas, 1906 ISS
             Syn.: Onychiurus fimentarius(sic) Wahlgren, 1906 ISS
             Syn.: Lipura fimetarius(sic) Theobald, 1910 ISS
             Syn.: Onychiurus fimentarius(sic) Axelson, 1919 ISS
             Syn.: Onychiurus fimentarius(sic) Brown, 1919 ISS
             Syn.: Onychiurus fimentarius(sic) Wahlgren, 1919 ISS
             Syn.: Onychiurus fimetaria(sic) Griepenburg, 1935 ISS
             Syn.: Onychiurus fimetaria(sic) Tarsia in Curia, 1935 ISS
             Syn.: Onychiurus fimentarius(sic) Imms, AD, 1939:291 ISS
            Syn.: Onychiurus fimetarius Salmon, 1941
Onychiurus fimetarius from New Zealand
After Salmon, J.T., 1941 Pl.47 Fig.130.

             Syn.: Onychiurus fimatarius(sic) Lawrence, 1959 ISS
             Syn.: Onychiurus firmetarius(sic) Vannier, G, 1971:123 ISS
             Syn.: Onychiurus (Onychiurus) fimentarius(sic) Yayuk, 1989 ISS
Onychiurus sp. fimetarius gr. from Tasmania
Post-antennal organ with compound lobes
After Ireson JE, 1990 Fig.73

             Syn.: Onychiurus fimeitayius(sic) Chen, J-X, Ma, Z-C, Yan, H-J et Zhang, F, 2007:157 ISS
Remark by Dányi, L. & Traser, G. 2008:21: Deuteraphorura fimetaria (Linnaeus, 1766) is a "species inquirenda according to Gisin (1960)". [Janssens, F., 2008.11.15. Contrary to what Dányi & Traser (2008:21) claim, Gisin (1960:140) refers to "fimetarius" sensu Denis (1938), not to fimetaria von Linné, 1767. The latter is considered as a species dubia by Gisin (1960:151)].


            Syn.: Podura monura Schrank, 1781,sd
            Syn.: Lipura volvator Gervais in Walckanaër, 1844
Lipure volvaire from France
Lipura volvator
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.3

            Syn.: Anurophorus stillicidii Wright, 1857
            Syn.: Anurophorus coecus Joseph, 1882             Syn.: Lipura Wrightii(sic) Carpenter, 1895 IOS
            Syn.: Lipura bipunctata Moniez, 1889
            Syn.: Aphorura folsomi Schäffer, 1900
            Syn.: Aphorura spelaea Absolon, K, 1900:411
             Syn.: Aphorura spelea(sic) Willem, V, 1902:278,B ISS
            Syn.: Aphorura spelaea ssp. multipunctata Absolon, 1901
            Syn.: Onychiurus pseudinermis Börner, 1903
            Syn.: Onychiurus pseudofimetarius Folsom, 1917
            Syn.: Onychiurus camerunensis Schött, 1926, sensu Salmon, 1964:180
            Syn.: Onychiurus fimetarius var. denticulata Denis, 1929
            Syn.: Onychiurus subterraneus Stach, J, 1930:285
            Syn.: Onychiurus pygmaeus Bagnall, 1937
            Syn.: Onychiurus insubrarius Gisin, 1952
            Syn.: Onychiurus cebennarius Gisin, 1956
             Syn.: Paronychiurus cebannarius(sic) Salmon, JT, 1964:186 ISS
            Syn.: Onychiurus imperfectus Bödvarsson, 1963



Aphorura inermis from Japan
After Folsom JW, 1899 Pl.1 Fig.1
Aphorura inermis from Japan
After Folsom JW, 1899 Pl.1 Fig.2
Deuteraphorura inermis from the UK
Post-antennal organ
2002 © Hopkin, S.P.
Deuteraphorura inermis from the UK
2006 © Valentine., B
Deuteraphorura inermis from the UK
2009.04.15 © Robertson, A.
Deuteraphorura inermis from the UK
2009.04.15 © Robertson, A.
Deuteraphorura inermis from France
2009.04.14 © Raphael from Paris
Deuteraphorura inermis from the UK
2009.04.29 © Robertson, A.
Deuteraphorura inermis from the UK
2010.02.09 © Batten-Phelps, A.
Deuteraphorura inermis from the UK
2011.02.27 © Robertson, A.
Deuteraphorura inermis from the UK
2012.01.22 © Jones, A.
Deuteraphorura inermis from the UK
2012.01.29 © Kent, J.J.
Deuteraphorura inermis from the UK
2012.01.29 © Kent, J.J.
Deuteraphorura inermis from the UK
2012.11.04 © Jones, A.
Deuteraphorura inermis from the UK
Complex sense organ of 3rd antennal segment (A3O) with 2 sensilla and 5 guard papilla
2013.02.06 © Murray, A.
Deuteraphorura inermis from the UK
Long post-antennal organ (PAO) with ca. 15 sensilla clusters
2013.02.06 © Murray, A.
Deuteraphorura inermis from Tasmania
2016.07.16 © Zosterops.
Deuteraphorura inermis from France
2019.02.13 © Garcelon, P.

  Subspecies dentiferus Kos, F, 1939:316

Onychiurus fimetarius var. dentiferus from Yugoslavia
After Kos F ©, 1939 Fig.80
Onychiurus fimetarius var. dentiferus from Yugoslavia
Infested with Gordius larva
After Kos F ©, 1939 Fig.81




             Syn.: Onychiurus fimetarius var. dentifera(sic) Kos, F, 1939:323 ISS
             Syn.: Onychiurus fimatarius(sic) var. dentifera(sic) Gisin, H, 1960:151,sd ISS



Species inferni (Gisin, H, 1956:351) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:66

Species insubraria (Gisin, 1952) Pomorski, RJ in Christiansen, K, 2001:il

Deuteraphorura insubraria
2006 © Boldsystems.org

Species jana (Christiansen, K & Bellinger, P, 1980:463) Pomorski, RJ, 2002:il

Species jitkae (Rusek, 1964) Pomorski, RJ in Christiansen, K, 2001:il

Species koreana Arbea, JI et Park, K-H, 2015:79

Deuteraphorura koreana from South Korea
After Arbea JI & Park K-H, 2015 Fig.1-2
            Syn.: Onychiurus (Deuteraphorura) izuruensis Lee & Park, 1986


Species kosarovi (Zonev, 1973) Pomorski, RJ in Christiansen, K, 2001:il

             Syn.: Deuteraphorura kosavori(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:67 ISS

Species kratochvili (Nosek, 1963) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Deuteraphorura fimetaria Kovac, 1998 cf
            Syn.: Deuteraphorura fimetaria Kovac, 1999 cf
            Syn.: Deuteraphorura fimetaria Kovac, 2000 cf
            Syn.: Deuteraphorura hussoni Kovac & Kosel, 1998 cf
            Syn.: Deuteraphorura hussoni Kovac, 1999 cf
            Syn.: Deuteraphorura kratochvili Papác, 2008 cf =?
            Syn.: Deuteraphorura sp.1 Kovac & al, 1999
            Syn.: Deuteraphorura sp.2 Kovac & al, 1999
            Syn.: Deuteraphorura sp. Kovac & al, 2003


Deuteraphorura kratochvili from Slovakia
After Kovác L & L'uptácik P in Parimuchová A & Kovác L, 2016 Fig.1

Species kruberaensis Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:60

Species labainensis (Beruete, E, Arbea, JI & Jordana, R, 2001:21) Pomorski, RJ, 2002:il

             Syn.: Deuteraphorura lebainensis(sic) Jordana R et Baquero E in Jordana, R, Baquero, E, Reboleira, S et Sendra, A, 2012:69 ISS

Species leitzaensis (Beruete, E, Arbea, JI & Jordana, R, 2001:13) Pomorski, RJ, 2002:il

Species lusa (Christiansen, K & Bellinger, P, 1980:465) Pomorski, RJ, 2002:il

Species mangazeya Babenko, AB, 2007:372

Species meziadica (Gruia, 1972) Pomorski, RJ in Christiansen, K, 2001:il

Species mildneri (Christian, E, 1985:146) Pomorski, RJ in Christiansen, K, 2001:il

Species montagudi Arbea, JI, 2015:237

Deuteraphorura montagudi from Spain
After Arbea JI, 2015 Fig.15-16

Species nervosa (Stach, J, 1954:167) Pomorski, RJ in Christiansen, K, 2001:il

Species nevoi (Gruia, Poliakov & Broza, 2000) Pomorski, RJ in Christiansen, K, 2001:il

Species opa (Christiansen, K & Bellinger, P, 1980:466) Pomorski, RJ, 2002:il

Species oregonensis (Denis, 1929) Pomorski, RJ in Christiansen, K, 2001:il

Species ossaria (Gisin, H, 1964:395) Pomorski, RJ in Christiansen, K, 2001:il

Species paro (Christiansen, K & Bellinger, P, 1980:468) Pomorski, RJ in Christiansen, K, 2001:il

Species pseudobosnaria (Dallai, R, 1970:447) Pomorski, RJ in Christiansen, K, 2001:il

Species pseudofimetaria (Folsom, 1917) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Aphorura spelaea Absolon, 1901
            Syn.: Aphorura inermis Guthrie, 1903
            Syn.: Onychiurus pseudinermis Börner, 1903


             Syn.: Onychiurus fimetarius-pseudofimetarius(sic) Denis, 1924 ISS
             Syn.: Onychiurus fimetarius-pseudofimetarius(sic) Paulian, R et Delamare Deboutteville, C, 1946:16 ISS
             Syn.: Onychiurus pseudofimeratus(sic) Salmon, 1964 ISS

Species pseudoghidinii (Dallai, 1969) Pomorski, RJ in Christiansen, K, 2001:il

Species pseudoinsubraria (Dallai, R, 1970:447) Pomorski, RJ in Christiansen, K, 2001:il

Species quadrisilvaria (Gisin, H, 1962:16) Christian, E, 2002:264

Species rendsinae (Haybach, G, 1962:94) Pomorski, RJ in Christiansen, K, 2001:il

Species romanica (Gruia, 1965) Pomorski, RJ in Christiansen, K, 2001:il

Species schoenviszkyi (Loksa, I, 1967:292) Parimuchová, A & Kovác, L, 2016:333

             Syn.: Onychiurus schöenviszkyi(sic) Bajomi, 1968 ISS

Species scotaria (Gisin, 1954) Pomorski, RJ, 1996:49

Deuteraphorura scotaria from the UK
Male genital plate
2002 © Hopkin, S.P.
Deuteraphorura scotaria from the UK
Female genital plate
2002 © Hopkin, S.P.

Species sexpunctata (Schäffer, 1897) Pomorski, RJ, 2002:il

Species silesiaca (Dunger, W, 1977:10) Pomorski, RJ, 1998:163

Species silvaria (Gisin, 1952) Weiner, WM, 1996:185

            Syn.: Onychiurus insubrarius da Gama, 1964
            Syn.: Onychiurus sp. Arbea & Baena, 2004
            Syn.: Onychiurus sp. Pérez Fernández, 2004
            Syn.: Onychiurus sp. Pérez Fernández & Tinaut, 2005
            Syn.: Onychiurus sp. Pérez Fernández, 2008
            Syn.: Onychiurus subargus Luciáñez & Gómez-Silgado, 2007 partim


             Syn.: Deuteraphorura silvarius(sic) Weiner, WM, 1996:185 ISS
             Syn.: Deuteraphorura sylvaria(sic) Babenko, AB, 2007:374 ISS
Deuteraphorura silvaria from Spain
On guano de murciélago
After Pérez T in Arbea JI, 2013 Fig.1
Deuteraphorura silvaria from Spain
After Arbea JI, 2013 Fig.5



Species simoni Arbea, JI, 2017:366

Deuteraphorura simoni from Spain
After Arbea JI, 2017 Fig.1
Deuteraphorura simoni from Spain
After Arbea JI, 2017 Fig.6

Species spipolae (Massera, 1949) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:68

Species traiani Gruia & Popa, 2005

Species trisilvaria (Gisin, H, 1962:15) Christian, E, 2002:264

Species troglocarpathica (Absolon, K, 1901:381) Jordana R & Baquero E in Jordana, R, Baquero, E, Reboleira, S & Sendra, A, 2012:68

Remark by Absolon, K. 1901:381: "...welche ich später beschreiben werde."

             Syn.: Aphorura troglocarpatica(sic) Stach, J, 1954:79 nn ISS
Remark by Stach, J. 1954:79: Nomen nudum.

Remark by Salmon, J.T. 1964:189: Species indeterminatae.



Species variabilis (Stach, J, 1954:164) Pomorski, RJ, 1996:53

Species vercoraria (Gisin, 1963) Pomorski, RJ in Christiansen, K, 2001:il

Species vinuensis (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:67) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus insubrarius Selga, 1971


Species zalbidensis (Beruete, E, Arbea, JI & Jordana, R, 2001:20) Pomorski, RJ, 2002:il


Genus Orthonychiurus Stach, J, 1954:26

  Syn.: Collembola Onychiums(sic) da Gama, MM, 1988:76 ISS

Species afghanistanensis (Stach, J, 1960:512) Pomorski, RJ in Christiansen, K, 2001:il

Species ancae (Gruia, M, 1971:285) Pomorski, RJ in Christiansen, K, 2001:il

Species argaensis (Arbea, JI & Jordana, R in Jordana, R & Ariño, AH, 1994:40) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus justi Arbea & Jordana, 1991


Species azoricus (Jacquemart, S, 1974:1) Pomorski, RJ, 2002:il

Species bunsteri (Loksa & Rubio, 1966) Pomorski, RJ in Christiansen, K, 2001:il

Species camerunensis (Schött, 1926) Pomorski, RJ in Christiansen, K, 2001:il

Species cunhai (Arlé, R, 1970:1) Pomorski, RJ, 2002:il

Species doinae (Gruia, 1972) Pomorski, RJ in Christiansen, K, 2001:il

Species durkeei (Christiansen, K & Bellinger, P, 1980:458) Pomorski, RJ in Christiansen, K, 2001:il

Species folsomi (Schäffer, 1900) Jordana & al, 1997

            Syn.: Aphorura inermis Folsom, 1899
            Syn.: Aphorura inermis Yosii, R, 1977:152
            Syn.: Onychiurus fimetarius Womersley, 1933
            Syn.: Onychiurus fimetarius Greenslade P & Ireson, 1986
            Syn.: Onychiurus sinensis Stach, 1954
            Syn.: Onychiurus sinensis Yosii, R, 1977:152
            Syn.: Onychiurus gotoi Choudhuri, DK, 1958:157
            Syn.: Onychiurus musae Selga, 1962
            Syn.: Tullbergia neomexicana Scott, 1961 partim holotype
            Syn.: Onychiurus justi Snider, 1971
            Syn.: Onychiurus justi Snider & Butcher, 1972


             Syn.: Aphorura Folsomi(sic) Absolon, K, 1901:381 ISS
             Syn.: Aphorura Folsomi(sic) Absolon, K, 1901:387 ISS

Species gloriensis (Gruia, M, 1987:151) Pomorski, RJ in Christiansen, K, 2001:il

Species gurjakhanii Choudhuri, DK, 1958:150

Species himalayensis (Choudhuri, DK, 1958:147) Pomorski, RJ in Christiansen, K, 2001:il

Species izuruensis (Yosii, R, 1956:51) Yoshii, 1995

Species kowalskii (Stach, J, 1964:12) Pomorski, RJ in Christiansen, K, 2001:il

Species lampedusae (Fanciulli & Dallai, 1995) Pomorski, RJ in Christiansen, K, 2001:il

Species neocaledonicus Najt, J et Weiner, WM in Najt, J et Matile, L, 1997:18


Onychiurus novae-zealandiae(sic) from New Zealand
After Salmon, J.T. 1942 Pl.12 Fig.1.
Species novaezealandiae (Salmon, JT, 1942:158) Pomorski, RJ in Christiansen, K, 2001:il

            Syn.: Onychiurus novae-zealandiae(sic) Salmon, JT, 1942:158 IOS


             Syn.: Onychiurus novae-zealandiae(sic) Salmon, JT, 1964:183 ISS



Species pseudostachianus (Gisin, H, 1956:346) Pomorski, RJ, 1998:182

Remark by Pomorski, R.J. 1998:182: The separateness of Orthonychiurus pseudostachianus [from Orthonychiurus stachianus (Bagnall, 1939)] should be confirmed by a detailed examination.

Species rectopapillatus (Stach, 1933) Pomorski, RJ, 1996:56, t.t.

            Syn.: Onychiurus pavlovskyi Nosek & Vysockaya, 1965


             Syn.: Orthonychiurus rectopapilatus(sic) Fiera, C, 2007:17 ISS
             Syn.: Onychiurus rectapapillatus(sic) Falahati HosseinAbad, A, Mehr, M S et Khyroodin, A, 2013:260 ISS

Species richardsi (Choudhuri, DK, 1958:150) Pomorski, RJ, 2002:il

Species saasveldensis (Weiner & Najt, 1991) Weiner, WM, 1996:187

Species selgae (Simón-Benito, JC & Luciáñez, MJ in Jordana, R & Ariño, AH, 1994:65) Pomorski, RJ in Christiansen, K, 2001:il

Species stachianus (Bagnall, 1939) Pomorski, RJ, 1998:179

            Syn.: Onychiurus stachi Denis, 1938, nec Bagnall, 1935


Species subantarcticus (Salmon, JT, 1949:15) Pomorski, RJ in Christiansen, K, 2001:il

Species subcadaverinus (Denis, 1931) Pomorski, RJ, 2002:il

            Syn.: Onychiurus fimetarius Mari Mutt, 1977


Species sublapidarius (Stach, 1963) Pomorski, RJ in Christiansen, K, 2001:il

Species ugandensis (Choudhuri, DK, 1958:152) Pomorski, RJ, 2002:il


Genus Bionychiurus Pomorski, RJ, 1996:92

  Syn.: Onychiurini Bagnallophorus Weiner, WM, 1996:183


Species changbaiensis Sun & Wu, 2012

Species normalis (Gisin, 1949) Pomorski, RJ, 1996:92, t.t., B

Species oblongatus (Lee & Park, 1986) Pomorski, RJ, 2002:il

Species orghidani (Gruia, M, 1967:103) Arbea, JI, 2018:106

Species qinglongensis Sun, X et Wu, D-H, 2014:102

Species yongyeonensis (Yosii, R, 1966:550) Pomorski, RJ, 2002:il


Genus Leeonychiurus Sun, X et Arbea, J, 2014:116 key

Species analis Lee, I et Park, K-H, 2017:295

Leeonychiurus analis from Korea
After Lee I & Park K-H, 2017 Fig.3A

Species antennalis (Sun, X & Zhang, F, 2012:1900) Sun, X & Arbea, J, 2014:117

Species fusongensis Sun, X et Arbea, J, 2014:117, t.t.

Leeonychiurus fusongensis from China
After Sun X & Arbea J, 2014 Fig.1

Species gulinensis (Sun, X & Zhang, F, 2012:1896) Sun, X & Arbea, J, 2014:117

Onychiurus gulinensis from China
Upper: ventral aspect
After Sun X & Zhang F, 2012 Fig.1A,B

Species koreanus Lee, I et Park, K-H, 2017:293

Leeonychiurus koreanus from Korea
After Lee I & Park K-H, 2017 Fig.1A
            Syn.: Leeonychiurus sp.1 Lee et al, 2014


Species mai (Wray, 1950) Bernard, EC, 2015:332

            Syn.: Onychiurus oreadis Mills, HB, 1935:133
            Syn.: Onychiurus flavescens Bernard, EC, 2015:332 =?
            Syn.: Onychiurus millsi Bernard, EC, 2015:332 =?


Species polychaetosus (Lee & Park, 1986) Sun, X & Arbea, J, 2014:120

Species saphianus (Nguyen, 2001) Sun, X & Arbea, J, 2014:117

Remark by Sun, X. &Arbea, J. 2014:117: Species assignment requires confirmation. We do not have information on the type material, in spite of attempts to contact the author of the species.


Genus Deharvengiurus Weiner, WM, 1996:183

  Syn.: Onychiurus argus-subgroup Deharveng, L et Gouze, A, 1984:389
  Syn.: Onychiuridae Aphorura Arbea, JI, 2018:106
  Syn.: Onychiuridae Onychiurus Arbea, JI, 2018:105


Deharvengiurus sp. from Spain
Dorsal aspect
After Arbea JI, 2018 Fig.1
Deharvengiurus sp. from Spain
Ventral aspect
After Arbea JI, 2018 Fig.2

Species argus (Denis, JR, 1924:197) Weiner, WM, 1996:183

            Syn.: Onychiurus Argus(sic) Denis, JR, 1924:197 IOS
            Syn.: Onychiurus severini Stach, 1934
            Syn.: Onychiurus severini Gisin, 1944
            Syn.: Aphorura severini Stach, J, 1954:54 =?
            Syn.: Onychiurus cantabricus Steiner, 1958




Species denisi (Stach, J, 1934:191) Pomorski, RJ, 1998:190

            Syn.: Onychiurus paxi Stach, 1936


Species lopezi (Deharveng, L & Gouze, A, 1984:395) Weiner, WM, 1996:184

Species lucianezae Arbea, JI, 2018:110

Deharvengiurus lucianezae from Spain
After Arbea JI, 2018 Fig.28

            Syn.: Onychiurus subargus Luciáñez & Gómez-Silgado, 2007 partim
            Syn.: Deharvengiurus steineri Arbea, JI, 2018:110 lapsus


Species microchaetosus (Loksa, 1959) Pomorski, RJ, 2002:il

Species pseudocantabricus (Gouze, 1983) Pomorski, RJ in Christiansen, K, 2001:il

Species severini (Willem, V, 1902:277) Pomorski, RJ in Christiansen, K, 2001:il, B

            Syn.: Onychiurus argus Marlier, GJ, 1942:8,B
            Syn.: Onychiurus (Onychiurus) argus Gisin, H, 1960:136,B
            Syn.: Onychiurus (Paronychiurus) argus Palissa, A in Brohmer, P, Ehrmann, P et Ulmer, G, 1964:120,B
            Syn.: Paronychiurus argus Salmon, JT, 1964:185,B
            Syn.: Onychiurus argus Thibaud, J-M et Massoud, Z, 1977:6,B
            Syn.: Onychiurus argus Hubart, J-M, 1982:343,B
            Syn.: Deharvengiurus argus Dethier, M et Hubart, J-M, 2000:28,B


Deharvengiurus severini from Belgium
Short fine anal spines not on papillae
2017.08.12 © Janssens, F.
Prep.: 2017.06-07 Dethier, M.

Species steineri Arbea, JI, 2018:109

Deharvengiurus steineri from Spain
After Arbea JI, 2018 Fig.20

            Syn.: Onychiurus cantabricus Arbea, JI, 2018:109 partim
            Syn.: Onychiurus subargus Simón JC in Arbea, JI, 2018:109


Species subargus (Deharveng, L & Gouze, A, 1984:395) Weiner, WM, 1996:184

            Syn.: Onychiurus argus Simón JC in Arbea, JI, 2018:109
            Syn.: Onychiurus cantabricus Arbea, JI, 2018:109 partim
            Syn.: Onychiurus pseudocantabricus Simón JC in Arbea, JI, 2018:109


Species xerophilus (Loksa, I, 1964:86) Pomorski, RJ in Christiansen, K, 2001:il


Genus Onychiuroides Bagnall, 1948

Species anellii (Denis, 1938) Weiner, WM, 1996:186

             Syn.: Onychiuroides annelli(sic) Weiner, WM, 1996:186 ISS
             Syn.: Onychiuroides anelli(sic) Pomorski, 2006 ISS

Species beroni (Gruia, 1972) Pomorski, 2006

Species bojani Pomorski, 2006

Species bureschi (Handschin, 1928) Pomorski, 2006

Species canzianus (Stach, J, 1934:200) Pomorski, 2006

             Syn.: Onychiuroides canzianits(sic) Pomorski, 2006 ISS

Species cavernicolus (Stach, J, 1934:212) Weiner, WM, 1996:186

             Syn.: Onychiurus cavernicola(sic) Gisin, H, 1962:10 ISS

Species cerberus Pomorski, 2006

Species granulosus (Stach, J, 1930:285) Pomorski, RJ, 1996:62

            Syn.: Onychiurus granulosus f. denticulata Stach, 1934
            Syn.: Onychiurus pseudogranulosus Delamare Deboutteville & Gisin, 1951


             Syn.: Onychiurus granulossus(sic) Gruia, M, 1994:73 ISS

Species hauseri (Dallai, 1975) Pomorski, RJ in Christiansen, K, 2001:il

Species hrabei (Rusek, 1963) Pomorski, 2006

Species igori Pomorski, 2006

Species longisetosus (Stach, J, 1954:145) Pomorski, 2006 si

             Syn.: Onychiurus (Onychiurus) longisetus(sic) Gisin, H, 1960:144 ISS
             Syn.: Onychiurus longisetus(sic) Rusek, J in Dlabola, J, 1977:13 ISS

Species multisetis (Gruia, M, 1971:287) Gruia, 2003

Species novus (Stach, J, 1960:511) Pomorski, RJ in Christiansen, K, 2001:il

Species occultus Pomorski, 2006

Species paoloi Pomorski, 2006

Species papillaeferus (Stach, J, 1946:20) Pomorski, 2006

Species paucituberculatus (Stach, J, 1930:285) Pomorski, 2006

            Syn.: Onychiurus paucotuberculatus(sic) Stach, J, 1930:271 IOS
Onychiuroides paucituberculatus from Croatia
2010.11.19 © Bilandzija, H.



Species peteri Pomorski, 2006

Species pirinensis Pomorski, 2006

Species postumicus (Bonet, 1931) Pomorski, RJ, 1996:95, t.t.

Species pseudogranulosus (Delamare Deboutteville & Gisin, 1951) Weiner, WM, 1996:186, B

Onychiuroides pseudogranulosus from Turkey
After Özata MA, Sevgili H & Kaprus IJ, 2017 Fig.10

Species quadripapillatus (Rusek, 1965) Pomorski, 2006

Species subgranulosus (da Gama, 1964) Pomorski, 2006

             Syn.: Onychiuroides subgranulostis(sic) Pomorski, 2006 ISS

Species vornatscheri (Stach, J, 1946:23) Pomorski, 2006


Genus Absolonia Börner, 1901

Species gigantea (Absolon, K, 1901:85) Börner, 1901, t.t.

Absolonia gigantea from Croatia
2010.11.19 © Bedek, J.
Absolonia gigantea from Croatia
2010.11.19 © Lukic, M.


Genus Argonychiurus Bagnall, 1949

Species bertrandi (Denis, 1936) Pomorski, RJ in Christiansen, K, 2001:il

Species bogheani (Gruia, M, 1989:104) Pomorski, RJ in Christiansen, K, 2001:il

Species fistulosus (Gisin, H, 1956:345) Weiner, WM, 1996:183

            Syn.: Onychiurus perforatus Gisin, 1943, nec Handschin, 1921

Species lenticularius (Gisin, H, 1962:5) Djanashvili, Barjadze, Jordana & Burkhardt, 2014

             Syn.: Onychiurus lenticularis(sic) Salmon, JT, 1964:183 ISS
             Syn.: Onychiurus lenticularis(sic) Dallai, R, Malatesta, E et Ramellini, P in Minelli, A, Ruffo, S et La Posta, S, 1995:7 ISS

Species multiocellatus Djanashvili, Barjadze, Jordana & Burkhardt, 2014

Species papulosus (Gisin, H, 1964:392) Djanashvili, Barjadze, Jordana & Burkhardt, 2014

Species perforatus (Handschin, 1920) Pomorski, RJ, 1996:94


Genus Ongulonychiurus Thibaud & Massoud, 1986

Species colpus Thibaud & Massoud, 1986, t.t.

Ongulonychiurus colpus from Spain
After Thibaud J-M & D'Haese C, 2010 Fig.20


Genus Vibronychiurus Pomorski, 1998 key

Species aestimabilis Pomorski, RJ, 2006:674

Species archivari (Christiansen, K, 1956:128) Pomorski, RJ, 2006:673

Species caucasicus Pomorski, RJ, 2006:675

Species hermonicus (Gruia, Poliakov & Broza, 2000) Pomorski, RJ, 2006:676

Species vinolentus Pomorski, 1998 tt


Genus Pilonychiurus Pomorski, RJ, 2007:56

Species delhezi (Stomp, 1983) Pomorski, RJ, 2007:56, t.t.


Genus Uralaphorura Martynova, EF, 1978:63

Species sp.1 Babenko, AB, 2007:372

Species liangshuiensis Xie, Sun & Weiner, 2017

Species schilovi (Martynova, 1976) Martynova, 1978, t.t.

Species tunguzica Babenko, AB, 2007:369

Species varicellata Babenko, 2009

Species yanensis Babenko, 2009


Genus Formosanonychiurus Weiner, 1986

  Syn.: Onychiurini Formosanochiurus(sic) Sun, X et Arbea, J, 2014:115 ISS

Species formosanus (Denis, 1929) Weiner in Dallai, 1986

            Syn.: Onychiurus (Paronychiurus) formosanus f. japonica Yosii, 1956


Species humboldtianus (Weiner, 1986) Weiner, WM, 1996:185

             Syn.: Formosanonychiurus humboltianus(sic) Weiner, WM, 1996:185 ISS



Tribus Cribrochiurini Weiner, 1996


Genus Cribrochiurus Weiner, WM, 1996:189

Species cribrosus (Gisin, 1957) Weiner, WM, 1996:189, t.t.

Species subcribrosus (Gisin, 1957) Weiner, WM, 1996:190


References